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<document ID-CLB-Dataset="6710" ID-DOI="http://dx.doi.org/10.3897/dez.63.10241" ID-GBIF-Dataset="0a6965a7-b124-4871-95d4-9e35c49073de" ID-Pensoft-Pub="1860-1324-2-211" ID-Pensoft-UUID="51243BBBE9F158F58BBC9BF3EA964017" ID-Zenodo-Dep="167458" ID-ZooBank="45552C4EC6AE4F9499980C2D492333B3" ModsDocID="1860-1324-63-2-211" checkinTime="1621372567443" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2016" docId="864E8437A1CEF89B89CD1B84D88A1FEF" docLanguage="en" docName="DeutEntomolZeit 63(2): 211-270" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docPubDate="2016-11-18" docSource="http://dx.doi.org/10.3897/dez.63.10241" docTitle="Cyphocoleus cychroides Chaudoir 1877" docType="treatment" docVersion="5" id="51243BBBE9F158F58BBC9BF3EA964017" lastPageNumber="244" masterDocId="51243BBBE9F158F58BBC9BF3EA964017" masterDocTitle="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia" masterLastPageNumber="270" masterPageNumber="211" pageNumber="244" updateTime="1691866435949" updateUser="ExternalLinkService">
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<mods:title id="28BE0E1FD8220E5596B72E497820A777">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="1A36148F0D9288C851527C18A45CD284">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="9E41C322B088B0829D2596025A2E29A9">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:identifier id="0844612AE2686F98AB83FD36D9E04988" type="DOI">http://dx.doi.org/10.3897/dez.63.10241</mods:identifier>
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<treatment id="864E8437A1CEF89B89CD1B84D88A1FEF" ID-GBIF-Taxon="182223720" LSID="urn:lsid:plazi:treatment:864E8437A1CEF89B89CD1B84D88A1FEF" httpUri="http://treatment.plazi.org/id/864E8437A1CEF89B89CD1B84D88A1FEF" lastPageNumber="244" pageId="33" pageNumber="244">
<subSubSection id="A01D53895159384C615AC013803684E6" pageId="33" pageNumber="244" type="nomenclature">
<paragraph id="4E01A74039B38476654DC4FB5A092BB6" pageId="33" pageNumber="244">
14.
<taxonomicName id="6322B29A6BF2966A1DD23FEF4165DB52" ID-CoL="6C8Z9" authority="Chaudoir, 1877" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus cychroides" order="Coleoptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="cychroides">Cyphocoleus cychroides Chaudoir, 1877</taxonomicName>
<figureCitation id="D3855E801354B946297620902FF119F5" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="33" pageNumber="244">Figures 107</figureCitation>
<figureCitation id="1BB54CA84D8DA7DCDDCC52C43758A7CD" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="33" pageNumber="244">, 114</figureCitation>
<figureCitation id="C71E0D97ED49AFB9167F493CDD4E992D" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="33" pageNumber="244">, 121</figureCitation>
<figureCitation id="CCE2378C1314D0D71628CA5ED73963FE" captionStart="Figures 137–140" captionStartId="F22" captionText="Figures 137 - 140. Distribution maps of Cyphocoleus spp.: 137, Cyphocoleus cychroides, Cyphocoleus cordatus, Cyphocoleus cardiopterus; 138, Cyphocoleus latipennis; 139, Cyphocoleus globulicollis; 140 Cyphocoleus flavipes, Cyphocoleus bourailensis." figureDoi="10.3897/dez.63.10241.figures137-140" httpUri="https://binary.pensoft.net/fig/111561" pageId="33" pageNumber="244">, 137</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="54C636A1436A53DD3A43E936CACDAE61" pageId="33" pageNumber="244" type="diagnosis">
<paragraph id="D7A4F68ACB52E4B4B5B33C2FBF69D7C7" pageId="33" pageNumber="244">Diagnosis.</paragraph>
<paragraph id="8A9CAE7D4E66332C47F77F891EB95B7F" pageId="33" pageNumber="244">
Among all the smaller-bodied species with orbicular pronotum and cordate elytra (Figs
<figureCitation id="06F63F4F2E8571C1D02ACA4976F68C6B" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="33" pageNumber="244">121-125</figureCitation>
,
<figureCitation id="ACE9CB0083AB33E0AB0FE7735B68A6E1" captionStart="Figures 161–164" captionStartId="F24" captionText="Figures 161 - 164. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 161, Cyphocoleus flavipes Yahoue; 162, Cyphocoleus bourailensis, Bourail; 163, Cyphocoleus subulatus, Mt. Koghi; 124, Cyphocoleus iledespinsensis, Ile des Pins." figureDoi="10.3897/dez.63.10241.figures161-164" httpUri="https://binary.pensoft.net/fig/111563" pageId="33" pageNumber="244">161-164</figureCitation>
), this species stands out due to the narrowly orbicular pronotum, MPW/PL = 0.88-0.94, and pronotal disc that is evenly traversed by wrinkles but without longitudinal parasagittal impressions. The pronotal lateral marginal beads arcuately extend to the narrow median base (Fig.
<figureCitation id="13B50C5DE8B63E4CA3C9F8D0F2111AE2" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="33" pageNumber="244">121</figureCitation>
). Some individuals of
<taxonomicName id="6BF88267F1E3F69E90CEEAC7450D4AD9" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus latipennis" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="F5CF7C5C7821D568DF0A439CF7A13E2A" italics="true" pageId="33" pageNumber="244">Cyphocoleus latipennis</emphasis>
</taxonomicName>
have the pronotum nearly as narrow, MPW/PL = 0.92-1.07, but beetles comprising
<taxonomicName id="18DFB23D248B5863F002720EC8D09CD2" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus cychroides" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="cychroides">
<emphasis id="03473BE8C312869AF7B0B7D019733225" italics="true" pageId="33" pageNumber="244">Cyphocoleus cychroides</emphasis>
</taxonomicName>
differ by their lack of supraorbital setae, whereas
<taxonomicName id="DABD1E265EE5FDB1354A13F1D5701E97" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus latipennis" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="B790C733C6E43AD4C987F570987C1EE5" italics="true" pageId="33" pageNumber="244">Cyphocoleus latipennis</emphasis>
</taxonomicName>
is characterized by presence of the posterior seta, though it is located well behind the eye just anterad the constricted neck. The pronotal disc of
<taxonomicName id="DBF45DA707E67739660BB6CE9D9DF31B" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus latipennis" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="621C20BB71971D24C5EA0659F547ED1A" italics="true" pageId="33" pageNumber="244">Cyphocoleus latipennis</emphasis>
</taxonomicName>
also differs by the presence of parasagittal impressions, and median pronotal base not arcuately and continuously margined as in
<taxonomicName id="D265E7C83B94710FE4E8F571402A30F8" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus cychroides" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="cychroides">
<emphasis id="3F39D372E7C357AD26FCFBA97F007669" italics="true" pageId="33" pageNumber="244">Cyphocoleus cychroides</emphasis>
</taxonomicName>
. Standardized body length 6.0-7.2 mm.
</paragraph>
</subSubSection>
<subSubSection id="FB031FC7ADD6F0C76FBFA1668ED5DA23" pageId="33" pageNumber="244" type="description">
<paragraph id="2EA217561A8559072209DE054DAB5471" pageId="33" pageNumber="244">Description</paragraph>
<paragraph id="814E98B4FA9EE5319B1254FD3D080473" pageId="33" pageNumber="244">
(n = 5). Head capsule narrowly ovoid, genae gradually narrowed to moderately constricted neck; eyes only slightly convex, oriented toward upper surface of head, ocular ratio quite variable, 1.20-1.56, about 18 ommatidia across horizontal diameter of eye; frons with paired, longitudinal depressions that surround a low median ridge; frontal grooves broad, deep, parallel, extended from just anterad hind margin of eye onto clypeus; mandibles elongate, length 2.1
<normalizedToken id="9E53135129A5FB177D3E8456E35EFBCE" originalValue="×">x</normalizedToken>
distance from antennal articulatory socket to anterolateral margin of labrum; antennae moderately elongate, scape length 2.75
<normalizedToken id="92B483504AC378D39C7A5CAAF968B00D" originalValue="×">x</normalizedToken>
maximal breadth; gena setose, subgenal seta present. Pronotum with front angles rounded, only slightly protruded; median base continuously margined, but bead straight along base; median longitudinal impression finely incised, slightly irregular as it traverses transverse wrinkles; anterior transverse impression a deep, oblique depression, defining a flat anterior callosity with indistinct longitudinal wrinkles; proepisternum slightly extended beyond marginal bead, narrowly visible in dorsal view; proepisternum mostly smooth, with indistinct vertical wrinkles; prosternal process broadly, medially depressed anteriorly, with paired, shallow longitudinal depressions ventrally, posterior face broadly convex. Elytra broadly cordate, basal groove sinuously incurved near depressed scutellum, continued in broad curve laterally around humerus, a slight hitch in the curve near base of third stria; discal intervals moderately convex, all intervals of subequal convexity; elytral striae deep, punctate, the punctures larger basally where they expand strial breadth; parascutellar seta present, articulatory socket papillate; dorsal elytral setae absent; lateral elytral setae arranged as 6 + 7, the articulatory sockets papillate; both subapical and apical elytral setae present, the articulatory socket for apical seta smaller; subapical sinuation well developed, lateral portion of concavity joined to depressed lateral margin in broad curve; elytral apices conjoined, no invagination of apical margin at fused suture. Mesepisternum impunctate; metepisternal dorsal length 1.1
<normalizedToken id="454307C71E116DB7127B288D6D2D8020" originalValue="×">x</normalizedToken>
diagonal width; metasternum with broad, shallow median fossa at base of metasternal process. Abdomen with apical margin of apical ventrite shallowly concave medially in both males and females; males with one seta each side of apical margin of apical ventrite, females with two setae each side. Legs moderately elongate; metacoxae bisetose; mt1 length/tibial length = 0.27; mt4 length to apex of outer lobe 1.6
<normalizedToken id="C3C6CE669022B88BAAB681EB179C716A" originalValue="×">x</normalizedToken>
median tarsomere length, 3-4 ventrolateral setae each side. Microsculpture on vertex stretched isodiametric mesh, the head capsule glossy; pronotal disc with evident transverse mesh visible within transverse wrinkles, the surface matte; elytral intervals covered with transverse-line microsculpture resulting in silvery iridescence; pelage little developed, glossy head capsule glabrous, pronotum and elytra with sparse distribution of exceedingly short microsetae, visible only in lateral view; ventrites and legs glabrous except for macrosetae. Coloration of body dorsum piceous, mouthparts, scutellum, and apical elytral margin rufopiceous; body venter including elytral epipleura rufopiceous; femora dark rufous; tibiae piceous medially, rufoflavous at base and apex; tarsi rufoflavous.
</paragraph>
<paragraph id="048EC3445F7676321ABE6C0FC030F225" pageId="33" pageNumber="244">
<emphasis id="955375B19A5C3D125D244F2B1C5DB704" italics="true" pageId="33" pageNumber="244">Male genitalia</emphasis>
(n = 10). Male aedeagal median lobe robust, broadly parallel sided over basal 2/3 of length, the lobe shaft angled ventrally ventral ostial opening, dorsoventral breadth at midlength 0.20-0.25
<normalizedToken id="298990F9EA57530ED3EF1E77589C7D19" originalValue="×">x</normalizedToken>
distance from tip to base of closed basal bulb (Figs
<figureCitation id="CC211C3202633A0FDC746DF8C71B3666" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">91-98</figureCitation>
); lobe apex sinuously curved, tip may be knob-like with a constriction distad ostial opening (Figs
<figureCitation id="C31303F10A0F0C86B0588ACED6195AC2" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">91</figureCitation>
,
<figureCitation id="6B2F94FF28AA7959A9810B464E5DBD40" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">92</figureCitation>
) or abruptly narrowed distad ostial opening to a parallel-sided apex (Figs
<figureCitation id="26B7BBB9113C6695C81F55CD2AF2C9EA" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">93-98</figureCitation>
); internal sac membranous without melanized microtrichia, length slightly greater than breadth (Figs
<figureCitation id="A26F599B2F6C4C89745E4AE9157E084F" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">91</figureCitation>
,
<figureCitation id="7406CCCA68CB4593F0395413F555AD8A" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">93</figureCitation>
,
<figureCitation id="D8137CECAF50F748522E1554ABF981E3" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">94</figureCitation>
,
<figureCitation id="065EE30607FD88D4F8B58A0C9019718E" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="33" pageNumber="244">96</figureCitation>
). Although the male aedeagi of individuals assigned to
<taxonomicName id="22319845E89204B4E7BB338D82899C67" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus globulicollis" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="globulicollis">
<emphasis id="F760B17246F7BBA4CE7C38917A7FD539" italics="true" pageId="33" pageNumber="244">Cyphocoleus globulicollis</emphasis>
</taxonomicName>
exhibit substantial variation in the configuration of the apex, all of this variation is exhibited among individuals from the same well-sampled site;
<normalizedToken id="D0880725529A45BC4D88A2984F582D5F" originalValue="Mandjélia">Mandjelia</normalizedToken>
. All individuals from this site share other external attributes, as well as the subangulate apex to the aedeagal median lobe, and so this variation is interpreted as infraspecific.
</paragraph>
<paragraph id="92403C8E4AF51FB64851CBAC7A1607BD" pageId="33" pageNumber="244">
<emphasis id="010F3C730403C94FBB75EE44A02033CB" italics="true" pageId="33" pageNumber="244">Female reproductive tract</emphasis>
(n = 2). Bursa copulatrix vase-shaped, abruptly narrowed distally near spermatheca, distance from base of gonocoxites to spermathecal basal sclerite 3
<normalizedToken id="9B0D2E3AB1A74F01CA41E30581F9A88E" originalValue="×">x</normalizedToken>
maximum breadth (dissection compressed under cover slip) (Fig.
<figureCitation id="F47D8D5EA1399250A4AB378096338731" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="33" pageNumber="244">107</figureCitation>
); bursal walls thin, translucent, lumenal surface smooth; basal gonocoxite with apical fringe of five to six setae (Fig.
<figureCitation id="D10A6246BBE298B5FF9C6CD7E4497CA0" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="33" pageNumber="244">114</figureCitation>
); apical gonocoxite broadly triangular with pointed apex and two lateral ensiform setae.
</paragraph>
</subSubSection>
<subSubSection id="700AEFFB76702D973D8AD9E3BD2974F9" pageId="33" pageNumber="244" type="types">
<paragraph id="31FA04FC09639E68451C4EAF80029D33" pageId="33" pageNumber="244">Types.</paragraph>
<paragraph id="7944BA9271F2B831F9641C2D29845C1B" pageId="33" pageNumber="244">
Holotype male (MNHN): Ex
<normalizedToken id="4DA865292634D40BCD6229C1FD39C7D0" originalValue="Musæo">Musaeo</normalizedToken>
/ Chaudoir (red-inked white label with red border) // HOLOTYPE /
<taxonomicName id="7895AF4F83121AFC9E4626BAED2E6C8E" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="47223BEB607F97C6AAFDCE5D709CEE59" italics="true" pageId="33" pageNumber="244">Cyphocoleus</emphasis>
</taxonomicName>
/
<taxonomicName id="14813D47DB73FDA7833EA3C989D75BDB" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="cychroides" order="Diptera" pageId="33" pageNumber="244" phylum="Arthropoda" rank="species" species="cychroides">
<emphasis id="392FCBEC760FD31B0A7CFEA270F012FA" italics="true" pageId="33" pageNumber="244">cychroides</emphasis>
</taxonomicName>
/
<bibRefCitation id="CE6DF3DF6CC336E4F31763B2C80B41AA" author="Chaudoir, M de" journalOrPublisher="Bulletin de la Societe Imperiale des naturalistes de Moscou" pageId="48" pageNumber="259" pagination="1 - 61" refId="B12" refString="Chaudoir, M de, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des naturalistes de Moscou 52: 1 - 61" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
(black-bordered red label). Holotype status is based on
<bibRefCitation id="8551F280182A3125F8CEB7CA9E422B15" author="Chaudoir, M de" journalOrPublisher="Bulletin de la Societe Imperiale des naturalistes de Moscou" pageId="48" pageNumber="259" pagination="1 - 61" refId="B12" refString="Chaudoir, M de, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des naturalistes de Moscou 52: 1 - 61" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">
<normalizedToken id="3DC151E0F16F9A87D7AA3E61DE621472" originalValue="Chaudoir’s">Chaudoir's</normalizedToken>
(1877
</bibRefCitation>
: 197) statement: &quot;Il
<normalizedToken id="07861519C5CCFC0A00EA0097E0ED189A" originalValue="m’a">m'a</normalizedToken>
<normalizedToken id="5A0FE42AEBFDCE6EFF28AF87FD7DA071" originalValue="été">ete</normalizedToken>
vendu par H.M. Deyrolle.&quot;
<normalizedToken id="8DB04E3954831E3CAD8753C713A428F8" originalValue="Mandjélia">Mandjelia</normalizedToken>
summit, 750 m el. hereby designated type locality.
</paragraph>
</subSubSection>
<subSubSection id="45B6C023B53EA1E04CD56C51EB0B5412" pageId="33" pageNumber="244" type="distribution">
<paragraph id="85C9F1451F893C854603E7FBB17B20A4" pageId="33" pageNumber="244">Distribution and habitat.</paragraph>
<paragraph id="7C0B1C4A8851F5DB0C9CBAE3BB6B9902" pageId="33" pageNumber="244">
Known from Province Nord. The recorded distribution is bounded by Canala on the south, and Col
<normalizedToken id="BC97FD85C78EC199C1979D41E3DB7AF3" originalValue="d’Amoss">d'Amoss</normalizedToken>
along the northeastern coast (Fig.
<figureCitation id="6AF60081C29BDC00E3D8B192324D2404" captionStart="Figures 137–140" captionStartId="F22" captionText="Figures 137 - 140. Distribution maps of Cyphocoleus spp.: 137, Cyphocoleus cychroides, Cyphocoleus cordatus, Cyphocoleus cardiopterus; 138, Cyphocoleus latipennis; 139, Cyphocoleus globulicollis; 140 Cyphocoleus flavipes, Cyphocoleus bourailensis." figureDoi="10.3897/dez.63.10241.figures137-140" httpUri="https://binary.pensoft.net/fig/111561" pageId="33" pageNumber="244">137</figureCitation>
, Suppl. material 3). Beetles have been collected from logs and trunks via the application of pyrethrin fog.
</paragraph>
</subSubSection>
</treatment>
</document>
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