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<document id="6F294E236E8A1C2E4BDF791C28B78DC5" ID-DOI="10.11646/zootaxa.3068.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5279932" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629743751952" checkinUser="felipe" docAuthor="Han, H. L. &amp; Kononenko, V. S." docDate="2011" docId="3B6C87C1AB5CE25783CEFAF74F893C23" docLanguage="en" docName="zt03068p068.pdf" docOrigin="Zootaxa 3068 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3068.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Athetis bicornuta Han &amp; Kononenko 2011, sp. n." docType="treatment" docVersion="3" lastPageNumber="67" masterDocId="C755FFB9AB4DE2458359FFA94B463E30" masterDocTitle="Twelve new species of Athetis Hübner, [1821] 1816 from China (Lepidoptera, Noctuidae)" masterLastPageNumber="68" masterPageNumber="49" pageNumber="66" updateTime="1699105602421" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="02E9D80A1F3AF5314A111CC4BCCE8F4E">Twelve new species of Athetis Hübner, [1821] 1816 from China (Lepidoptera, Noctuidae)</mods:title>
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<mods:name id="CA65CC6123399B44FC69BD0A9DB473DC" type="personal">
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<mods:namePart id="4E903DC0D95815D24E770B1D4E1794E4">Han, H. L.</mods:namePart>
<mods:affiliation id="9A6650DDEB93A6A062A86619DD336881">School of Forestry, Northeast Forestry University, Harbin, CH- 150014 China. E-mail: hanhuilin @ yahoo. com. cn</mods:affiliation>
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<mods:namePart id="187FEA10CBB266E6525CCB2DCBC536E8">Kononenko, V. S.</mods:namePart>
<mods:affiliation id="7259E19A4EFA9C6D8C7999BFBD73D5F9">Laboratory of Entomology, Institute of Biology and Soil Science Far Eastern Branch of Russian Academy of Sciences, RF- 690022 Vladivostok, Russia. E-mail: kononenko @ ibss. dvo. ru</mods:affiliation>
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<mods:title id="472662262453ABBF4AEAAA9D2D34DAA3">Zootaxa</mods:title>
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<mods:date id="D60D3ACF27F096381FA2BBA93493F0B6">2011</mods:date>
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<mods:number id="69F7367F9C6201B58185D695714085D6">2011-10-26</mods:number>
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<mods:url id="5C848F34AF1D07BF8B712692064AAEA1">https://biotaxa.org/Zootaxa/article/view/zootaxa.3068.1.2</mods:url>
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<mods:identifier id="0A6AB760EEE2702196DEB9629012A8D1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3B6C87C1AB5CE25783CEFAF74F893C23" ID-DOI="http://doi.org/10.5281/zenodo.5282571" ID-Zenodo-Dep="5282571" LSID="urn:lsid:plazi:treatment:3B6C87C1AB5CE25783CEFAF74F893C23" httpUri="http://treatment.plazi.org/id/3B6C87C1AB5CE25783CEFAF74F893C23" lastPageId="18" lastPageNumber="67" pageId="17" pageNumber="66">
<subSubSection id="FBDF655CAB5CE25483CEFAF749D23B47" box="[151,660,1374,1399]" pageId="17" pageNumber="66" type="nomenclature">
<paragraph id="B37A36D7AB5CE25483CEFAF749D23B47" blockId="17.[151,660,1374,1434]" box="[151,660,1374,1399]" pageId="17" pageNumber="66">
<heading id="E83281BBAB5CE25483CEFAF749D23B47" bold="true" box="[151,660,1374,1399]" fontSize="10" level="2" pageId="17" pageNumber="66" reason="2">
<emphasis id="81B1EAC5AB5CE25483CEFAF749D23B47" bold="true" box="[151,660,1374,1399]" pageId="17" pageNumber="66">
<taxonomicName id="74C54D54AB5CE25483CEFAF7490C3B47" authority="Han &amp; Kononenko" authorityName="Han &amp; Kononenko" authorityYear="2011" box="[151,586,1374,1399]" class="Insecta" family="Noctuidae" genus="Athetis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="66" phylum="Arthropoda" rank="species" species="bicornuta" status="sp. nov.">
<emphasis id="81B1EAC5AB5CE25483CEFAF74A183B47" bold="true" box="[151,350,1374,1399]" italics="true" pageId="17" pageNumber="66">Athetis bicornuta</emphasis>
Han &amp; Kononenko
</taxonomicName>
,
<taxonomicNameLabel id="9A8257BEAB5CE254810FFAF649D23B47" box="[598,660,1375,1399]" pageId="17" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5CE25483CEFA2B4A113BAA" box="[151,343,1410,1434]" pageId="17" pageNumber="66" type="description">
<paragraph id="B37A36D7AB5CE25483CEFA2B4A113BAA" blockId="17.[151,660,1374,1434]" box="[151,343,1410,1434]" pageId="17" pageNumber="66">
(
<figureCitation id="2BFE2A52AB5CE25483C6FA2B4BBB3BAA" box="[159,253,1410,1434]" captionStart="FIGURES 916" captionStartId="9.[151,263,1877,1900]" captionTargetBox="[171,1418,193,1842]" captionTargetId="figure-101@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 916. Athetis spp., adults. 9. A. flavicolor sp. n., male, holotype, China (ZFMK); 10. A. flavicolor sp. n., male, paratype, Thailand (AB); 11. A. biserrata sp. n., male, holotype, China (ZFMK); 12. A. minivalva sp. n., male, holotype, China (ZFMK); 13. A. longiharpe sp.n., male, holotype, China (NEFU); 14. A. bicornuta sp. n., male, holotype, China (ZFMK); 15. A. furcatula sp. n., male, holotype, China (ZFMK); 16. A. furcatula sp. n., male, paratype, China (NEFU)." figureDoi="http://doi.org/10.5281/zenodo.5279936" httpUri="https://zenodo.org/record/5279936/files/figure.png" pageId="17" pageNumber="66">Figs. 14</figureCitation>
,
<figureCitation id="2BFE2A52AB5CE2548251FA2B4A603BAA" box="[264,294,1410,1434]" captionStart="FIGURES 2529" captionStartId="12.[151,263,1877,1900]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-70@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 2529. Athetis spp., male genitalia. 25. A. biserrata sp. n., paratype, China, 1731ZFMK; 26. A. minivalva sp. n. Holotype, China, 1736ZFMK; 27. A. longiharpe sp.n., holotype, China, HHL-1976-1; 28. A. bicornuta sp. n., holotype, Hö1(Varga) ZFMK; 29. A. furcatula sp. n., paratype, China, HHL-1972-1." figureDoi="http://doi.org/10.5281/zenodo.5279942" httpUri="https://zenodo.org/record/5279942/files/figure.png" pageId="17" pageNumber="66">28</figureCitation>
,
<figureCitation id="2BFE2A52AB5CE254826BFA2B4A093BAA" box="[306,335,1410,1434]" captionStart="FIGURES 3541" captionStartId="14.[151,263,1877,1900]" captionTargetBox="[160,1410,193,1856]" captionTargetId="figure-89@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 3541. Athetis spp., female genitalia. 35. A. flavicolor sp. n., paratype, Thailand, 4-090311VK; 36. A. biserrata sp. n., paratype, China, 1729ZFMK; 37. A. minivalva sp. n., paratype, China, 1726VK; 38. A. longiharpe sp.n., paratype, China, HHL-1975-2; 39. A. bicornuta sp. n., paratype, China, 1734ZFMK; 40. A. furcatula sp. n., paratype, China, 1726VK; 41. A. furcatula sp. n., paratype, China, HHL-1973-2." figureDoi="http://doi.org/10.5281/zenodo.5279946" httpUri="https://zenodo.org/record/5279946/files/figure.png" pageId="17" pageNumber="66">39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5CE25483CEFA6D48A33812" pageId="17" pageNumber="66" type="materials_examined">
<paragraph id="B37A36D7AB5CE25483CEFA6D48A33812" blockId="17.[151,1437,1475,2032]" pageId="17" pageNumber="66">
Type material.
<materialsCitation id="03AD3C8AAB5CE2548214FA6D49443BCF" collectingDate="1935-05-12" collectionCode="ZFMK" collectorName="H. Hone" country="China" location="Provinz Nord-Yuennan" municipality="Likiang" pageId="17" pageNumber="66" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="81B1EAC5AB5CE2548214FA6D4A853BEC" bold="true" box="[333,451,1476,1500]" pageId="17" pageNumber="66">
<typeStatus id="6C7E8875AB5CE2548214FA6D4AF83BEC" box="[333,446,1476,1500]" pageId="17" pageNumber="66" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="CBD27647AB5CE2548292FA6D49543BEC" box="[459,530,1476,1500]" name="China" pageId="17" pageNumber="66">China</collectingCountry>
: male, [Prov.
<collectingRegion id="7101F835AB5CE25481F7FA6D484F3BEB" box="[686,777,1476,1499]" country="China" name="Yunnan" pageId="17" pageNumber="66">Yunnan</collectingRegion>
], &quot;
<collectingMunicipality id="531EACADAB5CE254807DFA6D48383BEC" box="[804,894,1476,1500]" pageId="17" pageNumber="66">Likiang</collectingMunicipality>
(
<collectingCountry id="CBD27647AB5CE25480D7FA6D48903BEC" box="[910,982,1476,1500]" name="China" pageId="17" pageNumber="66">China</collectingCountry>
),
<location id="B61A600CAB5CE25480B3FA6D4FB73BEC" LSID="urn:lsid:plazi:treatment:3B6C87C1AB5CE25783CEFAF74F893C23:B61A600CAB5CE25480B3FA6D4FB73BEC" box="[1002,1265,1476,1500]" country="China" municipality="Likiang" name="Provinz Nord-Yuennan" pageId="17" pageNumber="66" stateProvince="Yunnan">Provinz Nord-Yuennan</location>
&quot;
<date id="C77B1017AB5CE2548658FA6D4E363BEC" box="[1281,1392,1476,1500]" pageId="17" pageNumber="66" value="1935-05-12">
<collectingDate id="D73FE9FFAB5CE2548658FA6D4E363BEC" box="[1281,1392,1476,1500]" pageId="17" pageNumber="66" value="1935-05-12">12.v.1935</collectingDate>
</date>
(
<collectorName id="1E305301AB5CE25486D9FA6D4B9E3BCF" pageId="17" pageNumber="66">H. Höne</collectorName>
), genit. prep. H1.(
<collectionCode id="D5D4AE12AB5CE25482F8FA4E4ABC3BCF" box="[417,506,1511,1535]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="66" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03AD3C8AAB5CE2548156FA414B943813" collectingDate="1935-05-08" collectionCode="ZFMK" collectorName="H. Hone" country="China" location="Provinz Nord-Yuennan" municipality="Likiang" pageId="17" pageNumber="66" specimenCode="ZFMK 1743" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="81B1EAC5AB5CE2548156FA4149D63830" bold="true" box="[527,656,1512,1536]" pageId="17" pageNumber="66">
<typeStatus id="6C7E8875AB5CE2548156FA4149CD3830" box="[527,651,1512,1536]" pageId="17" pageNumber="66" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="A5C3FD5EAB5CE25481C0FA4E484C3BCF" box="[665,778,1511,1535]" pageId="17" pageNumber="66" type="female">2 females</specimenCount>
, same locality, 7,
<date id="C77B1017AB5CE2548080FA4E4F7F3BCF" box="[985,1081,1511,1535]" pageId="17" pageNumber="66" value="1935-05-08">
<collectingDate id="D73FE9FFAB5CE2548080FA4E4F7F3BCF" box="[985,1081,1511,1535]" pageId="17" pageNumber="66" value="1935-05-08">8.v.1935</collectingDate>
</date>
(
<collectorName id="1E305301AB5CE2548712FA4E4FF63BCF" box="[1099,1200,1511,1535]" pageId="17" pageNumber="66">H. Höne</collectorName>
), genit. prep.
<specimenCode id="E3639EACAB5CE2548616FA414B943813" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="66" type="Museum">ZFMK 1743</specimenCode>
</materialsCitation>
.
<materialsCitation id="03AD3C8AAB5CE2548384F9A248A63812" box="[221,992,1547,1571]" collectionCode="ZFMK" country="China" pageId="17" pageNumber="66" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
The
<typeStatus id="6C7E8875AB5CE2548256F9A24A373813" box="[271,369,1547,1571]" pageId="17" pageNumber="66" type="holotype">holotype</typeStatus>
and
<typeStatus id="6C7E8875AB5CE25482F1F9A249553813" box="[424,531,1547,1571]" pageId="17" pageNumber="66" type="paratype">paratypes</typeStatus>
are deposited in the collection of
<collectionCode id="D5D4AE12AB5CE25480D6F9A248A63812" box="[911,992,1547,1570]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="66" type="Museum">ZFMK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5CE254839CF9874E5738BD" pageId="17" pageNumber="66" type="diagnosis">
<paragraph id="B37A36D7AB5CE254839CF9874E5738BD" blockId="17.[151,1437,1475,2032]" pageId="17" pageNumber="66">
<emphasis id="81B1EAC5AB5CE254839CF9874A793877" bold="true" box="[197,319,1582,1607]" inLineHeading="true" pageId="17" pageNumber="66" reason="1">Diagnosis.</emphasis>
Externally the species is characterized by the pale yellowish-grey coloration of the forewing, with contrasting and well expressed dark reniform and orbicular. In the male genitalia it differs by the abrupt apical margin of valva, simple, curved harpe, absence of the digitus and by presence of two cornuti in the vesica.
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5CE257839CF9314A193FFB" lastPageId="18" lastPageNumber="67" pageId="17" pageNumber="66" type="description">
<paragraph id="B37A36D7AB5CE257839CF9314A193FFB" blockId="17.[151,1437,1475,2032]" lastBlockId="18.[151,1436,151,531]" lastPageId="18" lastPageNumber="67" pageId="17" pageNumber="66">
<emphasis id="81B1EAC5AB5CE254839CF9314A113881" bold="true" box="[197,343,1688,1713]" inLineHeading="true" pageId="17" pageNumber="66" reason="1">Description.</emphasis>
External appearance (
<figureCitation id="2BFE2A52AB5CE254810AF93149E13880" box="[595,679,1688,1712]" captionStart="FIGURES 916" captionStartId="9.[151,263,1877,1900]" captionTargetBox="[171,1418,193,1842]" captionTargetId="figure-101@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 916. Athetis spp., adults. 9. A. flavicolor sp. n., male, holotype, China (ZFMK); 10. A. flavicolor sp. n., male, paratype, Thailand (AB); 11. A. biserrata sp. n., male, holotype, China (ZFMK); 12. A. minivalva sp. n., male, holotype, China (ZFMK); 13. A. longiharpe sp.n., male, holotype, China (NEFU); 14. A. bicornuta sp. n., male, holotype, China (ZFMK); 15. A. furcatula sp. n., male, holotype, China (ZFMK); 16. A. furcatula sp. n., male, paratype, China (NEFU)." figureDoi="http://doi.org/10.5281/zenodo.5279936" httpUri="https://zenodo.org/record/5279936/files/figure.png" pageId="17" pageNumber="66">Fig. 14</figureCitation>
). Wingspan
<quantity id="743D9B32AB5CE254806CF93148F43880" box="[821,946,1688,1712]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="17" pageNumber="66" unit="mm" value="32.5" valueMax="33.0" valueMin="32.0">3233 mm</quantity>
. Male antennae filiform; palps pressed, laterally dark brown; 3rd segment short, less than half length of 2nd; eyes surrounded with brown lashes; head and thorax covered with brown hair-like scales, head with erected scales. Forewing pale brownish grey; basal line as dark spot or diffused line or black streak in costal field; subbasal line weakly expressed thin, dark, broken on few streaks, or not expressed; orbicular as dark blackish spot; reniform dark, small; postmedial line thin, dentate; terminal field somewhat darker than subterminal; terminal line as row of spots; cilia pale brownish with yellowish borderline. Hindwing pale, greyish-brown; discal spot traceable; cilia pale greyish with yellowish borderline.
<emphasis id="81B1EAC5AB5CE2548606F8C74A433999" bold="true" pageId="17" pageNumber="66">Male genitalia.</emphasis>
(
<figureCitation id="2BFE2A52AB5CE254824AF8384A213999" box="[275,359,1937,1961]" captionStart="FIGURES 2529" captionStartId="12.[151,263,1877,1900]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-70@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 2529. Athetis spp., male genitalia. 25. A. biserrata sp. n., paratype, China, 1731ZFMK; 26. A. minivalva sp. n. Holotype, China, 1736ZFMK; 27. A. longiharpe sp.n., holotype, China, HHL-1976-1; 28. A. bicornuta sp. n., holotype, Hö1(Varga) ZFMK; 29. A. furcatula sp. n., paratype, China, HHL-1972-1." figureDoi="http://doi.org/10.5281/zenodo.5279942" httpUri="https://zenodo.org/record/5279942/files/figure.png" pageId="17" pageNumber="66">Fig. 28</figureCitation>
). The description of male genitalia is done on the base of old genitalia preparation in canada balsam: the presence or absence of Abdominal hair brushes is not clear, the vesica is not everted. Uncus missing, tegumen high, with narrow lobes, vinculum relatively narrow, U-shaped; juxta deltoid with long handle; valva elongated, relatively short, with about parallel margins, somewhat expanded apically, its distal part abrupt; clavus expressed as wide conical basal extension; harpe curved basally, situated at distal part of valva, near distal margin; costa without digitus. Aedeagus straigth, rather thin, somewhat longer than valva in length; vesica armed with two cornuti, one of them short, spine-like, another one needle-like.
<emphasis id="81B1EAC5AB5FE2578030FEAB4F713F2B" bold="true" box="[873,1079,258,283]" pageId="18" pageNumber="67">Female genitalia.</emphasis>
(
<figureCitation id="2BFE2A52AB5FE2578711FEAB4FD83F2A" box="[1096,1182,258,282]" captionStart="FIGURES 3541" captionStartId="14.[151,263,1877,1900]" captionTargetBox="[160,1410,193,1856]" captionTargetId="figure-89@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURES 3541. Athetis spp., female genitalia. 35. A. flavicolor sp. n., paratype, Thailand, 4-090311VK; 36. A. biserrata sp. n., paratype, China, 1729ZFMK; 37. A. minivalva sp. n., paratype, China, 1726VK; 38. A. longiharpe sp.n., paratype, China, HHL-1975-2; 39. A. bicornuta sp. n., paratype, China, 1734ZFMK; 40. A. furcatula sp. n., paratype, China, 1726VK; 41. A. furcatula sp. n., paratype, China, HHL-1973-2." figureDoi="http://doi.org/10.5281/zenodo.5279946" httpUri="https://zenodo.org/record/5279946/files/figure.png" pageId="18" pageNumber="67">Fig. 39</figureCitation>
). Ovipositor moderate long, papillae anales relatively short, somewhat tapering, apophyses anteriores and posteriores long, slender, apophyses posteriores about 1,3 times shorter than anterior ones. Antrum large, wide and shallowly cut in apical margin, constricted to join ductus; ductus bursae relatively short, flat, folded in join with antrum, extended to join with bursa; appendix bursae small, membranous; corpus bursae relatively large, membranous, with fine scobination in caudal two thirds.
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5FE257839CFE7E4FDF3FDF" box="[197,1177,471,495]" pageId="18" pageNumber="67" type="etymology">
<paragraph id="B37A36D7AB5FE257839CFE7E4FDF3FDF" blockId="18.[151,1436,151,531]" box="[197,1177,471,495]" pageId="18" pageNumber="67">
<emphasis id="81B1EAC5AB5FE257839CFE7E4A0D3FDF" bold="true" box="[197,331,471,495]" inLineHeading="true" pageId="18" pageNumber="67" reason="1">Etymology.</emphasis>
The name
<taxonomicName id="74C54D54AB5FE2578291FE7E49733FDF" authorityName="Han &amp; Kononenko" authorityYear="2011" box="[456,565,471,495]" class="Insecta" family="Noctuidae" genus="Athetis" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="67" phylum="Arthropoda" rank="species" species="bicornuta">
<emphasis id="81B1EAC5AB5FE2578291FE7E49733FDF" box="[456,565,471,495]" italics="true" pageId="18" pageNumber="67">bicornuta</emphasis>
</taxonomicName>
refers to the two cornuti in the vesica of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="FBDF655CAB5FE257839CFE534F893C23" box="[197,1231,506,531]" pageId="18" pageNumber="67" type="distribution">
<paragraph id="B37A36D7AB5FE257839CFE534F893C23" blockId="18.[151,1436,151,531]" box="[197,1231,506,531]" pageId="18" pageNumber="67">
<emphasis id="81B1EAC5AB5FE257839CFE534A183C23" bold="true" box="[197,350,506,531]" inLineHeading="true" pageId="18" pageNumber="67" reason="1">Distribution.</emphasis>
South West
<collectingCountry id="CBD27647AB5FE25782B2FE5249683C23" box="[491,558,507,531]" name="China" pageId="18" pageNumber="67">China</collectingCountry>
(North of prov.
<collectingRegion id="7101F835AB5FE25781BCFE5248063C23" box="[741,832,507,531]" country="China" name="Yunnan" pageId="18" pageNumber="67">Yunnan</collectingRegion>
). The moths were collected in May.
</paragraph>
</subSubSection>
</treatment>
</document>