331 lines
32 KiB
XML
331 lines
32 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7818FFE4FF013FD2AD32E96B" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus litoralis Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="A031F0AE-D22A-42D8-BFDF-A34F137425AA" docUuidSource="ZooBank" docVersion="11" lastPageNumber="35" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="32" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>ZooKeys</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791386" ID-GBIF-Taxon="163747658" ID-Zenodo-Dep="3791386" LSID="urn:lsid:zoobank.org:act:A031F0AE-D22A-42D8-BFDF-A34F137425AA" httpUri="http://treatment.plazi.org/id/1772122E7818FFE4FF013FD2AD32E96B" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32">
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||
<subSubSection box="[140,575,166,193]" pageId="31" pageNumber="32" type="nomenclature">
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||
<paragraph blockId="31.[140,984,166,229]" box="[140,575,166,193]" pageId="31" pageNumber="32">
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||
<heading box="[140,575,166,193]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="3">
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||
<emphasis bold="true" box="[140,575,166,193]" pageId="31" pageNumber="32">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,498,166,193]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,390,166,193]" italics="true" pageId="31" pageNumber="32">Colossobolus litoralis</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[510,575,167,193]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,984,202,229]" pageId="31" pageNumber="32" type="description">
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||
<paragraph blockId="31.[140,984,166,229]" box="[140,984,202,229]" pageId="31" pageNumber="32">
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||
<uri box="[140,984,202,229]" pageId="31" pageNumber="32">
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||
urn:lsid:zoobank.org:act:
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||
<uuid box="[413,984,202,229]" pageId="31" pageNumber="32">A031F0AE-D22A-42D8-BFDF-A34F137425AA</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="materials_examined">
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<paragraph blockId="31.[140,1108,272,651]" pageId="31" pageNumber="32">
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<emphasis bold="true" box="[140,375,272,298]" pageId="31" pageNumber="32">
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<emphasis box="[140,241,272,298]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<specimenCount box="[386,434,272,299]" pageId="31" pageNumber="32" type="male">4 ♂</specimenCount>
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,
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<specimenCount box="[451,499,272,299]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
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<materialsCitation ID-GBIF-Occurrence="2620669321" collectingDate="2008-02-20" collectingMethod="pitfall trap" collectorName="S. Megson" country="Madagascar" elevation="10" latitude="-12.25" location="Foret d'Orangea" longLatPrecision="1293" longitude="49.366665" municipality="Province" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
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<emphasis box="[510,613,272,298]" italics="true" pageId="31" pageNumber="32">
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<typeStatus box="[510,607,272,298]" pageId="31" pageNumber="32">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[624,672,272,299]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
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(
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<quantity box="[691,794,272,299]" metricMagnitude="-1" metricUnit="m" metricValue="1.3199999999999998" pageId="31" pageNumber="32" unit="mm" value="132.0">132 mm</quantity>
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long), ZSM FGZC 1705,
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<collectingCountry box="[140,274,307,333]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
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,
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<collectingMunicipality box="[287,385,307,333]" pageId="31" pageNumber="32">Province</collectingMunicipality>
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<collectingRegion box="[393,532,307,333]" country="Madagascar" name="Antsiranana" pageId="31" pageNumber="32">Antsiranana</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E7818FFE4FF013FD2AD32E96B:9A04F5E37818FFD9FDAC3E40AD87EB1C" box="[545,734,307,334]" country="Madagascar" latitude="-12.25" longLatPrecision="1293" longitude="49.366665" municipality="Province" name="Foret d'Orangea" pageId="31" pageNumber="32" stateProvince="Antsiranana">Forêt d’Orangea</location>
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,
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<quantity box="[748,812,308,334]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="10.0">
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<elevation box="[748,812,308,334]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="10.0">10 m</elevation>
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</quantity>
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, littoral rainforest,
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<geoCoordinate degrees="12" direction="south" minutes="15" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="-12.25">12°15’ S</geoCoordinate>
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,
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<geoCoordinate box="[168,268,343,369]" degrees="49" direction="east" minutes="22" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="49.366665">49°22’ E</geoCoordinate>
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, leg.
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<collectorName box="[324,445,343,369]" pageId="31" pageNumber="32">S. Megson</collectorName>
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,
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<date box="[455,581,343,369]" pageId="31" pageNumber="32" value="2008-02-20">
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<collectingDate box="[455,581,343,369]" pageId="31" pageNumber="32" value="2008-02-20">20.II.2008</collectingDate>
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</date>
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,
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<collectingMethod box="[591,712,343,369]" pageId="31" pageNumber="32">pitfall trap</collectingMethod>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2620669326" collectingDate="2001-02-22" collectingDateMax="2001-02-28" collectingDateMin="2001-02-22" collectorName="Fisher & Griswold" country="Madagascar" elevation="90" latitude="-12.258889" location="Foret d'Orangea" longLatPrecision="21" longitude="49.37472" pageId="31" pageNumber="32" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis box="[722,829,343,368]" italics="true" pageId="31" pageNumber="32">
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<typeStatus box="[722,824,343,368]" pageId="31" pageNumber="32">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount box="[835,879,343,369]" pageId="31" pageNumber="32" type="male">3 ♂</specimenCount>
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,
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<specimenCount box="[890,932,343,369]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
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, FMMC 3919,
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||
<location LSID="urn:lsid:plazi:treatment:1772122E7818FFE4FF013FD2AD32E96B:9A04F5E37818FFD9FF013E0EAE1EEBC5" box="[140,327,378,404]" country="Madagascar" latitude="-12.258889" longLatPrecision="21" longitude="49.37472" name="Foret d'Orangea" pageId="31" pageNumber="32" stateProvince="Antsiranana">Forêt d’Orangea</location>
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,
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<quantity box="[338,401,378,404]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="m" value="90.0">
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||
<elevation box="[338,401,378,404]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="m" value="90.0">90 m</elevation>
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</quantity>
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, littoral rainforest,
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<geoCoordinate box="[616,757,378,405]" degrees="12" direction="south" minutes="15" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="32" value="-12.258889">12°15’32’’ S</geoCoordinate>
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,
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<geoCoordinate box="[768,911,378,404]" degrees="49" direction="east" minutes="22" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="29" value="49.37472">49°22’29’’ E</geoCoordinate>
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, leg.
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<collectorName box="[969,1039,378,404]" pageId="31" pageNumber="32">Fisher</collectorName>
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,
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<collectorName pageId="31" pageNumber="32">Griswold</collectorName>
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et al.,
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<date box="[272,443,413,439]" pageId="31" pageNumber="32" value="2001-02-22" valueMax="2001-02-28" valueMin="2001-02-22">
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<collectingDate box="[272,443,413,439]" pageId="31" pageNumber="32" value="2001-02-22" valueMax="2001-02-28" valueMin="2001-02-22">22–28.II.2001</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="diagnosis">
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<paragraph blockId="31.[140,1108,272,651]" pageId="31" pageNumber="32">
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||
<emphasis bold="true" box="[185,452,448,474]" pageId="31" pageNumber="32">
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<emphasis box="[185,324,448,474]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Differential</emphasis>
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diagnosis:
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</emphasis>
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the special-shaped anterior and posterior gonopods of
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species">
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<emphasis italics="true" pageId="31" pageNumber="32">C. litoralis</emphasis>
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</taxonomicName>
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are a unique character.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
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<paragraph blockId="31.[140,1108,272,651]" pageId="31" pageNumber="32">
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<emphasis bold="true" box="[185,335,518,544]" pageId="31" pageNumber="32">
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<emphasis box="[185,330,518,544]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Description</emphasis>
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.
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</emphasis>
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<emphasis box="[343,495,519,544]" italics="true" pageId="31" pageNumber="32">Measurements</emphasis>
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: males with 51 body rings,
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<quantity box="[812,968,519,545]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.38" metricValueMin="1.3199999999999998" pageId="31" pageNumber="32" unit="mm" value="135.0" valueMax="138.0" valueMin="132.0">132–138 mm</quantity>
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long, 11.0–
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<quantity box="[140,245,554,580]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="31" pageNumber="32" unit="mm" value="11.6">11.6 mm</quantity>
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wide. Female with 51 body rings,
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<quantity box="[637,734,554,580]" metricMagnitude="-1" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="32" unit="mm" value="128.0">128 mm</quantity>
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||
long,
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<quantity box="[806,911,554,580]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="31" pageNumber="32" unit="mm" value="12.9">12.9 mm</quantity>
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wide.
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</paragraph>
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<paragraph blockId="31.[140,1108,272,651]" pageId="31" pageNumber="32">
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<emphasis box="[185,299,589,615]" italics="true" pageId="31" pageNumber="32">Coloration</emphasis>
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affected by alcohol. Head, antennae, legs and telson red (
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<figureCitation box="[954,1037,589,615]" captionStart="Figure 17" captionStartId="31.[140,212,1622,1644]" captionTargetBox="[140,1108,768,1602]" captionTargetId="figure@31.[140,1109,768,1602]" captionTargetPageId="31" captionText="Figure 17. Colossobolus litoralis sp. n., living specimen, photographed by Angelika Knoll, February 2007 at Forêt Orangea (all rights reserved)." figureDoi="http://doi.org/10.5281/zenodo.3820808" httpUri="https://zenodo.org/record/3820808/files/figure.png" pageId="31" pageNumber="32">Fig. 17</figureCitation>
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). Pro- and anterior part of mesozona dark brown, posterior part of meso- and metazonites red.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3820808" ID-Zenodo-Dep="3820808" httpUri="https://zenodo.org/record/3820808/files/figure.png" pageId="31" pageNumber="32" startId="31.[140,212,1622,1644]" targetBox="[140,1108,768,1602]" targetPageId="31">
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<paragraph blockId="31.[140,1107,1622,1676]" pageId="31" pageNumber="32">
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<emphasis bold="true" box="[140,251,1622,1644]" pageId="31" pageNumber="32">Figure 17.</emphasis>
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[256,436,1622,1643]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[256,436,1622,1643]" italics="true" pageId="31" pageNumber="32">Colossobolus litoralis</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel box="[441,492,1623,1644]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
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||
, living specimen, photographed by Angelika Knoll,
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||
<date box="[971,1107,1622,1644]" pageId="31" pageNumber="32" value="2007-02">February 2007</date>
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at Forêt Orangea (all rights reserved).
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||
</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3820810" ID-Zenodo-Dep="3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="32" pageNumber="33" startId="32.[140,212,1558,1580]" targetBox="[179,1067,165,1538]" targetPageId="32">
|
||
<paragraph blockId="32.[140,1109,1558,1676]" pageId="32" pageNumber="33">
|
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<emphasis bold="true" box="[140,250,1558,1580]" pageId="32" pageNumber="33">Figure 18.</emphasis>
|
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[255,434,1558,1579]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[255,434,1558,1579]" italics="true" pageId="32" pageNumber="33">Colossobolus litoralis</emphasis>
|
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</taxonomicName>
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||
<taxonomicNameLabel box="[440,490,1559,1580]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
|
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, male
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<typeStatus box="[549,633,1558,1580]" pageId="32" pageNumber="33">holotype</typeStatus>
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||
.
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<emphasis bold="true" box="[641,660,1558,1580]" pageId="32" pageNumber="33">A</emphasis>
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||
head, lateral
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||
<emphasis bold="true" box="[781,798,1558,1580]" pageId="32" pageNumber="33">B</emphasis>
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||
telson
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||
<emphasis bold="true" box="[863,882,1558,1580]" pageId="32" pageNumber="33">C</emphasis>
|
||
legs 1–7, ventral view
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||
<emphasis bold="true" box="[1088,1107,1558,1580]" pageId="32" pageNumber="33">D</emphasis>
|
||
coxae of midbody leg, posterior view
|
||
<emphasis bold="true" box="[489,504,1590,1612]" pageId="32" pageNumber="33">E</emphasis>
|
||
anterior gonopods, anterior view
|
||
<emphasis bold="true" box="[821,835,1590,1612]" pageId="32" pageNumber="33">F</emphasis>
|
||
anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Tp = telopodite process. Not to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820812" ID-Zenodo-Dep="3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="33" pageNumber="34" startId="33.[140,212,694,716]" targetBox="[140,1108,165,674]" targetPageId="33">
|
||
<paragraph blockId="33.[140,1108,694,780]" pageId="33" pageNumber="34">
|
||
<emphasis bold="true" box="[140,252,694,716]" pageId="33" pageNumber="34">Figure 19.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[259,440,694,715]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[259,440,694,715]" italics="true" pageId="33" pageNumber="34">Colossobolus litoralis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[447,499,695,716]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male
|
||
<typeStatus box="[561,645,694,716]" pageId="33" pageNumber="34">holotype</typeStatus>
|
||
, posterior gonopods.
|
||
<emphasis bold="true" box="[847,866,694,716]" pageId="33" pageNumber="34">A</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" box="[1002,1019,694,716]" pageId="33" pageNumber="34">B</emphasis>
|
||
posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767296" ID-Zenodo-Dep="3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="33" pageNumber="34" startId="33.[140,212,1494,1516]" targetBox="[140,1108,834,1475]" targetPageId="33">
|
||
<paragraph blockId="33.[140,1108,1494,1676]" pageId="33" pageNumber="34">
|
||
<emphasis bold="true" box="[140,250,1494,1516]" pageId="33" pageNumber="34">Figure 20.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[258,454,1494,1515]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[258,454,1494,1515]" italics="true" pageId="33" pageNumber="34">Colossobolus aculeatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[461,514,1495,1516]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis bold="true" box="[671,690,1494,1516]" pageId="33" pageNumber="34">A</emphasis>
|
||
head, lateral
|
||
<emphasis bold="true" box="[817,834,1494,1516]" pageId="33" pageNumber="34">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[1051,1070,1494,1516]" pageId="33" pageNumber="34">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[179,198,1526,1548]" pageId="33" pageNumber="34">D</emphasis>
|
||
anterior gonopods, anterior view
|
||
<emphasis bold="true" box="[519,534,1526,1548]" pageId="33" pageNumber="34">E</emphasis>
|
||
right posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[909,923,1526,1548]" pageId="33" pageNumber="34">F</emphasis>
|
||
anterior gonopods, posterior view
|
||
<emphasis bold="true" box="[277,297,1558,1580]" pageId="33" pageNumber="34">G</emphasis>
|
||
right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="34.[140,1108,167,827]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,288,167,193]" italics="true" pageId="34" pageNumber="35">Antennae</emphasis>
|
||
protruding back to ring 3 (
|
||
<figureCitation box="[620,726,167,193]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18A</figureCitation>
|
||
).
|
||
<emphasis box="[750,969,167,193]" italics="true" pageId="34" pageNumber="35">Male coxal processes</emphasis>
|
||
on coxae 3 and 4 weakly developed, on coxae 5–7 barely visible (
|
||
<figureCitation box="[754,860,202,228]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18C</figureCitation>
|
||
).
|
||
<emphasis box="[882,1044,202,228]" italics="true" pageId="34" pageNumber="35">Preanal process</emphasis>
|
||
wellrounded, not projecting (
|
||
<figureCitation box="[426,528,237,263]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="34.[140,1108,167,827]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,372,272,299]" italics="true" pageId="34" pageNumber="35">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly rounded lobe (
|
||
<figureCitation box="[992,1095,272,298]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18E</figureCitation>
|
||
). Mesal process of coxite prominent, protruding into a long wide lobe with a wellrounded tip, lobe only slightly shorter than telopodite process (
|
||
<figureCitation box="[863,966,343,369]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18E</figureCitation>
|
||
). Telopodite process long and well-rounded. Mesal margin laterally sharp, at mid-length conspicuously protruding into a long and well-rounded process (
|
||
<figureCitation box="[771,871,413,439]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="34.[140,1108,167,827]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,379,448,474]" italics="true" pageId="34" pageNumber="35">Posterior gonopod</emphasis>
|
||
telopodites positioned parallel to one another. Coxite branch absent (
|
||
<figureCitation box="[231,337,483,510]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="34" pageNumber="35">Fig. 19A</figureCitation>
|
||
). Telopodite lateral branch very slender, straight, tapering towards pointed tip (
|
||
<figureCitation box="[280,382,519,545]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="34" pageNumber="35">Fig. 18A</figureCitation>
|
||
). Central membranous area present as a short, wide, well-round- ed knob (
|
||
<figureCitation box="[254,360,554,580]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="34" pageNumber="35">Fig. 19B</figureCitation>
|
||
). Main branch much longer than lateral branch, basally divided into two parts. Inner part developed as a short, slender, sclerotized process, apically tapering, with a pointed tip. Lateral part prominently elongated into large, flag-like membranous areas, extending beyond telopodite (
|
||
<figureCitation box="[703,840,659,686]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="34" pageNumber="35">Figs 19A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph blockId="34.[140,1108,167,827]" pageId="34" pageNumber="35">
|
||
<emphasis bold="true" box="[185,492,694,721]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,336,694,720]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[510,627,695,721]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[510,627,695,721]" italics="true" pageId="34" pageNumber="35">C. litoralis</emphasis>
|
||
</taxonomicName>
|
||
is only known from the littoral forest of Forêt d’Orangea (
|
||
<figureCitation box="[340,423,730,756]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="34" pageNumber="35" type="etymology">
|
||
<paragraph blockId="34.[140,1108,167,827]" pageId="34" pageNumber="35">
|
||
<emphasis bold="true" box="[185,323,765,791]" pageId="34" pageNumber="35">
|
||
<emphasis box="[185,318,765,791]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[331,412,765,791]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[331,412,765,791]" italics="true" pageId="34" pageNumber="35">litoralis</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the only known site were this species occurs, the littoral forest of Forêt d’Orangea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |