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<document id="AA717E738E9D1D9FC27DF3D79EA0C251" ID-DOI="10.5281/zenodo.275291" ID-GBIF-Dataset="1a88150f-f510-49d1-b9a3-6a8c4950511c" ID-ISSN="1175-5326" ID-Zenodo-Dep="275291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461127025734" checkinUser="plazi" docAuthor="Cruz, Estevam L., Silva, Da &amp; Lise, Arno A." docDate="2009" docId="41735161FFD06412FF078370E370FE5B" docLanguage="en" docName="zt02289p054.pdf" docOrigin="Zootaxa 2289" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enna segredo Cruz, Silva &amp; Lise, 2009, new species" docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="BD4A2919FFD16410FF908619E079FF9B" masterDocTitle="Four new species of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae) from Brazil" masterLastPageNumber="54" masterPageNumber="48" pageNumber="49" updateTime="1698597016831" updateUser="plazi">
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<mods:title id="A22082072D3C215D5806C707F77EFDC6">Four new species of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae) from Brazil</mods:title>
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<mods:name id="EAA21601C2DC28C4A5F598EF90FCAE6E" type="personal">
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<mods:namePart id="C4522F348DC765B3EAA8B0389DCDC186">Cruz, Estevam L.</mods:namePart>
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<mods:namePart id="7644C48D61414CE64623AF36A48A16C5">Silva, Da</mods:namePart>
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<mods:namePart id="40159B212AA5D708B0E09FA578968D15">Lise, Arno A.</mods:namePart>
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<mods:typeOfResource id="EC1B67DADD3443BA2BB95CA43FAAE574">text</mods:typeOfResource>
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<mods:title id="2AA73CBF8BE4FF2D0A53E7BE637BE66F">Zootaxa</mods:title>
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<mods:part id="B017E5F3348E2441AB1DADBCC102004E">
<mods:date id="54DF41075AEC78762359E71E5EA6E3C0">2009</mods:date>
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<treatment id="41735161FFD06412FF078370E370FE5B" ID-DOI="http://doi.org/10.5281/zenodo.6220117" ID-GBIF-Taxon="119626321" ID-Zenodo-Dep="6220117" LSID="urn:lsid:plazi:treatment:41735161FFD06412FF078370E370FE5B" httpUri="http://treatment.plazi.org/id/41735161FFD06412FF078370E370FE5B" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
<subSubSection id="81C0B3FCFFD06411FF078370E149FA3D" pageId="1" pageNumber="49" type="nomenclature">
<paragraph id="C965E077FFD06411FF078370E1ADFA19" blockId="1.[151,468,1384,1446]" box="[151,468,1384,1411]" pageId="1" pageNumber="49">
<heading id="922D571BFFD06411FF078370E1ADFA19" bold="true" box="[151,468,1384,1411]" fontSize="11" level="1" pageId="1" pageNumber="49" reason="1">
<emphasis id="FBAE3C65FFD06411FF078370E1ADFA19" bold="true" box="[151,468,1384,1411]" pageId="1" pageNumber="49">
<taxonomicName id="0EDA9BF4FFD06411FF078370E147FA18" ID-CoL="39VKY" box="[151,318,1385,1411]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="segredo" status="sp. nov.">
<emphasis id="FBAE3C65FFD06411FF078370E147FA18" bold="true" box="[151,318,1385,1411]" italics="true" pageId="1" pageNumber="49">Enna segredo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E09D811EFFD06411FED58371E1ADFA19" box="[325,468,1384,1410]" pageId="1" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C965E077FFD06411FF078395E149FA3D" blockId="1.[151,468,1384,1446]" box="[151,304,1420,1446]" pageId="1" pageNumber="49">
<figureCitation id="51E1FCF2FFD06411FF078395E090FA3D" box="[151,233,1420,1446]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1861,1885]" captionTargetBox="[151,1436,191,1840]" captionTargetId="figure@3.[151,1436,191,1840]" captionTargetPageId="3" captionText="FIGURES 1 4. Enna spp., female habitus, dorsal view. 1. Enna trivittata sp. n. 2. E. segredo sp. n. 3. E. meridionalis sp. n. 4. E. caparao sp. n." httpUri="https://zenodo.org/record/275292/files/figure.png" pageId="1" pageNumber="49">Figs. 2</figureCitation>
,
<figureCitation id="51E1FCF2FFD06411FF658395E149FA3D" box="[245,304,1420,1446]" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="1" pageNumber="49">810</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="81C0B3FCFFD06411FF0783C1E3F7F9DB" pageId="1" pageNumber="49" type="materials_examined">
<paragraph id="C965E077FFD06411FF0783C1E4D9F981" blockId="1.[151,1437,1496,2026]" pageId="1" pageNumber="49">
<emphasis id="FBAE3C65FFD06411FF0783C1E1A1FA69" bold="true" box="[151,472,1496,1522]" pageId="1" pageNumber="49">
<typeStatus id="16615ED5FFD06411FF0783C1E0ACFA69" box="[151,213,1496,1522]" pageId="1" pageNumber="49">Type</typeStatus>
material:
<typeStatus id="16615ED5FFD06411FECB83C1E1AAFA69" box="[347,467,1496,1522]" pageId="1" pageNumber="49" type="holotype">Holotype</typeStatus>
:
</emphasis>
female from Usina Hidrelétrica de
<taxonomicName id="0EDA9BF4FFD06411FC1383C0E39DFA68" box="[899,996,1497,1523]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="segredo" status="sp. nov.">Segredo</taxonomicName>
(Reservatório do rio Jordão/Candói/ Mangueirinha), Paraná,
<collectingCountry id="B1CDA0E7FFD06411FE228019E183F981" box="[434,506,1536,1562]" name="Brazil" pageId="1" pageNumber="49">Brazil</collectingCountry>
[
<geoCoordinate id="ACEE86B0FFD06411FD988019E20DF981" box="[520,628,1536,1562]" direction="south" orientation="latitude" pageId="1" pageNumber="49" precision="925" value="-23.766666">23° 46S</geoCoordinate>
,
<geoCoordinate id="ACEE86B0FFD06411FD118019E281F981" box="[641,760,1536,1562]" direction="west" orientation="longitude" pageId="1" pageNumber="49" precision="925" value="-46.3">46° 18W</geoCoordinate>
],
<date id="BD64C6B7FFD06411FC9C8019E315F981" box="[780,876,1536,1562]" pageId="1" pageNumber="49" value="1996-04">IV.1996</date>
, R. Bertani (
<collectionCode id="AFCB78B2FFD06411FB908019E446F981" box="[1024,1087,1536,1562]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="1" pageNumber="49">IBSP</collectionCode>
11678).
</paragraph>
<paragraph id="C965E077FFD06411FF55803FE3F7F9DB" blockId="1.[151,1437,1496,2026]" box="[197,910,1574,1600]" pageId="1" pageNumber="49">
<emphasis id="FBAE3C65FFD06411FF55803FE135F9DB" bold="true" box="[197,332,1574,1600]" pageId="1" pageNumber="49">
<typeStatus id="16615ED5FFD06411FF55803FE13EF9DB" box="[197,327,1574,1600]" pageId="1" pageNumber="49" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2Ƥ, same data as
<typeStatus id="16615ED5FFD06411FDB2803FE2F1F9DB" box="[546,648,1574,1600]" pageId="1" pageNumber="49" type="holotype">holotype</typeStatus>
(
<collectionCode id="AFCB78B2FFD06411FD08803FE2A1F9DB" box="[664,728,1574,1600]" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="1" pageNumber="49">IBSP</collectionCode>
11650, 11169).
</paragraph>
</subSubSection>
<subSubSection id="81C0B3FCFFD06411FF558054E4EFF9FC" box="[197,1174,1613,1639]" pageId="1" pageNumber="49" type="etymology">
<paragraph id="C965E077FFD06411FF558054E4EFF9FC" blockId="1.[151,1437,1496,2026]" box="[197,1174,1613,1639]" pageId="1" pageNumber="49">
<emphasis id="FBAE3C65FFD06411FF558054E128F9FC" bold="true" box="[197,337,1613,1639]" pageId="1" pageNumber="49">Etymology.</emphasis>
The specific name is a noun in apposition referring to the
<typeStatus id="16615ED5FFD06411FC6F8054E44BF9FC" box="[1023,1074,1613,1639]" pageId="1" pageNumber="49">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="81C0B3FCFFD06411FF55806AE27AF899" pageId="1" pageNumber="49" type="diagnosis">
<paragraph id="C965E077FFD06411FF55806AE27AF899" blockId="1.[151,1437,1496,2026]" pageId="1" pageNumber="49">
<emphasis id="FBAE3C65FFD06411FF55806AE13DF916" bold="true" box="[197,324,1651,1677]" pageId="1" pageNumber="49">Diagnosis.</emphasis>
The female of
<taxonomicName id="0EDA9BF4FFD06411FE66806DE20BF916" box="[502,626,1652,1677]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="segredo" status="sp. nov.">
<emphasis id="FBAE3C65FFD06411FE66806DE20BF916" box="[502,626,1652,1677]" italics="true" pageId="1" pageNumber="49">E. segredo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E09D811EFFD06411FDE9806AE2C2F916" box="[633,699,1651,1677]" pageId="1" pageNumber="49" rank="species">
<emphasis id="FBAE3C65FFD06411FDE9806AE2C2F916" bold="true" box="[633,699,1651,1677]" pageId="1" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
is similar to the one of
<taxonomicName id="0EDA9BF4FFD06411FC5E806DE54BF915" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[974,1330,1652,1678]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="49" phylum="Arthropoda" rank="species" species="estebanensis">
<emphasis id="FBAE3C65FFD06411FC5E806DE4FAF916" box="[974,1155,1652,1677]" italics="true" pageId="1" pageNumber="49">E. estebanensis</emphasis>
(Simon, 1898)
</taxonomicName>
(
<bibRefCitation id="AD4B9D86FFD06411FAD2806DE17BF92F" author="Silva" pageId="1" pageNumber="54" refString="Silva, E. L. C.; A. A. Lise &amp; J. E. Carico. (2008) Revision of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae). Journal of Arachnology, 36, 76 - 110." type="journal article" year="2008">
Silva
<emphasis id="FBAE3C65FFD06411FA17806DE0D7F92F" italics="true" pageId="1" pageNumber="49">et al</emphasis>
., 2008
</bibRefCitation>
, p. 92, figs. 6870) by the shape of the anterior margin of the middle field of epigynum (
<figureCitation id="51E1FCF2FFD06411FA888083E526F92F" box="[1304,1375,1690,1716]" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="1" pageNumber="49">Fig. 8</figureCitation>
), but can be distinguished by an elevation on the central area of middle field (MF) (
<figureCitation id="51E1FCF2FFD06411FBB580D8E4EAF940" box="[1061,1171,1729,1755]" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="1" pageNumber="49">Figs. 8, 9</figureCitation>
) and by the prominent median septum (MS) (
<figureCitation id="51E1FCF2FFD06411FE0E80F1E18FF899" box="[414,502,1768,1794]" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="1" pageNumber="49">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81C0B3FCFFD06412FF558117E0BCFE01" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49" type="description">
<paragraph id="C965E077FFD06412FF558117E0BCFE01" blockId="1.[151,1437,1496,2026]" lastBlockId="2.[151,1437,152,448]" lastPageId="2" lastPageNumber="50" pageId="1" pageNumber="49">
<emphasis id="FBAE3C65FFD06411FF558117E11BF8B3" bold="true" box="[197,354,1806,1832]" pageId="1" pageNumber="49">Description.</emphasis>
Female (
<typeStatus id="16615ED5FFD06411FE448117E230F8B3" box="[468,585,1806,1832]" pageId="1" pageNumber="49" type="holotype">Holotype</typeStatus>
, IBSP 11678). Total length 9.70. Carapace 4.92 long, 4.38 wide, dark brown, with three light brown one median and lateral stripes, darker laterally (
<figureCitation id="51E1FCF2FFD06411FBD0812CE4F3F8D4" box="[1088,1162,1845,1871]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1861,1885]" captionTargetBox="[151,1436,191,1840]" captionTargetId="figure@3.[151,1436,191,1840]" captionTargetPageId="3" captionText="FIGURES 1 4. Enna spp., female habitus, dorsal view. 1. Enna trivittata sp. n. 2. E. segredo sp. n. 3. E. meridionalis sp. n. 4. E. caparao sp. n." httpUri="https://zenodo.org/record/275292/files/figure.png" pageId="1" pageNumber="49">Fig. 2</figureCitation>
). Clypeus light brown, bristly, 0.40 high. Anterior eye row straight, 1.04 wide; posterior 2.03 wide, recurved. Eye diameters, interdistances, and median ocular quadrangle: AME 0.16, ALE 0.14, PME 0.22, PLE 0.26; AME-AME 0.20, AME-ALE 0.10, PME-PME 0.50, PME-PLE 0.42, OQA 0.48, OQP 0.99, OQH 0.63. Chelicerae dark brown, bristly; promargin and retromargin of fang furrow with three teeth equidistant and equal in size. Sternum yellowish, bristly; 4.39 long, 4.89 wide. Labium dark brown, lighter posteriorly, 1.99 long, 1.90 wide. Legs brownish with light brown annuli on femora, relative length: IV-I-II-III, I femur 4.23/ tibia-patella 5.56/ metatarsus 3.73/ tarsus 1.82/ total 15.34; II 4.31/ 5.47/ 3.57/ 1.74/ 15.09; III 3.48/ 4.15/ 3.90/ 1.41/ 12.94; IV 4.33/ 5.22/ 3.98/ 1.90/ 15.43. Ventral pairs of macrosetae on tibiae:
<date id="BD64C6B7FFD36412FC7F8715E46DFEBD" box="[1007,1044,268,294]" pageId="2" pageNumber="50">I-4</date>
;
<date id="BD64C6B7FFD36412FBB18715E428FEBD" box="[1057,1105,268,294]" pageId="2" pageNumber="50">II-4</date>
;
<date id="BD64C6B7FFD36412FBCD8715E4EFFEBD" box="[1117,1174,268,294]" pageId="2" pageNumber="50">III-3</date>
;
<date id="BD64C6B7FFD36412FB328715E4A6FEBD" box="[1186,1247,268,294]" pageId="2" pageNumber="50">IV-3.</date>
Abdomen, 4.78 long, grayish, bristly (
<figureCitation id="51E1FCF2FFD36412FE33872BE196FED7" box="[419,495,306,332]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1861,1885]" captionTargetBox="[151,1436,191,1840]" captionTargetId="figure@3.[151,1436,191,1840]" captionTargetPageId="3" captionText="FIGURES 1 4. Enna spp., female habitus, dorsal view. 1. Enna trivittata sp. n. 2. E. segredo sp. n. 3. E. meridionalis sp. n. 4. E. caparao sp. n." httpUri="https://zenodo.org/record/275292/files/figure.png" pageId="2" pageNumber="50">Fig. 2</figureCitation>
). Venter grayish, with scattered setae. Middle field of epigynum elevated and scape slightly projected (
<figureCitation id="51E1FCF2FFD36412FE538740E24EFEE8" box="[451,567,345,371]" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="2" pageNumber="50">Figs. 8, 9</figureCitation>
); spermathecae short and elliptical with long copulatory duct (CD) (
<figureCitation id="51E1FCF2FFD36412FAFC8740E0C1FE01" captionStart="FIGURES 5 16. 5 7" captionStartId="4.[151,269,1647,1671]" captionTargetBox="[155,1424,203,1623]" captionTargetId="figure@4.[151,1436,194,1623]" captionTargetPageId="4" captionText="FIGURES 5 16. 5 7. Enna trivittata sp. n., female epigynum (5 ventral, 6 posterior, 7 dorsal). 8 10. E. segredo sp. n., female epigynum (8 ventral, 9 posterior, 10 dorsal). 11 13. E. meridionalis sp. n., female epigynum (11 ventral, 12 posterior, 13 dorsal). 14 16. E. caparao sp. n., female epigynum (14 ventral, 15 posterior, 16 dorsal). (AS = accessory spermathecae; CD = copulatory duct; FD = fertilization duct; HS = head of spermathecae, MF = middle field of epigynum; S = spermathecae, SC = scape)." httpUri="https://zenodo.org/record/275293/files/figure.png" pageId="2" pageNumber="50">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81C0B3FCFFD36412FF5587BFE370FE5B" box="[197,777,422,448]" pageId="2" pageNumber="50" type="distribution">
<paragraph id="C965E077FFD36412FF5587BFE370FE5B" blockId="2.[151,1437,152,448]" box="[197,777,422,448]" pageId="2" pageNumber="50">
<emphasis id="FBAE3C65FFD36412FF5587BFE11CFE5B" bold="true" box="[197,357,422,448]" pageId="2" pageNumber="50">Distribution.</emphasis>
Known only from the
<typeStatus id="16615ED5FFD36412FDE187BFE2DAFE5B" box="[625,675,422,448]" pageId="2" pageNumber="50">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>