231 lines
20 KiB
XML
231 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. & Dowling, Ashley P. G." docDate="2017" docId="2DFA012C5FE5127F31DB73395C4E76C3" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola keesdavidsi Cramer 1992" docType="treatment" docVersion="4" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Danielle M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skvarla, Michael J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nelson, Whitney A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>701</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>496</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
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<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134839666" LSID="urn:lsid:plazi:treatment:2DFA012C5FE5127F31DB73395C4E76C3" httpUri="http://treatment.plazi.org/id/2DFA012C5FE5127F31DB73395C4E76C3" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="Cramer, 1992" authorityName="Cramer" authorityYear="1992" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi Cramer, 1992</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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Cramer, 1992: 17.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(6 ♀; 10 ♂). Arizona, USA: 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; beside Forest Road 42 near junction with Forest Road 42B, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="31.931944">31°55'55"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-109.254166">109°15'15"W</geoCoordinate>
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), 16 July 1987, by IM Smith, IMS870093A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; Sycamore Campground east of Sunizona, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="31.881111">31°52'52"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-109.33889">109°20'20"W</geoCoordinate>
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), 15 July 1987, by IM Smith, IMS870091
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<normalizedToken originalValue="•">*</normalizedToken>
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3 ♂ from Cochise County, Chiricahua Mountains; Cave Creek Recreation Area; John Hands Picnic Area off Forest Road 42A west of Portal, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="31.898056">31°53'53"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-109.220276">109°13'13"W</geoCoordinate>
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), 15 July 1987, by IM Smith, IMS870092A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Coconino County, Oak Creek Canyon; beside Rt. 89A just north of Pine Flat Campground, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="35.016945">35°1'1"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.74555">111°44'44"W</geoCoordinate>
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), 21 July 1987, by IM Smith, IMS870099A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Coconino County, Oak Creek Canyon; beside Rt. 89A between Bootlegger and Banjo Bill Campgrounds, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="34.982777">34°58'58"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.7625">111°45'45"W</geoCoordinate>
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), 21 July 1987, by IM Smith, IMS870100B
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Coconino County, Oak Creek Canyon; beside Rt. 89A just north of Pine Flat Campground, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="35.016945">35°1'1"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.74555">111°44'44"W</geoCoordinate>
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), 21 July 1987, by IM Smith, IMS870099B
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<normalizedToken originalValue="•">*</normalizedToken>
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New Mexico, USA: 2 ♀ from Catron County, Glenwood; Whitewater Picnic Area 8 km east of Rt. 180, (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.372776">33°22'22"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-108.84722">108°50'50"W</geoCoordinate>
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), 12 July 1987, by IM Smith, IMS870084
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<normalizedToken originalValue="•">*</normalizedToken>
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2 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180 (Silver City), (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.203335">33°12'12"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-108.220276">108°13'13"W</geoCoordinate>
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), 10 July 1987, by IM Smith, IMS870081A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180 (Silver City), (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.203335">33°12'12"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-108.220276">108°13'13"W</geoCoordinate>
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), 10 July 1987, by IM Smith, IMS870081B.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type deposition">
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<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype (♀) and allotype (♂) deposited in coll. Cristina Cramer, Instituto de
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<normalizedToken originalValue="Biología">Biologia</normalizedToken>
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, UNAM.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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are similar to other members of the Rala Group (
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<taxonomicName lsidName="T. rala" pageId="0" pageNumber="1" rank="species" species="rala">T. rala</taxonomicName>
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,
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<taxonomicName lsidName="T. lamellipalpis" pageId="0" pageNumber="1" rank="species" species="lamellipalpis">T. lamellipalpis</taxonomicName>
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,
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<taxonomicName lsidName="T. boettgeri" pageId="0" pageNumber="1" rank="species" species="boettgeri">T. boettgeri</taxonomicName>
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,
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<taxonomicName lsidName="T. kurtvietsi" pageId="0" pageNumber="1" rank="species" species="kurtvietsi">T. kurtvietsi</taxonomicName>
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,
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<taxonomicName lsidName="T. dolichodactyla" pageId="0" pageNumber="1" rank="species" species="dolichodactyla">T. dolichodactyla</taxonomicName>
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, and
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<taxonomicName lsidName="T. anoplopalpa" pageId="0" pageNumber="1" rank="species" species="anoplopalpa">T. anoplopalpa</taxonomicName>
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) by being colorless, having incomplete hind coxal margins and being distributed in the southwest.
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<taxonomicName lsidName="T. keesdavidsi" pageId="0" pageNumber="1" rank="species" species="keesdavidsi">T. keesdavidsi</taxonomicName>
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can be differentiated from all other Rala Group by having a shorter dorsum (length ♀ = 555-605 in
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<taxonomicName lsidName="T. keesdavidsi" pageId="0" pageNumber="1" rank="species" species="keesdavidsi">T. keesdavidsi</taxonomicName>
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, 630-800 in others; ♂ = 500-590 in
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<taxonomicName lsidName="T. keesdavidsi" pageId="0" pageNumber="1" rank="species" species="keesdavidsi">T. keesdavidsi</taxonomicName>
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, 600-780 in others) and dentate, flanged ventral extensions on the femora (others are lacking extensions, have tuberculate extensions, or flat, wide lamellate extensions). Additionally,
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<taxonomicName lsidName="T. keesdavidsi" pageId="0" pageNumber="1" rank="species" species="keesdavidsi">T. keesdavidsi</taxonomicName>
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can be differentiated from all other Rala Group by having more elongate pedipalpal tibiae (length/width = 4.50-5.20 in
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<taxonomicName lsidName="T. keesdavidsi" pageId="0" pageNumber="1" rank="species" species="keesdavidsi">T. keesdavidsi</taxonomicName>
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, 1.75-3.38 in others), except
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<taxonomicName lsidName="T. lamellipalpis" pageId="0" pageNumber="1" rank="species" species="lamellipalpis">T. lamellipalpis</taxonomicName>
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(4.32-4.94).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Re-description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Female (Figure 112) (n = 5) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (555-605 long; 450-495 wide) circular and colorless. Anterio-medial platelets (135-137.5 long; 50-60 wide). Anterio-lateral platelets (180-192.5 long; 70-80 wide) free from dorsal plate. Dgl-4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 315-335). Dorsal plate proportions: dorsum length/width 1.21-1.31; dorsal width/distance between Dgl-4 1.41-1.48; anterio-medial platelet length/width 2.25-2.70; anterio-lateral platelet length/width 2.38-2.75; anterio-lateral/anterio-medial length 1.33-1.43.</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (312.5-327.5 long (ventral); 222.5-240 long (dorsal); 97.5-105 tall) colorless. Rostrum (120-130 long; 35-37.5 wide). Chelicerae (295-310 long) with curved fangs (45-50 long). Subcapitular proportions: ventral length/height 3.07-3.21; rostrum length/width 3.33-3.57. Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (35-37.5 long); femur (105-110 long); genu (55-57.5 long); tibia (90-95 long; 18.75-20 wide); tarsus (15-17.5 long). Palpomere proportions: femur/genu 1.91-2.00; tibia/femur 0.84-0.88; tibia length/width 4.50-4.93.</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (695-755 long; 510-560 wide) colorless. Gnathosomal bay (145-150 long; 67.5-80 wide). Cxgl-4 far from apex. Medial suture (35-55 long). Genital plates (162.5-175 long; 152.5-160 wide). Additional measurements: Cx-1 (285-295 long (total); 140-150 long (medial)); Cx-3 (320-330 wide); anterior venter (192.5-222.5 long). Ventral proportions: gnathosomal bay length/width 1.88-2.19; anterior venter/genital field length 1.18-1.29; anterior venter length/genital field width 1.26-1.44; anterior venter/medial suture 4.05-5.50.</paragraph>
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<paragraph pageId="0" pageNumber="1">Male (Figure 113) (n = 5) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (500-590 long; 410-450 wide) circular and colorless. Anterio-medial platelets (120-130 long; 55-60 wide). Anterio-lateral platelets (165-190 long; 65-75 wide) free from dorsal plate. Dgl-4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 280-320). Dorsal plate proportions: dorsum length/width 1.21-1.33; dorsal width/distance between Dgl-4 1.40-1.48; anterio-medial platelet length/width 2.09-2.27; anterio-lateral platelet length/width 2.33-2.71; anterio-lateral/anterio-medial length 1.32-1.48.</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (280-310 long (ventral); 202.5-230 long (dorsal); 85-105 tall) colorless. Rostrum (107.5-125 long; 32.5-35 wide). Chelicerae (260-297.5 long) with curved fangs (40-50 long). Subcapitular proportions: ventral length/height 2.81-3.29; rostrum length/width 3.29-3.57. Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (32.5-35 long); femur (97.5-110 long); genu (50-56.25 long); tibia (87.5-97.5 long; 17.5-20 wide); tarsus (15-17.5 long). Palpomere proportions: femur/genu 1.84-1.96; tibia/femur 0.86-0.95; tibia length/width 4.81-5.20.</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (630-720 long; 460-510 wide) colorless. Gnathosomal bay (130-152.5 long; 60-75 wide). Cxgl-4 far from apex. Medial suture (80-95 long). Genital plates (140-155 long; 115-125 wide). Additional measurements: Cx-1 (260-280 long (total); 130-150 long (medial)); Cx-3 (280-340 wide); anterior venter (225-260 long). Ventral proportions: gnathosomal bay length/width 1.93-2.17; anterior venter/genital field length 1.55-1.86; anterior venter length/genital field width 1.96-2.17; anterior venter/medial suture 2.61-2.89.</paragraph>
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<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Cramer, C" pageId="0" pageNumber="1" url="http://revistas.unam.mx/index.php/zoo/article/view/7056" year="1992">Cramer (1992)</bibRefCitation>
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named this species in honor of Kees Davids "as an acknowledgement of his great contributions to acarology and aquatic ecology" (translated from Spanish).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Southwest (Arizona and New Mexico), extending southward into Mexico (Figure 111).</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 111.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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distribution. Blue star represents type locality (
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<bibRefCitation author="Cramer, C" pageId="0" pageNumber="1" url="http://revistas.unam.mx/index.php/zoo/article/view/7056" year="1992">Cramer 1992</bibRefCitation>
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); and red circles represent new records and material examined.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 112.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 113.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Unfortunately, we were unable to acquire fresh material of
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola keesdavidsi" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keesdavidsi">Torrenticola keesdavidsi</taxonomicName>
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and therefore this species is not included in our phylogenetic analyses. We were also unable to examine type material. However, we were able to examine new material from Arizona and New Mexico that fits well within the species hypothesis proposed by
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<bibRefCitation author="Cramer, C" pageId="0" pageNumber="1" url="http://revistas.unam.mx/index.php/zoo/article/view/7056" year="1992">Cramer (1992)</bibRefCitation>
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. The overall appearance, incomplete hind coxal margins, distribution, and lack of color place this species in the Rala Group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |