229 lines
23 KiB
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229 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jor.31.73800" ID-GBIF-Dataset="34ac9f69-6c5b-42a3-a3f6-37c26a0f52bf" ID-GBIF-Taxon="193418333" ID-Pensoft-Pub="1937-2426-1-9" ID-Pensoft-UUID="DF53616E0253557E849F8362F9D7BA1E" ID-ZooBank="3906D11118494F9B87FDF70673B1B60E" LSID="urn:lsid:zoobank.org:act:D074F7D3-D6C5-463C-8D3E-4EFD304AD4C9" ModsDocID="1937-2426-31-1-9" checkinTime="1645432308379" checkinUser="pensoft" docAuthor="Tan, Ming Kai & Robillard, Tony" docDate="2022" docId="44CDBB7378FF50098B80EE7AB710F0B2" docLanguage="en" docName="JourOrthoptRes 31(1): 9-40" docOrigin="Journal of Orthoptera Research 31 (1)" docPubDate="2022-02-21" docSource="http://dx.doi.org/10.3897/jor.31.73800" docTitle="Rugabinthus biakis Tan & Robillard 2022, sp. nov." docType="treatment" docUuid="D074F7D3-D6C5-463C-8D3E-4EFD304AD4C9" docUuidSource="ZooBank" docVersion="4" id="DF53616E0253557E849F8362F9D7BA1E" lastPageNumber="9" masterDocId="DF53616E0253557E849F8362F9D7BA1E" masterDocTitle="Rugabinthus, a new genus of Lebinthina (Orthoptera, Gryllidae, Eneopterinae) from New Guinea" masterLastPageNumber="40" masterPageNumber="9" pageId="0" pageNumber="9" updateTime="1645436839305" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Rugabinthus, a new genus of Lebinthina (Orthoptera, Gryllidae, Eneopterinae) from New Guinea</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tan, Ming Kai</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4324-6305</mods:nameIdentifier>
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<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:nameIdentifier type="email">orthoptera.mingkai@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Robillard, Tony</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2177-9549</mods:nameIdentifier>
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<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Orthoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>31</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>9</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jor.31.73800</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jor.31.73800</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1937-2426-1-9</mods:identifier>
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<mods:identifier type="ZooBank">3906D11118494F9B87FDF70673B1B60E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">DF53616E0253557E849F8362F9D7BA1E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="193418333" LSID="urn:lsid:zoobank.org:act:D074F7D3-D6C5-463C-8D3E-4EFD304AD4C9" httpUri="http://treatment.plazi.org/id/44CDBB7378FF50098B80EE7AB710F0B2" lastPageNumber="9" pageId="0" pageNumber="9">
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<subSubSection pageId="0" pageNumber="9" type="nomenclature">
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<paragraph pageId="0" pageNumber="9">
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<taxonomicName LSID="http://zoobank.org/D074F7D3-D6C5-463C-8D3E-4EFD304AD4C9" authority="Tan & Robillard, 2022" authorityName="Tan & Robillard" authorityYear="2022" family="Gryllidae" genus="Rugabinthus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rugabinthus biakis" order="Orthoptera" pageId="0" pageNumber="9" rank="species" species="biakis" status="sp. nov.">Rugabinthus biakis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="9">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="description">
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<paragraph pageId="0" pageNumber="9">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. Map of New Guinea island showing the distribution of Rugabinthus species; B. Female Rugabinthus species in their natural habitats in Acemo, South Manokwari (https: // www. inaturalist. org / observations / 73044465) and C. Malagufuk (https: // www. inaturalist. org / observations / 72638321). Photo credit: Benoit Segerer." figureDoi="10.3897/jor.31.73800.figure3" httpUri="https://binary.pensoft.net/fig/647924" pageId="0" pageNumber="9">(Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">, 4H</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Face: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure5" httpUri="https://binary.pensoft.net/fig/647926" pageId="0" pageNumber="9">, 5H</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and pronotum in lateral views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure6" httpUri="https://binary.pensoft.net/fig/647927" pageId="0" pageNumber="9">, 6H</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure7" httpUri="https://binary.pensoft.net/fig/647928" pageId="0" pageNumber="9">, 7H</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia in dorsal view; posterior apex of pseudepiphallus: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure8" httpUri="https://binary.pensoft.net/fig/647929" pageId="0" pageNumber="9">, 8H</figureCitation>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. R. biakis sp. nov. male habitus in dorsal (A) and lateral (B) views. Scale bar: 10 mm." figureDoi="10.3897/jor.31.73800.figure19" httpUri="https://binary.pensoft.net/fig/647940" pageId="0" pageNumber="9">, 19</figureCitation>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. biakis sp. nov.; C, D. R. nabire sp. nov.; E, F. R. tariku sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure20" httpUri="https://binary.pensoft.net/fig/647941" pageId="0" pageNumber="9">, 20A</figureCitation>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. biakis sp. nov.; C, D. R. nabire sp. nov.; E, F. R. tariku sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure20" httpUri="https://binary.pensoft.net/fig/647941" pageId="0" pageNumber="9">, 20B)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="materials_examined">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Material examined.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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<materialsCitation collectingDate="2004-11-14" collectingDateMax="2004-11-15" collectingDateMin="2004-11-14" collectorName="W New, A. Gorochov" country="INDONESIA" location="Corem" municipality="Biak Island" specimenCount="♂" stateProvince="West Papua" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="9">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingRegion country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
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,
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<collectingMunicipality>Biak Island</collectingMunicipality>
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(not far from N. coast of
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<collectorName>W New</collectorName>
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Guinea), secondary forest on hill near vill.
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<location LSID="urn:lsid:plazi:treatment:44CDBB7378FF50098B80EE7AB710F0B2:1F65E14416FD3ABCC8813205D5CFE8BA" country="INDONESIA" municipality="Biak Island" name="Corem" stateProvince="West Papua">Corem</location>
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;
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<collectingDate value="2004-11-14" valueMax="2004-11-15" valueMin="2004-11-14">14-15 November 2004</collectingDate>
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;
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<collectorName>A. Gorochov</collectorName>
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leg.; ZIN
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</materialsCitation>
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.
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="INDONESIA" location="INDONESIA" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="9">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
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•
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<specimenCount type="male">1♂</specimenCount>
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; same information as
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<typeStatus>holotype</typeStatus>
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; molecular sample L86; MZB
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</materialsCitation>
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•
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<materialsCitation collectingDate="2004-11-14" collectingDateMax="2004-11-15" collectingDateMin="2004-11-14" collectorName="W New, A. Gorochov" country="Guinea" location="Biak Island" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount type="male">1♂</specimenCount>
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;
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<location LSID="urn:lsid:plazi:treatment:44CDBB7378FF50098B80EE7AB710F0B2:2AF8761E2575380053C51133927BAE87" country="Guinea" name="Biak Island">Biak Island</location>
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(not far from N. coast of
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<collectorName>W New</collectorName>
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<collectingCountry name="Guinea-Bissau">Guinea</collectingCountry>
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), secondary forest on hill near vill. Corem;
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<collectingDate value="2004-11-14" valueMax="2004-11-15" valueMin="2004-11-14">14-15 November 2004</collectingDate>
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;
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<collectorName>A. Gorochov</collectorName>
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leg.; MNHN-EO-ENSIF1745
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="materials_examined">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Other material.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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<materialsCitation collectingDate="2012-01-17" collectingDateMax="2012-01-20" collectingDateMin="2012-01-17" collectorName="A. Gorochov" country="INDONESIA" location="Biak Town" municipality="Biak Island" specimenCount="1" specimenCount-juvenile="1" stateProvince="West Papua">
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<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
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•
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<specimenCount type="juvenile">1 juvenile</specimenCount>
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;
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<collectingRegion country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
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,
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<collectingMunicipality>Biak Island</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:44CDBB7378FF50098B80EE7AB710F0B2:3CC9E2143CA4BD6D499D3122FCA658A0" country="INDONESIA" municipality="Biak Island" name="Biak Town" stateProvince="West Papua">Biak Town</location>
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, forest on not high hill near airport;
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<collectingDate value="2012-01-17" valueMax="2012-01-20" valueMin="2012-01-17">17-20 January 2012</collectingDate>
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; leg.
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<collectorName>A. Gorochov</collectorName>
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; molecular sample L85; ZIN
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="type locality">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Type locality.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">INDONESIA: West Papua: Biak.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="etymology">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Etymology.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">The species is named after the type locality: Biak Island.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="diagnosis">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Diagnosis.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">This new species differs from all congeners by its small size; dorsum of head yellow brown with five well-separated and well-defined dark brown longitudinal bands; pronotal disk yellow brown with a median dark brown longitudinal band and sparse but well defined dark brown spots of different sizes; lateral lobe contrasting dark in coloration. It also differs from all congeners by male genitalia with pseudepiphallus rectangular, stouter than congeners, apex truncated forming a short posterior plate with a small median nodule.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="description">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Description.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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Small sized among congeners (Fig.
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. R. biakis sp. nov. male habitus in dorsal (A) and lateral (B) views. Scale bar: 10 mm." figureDoi="10.3897/jor.31.73800.figure19" httpUri="https://binary.pensoft.net/fig/647940" pageId="0" pageNumber="9">19</figureCitation>
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). Dorsum of head yellow brown with five well-separated and well-defined dark brown longitudinal bands, middle band widest resulting from fusion of two bands, partly split anteriorly (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">4H</figureCitation>
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). Fastigium yellow brown with dark brown patterns. Scapes brown. Antennae yellow brown basally, distally brown with yellow rings. Fastigium verticis and frons black, frons with two faint yellow brown spots, eyes underlined with yellow; clypeus and mouthparts dark brown to black, labrum distinctly cream-colored (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Face: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure5" httpUri="https://binary.pensoft.net/fig/647926" pageId="0" pageNumber="9">5H</figureCitation>
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). Maxillary palpi cream-colored with brown patterns. Pronotal disk yellow brown with a median dark brown longitudinal band and sparse but well defined dark brown spots of different size (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">4H</figureCitation>
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). Lateral lobes of pronotum entirely dark brown, distinctly darker than disk (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and pronotum in lateral views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure6" httpUri="https://binary.pensoft.net/fig/647927" pageId="0" pageNumber="9">6H</figureCitation>
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). Legs pale yellow brown with well-defined brown spots and rings. FIs and FIIs mostly cream-colored with a few well-defined brown spots, knees brown; TIs and TIIs dark brown with a cream-colored ring in middle. FIIIs cream-colored with numerous oblique dark brown bands, knees dark brown. Tergites unicolorous dark brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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<emphasis bold="true" pageId="0" pageNumber="9">Male.</emphasis>
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FWs reaching middle of fourth abdominal tergite. FW coloration (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure7" httpUri="https://binary.pensoft.net/fig/647928" pageId="0" pageNumber="9">7H</figureCitation>
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): Dorsal field cells and veins mostly brown; some veins near basal area cream-colored; M vein orange brown; area between M and R infumate cream-colored; basal area with a large cream-colored spot on anterior corner and another one near 1A and 2A. Lateral field dark red brown in dorsal half, gray brown in ventral half. FW venation typical of genus, 1A bisinuate anterior to angle; oblique vein base faded, both its posterior and anterior branches almost straight. Apex of dorsal field obliquely rounded.
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Male genitalia</emphasis>
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: (Figs
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia in dorsal view; posterior apex of pseudepiphallus: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure8" httpUri="https://binary.pensoft.net/fig/647929" pageId="0" pageNumber="9">8H</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. biakis sp. nov.; C, D. R. nabire sp. nov.; E, F. R. tariku sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure20" httpUri="https://binary.pensoft.net/fig/647941" pageId="0" pageNumber="9">20A</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. biakis sp. nov.; C, D. R. nabire sp. nov.; E, F. R. tariku sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure20" httpUri="https://binary.pensoft.net/fig/647941" pageId="0" pageNumber="9">20B</figureCitation>
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) Pseudepiphallus rectangular, stouter, and shorter than in congeners, very wide laterally at base of rami, its basal anterior margin almost straight, lateral margin generally parallel, apex truncated, forming a short posterior plate, slightly concave dorsally, with a small median nodule. Rami short, but longer than half of pseudepiphallus length. Pseudepiphallic parameres strongly bent 90° in the middle, apex strongly sclerotized and forming two stout lobules. Ectophallic apodemes surpassing anterior margin of pseudepiphallic sclerite. Endophallic sclerite short, barely reaching anterior margin of pseudepiphallus, with thin posterior lateral arms and a very small medio-posterior expansion.
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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<emphasis bold="true" pageId="0" pageNumber="9">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="9" type="description">
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<paragraph pageId="0" pageNumber="9">
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<emphasis italics="true" pageId="0" pageNumber="9">Measurements.</emphasis>
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-
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</paragraph>
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<paragraph pageId="0" pageNumber="9">
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See Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm)." httpUri="http://table.plazi.org/id/1CD01F98C3B7E3BF17EF858517C029E3" pageId="0" pageNumber="9" tableUuid="1CD01F98C3B7E3BF17EF858517C029E3">1</tableCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |