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<document id="9BDDDD43A3C58F42CF8DD629636714D4" ID-DOI="10.1649/0010-065X-73.1.127" ID-ISSN="1938-4394" ID-Zenodo-Dep="5333118" ID-ZooBank="urn:lsid:zoobank.org:pub:A5E82F63-4AC3-41BD-B1D7-78430C22A742" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173649495" checkinUser="valdenar" docAuthor="López-García, Margarita M. &amp; Deloya, Cuauhtémoc" docDate="2019" docId="6F6ED021D62FFFC9FF7DFEF47F4BFC43" docLanguage="en" docName="0010-065X-73.1.127.pdf" docOrigin="The Coleopterists Bulletin 73 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-73.1.127" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Tomarus paranaensis López-García &amp; Deloya, 2019, new species" docType="treatment" docVersion="4" lastPageNumber="133" masterDocId="9357A859D628FFCEFFD0FFE87C4AFFFC" masterDocTitle="Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species" masterLastPageNumber="141" masterPageNumber="127" pageNumber="133" updateTime="1699174053468" updateUser="plazi">
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<mods:title id="2F8E20843A0C625C2A4EBD4D46F1B1EE">Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species</mods:title>
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<mods:namePart id="4FD48048DC525525F18C1D0D4292E25C">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="602720983D487374302BE6DF97B45EF7">Deloya, Cuauhtémoc</mods:namePart>
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<mods:title id="CA530ED6D9FDABA8CA35106DD1BCE68F">The Coleopterists Bulletin</mods:title>
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<mods:date id="CEAFADCBA9549689490AEDCE65B4B432">2019</mods:date>
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<treatment id="6F6ED021D62FFFC9FF7DFEF47F4BFC43" ID-DOI="http://doi.org/10.5281/zenodo.5412053" ID-Zenodo-Dep="5412053" LSID="urn:lsid:plazi:treatment:6F6ED021D62FFFC9FF7DFEF47F4BFC43" httpUri="http://treatment.plazi.org/id/6F6ED021D62FFFC9FF7DFEF47F4BFC43" lastPageNumber="133" pageId="7" pageNumber="133">
<subSubSection id="AFDD32BCD62FFFC9FF7DFEF47DAAFE67" pageId="7" pageNumber="133" type="nomenclature">
<paragraph id="E7786137D62FFFC9FF7DFEF47D90FEB0" blockId="7.[173,659,283,412]" pageId="7" pageNumber="133">
<heading id="BC30D65BD62FFFC9FF7DFEF47D90FEB0" centered="true" fontSize="9" level="2" pageId="7" pageNumber="133" reason="2">
<emphasis id="D5B3BD25D62FFFC9FF7DFEF47D90FEB0" bold="true" pageId="7" pageNumber="133">
<taxonomicName id="20C71AB4D62FFFC9FF7DFEF47DC9FECD" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2019" box="[173,387,284,305]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="paranaensis">
<emphasis id="D5B3BD25D62FFFC9FF7DFEF47DC9FECD" bold="true" box="[173,387,284,305]" italics="true" pageId="7" pageNumber="133">Tomarus paranaensis</emphasis>
</taxonomicName>
López-Garc´ ıa and Deloya,
<taxonomicNameLabel id="CE80005ED62FFFC9FEB5FEDF7D90FEB0" box="[357,474,311,332]" pageId="7" pageNumber="133" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E7786137D62FFFC9FF39FEB97DAAFE67" blockId="7.[173,659,283,412]" pageId="7" pageNumber="133">
Zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="93615BE2D62FFFC9FF6FFE847ECBFE7D" box="[191,641,364,385]" pageId="7" pageNumber="133">5FF67FC3-A050-475F-9541-D42E2CCFFA26</uuid>
</uri>
(
<figureCitation id="7FFC7DB2D62FFFC9FEB6FE6E7D93FE67" box="[358,473,390,412]" captionStart="Figs" captionStartId="7.[197,236,1687,1706]" captionTargetBox="[178,1187,1044,1667]" captionTargetId="figure-407@7.[175,1190,1040,1670]" captionTargetPageId="7" captionText="Figs. 1719. Tomarus paranaensis. 17) Habitus; 18) Parameres, frontal view; 19) Parameres, lateral view. Scale bars = 2 mm (Fig. 17), 0.5 mm (Figs. 1819)." figureDoi="http://doi.org/10.5281/zenodo.5333132" httpUri="https://zenodo.org/record/5333132/files/figure.png" pageId="7" pageNumber="133">Figs. 1719</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="AFDD32BCD62FFFC9FF15FE597E65FDE9" pageId="7" pageNumber="133" type="materials_examined">
<paragraph id="E7786137D62FFFC9FF15FE597E65FDE9" blockId="7.[173,660,432,959]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FF15FE597D2BFE3A" bold="true" box="[197,353,433,454]" pageId="7" pageNumber="133">Type Material.</emphasis>
<materialsCitation id="57AF6B6AD62FFFC9FEBEFE597E65FDE9" collectingDate="1965-02-03" collectionCode="USNM" country="Brazil" determinerName="Halik" location="Ponta Grossa" pageId="7" pageNumber="133" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
16.
<typeStatus id="387CDF95D62FFFC9FE4CFE597DBEFE3A" box="[412,500,433,454]" pageId="7" pageNumber="133" type="holotype">Holotype</typeStatus>
labeled “
<location id="E21837ECD62FFFC9FD8DFE597CA4FE1C" LSID="urn:lsid:plazi:treatment:6F6ED021D62FFFC9FF7DFEF47F4BFC43:E21837ECD62FFFC9FD8DFE597CA4FE1C" country="Brazil" name="Ponta Grossa" pageId="7" pageNumber="133" stateProvince="Parana">Ponta Grossa</location>
/
<collectingRegion id="2503AFD5D62FFFC9FF2AFE237D70FE1C" box="[250,314,459,480]" country="Brazil" name="Parana" pageId="7" pageNumber="133">Paraná</collectingRegion>
/
<date id="937947F7D62FFFC9FE9AFE237DD4FE1C" box="[330,414,459,480]" pageId="7" pageNumber="133" value="1965-02-03">
<collectingDate id="833DBE1FD62FFFC9FE9AFE237DD4FE1C" box="[330,414,459,480]" pageId="7" pageNumber="133" value="1965-02-03">3.ii 1965</collectingDate>
</date>
/Moses / 26438 //
<taxonomicName id="20C71AB4D62FFFC9FD99FE237D1CFE07" authority="DeGeer" authorityName="DeGeer" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="gibbosus">Ligyrus gibbosus DeGeer</taxonomicName>
//
<determinerName id="12A48DF3D62FFFC9FEABFE0E7DE5FE07" box="[379,431,486,507]" pageId="7" pageNumber="133">Halik</determinerName>
det. 1965 //
<collectingCountry id="9FD021A7D62FFFC9FDEEFE0E7ED8FE07" box="[574,658,486,507]" name="Brazil" pageId="7" pageNumber="133">BRASIL</collectingCountry>
Halik / 1966 / Collection” (16
<collectionCode id="81D6F9F2D62FFFC9FE0CFDE87E6FFDE9" box="[476,549,512,533]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="133" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AFDD32BCD62FFFC9FF15FDF378EDFE3A" pageId="7" pageNumber="133" type="description">
<paragraph id="E7786137D62FFFC9FF15FDF378EDFE3A" blockId="7.[173,660,432,959]" lastBlockId="7.[707,1194,194,959]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FF15FDF37DE6FDCC" bold="true" box="[197,428,539,560]" pageId="7" pageNumber="133">
Description.
<typeStatus id="387CDF95D62FFFC9FE9AFDF37DE2FDCC" box="[330,424,539,560]" pageId="7" pageNumber="133" type="holotype">Holotype</typeStatus>
.
</emphasis>
Habitus as in
<figureCitation id="7FFC7DB2D62FFFC9FD95FDF37EC5FDCC" box="[581,655,539,560]" captionStart="Figs" captionStartId="7.[197,236,1687,1706]" captionTargetBox="[178,1187,1044,1667]" captionTargetId="figure-407@7.[175,1190,1040,1670]" captionTargetPageId="7" captionText="Figs. 1719. Tomarus paranaensis. 17) Habitus; 18) Parameres, frontal view; 19) Parameres, lateral view. Scale bars = 2 mm (Fig. 17), 0.5 mm (Figs. 1819)." figureDoi="http://doi.org/10.5281/zenodo.5333132" httpUri="https://zenodo.org/record/5333132/files/figure.png" pageId="7" pageNumber="133">Fig. 17</figureCitation>
. Length
<quantity id="203FCCD2D62FFFC9FF2CFDDE7D13FDB7" box="[252,345,566,587]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="133" unit="mm" value="15.5">15.5 mm</quantity>
; width across humeri
<quantity id="203FCCD2D62FFFC9FDEFFDDE7EDAFDB7" box="[575,656,566,587]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="133" unit="mm" value="8.7">8.7 mm</quantity>
. Color reddish brown.
<emphasis id="D5B3BD25D62FFFC9FE53FDB87D8AFD99" bold="true" box="[387,448,592,613]" pageId="7" pageNumber="133">Head:</emphasis>
Frons coarsely punctate, area between eyes nearly impunctate. Frontoclypeal region with 2 cariniform tubercles separated by less than 1 tubercle diameter. Clypeus slightly narrowed towards apex, base 2 times as wide as apex. Clypeal teeth triangular, separated by a tooth diameter. Mandibles with 2 apical, acute teeth and a widely rounded basal lobe. Interocular distance equal to 5 times eye width. Antennal club long, 1.5 times longer than antennomeres 27.
<emphasis id="D5B3BD25D62FFFC9FF7DFCB27D51FC93" bold="true" box="[173,283,858,879]" pageId="7" pageNumber="133">Pronotum:</emphasis>
Surface with small, dense punctures. Apical tubercle and fovea absent. Margin of apical angles entire.
<emphasis id="D5B3BD25D62FFFC9FE97FC677DDFFC58" bold="true" box="[327,405,911,932]" pageId="7" pageNumber="133">Elytra:</emphasis>
First interval punctate, punctures similar to those on other intervals.
<emphasis id="D5B3BD25D62FFFC9FD13FF2A7F61FF2B" bold="true" box="[707,811,194,215]" pageId="7" pageNumber="133">Pygidium:</emphasis>
Surface with small, evenly distributed punctures; not rugose Punctures small, evenly distributed. Strongly convex in lateral view. Apex regularly rounded.
<emphasis id="D5B3BD25D62FFFC9FCA6FEF97FE7FEDA" bold="true" box="[886,941,273,294]" pageId="7" pageNumber="133">Legs:</emphasis>
Protibia tridentate without an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia strongly narrowed before apex. Apex of metatibia entire, with 19 spinules.
<emphasis id="D5B3BD25D62FFFC9FBE4FE9378E3FE6C" bold="true" box="[1076,1193,379,400]" pageId="7" pageNumber="133">Parameres:</emphasis>
With a medial tooth on each side; apex widened but ending in an acute, triangular point (
<figureCitation id="7FFC7DB2D62FFFC9FBF5FE5978D0FE3A" box="[1061,1178,432,454]" captionStart="Figs" captionStartId="7.[197,236,1687,1706]" captionTargetBox="[178,1187,1044,1667]" captionTargetId="figure-407@7.[175,1190,1040,1670]" captionTargetPageId="7" captionText="Figs. 1719. Tomarus paranaensis. 17) Habitus; 18) Parameres, frontal view; 19) Parameres, lateral view. Scale bars = 2 mm (Fig. 17), 0.5 mm (Figs. 1819)." figureDoi="http://doi.org/10.5281/zenodo.5333132" httpUri="https://zenodo.org/record/5333132/files/figure.png" pageId="7" pageNumber="133">Figs. 1819</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AFDD32BCD62FFFC9FD0BFE237866FE07" pageId="7" pageNumber="133" type="etymology">
<paragraph id="E7786137D62FFFC9FD0BFE237866FE07" blockId="7.[707,1194,194,959]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FD0BFE237F07FE1C" bold="true" box="[731,845,459,480]" pageId="7" pageNumber="133">Etymology.</emphasis>
The species name refers to the
<typeStatus id="387CDF95D62FFFC9FB50FE2378E3FE1C" box="[1152,1193,459,480]" pageId="7" pageNumber="133">type</typeStatus>
locality in the state of
<collectingRegion id="2503AFD5D62FFFC9FC4EFE0E7FABFE07" box="[926,993,486,507]" country="Brazil" name="Parana" pageId="7" pageNumber="133">Paraná</collectingRegion>
,
<collectingCountry id="9FD021A7D62FFFC9FC3CFE0E7863FE07" box="[1004,1065,486,507]" name="Brazil" pageId="7" pageNumber="133">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="AFDD32BCD62FFFC9FD0BFDE87806FD66" pageId="7" pageNumber="133" type="distribution">
<paragraph id="E7786137D62FFFC9FD0BFDE878CFFD7C" blockId="7.[707,1194,194,959]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FD0BFDE87F16FDE9" bold="true" box="[731,860,512,533]" pageId="7" pageNumber="133">Distribution.</emphasis>
The
<typeStatus id="387CDF95D62FFFC9FC4EFDE87FBBFDE9" box="[926,1009,512,533]" pageId="7" pageNumber="133" type="holotype">holotype</typeStatus>
was collected in Ponta Grossa (
<figureCitation id="7FFC7DB2D62FFFC9FC9BFDF37FDAFDCC" box="[843,912,539,560]" captionStart="Fig" captionStartId="9.[197,228,918,937]" captionTargetBox="[177,1188,196,901]" captionTargetId="figure-402@9.[175,1190,194,903]" captionTargetPageId="9" captionText="Fig. 23. Known localities of five newly described species of Tomarus from Central and South America." figureDoi="http://doi.org/10.5281/zenodo.5333140" httpUri="https://zenodo.org/record/5333140/files/figure.png" pageId="7" pageNumber="133">Fig. 23</figureCitation>
), a municipality located in the Second Plateau of the state of
<collectingRegion id="2503AFD5D62FFFC9FC0AFDDD7850FDB6" box="[986,1050,565,586]" country="Brazil" name="Parana" pageId="7" pageNumber="133">Paraná</collectingRegion>
(
<collectingCountry id="9FD021A7D62FFFC9FBF5FDDD782BFDB6" box="[1061,1121,565,586]" name="Brazil" pageId="7" pageNumber="133">Brazil</collectingCountry>
) within the Atlantic Forest biome, represented by the Araucaria Forest ecosystem (
<bibRefCitation id="83561CC6D62FFFC9FC12FD837830FD7C" author="Mielke, O. H. H." box="[962,1146,619,640]" pageId="7" pageNumber="133" pagination="59 - 66" refId="ref9044" refString="Mielke, O. H. H., E. Carneiro, and M. M. Casagrande. 2012. Hesperiidae (Lepidoptera, Hesperioidea) from Ponta Grossa, Parana, Brazil: 70 years of records with special reference to faunal composition of Vila Velha State Park. Revista Brasileira de Entomolog'ia 56 (1): 59 - 66." type="journal article" year="2012">
Mielke
<emphasis id="D5B3BD25D62FFFC9FBDEFD837875FD7C" box="[1038,1087,619,640]" italics="true" pageId="7" pageNumber="133">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
<paragraph id="E7786137D62FFFC9FD0BFD6D7806FD66" blockId="7.[707,1194,194,959]" box="[731,1100,645,666]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FD0BFD6D7F8FFD66" bold="true" box="[731,965,645,666]" pageId="7" pageNumber="133">Temporal Distribution.</emphasis>
February (1).
</paragraph>
</subSubSection>
<subSubSection id="AFDD32BCD62FFFC9FD0BFD487F4BFC43" pageId="7" pageNumber="133" type="diagnosis">
<paragraph id="E7786137D62FFFC9FD0BFD487F4BFC43" blockId="7.[707,1194,194,959]" pageId="7" pageNumber="133">
<emphasis id="D5B3BD25D62FFFC9FD0BFD487F08FD49" bold="true" box="[731,834,672,693]" pageId="7" pageNumber="133">Diagnosis.</emphasis>
<taxonomicName id="20C71AB4D62FFFC9FC80FD48786EFD49" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2019" box="[848,1060,672,693]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="paranaensis">
<emphasis id="D5B3BD25D62FFFC9FC80FD48786EFD49" box="[848,1060,672,693]" italics="true" pageId="7" pageNumber="133">Tomarus paranaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="20C71AB4D62FFFC9FBB1FD487F45FD33" baseAuthorityName="Steinheil" baseAuthorityYear="1872" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="D5B3BD25D62FFFC9FBB1FD487F45FD33" italics="true" pageId="7" pageNumber="133">T. burmeisteri</emphasis>
</taxonomicName>
are similar in their external morphology. However, the metatibiae are strongly narrowed before the apex and have 19 spinules on the apical margin in
<taxonomicName id="20C71AB4D62FFFC9FCFCFCE27FF1FCE3" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2019" box="[812,955,778,799]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="paranaensis">
<emphasis id="D5B3BD25D62FFFC9FCFCFCE27FF1FCE3" box="[812,955,778,799]" italics="true" pageId="7" pageNumber="133">T. paranaensis</emphasis>
</taxonomicName>
, whereas they have the sides nearly parallel and the apical margin with 2527 spinules in
<taxonomicName id="20C71AB4D62FFFC9FCBEFCD77FBBFCA8" baseAuthorityName="Steinheil" baseAuthorityYear="1872" box="[878,1009,831,852]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="D5B3BD25D62FFFC9FCBEFCD77FBBFCA8" box="[878,1009,831,852]" italics="true" pageId="7" pageNumber="133">T. burmeisteri</emphasis>
</taxonomicName>
. The parameres are apically acute in
<taxonomicName id="20C71AB4D62FFFC9FCBAFCB27FB2FC93" authorityName="Lopez-Garcia &amp; Deloya" authorityYear="2019" box="[874,1016,858,879]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="paranaensis">
<emphasis id="D5B3BD25D62FFFC9FCBAFCB27FB2FC93" box="[874,1016,858,879]" italics="true" pageId="7" pageNumber="133">T. paranaensis</emphasis>
</taxonomicName>
(
<figureCitation id="7FFC7DB2D62FFFC9FBD7FCB27831FC93" box="[1031,1147,857,879]" captionStart="Figs" captionStartId="7.[197,236,1687,1706]" captionTargetBox="[178,1187,1044,1667]" captionTargetId="figure-407@7.[175,1190,1040,1670]" captionTargetPageId="7" captionText="Figs. 1719. Tomarus paranaensis. 17) Habitus; 18) Parameres, frontal view; 19) Parameres, lateral view. Scale bars = 2 mm (Fig. 17), 0.5 mm (Figs. 1819)." figureDoi="http://doi.org/10.5281/zenodo.5333132" httpUri="https://zenodo.org/record/5333132/files/figure.png" pageId="7" pageNumber="133">Figs. 1819</figureCitation>
) but rounded to subquadrate in
<taxonomicName id="20C71AB4D62FFFC9FC17FC9C7804FC75" baseAuthorityName="Steinheil" baseAuthorityYear="1872" box="[967,1102,884,905]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="D5B3BD25D62FFFC9FC17FC9C7804FC75" box="[967,1102,884,905]" italics="true" pageId="7" pageNumber="133">T. burmeisteri</emphasis>
</taxonomicName>
(Fig. 49; see also fig.
<quantity id="203FCCD2D62FFFC9FC9FFC677FCCFC58" box="[847,902,911,932]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="7" pageNumber="133" unit="in" value="18.0">18 in</quantity>
Neita-Moreno and Ratcliffe 2017).
</paragraph>
</subSubSection>
</treatment>
</document>