treatments-xml/data/6F/E3/87/6FE387255BF05CD483C01A0433C6E0DE.xml

<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.36.55090" ID-GBIF-Dataset="b552a045-8720-4dab-af04-b73db6a29873" ID-GBIF-Taxon="169091746" ID-Pensoft-Pub="1314-2615-36-35" ID-Pensoft-UUID="AA0514E353C65B57B08EF57BD89697C8" ID-ZooBank="E876880C80844882BA3B1AF7396FB4F2" LSID="urn:lsid:zoobank.org:act:B7836D85-CC52-4284-A34E-F1BA66BB8BC0" ModsDocID="1314-2615-36-35" checkinTime="1605571741355" checkinUser="pensoft" docAuthor="Nunez 1, Jorge, Glasby 2, Christopher J. &amp; Naranjo 3, Manuel" docDate="2020" docId="6FE387255BF05CD483C01A0433C6E0DE" docLanguage="en" docName="SubterBiol 36: 35-49" docOrigin="Subterranean Biology 36" docSource="http://dx.doi.org/10.3897/subtbiol.36.55090" docTitle="Namanereis canariarum Núñez 1 &amp; Glasby 2 &amp; Naranjo 3 2020, sp. nov." docType="treatment" docUuid="B7836D85-CC52-4284-A34E-F1BA66BB8BC0" docUuidSource="ZooBank" docVersion="3" id="AA0514E353C65B57B08EF57BD89697C8" lastPageNumber="35" masterDocId="AA0514E353C65B57B08EF57BD89697C8" masterDocTitle="Groundwater annelids from Gran Canaria and Fuerteventura (Canary Islands), with the description of two new species of Namanereis (Namanereidinae, Nereididae, Polychaeta)" masterLastPageNumber="49" masterPageNumber="35" pageId="0" pageNumber="35" updateTime="1643441998320" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Groundwater annelids from Gran Canaria and Fuerteventura (Canary Islands), with the description of two new species of Namanereis (Namanereidinae, Nereididae, Polychaeta)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nunez 1, Jorge</mods:namePart>
<mods:affiliation>Laboratorio de Bentos, Departamento de Biologia Animal, Edafologia y Geologia, Facultad de Ciencias, Universidad de La Laguna, 38200 La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glasby 2, Christopher J.</mods:namePart>
<mods:affiliation>Laboratorio de Bentos, Departamento de Biologia Animal, Edafologia y Geologia, Facultad de Ciencias, Universidad de La Laguna, 38200 La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naranjo 3, Manuel</mods:namePart>
<mods:affiliation>Laboratorio de Bentos, Departamento de Biologia Animal, Edafologia y Geologia, Facultad de Ciencias, Universidad de La Laguna, 38200 La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>36</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>35</mods:start>
<mods:end>49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/subtbiol.36.55090</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.36.55090</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2615-36-35</mods:identifier>
<mods:identifier type="ZooBank">E876880C80844882BA3B1AF7396FB4F2</mods:identifier>
<mods:identifier type="Pensoft-UUID">AA0514E353C65B57B08EF57BD89697C8</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="169091746" LSID="urn:lsid:zoobank.org:act:B7836D85-CC52-4284-A34E-F1BA66BB8BC0" httpUri="http://treatment.plazi.org/id/6FE387255BF05CD483C01A0433C6E0DE" lastPageNumber="35" pageId="0" pageNumber="35">
<subSubSection pageId="0" pageNumber="35" type="nomenclature">
<paragraph pageId="0" pageNumber="35">
<taxonomicName LSID="6FE38725-5BF0-5CD4-83C0-1A0433C6E0DE" authority="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3, 2020" authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum" status="sp. nov.">Namanereis canariarum</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. a Distribution in Canary Islands of Namanereis canariarum sp. nov. (yellow dot) and Namanereis llanetensis (red dot) b Namanereis canariarum sp. nov. live specimen, dorsal view c Namanereis llanetensis sp. nov. live specimen, dorsal view d Los Llanetes water mine (Valsequillo, Gran Canaria) type locality of N. llanetensis sp. nov." figureDoi="10.3897/subtbiol.36.55090.figure1" httpUri="https://binary.pensoft.net/fig/476122" pageId="0" pageNumber="35">Figs 1a, b</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">; 2</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namanereis canariarum sp. nov. a jaw pieces of the pharynx, ventral view b parapodium, detail of the acicular lobes c pseudospiniger blade tip d subneuroacicular heterogomph falciger, chaetiger 60. Namanereis llanetensis sp. nov. e Holotype, detail of the basal spinulation, hererogomph spiniger, chaetiger 10 f Holotype, detail of spinulation, subneuroacicular heterogomph falciger, chaetiger 10." figureDoi="10.3897/subtbiol.36.55090.figure4" httpUri="https://binary.pensoft.net/fig/476125" pageId="0" pageNumber="35">; 4a-d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="reference_group">
<paragraph pageId="0" pageNumber="35">
<taxonomicName baseAuthorityName="Augener" baseAuthorityYear="1933" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis hummelincki" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="hummelincki">Namanereis hummelincki</taxonomicName>
.- 
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
, 1988: 181-182. 
<emphasis italics="true" pageId="0" pageNumber="35">Non</emphasis>
Augener, 1933. 
<taxonomicName authorityName="Chamberlin" authorityYear="1919" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="genus">Namanereis</taxonomicName>
sp. nov. 1 Glasby, Fiege &amp; Van Damme, 2014: 28 (Table 1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="type locality">
<paragraph pageId="0" pageNumber="35">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="35">
Canary Islands, Fuerteventura, Las Playitas (
<geoCoordinate degrees="28" direction="north" minutes="13" orientation="latitude" precision="1" seconds="55.42" value="28.232061">28°13'55.42&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="west" minutes="59" orientation="longitude" precision="1" seconds="4.92" value="-13.9847">13°59'4.92&quot;W</geoCoordinate>
) 14 m above sea level, brackish water well.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="type material">
<paragraph pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
: TFMCBM-AN/240, coll. J. 
<normalizedToken originalValue="Núñez">Nunez</normalizedToken>
, 8-2-2002; 2 
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Paratypes</emphasis>
: TFMCBM-AN/241, 2 spec., Fuerteventura, Las Playitas, Canary Islands, 
<geoCoordinate degrees="28" direction="north" minutes="13" orientation="latitude" precision="1" seconds="56.73" value="28.232426">28°13'56.73&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="13" direction="west" minutes="59" orientation="longitude" precision="1" seconds="10.26" value="-13.986183">13°59'10.26&quot;W</geoCoordinate>
, 18 m above sea level, coll. L. Santos &amp; J.R. Docoito, 28-9-2011.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="materials_examined">
<paragraph pageId="0" pageNumber="35">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="35">
Fataga water mine (Barranco de Fataga, Gran Canaria, (
<geoCoordinate degrees="27" direction="north" minutes="47" orientation="latitude" precision="1" seconds="59.18" value="27.799772">27°47'59.18&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="15" direction="west" minutes="35" orientation="longitude" precision="1" seconds="12.57" value="-15.586824">15°35'12.57&quot;W</geoCoordinate>
, 160 m above sea level), 12 spec. (6 spec. TFMCBM-AN/242; 6 spec. DBAULL/2020), coll. P. 
<normalizedToken originalValue="Oromí">Oromi</normalizedToken>
&amp; M. Naranjo, 6-12-2013.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="materials_examined">
<paragraph pageId="0" pageNumber="35">Comparative material examined.</paragraph>
<paragraph pageId="0" pageNumber="35">
Material identified by G. 
<bibRefCitation author="Hartmann-Schroeder, G" journalOrPublisher="Bulletin Zooelogisch Museum Universiteit Van Amsterdam" pageId="0" pageNumber="35" pagination="177 - 184" refId="B11" refString="Hartmann-Schroeder, G, 1988. Stygofauna of the Canary Islands, 13 die Polychaeten der sammel reisen 1985 und 1987. Bulletin Zooelogisch Museum Universiteit Van Amsterdam 11 (22): 177 - 184" title="Stygofauna of the Canary Islands, 13 die Polychaeten der sammel reisen 1985 und 1987." volume="11" year="1988">
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
(1988)
</bibRefCitation>
as 
<taxonomicName baseAuthorityName="Augener" baseAuthorityYear="1933" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis hummelincki" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis hummelincki</emphasis>
</taxonomicName>
from Canary Islands, Fuerteventura, well S. of Betancuria, 400 m a.s.l., coll. J.H. Stock, 4-5-1987, 8 ex ZMA VPol 2953 (now in Naturalis, Leiden).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="description">
<paragraph pageId="0" pageNumber="35">Description.</paragraph>
<paragraph pageId="0" pageNumber="35">Holotype with 133 chaetigers, 46 mm long and 0.7 mm wide excluding parapodia and chaetae. Two paratypes, fragmented, one with 77 and the other with 88 chaetigers; one paratype female with gametes in the coelom.</paragraph>
<paragraph pageId="0" pageNumber="35">
Living individuals with depigmented epidermis, highlighting the red dorsal blood vessel over the fleshy pink coloured body. Body slender, with uniform width in the anterior and middle regions, abruptly tapering only near the posterior end; trunk convex dorsally and flattened ventrally (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. a Distribution in Canary Islands of Namanereis canariarum sp. nov. (yellow dot) and Namanereis llanetensis (red dot) b Namanereis canariarum sp. nov. live specimen, dorsal view c Namanereis llanetensis sp. nov. live specimen, dorsal view d Los Llanetes water mine (Valsequillo, Gran Canaria) type locality of N. llanetensis sp. nov." figureDoi="10.3897/subtbiol.36.55090.figure1" httpUri="https://binary.pensoft.net/fig/476122" pageId="0" pageNumber="35">1b</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="35">
Prostomium semi-circular in shape, slightly wider than long; cleft absent, shallow dorsal hollow present; two short conical frontal antennae, smooth, extending beyond tip of palpophore; palps broad and globose, biarticulated, with globular palpostyles; eyes absent (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2a</figureCitation>
). Tentacular segment as wide as first chaetiger, but half its length, bearing three pairs of short tentacular cirri, cirrophores inconspicuous, cirrostyles smooth and tapering. The longest pair of cirrostyles extends to reach the third chaetiger (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2a</figureCitation>
). Pharynx divided into a maxillary and an oral region, lacking paragnaths or papillae, but possessing a pair of delta-shaped jaws, dark, with only bifid terminal teeth (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2a, b</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namanereis canariarum sp. nov. a jaw pieces of the pharynx, ventral view b parapodium, detail of the acicular lobes c pseudospiniger blade tip d subneuroacicular heterogomph falciger, chaetiger 60. Namanereis llanetensis sp. nov. e Holotype, detail of the basal spinulation, hererogomph spiniger, chaetiger 10 f Holotype, detail of spinulation, subneuroacicular heterogomph falciger, chaetiger 10." figureDoi="10.3897/subtbiol.36.55090.figure4" httpUri="https://binary.pensoft.net/fig/476125" pageId="0" pageNumber="35">4a</figureCitation>
).
</paragraph>
<caption doi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figure 2.</emphasis>
<taxonomicName authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis canariarum</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="35">a</emphasis>
anterior end, everted pharynx, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="35">b</emphasis>
jaw pieces of the pharynx, frontal view 
<emphasis bold="true" pageId="0" pageNumber="35">c</emphasis>
parapodium from chaetiger 10 
<emphasis bold="true" pageId="0" pageNumber="35">d</emphasis>
parapodium from chaetiger 60 
<emphasis bold="true" pageId="0" pageNumber="35">e</emphasis>
supraneuroacicular falciger, chaetiger 30 
<emphasis bold="true" pageId="0" pageNumber="35">f</emphasis>
subneuroacicular falciger, chaetiger 10 
<emphasis bold="true" pageId="0" pageNumber="35">g</emphasis>
subneuroacicular pseudospiniger, chaetiger 30 
<emphasis bold="true" pageId="0" pageNumber="35">h</emphasis>
supraneuroacicular spiniger, blade not fully shown, chaetiger 10 
<emphasis bold="true" pageId="0" pageNumber="35">i</emphasis>
supraneuroacicular spiniger, blade fully shown, chaetiger 10 
<emphasis bold="true" pageId="0" pageNumber="35">j</emphasis>
subneuroacicular pseudospiniger, chaetiger 10 
<emphasis bold="true" pageId="0" pageNumber="35">k</emphasis>
supraneuroacicular falciger, chaetiger 30 
<emphasis bold="true" pageId="0" pageNumber="35">l</emphasis>
subneuroacicular falciger, chaetiger 30.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="35">
Parapodia with notopodial branch reduced to the inner acicula; neuropodium well-developed, acicular neuropodial ligule subconical (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2c, d</figureCitation>
) with prominent acicular lobes (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namanereis canariarum sp. nov. a jaw pieces of the pharynx, ventral view b parapodium, detail of the acicular lobes c pseudospiniger blade tip d subneuroacicular heterogomph falciger, chaetiger 60. Namanereis llanetensis sp. nov. e Holotype, detail of the basal spinulation, hererogomph spiniger, chaetiger 10 f Holotype, detail of spinulation, subneuroacicular heterogomph falciger, chaetiger 10." figureDoi="10.3897/subtbiol.36.55090.figure4" httpUri="https://binary.pensoft.net/fig/476125" pageId="0" pageNumber="35">4b</figureCitation>
). Dorsal cirri acuminated, similar in length throughout the body and clearly surpassing the parapodial lobe; ventral cirri short and thin, being about a third of the length of the dorsal cirri (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2c, d</figureCitation>
). Notochaetae absent, neurochaetae arrangement as in Type A (
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.25.1999.1354" author="Glasby, CJ" journalOrPublisher="Records of the Australian Museum, Supplement" pageId="0" pageNumber="35" pagination="1 - 146" refId="B6" refString="Glasby, CJ, 1999. The Namanereidinae (Polychaeta: Nereididae) Part 1 Taxonomy and Phylogeny, Part 2 Cladistic Biogeography. Records of the Australian Museum, Supplement 25: 1 - 146, DOI: https://doi.org/10.3853/j.0812-7387.25.1999.1354" title="The Namanereidinae (Polychaeta: Nereididae) Part 1 Taxonomy and Phylogeny, Part 2 Cladistic Biogeography." url="https://doi.org/10.3853/j.0812-7387.25.1999.1354" volume="25" year="1999">Glasby 1999</bibRefCitation>
), although with slight modifications. Overall, less than 7 chaetae on each parapodium; supraneuroacicular chaetae normally includes 1 sesquigomph spiniger chaeta in the postacicular fascicle (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2h, i</figureCitation>
) and 1 heterogomph falciger chaeta in the preacicular fascicle throughout the whole body (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2e, k</figureCitation>
). Subneuroacicular chaetae normally includes 1 heterogomph pseudospiniger chaeta (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2g, j</figureCitation>
; 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namanereis canariarum sp. nov. a jaw pieces of the pharynx, ventral view b parapodium, detail of the acicular lobes c pseudospiniger blade tip d subneuroacicular heterogomph falciger, chaetiger 60. Namanereis llanetensis sp. nov. e Holotype, detail of the basal spinulation, hererogomph spiniger, chaetiger 10 f Holotype, detail of spinulation, subneuroacicular heterogomph falciger, chaetiger 10." figureDoi="10.3897/subtbiol.36.55090.figure4" httpUri="https://binary.pensoft.net/fig/476125" pageId="0" pageNumber="35">4c</figureCitation>
) and 2-3 falciger chaetae in the preacicular fascicles (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namanereis canariarum sp. nov. a anterior end, everted pharynx, dorsal view b jaw pieces of the pharynx, frontal view c parapodium from chaetiger 10 d parapodium from chaetiger 60 e supraneuroacicular falciger, chaetiger 30 f subneuroacicular falciger, chaetiger 10 g subneuroacicular pseudospiniger, chaetiger 30 h supraneuroacicular spiniger, blade not fully shown, chaetiger 10 i supraneuroacicular spiniger, blade fully shown, chaetiger 10 j subneuroacicular pseudospiniger, chaetiger 10 k supraneuroacicular falciger, chaetiger 30 l subneuroacicular falciger, chaetiger 30." figureDoi="10.3897/subtbiol.36.55090.figure2" httpUri="https://binary.pensoft.net/fig/476123" pageId="0" pageNumber="35">2f, l</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namanereis canariarum sp. nov. a jaw pieces of the pharynx, ventral view b parapodium, detail of the acicular lobes c pseudospiniger blade tip d subneuroacicular heterogomph falciger, chaetiger 60. Namanereis llanetensis sp. nov. e Holotype, detail of the basal spinulation, hererogomph spiniger, chaetiger 10 f Holotype, detail of spinulation, subneuroacicular heterogomph falciger, chaetiger 10." figureDoi="10.3897/subtbiol.36.55090.figure4" httpUri="https://binary.pensoft.net/fig/476125" pageId="0" pageNumber="35">4d</figureCitation>
); less frequently 2 pseudospiniger chaetae, and exceptionally up to 3 falciger and no pseudospiniger chaetae might be found. In the posterior region, starting from chaetiger 60, pseudospiniger chaetae are generally longer, resembling spiniger chaetae. Supraneuroacicular falciger chaetae in chaetiger 10 with blades 4.90
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of shaft head (4.7-5.0), finely serrated, about 30 teeth, length of the teeth decreasing towards the apex. Subneuroacicular falciger chaetae in chaetiger 10 with blades 5.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than width of shaft head (5.2-5.7), finely serrated (similar to supraneuroacicular falcigers), length of the teeth decreasing towards the apex. Supraneuroacicular sesquigomph spiniger chaetae in chaetiger 10 with boss (vertical length of blade joint opposite collar) of the joint about 1.6
<normalizedToken originalValue="×">x</normalizedToken>
length of collar. Aciculae dark honey coloured. Pygidium with terminal anus, and two short ventrolateral anal cirri, slender and smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="remarks">
<paragraph pageId="0" pageNumber="35">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="35">
Morphologically, 
<taxonomicName authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis canariarum</emphasis>
</taxonomicName>
sp. nov. belongs to the bifid-jaw group of 
<taxonomicName authorityName="Chamberlin" authorityYear="1919" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis</emphasis>
</taxonomicName>
: this includes 
<taxonomicName genus="N." lsidName="N. araps" pageId="0" pageNumber="35" rank="species" species="araps">
<emphasis italics="true" pageId="0" pageNumber="35">N. araps</emphasis>
</taxonomicName>
Glasby, 1997 (Arabian Peninsula), 
<taxonomicName genus="N." lsidName="N. cavernicola" pageId="0" pageNumber="35" rank="species" species="cavernicola">
<emphasis italics="true" pageId="0" pageNumber="35">N. cavernicola</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Solís-Weiss">Solis-Weiss</normalizedToken>
&amp; Espinasa 1991) (Mexico, Caribbean side), 
<taxonomicName genus="N." lsidName="N. christopheri" pageId="0" pageNumber="35" rank="species" species="christopheri">
<emphasis italics="true" pageId="0" pageNumber="35">N. christopheri</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/subtbiol.23.13701" author="Conde-Vela, VM" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="35" pagination="19 - 46" refId="B3" refString="Conde-Vela, VM, 2017. The troglomorphic adaptations of Namanereidinae (Annelida, Nereididae) revisited, including a redescription of Namanereis cavernicola (Solis-Weiss &amp; Espinasa, 1991), and a new Caribbean species of Namanereis Chamberlin, 1919. Subterranean Biology 23: 19 - 46, DOI: https://doi.org/10.3897/subtbiol.23.13701" title="The troglomorphic adaptations of Namanereidinae (Annelida, Nereididae) revisited, including a redescription of Namanereis cavernicola (Solis-Weiss &amp; Espinasa, 1991), and a new Caribbean species of Namanereis Chamberlin, 1919." url="https://doi.org/10.3897/subtbiol.23.13701" volume="23" year="2017">Conde-Vela 2017</bibRefCitation>
(Saint Vincent, Caribbean), 
<taxonomicName genus="N." lsidName="N. hummelincki" pageId="0" pageNumber="35" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">N. hummelincki</emphasis>
</taxonomicName>
(Augener 1933) (Caribbean), 
<taxonomicName genus="N." lsidName="N. minuta" pageId="0" pageNumber="35" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="35">N. minuta</emphasis>
</taxonomicName>
Glasby, 1999 (Caribbean), 
<taxonomicName genus="N." lsidName="N. llanetensis" pageId="0" pageNumber="35" rank="species" species="llanetensis">
<emphasis italics="true" pageId="0" pageNumber="35">N. llanetensis</emphasis>
</taxonomicName>
sp. nov. (Canary Islands), 
<taxonomicName genus="N." lsidName="N. serratis" pageId="0" pageNumber="35" rank="species" species="serratis">
<emphasis italics="true" pageId="0" pageNumber="35">N. serratis</emphasis>
</taxonomicName>
Glasby, 1999 (Caribbean), 
<taxonomicName genus="N." lsidName="N. socotrensis" pageId="0" pageNumber="35" rank="species" species="socotrensis">
<emphasis italics="true" pageId="0" pageNumber="35">N. socotrensis</emphasis>
</taxonomicName>
Glasby, Fiege &amp; Van Damme, 2014 (Socotra Archipelago) and 
<taxonomicName genus="N." lsidName="N. stocki" pageId="0" pageNumber="35" rank="species" species="stocki">
<emphasis italics="true" pageId="0" pageNumber="35">N. stocki</emphasis>
</taxonomicName>
Glasby, 1999 (Caribbean). The new species can be distinguished from 
<taxonomicName genus="N." lsidName="N. cavernicola" pageId="0" pageNumber="35" rank="species" species="cavernicola">
<emphasis italics="true" pageId="0" pageNumber="35">N. cavernicola</emphasis>
</taxonomicName>
, 
<taxonomicName genus="N." lsidName="N. minuta" pageId="0" pageNumber="35" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="35">N. minuta</emphasis>
</taxonomicName>
and 
<taxonomicName genus="N." lsidName="N. stocki" pageId="0" pageNumber="35" rank="species" species="stocki">
<emphasis italics="true" pageId="0" pageNumber="35">N. stocki</emphasis>
</taxonomicName>
by the number and size of teeth on the blades of the supraneuroacicular falciger chaetae. 
<taxonomicName authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis canariarum</emphasis>
</taxonomicName>
has 20-30 moderate-sized teeth, compared to 35-80 very fine teeth in 
<taxonomicName genus="N." lsidName="N. cavernicola" pageId="0" pageNumber="35" rank="species" species="cavernicola">
<emphasis italics="true" pageId="0" pageNumber="35">N. cavernicola</emphasis>
</taxonomicName>
, 50-60 fine teeth in 
<taxonomicName genus="N." lsidName="N. minuta" pageId="0" pageNumber="35" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="35">N. minuta</emphasis>
</taxonomicName>
, and 9-14 moderate-sized teeth in 
<taxonomicName genus="N." lsidName="N. stocki" pageId="0" pageNumber="35" rank="species" species="stocki">
<emphasis italics="true" pageId="0" pageNumber="35">N. stocki</emphasis>
</taxonomicName>
. Furthermore, 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
has much shorter antennae and tentacular cirri than the three aforementioned species. On the other hand, 
<taxonomicName genus="N." lsidName="N. minuta" pageId="0" pageNumber="35" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="35">N. minuta</emphasis>
</taxonomicName>
and 
<taxonomicName genus="N." lsidName="N. stocki" pageId="0" pageNumber="35" rank="species" species="stocki">
<emphasis italics="true" pageId="0" pageNumber="35">N. stocki</emphasis>
</taxonomicName>
differ from 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
in having four pairs of tentacular cirri. 
<taxonomicName authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis canariarum</emphasis>
</taxonomicName>
has shorter tentacular cirri and smooth antennae, unlike 
<taxonomicName genus="N." lsidName="N. araps" pageId="0" pageNumber="35" rank="species" species="araps">
<emphasis italics="true" pageId="0" pageNumber="35">N. araps</emphasis>
</taxonomicName>
that has faintly jointed and longer tentacular cirri, with the posterodorsal one extending back to chaetiger 5; antennae in 
<taxonomicName genus="N." lsidName="N. araps" pageId="0" pageNumber="35" rank="species" species="araps">
<emphasis italics="true" pageId="0" pageNumber="35">N. araps</emphasis>
</taxonomicName>
slightly exceed the length of the prostomium. 
<taxonomicName authorityName="Núñez 1 &amp; Glasby 2 &amp; Naranjo 3" authorityYear="2020" class="Polychaeta" family="Nereididae" genus="Namanereis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Namanereis canariarum" order="Phyllodocida" pageId="0" pageNumber="35" phylum="Annelida" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">Namanereis canariarum</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName genus="N." lsidName="N. hummelincki" pageId="0" pageNumber="35" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">N. hummelincki</emphasis>
</taxonomicName>
, 
<taxonomicName genus="N." lsidName="N. christopheri" pageId="0" pageNumber="35" rank="species" species="christopheri">
<emphasis italics="true" pageId="0" pageNumber="35">N. christopheri</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="N." lsidName="N. llanetensis" pageId="0" pageNumber="35" rank="species" species="llanetensis">
<emphasis italics="true" pageId="0" pageNumber="35">N. llanetensis</emphasis>
</taxonomicName>
sp. nov. However, 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
differs from (1) 
<taxonomicName genus="N." lsidName="N. llanetensis" pageId="0" pageNumber="35" rank="species" species="llanetensis">
<emphasis italics="true" pageId="0" pageNumber="35">N. llanetensis</emphasis>
</taxonomicName>
sp. nov. in the longer tentacular and parapodial dorsal cirri, as well as the absence of pseudospiniger chaetae; (2) from 
<taxonomicName genus="N." lsidName="N. christopheri" pageId="0" pageNumber="35" rank="species" species="christopheri">
<emphasis italics="true" pageId="0" pageNumber="35">N. christopheri</emphasis>
</taxonomicName>
in the absence of long pseudospiniger chaetae, as well as the presence of shorter and most uniform spinulation in the falciger chaetae, with a longer blade with a greater number of denticles than in 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
; (3) and from 
<taxonomicName genus="N." lsidName="N. hummelincki" pageId="0" pageNumber="35" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">N. hummelincki</emphasis>
</taxonomicName>
by the shape of the jaws, in 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
they are delta-shaped in the toothless area, and 
<taxonomicName genus="N." lsidName="N. hummelincki" pageId="0" pageNumber="35" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">N. hummelincki</emphasis>
</taxonomicName>
is oblong in shape; as well as the greater number of pseudospiniger chaetae per parapodium in 
<taxonomicName genus="N." lsidName="N. hummelincki" pageId="0" pageNumber="35" rank="species" species="hummelincki">
<emphasis italics="true" pageId="0" pageNumber="35">N. hummelincki</emphasis>
</taxonomicName>
. Finally, 
<taxonomicName genus="N." lsidName="N. canariarum" pageId="0" pageNumber="35" rank="species" species="canariarum">
<emphasis italics="true" pageId="0" pageNumber="35">N. canariarum</emphasis>
</taxonomicName>
differs from 
<taxonomicName genus="N." lsidName="N. serratis" pageId="0" pageNumber="35" rank="species" species="serratis">
<emphasis italics="true" pageId="0" pageNumber="35">N. serratis</emphasis>
</taxonomicName>
and 
<taxonomicName genus="N." lsidName="N. socotrensis" pageId="0" pageNumber="35" rank="species" species="socotrensis">
<emphasis italics="true" pageId="0" pageNumber="35">N. socotrensis</emphasis>
</taxonomicName>
because these two species only have heterogomph falcigers setae in subneuroacicular fascicle.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="habitat">
<paragraph pageId="0" pageNumber="35">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="35">
The type material was collected from a brackish water well at 14 metres above sea level and about 400 metres from the coastline, located on the south of Fuerteventura. Additional material comes from the southern sector of Gran Canaria, collected in a water mine about 140 m a.s.l.. The mine had waterlogged sections rich in plant roots and slime. The stygobiont amphipod 
<taxonomicName authorityName="Stock" authorityYear="1988" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoniphargus fontinalis" order="Amphipoda" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="fontinalis">
<emphasis italics="true" pageId="0" pageNumber="35">Pseudoniphargus fontinalis</emphasis>
</taxonomicName>
and the diving beetle 
<taxonomicName baseAuthorityName="Germar" baseAuthorityYear="1824" class="Insecta" family="Dytiscidae" genus="Bidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bidessus minutissimus" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis italics="true" pageId="0" pageNumber="35">Bidessus minutissimus</emphasis>
</taxonomicName>
(Naranjo &amp; 
<normalizedToken originalValue="Martín">Martin</normalizedToken>
2016) were also found in this locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="distribution">
<paragraph pageId="0" pageNumber="35">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="35">Type locality: Canary Islands, Fuerteventura, Las Playitas. Other specimens from Fataga water mine, Barranco de Fataga, Gran Canaria.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="etymology">
<paragraph pageId="0" pageNumber="35">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="35">The species name derives from the Canary archipelago, in plural for being found in two islands.</paragraph>
</subSubSection>
</treatment>
</document>