300 lines
42 KiB
XML
300 lines
42 KiB
XML
<document id="DF84EDDB55E566E94C1F5F1165EE9F91" ID-CLB-Dataset="32555" ID-DOI="10.11646/zootaxa.4294.5.3" ID-GBIF-Dataset="3ca3fa05-39fd-4321-aeda-297dcc6c291e" ID-ISSN="1175-5326" ID-Zenodo-Dep="839194" ID-ZooBank="62D89C4D-72CB-4CF5-91D7-A828D0BD8C69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501931957015" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="3B39879CFF86FFE5FF1EFC8BE554FC22" docLanguage="en" docName="zootaxa.4294.5.3.pdf.imf" docOrigin="Zootaxa 4294 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cephaloconnus Franz" docType="treatment" docVersion="9" lastPageNumber="534" masterDocId="C700FFE4FF84FFE6FF89FC13E607FF87" masterDocTitle="Taxonomy of ' Euconnus complex'. Part XIII. Status of Afrotropical subgenera of Euconnus: Cephaloconnus, Cerviconnus, Ursoconnus and Nodoconnus (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="544" masterPageNumber="531" pageNumber="533" updateTime="1698466214692" updateUser="ExternalLinkService">
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<mods:title id="4A0FA18ADD16B671E2988A17E2F1D516">Taxonomy of ' Euconnus complex'. Part XIII. Status of Afrotropical subgenera of Euconnus: Cephaloconnus, Cerviconnus, Ursoconnus and Nodoconnus (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="D3230A32C62CA0A803E95D5B6169B24D">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="438DC8759384181DF9247A3938C5DBA6">Zootaxa</mods:title>
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<mods:date id="605AA32EC21D7F33ADE3FB178018C089">2017</mods:date>
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<mods:number id="6C1317CA4A91278DF29A572CB291970D">4294</mods:number>
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<subSection id="DA1F2F61FF86FFE5FF1EFC8BE554FC22" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533" type="multiple">
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<treatment id="3B39879CFF86FFE5FF1EFC8BE554FC22" ID-DOI="http://doi.org/10.5281/zenodo.6036316" ID-GBIF-Taxon="132630468" ID-Zenodo-Dep="6036316" LSID="urn:lsid:plazi:treatment:3B39879CFF86FFE5FF1EFC8BE554FC22" httpUri="http://treatment.plazi.org/id/3B39879CFF86FFE5FF1EFC8BE554FC22" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
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<subSubSection id="FB8A6501FF86FFE4FF1EFC8BE230FF35" box="[151,1079,151,178]" pageId="2" pageNumber="533" type="nomenclature">
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<paragraph id="B32F368AFF86FFE4FF1EFC8BE230FF35" blockId="2.[151,1079,151,178]" box="[151,1079,151,178]" pageId="2" pageNumber="533">
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<heading id="E86781E6FF86FFE4FF1EFC8BE230FF35" bold="true" box="[151,1079,151,178]" fontSize="11" level="1" pageId="2" pageNumber="533" reason="1">
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<emphasis id="81E4EA98FF86FFE4FF1EFC8BE230FF35" bold="true" box="[151,1079,151,178]" pageId="2" pageNumber="533">
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1. Morphological structures and taxonomic status of
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<taxonomicName id="74904D09FF86FFE4FCA0FC84E230FF35" authority="Franz" authorityName="Franz" box="[809,1079,151,178]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="81E4EA98FF86FFE4FCA0FC84E5E1FF36" bold="true" box="[809,998,151,177]" italics="true" pageId="2" pageNumber="533">Cephaloconnus</emphasis>
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Franz
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FB8A6501FF86FFE5FF1EFCCDE4B5FDE7" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533" type="description">
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<paragraph id="B32F368AFF86FFE4FF1EFCCDE547FE48" blockId="2.[151,1436,222,463]" pageId="2" pageNumber="533">
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The general body shape of
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<taxonomicName id="74904D09FF86FFE4FE44FCCCE5B2FF71" authorityName="Franz." authorityYear="1962" box="[461,949,222,247]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="hypertrophicus" subGenus="Cephaloconnus">
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<emphasis id="81E4EA98FF86FFE4FE44FCCCE43CFF71" box="[461,571,223,246]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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(
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<emphasis id="81E4EA98FF86FFE4FDC4FCCDE4FCFF71" box="[589,763,222,246]" italics="true" pageId="2" pageNumber="533">Cephaloconnus</emphasis>
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)
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<emphasis id="81E4EA98FF86FFE4FC84FCCDE5B2FF71" box="[781,949,222,246]" italics="true" pageId="2" pageNumber="533">hypertrophicus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="2BAB2A0FFF86FFE4FC4FFCCDE20FFF70" box="[966,1032,222,247]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dorsal habitus (1 – 4) and set of labels of lectotypes (5, 7 – 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="2" pageNumber="533">Fig. 1</figureCitation>
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) resembles that of many species of
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<taxonomicName id="74904D09FF86FFE4FF1EFD10E797FE9C" authority="(Napochus)" authorityName="Thomson" authorityYear="1859" baseAuthorityName="Napochus" box="[151,400,259,283]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="81E4EA98FF86FFE4FF1EFD10E702FE9D" box="[151,261,259,282]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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(
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<emphasis id="81E4EA98FF86FFE4FE9CFD10E782FE9C" box="[277,389,259,283]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
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)
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</taxonomicName>
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; especially in the antennae with 4-segmented and sharply demarcated club and the pronotum broadest behind middle and strongly narrowing anteriorly, with obtuse-angled but well-marked posterior and anterior corners and the anterior margin much shorter than posterior margin. The pronotum, however, is not broadest at base, as in the
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<typeStatus id="6C2B8828FF86FFE4FE33FD7CE7EDFE00" box="[442,490,367,391]" pageId="2" pageNumber="533">type</typeStatus>
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species of
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<taxonomicName id="74904D09FF86FFE4FDEFFD7DE4D1FE01" authorityName="Thomson" authorityYear="Thomson" box="[614,726,366,390]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="81E4EA98FF86FFE4FDEFFD7DE4D1FE01" box="[614,726,366,390]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
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</taxonomicName>
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, but between middle and posterior third. However, the shape of pronotum is variable within
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<taxonomicName id="74904D09FF86FFE4FE55FD80E44BFE2C" authorityName="Thomson" authorityYear="Thomson" box="[476,588,403,427]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="81E4EA98FF86FFE4FE55FD80E44BFE2C" box="[476,588,403,427]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
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</taxonomicName>
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; species assigned to this subgenus and with the pronota broadest between base and middle are known (e.g.,
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<bibRefCitation id="D7014B7BFF86FFE4FD99FDA5E4D8FE48" author="Jaloszynski" box="[528,735,438,463]" pageId="2" pageNumber="533" refString="Jaloszynski, P. (2015 a) Taxonomy of ' Euconnus complex'. Part III. Morphology of Euconnus subgenus Napochus and revision of Australian species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3925 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 3925.1.1" type="journal article" year="2015" yearSuffix="a">Jałoszyński 2015a</bibRefCitation>
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||
,
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||
<bibRefCitation id="D7014B7BFF86FFE4FD63FDA4E534FE48" author="Jaloszynski" box="[746,819,439,463]" pageId="2" pageNumber="533" refString="Jaloszynski, P. (2017 c) Taxonomy of ' Euconnus complex'. Part XI. Status of Euconophron and Scopophus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4236 (3), 450 - 460." type="journal article" year="2017" yearSuffix="c">2017c</bibRefCitation>
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).
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</paragraph>
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||
<caption id="E7EF6602FF86FFE4FF1EFAC7E4ABF8AE" httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="2" pageNumber="533" targetBox="[152,1430,523,1718]" targetPageId="2">
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<paragraph id="B32F368AFF86FFE4FF1EFAC7E4ABF8AE" blockId="2.[151,1436,1748,1833]" pageId="2" pageNumber="533">
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<emphasis id="81E4EA98FF86FFE4FF1EFAC7E73DF96D" bold="true" box="[151,314,1748,1771]" pageId="2" pageNumber="533">FIGURES 1–8.</emphasis>
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Dorsal habitus (1–4) and set of labels of lectotypes (5, 7–8) and a holotype (6) of
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<taxonomicName id="74904D09FF86FFE4FB0AFAC5E777F88D" authority="Franz" authorityName="Franz." authorityYear="1962" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="hypertrophicus" subGenus="Cephaloconnus">
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||
<emphasis id="81E4EA98FF86FFE4FB0AFAC5E2E0F96C" box="[1155,1255,1750,1771]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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(
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<emphasis id="81E4EA98FF86FFE4FB7FFAC6E394F96C" box="[1270,1427,1749,1771]" italics="true" pageId="2" pageNumber="533">Cephaloconnus</emphasis>
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)
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<emphasis id="81E4EA98FF86FFE4FF1EFAE7E728F88D" box="[151,303,1780,1802]" italics="true" pageId="2" pageNumber="533">hypertrophicus</emphasis>
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Franz
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</taxonomicName>
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(1, 5),
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<taxonomicName id="74904D09FF86FFE4FE35FAE6E55FF88D" authority="Franz" authorityName="Franz." authorityYear="1962" box="[444,856,1780,1802]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="bundukii" subGenus="Cerviconnus">
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<emphasis id="81E4EA98FF86FFE4FE35FAE6E427F88D" box="[444,544,1781,1802]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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(
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<emphasis id="81E4EA98FF86FFE4FDA6FAE6E4A8F88D" box="[559,687,1781,1802]" italics="true" pageId="2" pageNumber="533">Cerviconnus</emphasis>
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)
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<emphasis id="81E4EA98FF86FFE4FD36FAE7E51FF88D" box="[703,792,1780,1802]" italics="true" pageId="2" pageNumber="533">bundukii</emphasis>
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Franz
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</taxonomicName>
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(2, 6),
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<taxonomicName id="74904D09FF86FFE4FC2DFAE6E350F88D" authority="Franz" authorityName="Franz." authorityYear="1962" box="[932,1367,1780,1802]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="basilewskyi" subGenus="Ursoconnus">
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<emphasis id="81E4EA98FF86FFE4FC2DFAE6E20FF88D" box="[932,1032,1781,1802]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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(
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<emphasis id="81E4EA98FF86FFE4FB9EFAE6E296F88D" box="[1047,1169,1781,1802]" italics="true" pageId="2" pageNumber="533">Ursoconnus</emphasis>
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)
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<emphasis id="81E4EA98FF86FFE4FB28FAE7E311F88D" box="[1185,1302,1780,1802]" italics="true" pageId="2" pageNumber="533">basilewskyi</emphasis>
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Franz
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</taxonomicName>
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(3, 7), and
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<taxonomicName id="74904D09FF86FFE4FF4BFB07E46FF8AE" authority="Franz" authorityName="Franz. Left" authorityYear="1986" box="[194,616,1811,1833]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="gomyanus" subGenus="Nodoconnus">
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<emphasis id="81E4EA98FF86FFE4FF4BFB07E721F8AE" box="[194,294,1812,1833]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
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||
(
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<emphasis id="81E4EA98FF86FFE4FEBDFB00E7B4F8AE" box="[308,435,1811,1833]" italics="true" pageId="2" pageNumber="533">Nodoconnus</emphasis>
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)
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<emphasis id="81E4EA98FF86FFE4FE4BFB07E42EF8AE" box="[450,553,1812,1833]" italics="true" pageId="2" pageNumber="533">gomyanus</emphasis>
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Franz
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</taxonomicName>
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(4, 8).
|
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</paragraph>
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</caption>
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<paragraph id="B32F368AFF86FFE5FF4EFB46E5CCFE2A" blockId="2.[151,1436,1877,2010]" lastBlockId="3.[151,1437,151,933]" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
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||
Other characters shared by
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<taxonomicName id="74904D09FF86FFE4FD98FB46E4B8F8EA" box="[529,703,1877,1901]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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<emphasis id="81E4EA98FF86FFE4FD98FB46E4B8F8EA" box="[529,703,1877,1901]" italics="true" pageId="2" pageNumber="533">Cephaloconnus</emphasis>
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||
</taxonomicName>
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||
and
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||
<taxonomicName id="74904D09FF86FFE4FC88FB46E576F8EA" authorityName="Thomson" authorityYear="Thomson" box="[769,881,1877,1901]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
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||
<emphasis id="81E4EA98FF86FFE4FC88FB46E576F8EA" box="[769,881,1877,1901]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
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||
</taxonomicName>
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are: a short head with the vertex not bulging posterodorsally (
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<figureCitation id="2BAB2A0FFF86FFE4FEDAFB6AE79AF816" box="[339,413,1913,1937]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="2" pageNumber="533">Figs 9</figureCitation>
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¯10); the frontoclypeal groove faint or absent (
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<figureCitation id="2BAB2A0FFF86FFE4FC39FB6AE202F816" box="[944,1029,1913,1937]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="2" pageNumber="533">Fig. 10</figureCitation>
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); the hypomeral ridges (
|
||
<figureCitation id="2BAB2A0FFF86FFE4FA93FB6AE36AF816" box="[1306,1389,1913,1937]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="2" pageNumber="533">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF86FFE4FAF3FB6AE394F816" box="[1402,1427,1913,1937]" italics="true" pageId="2" pageNumber="533">hr</emphasis>
|
||
) nearly complete and typically strongly convergent posteriorly; the thick bristles on tempora and sides of prothorax (in some cases reduced in
|
||
<taxonomicName id="74904D09FF86FFE4FE4AFBD2E434F85E" authorityName="Thomson" authorityYear="Thomson" box="[451,563,1985,2009]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF86FFE4FE4AFBD2E434F85E" box="[451,563,1985,2009]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, but present in its
|
||
<typeStatus id="6C2B8828FF86FFE4FC8FFBD1E531F85D" box="[774,822,1986,2010]" pageId="2" pageNumber="533">type</typeStatus>
|
||
species and in
|
||
<taxonomicName id="74904D09FF86FFE4FC6FFBD1E2A9F85E" authorityName="Franz." authorityYear="1962" box="[998,1198,1985,2009]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="hypertrophicus">
|
||
<emphasis id="81E4EA98FF86FFE4FC6FFBD1E2A9F85E" box="[998,1198,1985,2009]" italics="true" pageId="2" pageNumber="533">E. hypertrophicus</emphasis>
|
||
</taxonomicName>
|
||
); an even number of antebasal pronotal pits (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FE31FC84E403FF37" box="[440,516,151,176]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF87FFE5FD91FC8BE445FF37" box="[536,578,152,176]" italics="true" pageId="3" pageNumber="534">abp</emphasis>
|
||
) (one or two pairs of small pits, variable within
|
||
<taxonomicName id="74904D09FF87FFE5FB22FC8BE31CFF37" authorityName="Thomson" authorityYear="Thomson" box="[1195,1307,152,176]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FB22FC8BE31CFF37" box="[1195,1307,152,176]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
), typically accompanied by short and indistinct sublateral carinae (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FC9FFCAFE559FF53" box="[790,862,188,213]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF87FFE5FCE5FCAFE58CFF53" box="[876,907,188,212]" italics="true" pageId="3" pageNumber="534">slc</emphasis>
|
||
) (variable within
|
||
<taxonomicName id="74904D09FF87FFE5FBD2FCAFE2CCFF53" authorityName="Thomson" authorityYear="Thomson" box="[1115,1227,188,212]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FBD2FCAFE2CCFF53" box="[1115,1227,188,212]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, but present in its
|
||
<typeStatus id="6C2B8828FF87FFE5FF1EFCF3E6C0FF7F" box="[151,199,224,248]" pageId="3" pageNumber="534">type</typeStatus>
|
||
species and in
|
||
<taxonomicName id="74904D09FF87FFE5FE08FCF2E44BFF7F" authorityName="Franz." authorityYear="1962" box="[385,588,224,248]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="hypertrophicus">
|
||
<emphasis id="81E4EA98FF87FFE5FE08FCF2E44BFF7F" box="[385,588,224,248]" italics="true" pageId="3" pageNumber="534">E. hypertrophicus</emphasis>
|
||
</taxonomicName>
|
||
); the basisternal part of prosternum (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FB8AFCCCE25FFF7F" box="[1027,1112,223,248]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF87FFE5FBE0FCF3E28DFF7F" box="[1129,1162,224,248]" italics="true" pageId="3" pageNumber="534">bst</emphasis>
|
||
) strikingly short, much shorter than the coxal part; a relatively narrow metaventral intercoxal process (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FB98FD17E262FE9B" box="[1041,1125,260,285]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF87FFE5FBFBFD16E2AFFE9B" box="[1138,1192,261,284]" italics="true" pageId="3" pageNumber="534">mtvp</emphasis>
|
||
) (its width and shape is variable within
|
||
<taxonomicName id="74904D09FF87FFE5FEFFFD3BE7E1FEC7" authorityName="Thomson" authorityYear="Thomson" box="[374,486,296,320]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FEFFFD3BE7E1FEC7" box="[374,486,296,320]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
); and the general shape of the aedeagus (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FC64FD3BE24AFEC7" box="[1005,1101,296,320]" captionStart="FIGURES 27 – 34" captionStartId="11.[151,264,1940,1963]" captionTargetBox="[163,1433,957,1905]" captionTargetId="figure@11.[151,1436,949,1920]" captionTargetPageId="11" captionText="FIGURES 27 – 34. Aedeagus in ventral (27, 29, 31, 33) and lateral (28, 30, 32, 34) view. Euconnus (Cephaloconnus) hypertrophicus Franz (27 – 28); Euconnus (Cerviconnus) bundukii Franz (29 – 30); Euconnus (Ursoconnus) basilewskyi Franz (31 – 32); Euconnus (Nodoconnus) gomyanus Franz (33 – 34)." httpUri="https://zenodo.org/record/844825/files/figure.png" pageId="3" pageNumber="534">Figs 27</figureCitation>
|
||
¯28), except for a distinctly emarginate apex of the median lobe in
|
||
<taxonomicName id="74904D09FF87FFE5FDC6FD5EE511FEE3" authorityName="Franz." authorityYear="1962" box="[591,790,332,356]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="hypertrophicus">
|
||
<emphasis id="81E4EA98FF87FFE5FDC6FD5EE511FEE3" box="[591,790,332,356]" italics="true" pageId="3" pageNumber="534">E. hypertrophicus</emphasis>
|
||
</taxonomicName>
|
||
, whereas the aedeagal apex in most species of
|
||
<taxonomicName id="74904D09FF87FFE5FAA5FD5FE39BFEE3" authorityName="Thomson" authorityYear="Thomson" box="[1324,1436,332,356]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FAA5FD5FE39BFEE3" box="[1324,1436,332,356]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
is not emarginate (but the shape of the aedeagus, and especially its apical portion, shows a high degree of variability within
|
||
<taxonomicName id="74904D09FF87FFE5FED6FD87E7C8FE2B" authorityName="Thomson" authorityYear="Thomson" box="[351,463,404,428]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FED6FD87E7C8FE2B" box="[351,463,404,428]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, e.g., see figs 35̄
|
||
<quantity id="74689B6FFF87FFE5FD11FD87E4D3FE2A" box="[664,724,404,429]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="3" pageNumber="534" unit="in" value="54.0">54 in</quantity>
|
||
<bibRefCitation id="D7014B7BFF87FFE5FD52FD87E5C0FE2A" author="Jaloszynski" box="[731,967,404,429]" pageId="3" pageNumber="534" refString="Jaloszynski, P. (2015 a) Taxonomy of ' Euconnus complex'. Part III. Morphology of Euconnus subgenus Napochus and revision of Australian species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3925 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 3925.1.1" type="journal article" year="2015" yearSuffix="a">Jałoszyński (2015a))</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B32F368AFF87FFE5FF4EFDABE4B5FDE7" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="534">
|
||
A striking character of
|
||
<taxonomicName id="74904D09FF87FFE5FE42FDABE47EFE57" box="[459,633,440,464]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FE42FDABE47EFE57" box="[459,633,440,464]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
, seen in
|
||
<taxonomicName id="74904D09FF87FFE5FD54FDAAE5A2FE57" authorityName="Franz." authorityYear="1962" box="[733,933,440,464]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="hypertrophicus">
|
||
<emphasis id="81E4EA98FF87FFE5FD54FDAAE5A2FE57" box="[733,933,440,464]" italics="true" pageId="3" pageNumber="534">E. hypertrophicus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="2BAB2A0FFF87FFE5FC3DFDA4E5FCFE57" box="[948,1019,439,464]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[152,1430,523,1718]" captionTargetId="figure@2.[151,1436,515,1727]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dorsal habitus (1 – 4) and set of labels of lectotypes (5, 7 – 8) and a holotype (6) of Euconnus (Cephaloconnus) hypertrophicus Franz (1, 5), Euconnus (Cerviconnus) bundukii Franz (2, 6), Euconnus (Ursoconnus) basilewskyi Franz (3, 7), and Euconnus (Nodoconnus) gomyanus Franz (4, 8)." httpUri="https://zenodo.org/record/844809/files/figure.png" pageId="3" pageNumber="534">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="2BAB2A0FFF87FFE5FB80FDABE21EFE57" box="[1033,1049,440,464]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">9</figureCitation>
|
||
̄10, 13̄14), is the modified head. The vertex is more convex than in most species of
|
||
<taxonomicName id="74904D09FF87FFE5FD46FDCFE539FE73" authorityName="Thomson" authorityYear="Thomson" box="[719,830,476,500]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FD46FDCFE539FE73" box="[719,830,476,500]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, the frons is bulging anterodorsally, and these regions are separated by a transverse groove and a rounded ridge (
|
||
<figureCitation id="2BAB2A0FFF87FFE5FCB2FDECE596FD9F" box="[827,913,511,536]" captionStart="FIGURES 9 – 12" captionStartId="4.[151,264,1860,1883]" captionTargetBox="[179,1403,193,1841]" captionTargetId="figure@4.[179,1403,193,1841]" captionTargetPageId="4" captionText="FIGURES 9 – 12. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and elytral base in dorsal view (9); head in anterior view (10); head and prothorax in ventral view (11); pterothorax in ventral view (12). Abbreviations: abp, antebasal pronotal pit; bst, basisternal part of prosternum; ce, compound eye; fr, frons; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; lbr, labrum; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; slc, sublateral carina; smn, submentum; vt, vertex." httpUri="https://zenodo.org/record/844813/files/figure.png" pageId="3" pageNumber="534">Fig. 10</figureCitation>
|
||
). This modification is restricted to males and small pits at each side of the transverse groove suggest a presence of large cephalic glands (the pits are here interpreted as glandular openings;
|
||
<figureCitation id="2BAB2A0FFF87FFE5FD93FE54E46AFDE7" box="[538,621,583,608]" captionStart="FIGURES 13 – 14" captionStartId="5.[151,264,1822,1845]" captionTargetBox="[151,1436,208,1772]" captionTargetId="figure@5.[151,1436,208,1772]" captionTargetPageId="5" captionText="FIGURES 13 – 14. Morphological structures of Euconnus (Cephaloconnus) hypertrophicus Franz. Head, prothorax and anterior portion of pterothorax in lateral (13) and slightly dorsolateral (14) views. Abbreviations: bef, basal elytral fovea; fr, frons; gop, glandular opening; gp, gular plate; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; occ, occipital constriction; pre, prepectus; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/844815/files/figure.png" pageId="3" pageNumber="534">Fig. 14</figureCitation>
|
||
;
|
||
<emphasis id="81E4EA98FF87FFE5FDF1FE5AE4A5FDE7" box="[632,674,585,608]" italics="true" pageId="3" pageNumber="534">gop</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB8A6501FF87FFE5FF4EFE7FE554FC22" pageId="3" pageNumber="534" type="discussion">
|
||
<paragraph id="B32F368AFF87FFE5FF4EFE7FE554FC22" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="534">
|
||
<emphasis id="81E4EA98FF87FFE5FF4EFE7FE758FD02" bold="true" box="[199,351,620,645]" pageId="3" pageNumber="534">Conclusions.</emphasis>
|
||
The only major morphological difference between the
|
||
<typeStatus id="6C2B8828FF87FFE5FC70FE7EE22EFD02" box="[1017,1065,621,645]" pageId="3" pageNumber="534">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="74904D09FF87FFE5FB31FE7FE361FD03" box="[1208,1382,620,644]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FB31FE7FE361FD03" box="[1208,1382,620,644]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="74904D09FF87FFE5FF1EFE83E700FD2F" authorityName="Thomson" authorityYear="Thomson" box="[151,263,656,680]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FF1EFE83E700FD2F" box="[151,263,656,680]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
is the presumably glandular modification of the head in males of the former; females of
|
||
<taxonomicName id="74904D09FF87FFE5FB67FE83E39BFD2F" box="[1262,1436,656,680]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FB67FE83E39BFD2F" box="[1262,1436,656,680]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
do not differ from those of
|
||
<taxonomicName id="74904D09FF87FFE5FE67FEA7E459FD4B" authorityName="Thomson" authorityYear="Thomson" box="[494,606,692,716]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FE67FEA7E459FD4B" box="[494,606,692,716]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, except for having slightly larger heads in relation to the pronotum.
|
||
<taxonomicName id="74904D09FF87FFE5FF1EFECBE7E8FD77" authority="Machulka, 1929" authorityName="Machulka" authorityYear="1929" box="[151,495,727,752]" class="Insecta" family="Scydmaenidae" genus="Neonapochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FF1EFECBE728FD77" box="[151,303,728,752]" italics="true" pageId="3" pageNumber="534">Neonapochus</emphasis>
|
||
<bibRefCitation id="D7014B7BFF87FFE5FEBFFEC4E7E8FD77" author="Machulka" box="[310,495,727,752]" pageId="3" pageNumber="534" refString="Machulka, I. V. (1929) Beitrag zur Kenntnis der Coleopteren-Fauna der ostlichen Slowakei. Pselaphidae et Scydmaenidae. Wiener Entomologische Zeitung, 46, 19 - 21." type="journal article" year="1929">Machulka, 1929</bibRefCitation>
|
||
</taxonomicName>
|
||
, defined solely on the basis of one apomorphy, the modified antennal club in males, was previously placed as a junior synonym of
|
||
<taxonomicName id="74904D09FF87FFE5FD2AFEEFE207FC92" authority="Jaloszynski 2016b" authorityName="Jaloszynski" authorityYear="2016" box="[675,1024,764,789]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FD2AFEEFE515FC93" box="[675,786,764,788]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
(
|
||
<bibRefCitation id="D7014B7BFF87FFE5FCABFEEFE5F0FC92" author="Jaloszynski" box="[802,1015,764,789]" pageId="3" pageNumber="534" refString="Jaloszynski, P. (2016 b) Taxonomy of ' Euconnus complex'. Part VI. Status of Protoscydmus and Euconnus subgenera Neonapochus, Paranapochus and Filonapochus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4067 (1), 35 - 48. https: // doi. org / 10.11646 / zootaxa. 4067.1.2" type="journal article" year="2016" yearSuffix="b">Jałoszyński 2016b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; the same solution seems reasonable in the case of
|
||
<taxonomicName id="74904D09FF87FFE5FEB3FF33E7EFFCBF" box="[314,488,800,824]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FEB3FF33E7EFFCBF" box="[314,488,800,824]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
, which also shows an apomorphy restricted to one sex, and additionally a variable one, as
|
||
<bibRefCitation id="D7014B7BFF87FFE5FF79FF56E783FCDA" author="Franz" box="[240,388,836,861]" pageId="3" pageNumber="534" refString="Franz, H. (1962) Coleoptera Scydmaenidae: Scydmaenus et Euconnus. Mission zoologique de l'I. R. S. A. C. en Afrique orientale. (P. Basilewsky et N. Leleup, 1957). Annales du Musee Royal de l'Afrique Centrale, Tervuren (series 8: Sciences Zoologiques), 107, 338 - 419." type="journal article" year="1962">Franz (1962)</bibRefCitation>
|
||
placed in
|
||
<taxonomicName id="74904D09FF87FFE5FE76FF57E4AAFCDB" box="[511,685,836,860]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FE76FF57E4AAFCDB" box="[511,685,836,860]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
species with the head bearing an impression, tooth or a tubercle. Such characters can be used to define species groups within
|
||
<taxonomicName id="74904D09FF87FFE5FC0AFF7BE5F4FC07" authorityName="Jaloszynski" authorityYear="2016" box="[899,1011,872,896]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FC0AFF7BE5F4FC07" box="[899,1011,872,896]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, but not subgenera.
|
||
<emphasis id="81E4EA98FF87FFE5FB7AFF74E554FC22" bold="true" pageId="3" pageNumber="534">
|
||
Consequently,
|
||
<taxonomicName id="74904D09FF87FFE5FF1EFF98E74CFC23" box="[151,331,907,932]" class="Insecta" family="Scydmaenidae" genus="Cephaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FF1EFF98E74CFC23" bold="true" box="[151,331,907,932]" italics="true" pageId="3" pageNumber="534">Cephaloconnus</emphasis>
|
||
</taxonomicName>
|
||
is placed as a junior synonym of
|
||
<taxonomicName id="74904D09FF87FFE5FD51FF98E54BFC23" authorityName="Jaloszynski" authorityYear="2016" box="[728,844,907,932]" class="Insecta" family="Scydmaenidae" genus="Napochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="81E4EA98FF87FFE5FD51FF98E54BFC23" bold="true" box="[728,844,907,932]" italics="true" pageId="3" pageNumber="534">Napochus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |