198 lines
27 KiB
XML
198 lines
27 KiB
XML
<document id="F320BD161AB6252E247CDBB0F53526DE" ID-CLB-Dataset="7002" ID-DOI="10.11646/zootaxa.4965.2.9" ID-GBIF-Dataset="73b8e644-f46f-4526-9b4e-b323f2205c8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4751849" ID-ZooBank="C0F65518-6EA9-4099-AEA2-51C5EECFFC78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619710709549" checkinUser="plazi" docAuthor="Allsopp, Peter G." docDate="2021" docId="0C628795FF85FF8BFF1B140DFA2CDEA4" docLanguage="en" docName="zootaxa.4965.2.9.pdf" docOrigin="Zootaxa 4965 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesystoechus Waterhouse 1878" docType="treatment" docVersion="6" lastPageNumber="367" masterDocId="F05BFFEDFF87FF8FFF8C1258FFB1DD73" masterDocTitle="The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)" masterLastPageNumber="374" masterPageNumber="363" pageNumber="365" updateTime="1698938014406" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8BB213E1A2A45F4CF5C5196485BD7804">The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)</mods:title>
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<mods:namePart id="10AE60EDBA72019E25812F75DBC44187">Allsopp, Peter G.</mods:namePart>
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<mods:title id="4DFF78BEC7B2C78E661C697518284815">Zootaxa</mods:title>
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<mods:part id="6B27D88CF39743364AAC2AD9F7F2BE85">
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<mods:date id="475253CAC8D43DEFB90550CFB079941E">2021</mods:date>
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<mods:number id="6AFFC84345360C8DB147BBF431A5DB22">2021-04-28</mods:number>
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<mods:number id="E72A3E6E0DA80921D9575E75CC80ED23">4965</mods:number>
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<mods:identifier id="B0B21A9E0461CA41CE0A0A15563F765A" type="DOI">10.11646/zootaxa.4965.2.9</mods:identifier>
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<treatment id="0C628795FF85FF8BFF1B140DFA2CDEA4" ID-DOI="http://doi.org/10.5281/zenodo.4727956" ID-GBIF-Taxon="181983389" ID-Zenodo-Dep="4727956" LSID="urn:lsid:plazi:treatment:0C628795FF85FF8BFF1B140DFA2CDEA4" httpUri="http://treatment.plazi.org/id/0C628795FF85FF8BFF1B140DFA2CDEA4" lastPageId="4" lastPageNumber="367" pageId="2" pageNumber="365">
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<subSubSection id="CCD16508FF85FF8DFF1B140DFD94DB1C" box="[151,549,1621,1648]" pageId="2" pageNumber="365" type="nomenclature">
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<paragraph id="84743683FF85FF8DFF1B140DFD94DB1C" blockId="2.[151,549,1621,1648]" box="[151,549,1621,1648]" pageId="2" pageNumber="365">
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<heading id="DF3C81EFFF85FF8DFF1B140DFD94DB1C" bold="true" box="[151,549,1621,1648]" fontSize="11" level="1" pageId="2" pageNumber="365" reason="1">
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<taxonomicName id="43CB4D00FF85FF8DFF1B140DFD94DB1C" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[151,549,1621,1648]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
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<emphasis id="B6BFEA91FF85FF8DFF1B140DFD94DB1C" bold="true" box="[151,549,1621,1648]" pageId="2" pageNumber="365">
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<emphasis id="B6BFEA91FF85FF8DFF1B140DFE8EDB03" bold="true" box="[151,319,1621,1648]" italics="true" pageId="2" pageNumber="365">Mesystoechus</emphasis>
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<bibRefCitation id="E05A4B72FF85FF8DFECA140DFD94DB1C" author="Waterhouse, C. O." box="[326,549,1621,1647]" pageId="2" pageNumber="365" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCD16508FF85FF8DFF1B14C2FDAADBC1" box="[151,539,1690,1714]" pageId="2" pageNumber="365" type="reference_group">
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<paragraph id="84743683FF85FF8DFF1B14C2FDAADBC1" blockId="2.[151,539,1690,1714]" box="[151,539,1690,1714]" pageId="2" pageNumber="365">
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<treatmentCitationGroup id="A4DB11ADFF85FF8DFF1B14C2FDAADBC1" box="[151,539,1690,1714]" pageId="2" pageNumber="365">
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<treatmentCitation id="056A1092FF85FF8DFF1B14C2FDA7DBC1" author="Waterhouse, C. O." box="[151,534,1690,1714]" page="227" pageId="2" pageNumber="365" year="1878">
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<taxonomicName id="43CB4D00FF85FF8DFF1B14C2FDA7DBC1" authority="Waterhouse, 1878: 227" authorityName="Waterhouse" authorityPageNumber="227" authorityYear="1878" box="[151,534,1690,1714]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
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<emphasis id="B6BFEA91FF85FF8DFF1B14C2FE95DBC1" box="[151,292,1690,1714]" italics="true" pageId="2" pageNumber="365">Mesystoechus</emphasis>
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<bibRefCitation id="E05A4B72FF85FF8DFEA614C2FDA7DBC1" author="Waterhouse, C. O." box="[298,534,1690,1714]" pageId="2" pageNumber="365" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878: 227</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCD16508FF85FF8DFF1B148FFCEDDB83" box="[151,860,1751,1776]" pageId="2" pageNumber="365" type="type_taxon">
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<paragraph id="84743683FF85FF8DFF1B148FFCEDDB83" blockId="2.[151,860,1751,1776]" box="[151,860,1751,1776]" pageId="2" pageNumber="365">
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<emphasis id="B6BFEA91FF85FF8DFF1B148FFE92DB9C" bold="true" box="[151,291,1751,1775]" pageId="2" pageNumber="365">
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<typeStatus id="5B708821FF85FF8DFF1B148FFF7DDB9C" box="[151,204,1751,1775]" pageId="2" pageNumber="365">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="43CB4D00FF85FF8DFEA6148FFD77DB9C" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[298,710,1751,1775]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="ciliatus">
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<emphasis id="B6BFEA91FF85FF8DFEA6148FFDB9DB9C" box="[298,520,1751,1775]" italics="true" pageId="2" pageNumber="365">Mesystoechus ciliatus</emphasis>
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<bibRefCitation id="E05A4B72FF85FF8DFD81148FFD77DB9C" author="Waterhouse, C. O." box="[525,710,1751,1775]" pageId="2" pageNumber="365" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCD16508FF85FF8BFF1B1545FEFBDCDE" lastPageId="4" lastPageNumber="367" pageId="2" pageNumber="365" type="diagnosis">
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<paragraph id="84743683FF85FF8BFF1B1545FC86DC32" blockId="2.[151,1437,1821,2028]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="367" pageId="2" pageNumber="365">
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<emphasis id="B6BFEA91FF85FF8DFF1B1545FED7DA44" bold="true" box="[151,358,1821,1847]" pageId="2" pageNumber="365">Diagnosis. Males.</emphasis>
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Anoplognathini: Schizognathina. Mentum strongly setose. Labial palp small, with 3 palpomeres. Maxillae with galeae small, not toothed; palps short, slender, with elongate, narrow dorsal sensorium on apical palpomere. Mandibles produced beyond clypeus, margins rounded. Antennae with 10 antennomeres, club slightly sinuate, much longer than shaft. Labrum produced beyond clypeus, setose, horizontal, transverse. Clypeus transverse, margin rounded in profile, weakly to moderately recurved. Clypeofrontal suture curved anteriorly, obscure in middle. Frons strongly punctate. Ocular canthi well developed, setose. Pronotum punctate, without or with only a trace of a median impressed line, basal ridge obsolete or very fine. Elytra heavily punctate, intervals costate, epipleurae nonmembranous, coarsely sculptured. Pygidium vertical, convex, with close, fine sculpturing. Thorax and abdomen with abundant long setae. Protibiae 3-dentate, without a spur, protarsi slender, tarsomeres 1–4 much longer than tarsomere 5, claws long, slender, sharp, subequal, larger with at least a trace of a minute tooth. Metatibiae 2-carinate, spurs long and sharp, metatarsi long and slender.
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</paragraph>
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<caption id="D0B4660BFF84FF8CFF1B1526FACCDAAD" ID-DOI="http://doi.org/10.5281/zenodo.4751853" ID-Zenodo-Dep="4751853" httpUri="https://zenodo.org/record/4751853/files/figure.png" pageId="3" pageNumber="366" startId="3.[151,264,1918,1942]" targetBox="[212,1364,181,1894]" targetPageId="3">
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<paragraph id="84743683FF84FF8CFF1B1526FACCDAAD" blockId="3.[151,1436,1918,2014]" pageId="3" pageNumber="366">
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<emphasis id="B6BFEA91FF84FF8CFF1B1526FE8BDAE5" bold="true" box="[151,314,1918,1942]" pageId="3" pageNumber="366">FIGURES 1–6.</emphasis>
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<taxonomicName id="43CB4D00FF84FF8CFECC1526FD0FDAE5" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[320,702,1918,1942]" class="Insecta" family="Rutelidae" genus="Amblochilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis id="B6BFEA91FF84FF8CFECC1526FDBCDAE5" box="[320,525,1918,1942]" italics="true" pageId="3" pageNumber="366">Amblochilus bicolor</emphasis>
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<bibRefCitation id="E05A4B72FF84FF8CFD9F1526FD0FDAE5" author="Blanchard, C. E." box="[531,702,1918,1942]" pageId="3" pageNumber="366" refId="ref7530" refString="Blanchard, C. E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des Insectes. Ordre des Coleopteres. Vol. 2. Museum d'Histoire Naturelle, Paris, pp. 129 - 240." type="book" year="1851">Blanchard, 1851</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="B6BFEA91FF84FF8CFD4B1526FD5CDAE5" bold="true" box="[711,749,1918,1942]" pageId="3" pageNumber="366">1–2</emphasis>
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, male lectotype:
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<emphasis id="B6BFEA91FF84FF8CFC161526FC16DAE5" bold="true" box="[922,935,1918,1942]" pageId="3" pageNumber="366">1</emphasis>
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, lateral view;
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<emphasis id="B6BFEA91FF84FF8CFBBA1526FBF2DAE5" bold="true" box="[1078,1091,1918,1942]" pageId="3" pageNumber="366">2</emphasis>
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, labels;
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<emphasis id="B6BFEA91FF84FF8CFB1B1526FB0CDAE5" bold="true" box="[1175,1213,1918,1942]" pageId="3" pageNumber="366">3–5</emphasis>
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, male:
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<emphasis id="B6BFEA91FF84FF8CFA8B1526FAA5DAE5" bold="true" box="[1287,1300,1918,1942]" pageId="3" pageNumber="366">3</emphasis>
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, dorsal view;
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<emphasis id="B6BFEA91FF84FF8CFF1B15FAFF15DAC9" bold="true" box="[151,164,1954,1978]" pageId="3" pageNumber="366">4</emphasis>
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, lateral view;
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<emphasis id="B6BFEA91FF84FF8CFEBF15FAFEF1DAC9" bold="true" box="[307,320,1954,1978]" pageId="3" pageNumber="366">5</emphasis>
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, ventral view;
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<emphasis id="B6BFEA91FF84FF8CFE5B15FAFE55DAC9" bold="true" box="[471,484,1954,1978]" pageId="3" pageNumber="366">6</emphasis>
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, known distribution of
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<taxonomicName id="43CB4D00FF84FF8CFD5C15FAFC2CDAC9" authorityName="Blanchard" authorityYear="1851" box="[720,925,1954,1978]" class="Insecta" family="Rutelidae" genus="Amblochilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis id="B6BFEA91FF84FF8CFD5C15FAFC2CDAC9" box="[720,925,1954,1978]" italics="true" pageId="3" pageNumber="366">Amblochilus bicolor</emphasis>
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</taxonomicName>
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in southeastern Queensland and northeastern New South Wales. Figs. 1–2 photographed by Muséum national d’Histoire naturelle. Figs. 3
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<emphasis id="B6BFEA91FF84FF8CFB8A159EFBA2DAAD" bold="true" box="[1030,1043,1990,2014]" pageId="3" pageNumber="366">–</emphasis>
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5 photographed by Matthias Seidel.
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</paragraph>
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</caption>
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<paragraph id="84743683FF83FF8BFF4B1313FEFBDCDE" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="367">
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<emphasis id="B6BFEA91FF83FF8BFF4B1313FE81DC16" bold="true" box="[199,304,331,357]" pageId="4" pageNumber="367">Females.</emphasis>
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Only that of
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<taxonomicName id="43CB4D00FF83FF8BFE451313FDFADC17" authorityName="Carne" authorityYear="1958" box="[457,587,331,356]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="costatus">
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<emphasis id="B6BFEA91FF83FF8BFE451313FDFADC17" box="[457,587,331,356]" italics="true" pageId="4" pageNumber="367">M. costatus</emphasis>
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</taxonomicName>
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is known and it has the antennal club and tarsi reduced relative to those of the male. Although he listed no females,
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<bibRefCitation id="E05A4B72FF83FF8BFDE11337FC8FDCFA" author="Carne, P. B." box="[621,830,366,393]" pageId="4" pageNumber="367" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne (1958: 198)</bibRefCitation>
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stated that the antennal club of males is much larger than in females.
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</paragraph>
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</subSubSection>
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<subSection id="ED442F68FF83FF8BFF1B13A7FA2CDEA4" pageId="4" pageNumber="367" type="key">
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<subSubSection id="CCD16508FF83FF8BFF1B13A7FE88DF6A" box="[151,313,511,537]" pageId="4" pageNumber="367" type="multiple">
|
||
<paragraph id="84743683FF83FF8BFF1B13A7FE88DF6A" blockId="4.[151,313,511,537]" box="[151,313,511,537]" pageId="4" pageNumber="367">
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||
<heading id="DF3C81EFFF83FF8BFF1B13A7FE88DF6A" bold="true" box="[151,313,511,537]" fontSize="11" level="1" pageId="4" pageNumber="367" reason="1">
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<emphasis id="B6BFEA91FF83FF8BFF1B13A7FE88DF6A" bold="true" box="[151,313,511,537]" pageId="4" pageNumber="367">Key to males</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCD16508FF83FF8BFF1B101CFA2CDEA4" pageId="4" pageNumber="367" type="key">
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<key id="F05A61B2FF83FF8BFF1B101CFA2CDEA4" pageId="4" pageNumber="367">
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<keyStep id="3F3F2E26FF83FF8BFF1B101CFA2DDE7A" pageId="4" pageNumber="367">
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<paragraph id="84743683FF83FF8BFF1B101CFA2DDFE7" blockId="4.[151,1437,580,983]" pageId="4" pageNumber="367">
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<keyLead id="3F3A95B6FF83FF8BFF1B101CFA2DDFE7" pageId="4" pageNumber="367">
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1 Pronotum brown to black with a green or coppery-green sheen; elytra with striae pale yellowish brown, costate intervals pronounced, deep brown (
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<figureCitation id="1CF02A06FF83FF8BFE311039FE49DF04" box="[445,504,609,631]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Fig. 9</figureCitation>
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); larger claws without a minute tooth; highlands of southeastern
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<collectingRegion id="460FF861FF83FF8BFBEB1039FB69DF04" box="[1127,1240,609,631]" country="Australia" name="Queensland" pageId="4" pageNumber="367">Queensland</collectingRegion>
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......................................................................................
|
||
<taxonomicName id="43CB4D00FF83FF8BFBCB1026FA2DDFE7" authority="Carne, 1958" authorityName="Carne" authorityYear="1958" box="[1095,1436,638,660]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="B6BFEA91FF83FF8BFBCB1026FA90DFE7" box="[1095,1313,638,660]" italics="true" pageId="4" pageNumber="367">Mesystoechus costatus</emphasis>
|
||
<bibRefCitation id="E05A4B72FF83FF8BFAAB1026FA2DDFE7" author="Carne, P. B." box="[1319,1436,638,660]" pageId="4" pageNumber="367" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="84743683FF83FF8BFF1B10C4FA2DDE7A" blockId="4.[151,1437,580,983]" pageId="4" pageNumber="367">
|
||
<keyLead id="3F3A95B6FF83FF8BFF1B10C4FA2DDE7A" pageId="4" pageNumber="367">- Pronotum with a variable pattern of dark yellowish black, fully developed consisting at least of a median longitudinal zone broadening posteriorly and a pair of symmetrical lateral curved bands and a lateral patch on each side; elytra with intervals not differentially pigmented, intervals weakly costate; larger claws of mid tarsi with at least a trace of a minute tooth at about half to two-thirds length................................................................................... 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="3F3F2E26FF83FF8BFF1B1149FA2CDEA4" pageId="4" pageNumber="367">
|
||
<paragraph id="84743683FF83FF8BFF1B1149FA2DDE0D" blockId="4.[151,1437,580,983]" pageId="4" pageNumber="367">
|
||
<keyLead id="3F3A95B6FF83FF8BFF1B1149FA2DDE0D" pageId="4" pageNumber="367">
|
||
2 Clypeus and V-shaped central section of frons yellowish brown, lateral frons dark brown to black (
|
||
<figureCitation id="1CF02A06FF83FF8BFB131149FB6ADE55" box="[1183,1243,785,807]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Fig. 7</figureCitation>
|
||
); mentum glabrous, anterior margin evenly convex; anterior margin of upper section of clypeus weakly recurved (
|
||
<figureCitation id="1CF02A06FF83FF8BFBD11176FB29DE37" box="[1117,1176,814,836]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Fig. 7</figureCitation>
|
||
); anterior margin of pronotum glabrous; coastal southeastern
|
||
<collectingRegion id="460FF861FF83FF8BFDBF1114FD15DE11" box="[563,676,844,866]" country="Australia" name="Queensland" pageId="4" pageNumber="367">Queensland</collectingRegion>
|
||
and northeastern
|
||
<collectingRegion id="460FF861FF83FF8BFCDC1114FC4ADE11" box="[848,1019,844,866]" country="Australia" name="New South Wales" pageId="4" pageNumber="367">New South Wales</collectingRegion>
|
||
.....................................................................................................
|
||
<taxonomicName id="43CB4D00FF83FF8BFB941131FA2DDE0D" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[1048,1436,873,895]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="ciliatus">
|
||
<emphasis id="B6BFEA91FF83FF8BFB941131FB5BDE0D" box="[1048,1258,873,894]" italics="true" pageId="4" pageNumber="367">Mesystoechus ciliatus</emphasis>
|
||
<bibRefCitation id="E05A4B72FF83FF8BFB7C1131FA2DDE0D" author="Waterhouse, C. O." box="[1264,1436,873,895]" pageId="4" pageNumber="367" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="84743683FF83FF8BFF1B11DFFA2CDEA4" blockId="4.[151,1437,580,983]" pageId="4" pageNumber="367">
|
||
<keyLead id="3F3A95B6FF83FF8BFF1B11DFFA2CDEA4" pageId="4" pageNumber="367">
|
||
- Clypeus and frons black, with indistinct browner median patch (
|
||
<figureCitation id="1CF02A06FF83FF8BFCD211DEFC17DEEF" box="[862,934,902,924]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Fig. 11</figureCitation>
|
||
); mentum with long setae, anterior margin slightly concave and notched in middle; anterior margin of upper section of clypeus strongly recurved (
|
||
<figureCitation id="1CF02A06FF83FF8BFB0211FCFB67DEC9" box="[1166,1238,932,954]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Fig. 11</figureCitation>
|
||
); anterior margin of pronotum with setae near angles; inland southeastern
|
||
<collectingRegion id="460FF861FF83FF8BFD691199FCE7DEA4" box="[741,854,961,983]" country="Australia" name="Queensland" pageId="4" pageNumber="367">Queensland</collectingRegion>
|
||
..........
|
||
<taxonomicName id="43CB4D00FF83FF8BFC681199FA91DEA4" authority="Allsopp" authorityName="Allsopp" authorityYear="2021" box="[996,1312,961,983]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="lithgowae" status="sp. nov.">
|
||
<emphasis id="B6BFEA91FF83FF8BFC681199FB7EDEA5" box="[996,1231,961,982]" italics="true" pageId="4" pageNumber="367">Mesystoechus lithgowae</emphasis>
|
||
Allsopp
|
||
</taxonomicName>
|
||
<emphasis id="B6BFEA91FF83FF8BFAAA1199FA2CDEA4" bold="true" box="[1318,1437,961,983]" pageId="4" pageNumber="367">
|
||
<taxonomicNameLabel id="AD8C57EAFF83FF8BFAAA1199FA2CDEA4" box="[1318,1437,961,983]" pageId="4" pageNumber="367" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |