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<document id="7F18F5CEDC86D51C65064D5ED9BC05AC" ID-CLB-Dataset="7002" ID-DOI="10.11646/zootaxa.4965.2.9" ID-GBIF-Dataset="73b8e644-f46f-4526-9b4e-b323f2205c8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4751849" ID-ZooBank="C0F65518-6EA9-4099-AEA2-51C5EECFFC78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619710709549" checkinUser="plazi" docAuthor="Allsopp, Peter G." docDate="2021" docId="0C628795FF83FF8AFF1B1671FCB3DE4A" docLanguage="en" docName="zootaxa.4965.2.9.pdf" docOrigin="Zootaxa 4965 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesystoechus ciliatus Waterhouse 1878" docType="treatment" docVersion="7" lastPageNumber="368" masterDocId="F05BFFEDFF87FF8FFF8C1258FFB1DD73" masterDocTitle="The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)" masterLastPageNumber="374" masterPageNumber="363" pageNumber="367" updateTime="1698938014406" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3EE096043AF23BBEB9E5FAC8D156ED91" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6383E0D86EE774C7436E87FA7A566C13">
<mods:title id="5DD142580343091016EB427F4623193C">The Australian endemic genera Mesystoechus Waterhouse, 1878, Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited (Coleoptera Scarabaeidae: Rutelinae: Anoplognathini)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="AA9A56E4257EA6A3CD40CD4292AE0966">Allsopp, Peter G.</mods:namePart>
</mods:name>
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<mods:title id="A4E755AACD0EBB39C6D5B64106A28C1C">Zootaxa</mods:title>
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<mods:part id="E4F7ECB5788AE29BDE0C10BE2113B664">
<mods:date id="5C09F3415B85C8FF45FAE1F08B4580F2">2021</mods:date>
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<mods:number id="8DDFD4F7A3AF5FC0F053BFF4973BA07D">2021-04-28</mods:number>
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<mods:number id="2F7324DD9258CCF971EE4A85F6586BB3">4965</mods:number>
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<mods:number id="28D26B43AB00E675B0EC9ABB52D09A1F">2</mods:number>
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<mods:classification id="1EFBA8320E6110AE80C22C08364EA203">journal article</mods:classification>
<mods:identifier id="B245F3997038182C81E664E15E1FEBD5" type="CLB-Dataset">7002</mods:identifier>
<mods:identifier id="A80CEE82DA68EC1E89B9F3232693E2C5" type="DOI">10.11646/zootaxa.4965.2.9</mods:identifier>
<mods:identifier id="BCF0EADD1DE9951D6B016009CC393DC0" type="GBIF-Dataset">73b8e644-f46f-4526-9b4e-b323f2205c8d</mods:identifier>
<mods:identifier id="1051B694E9C995B2FC570E7D47AB9883" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1E8B618C639D3E711974B3E5B60515C1" type="Zenodo-Dep">4751849</mods:identifier>
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</mods:mods>
<treatment id="0C628795FF83FF8AFF1B1671FCB3DE4A" ID-DOI="http://doi.org/10.5281/zenodo.4751769" ID-GBIF-Taxon="181983391" ID-Zenodo-Dep="4751769" LSID="urn:lsid:plazi:treatment:0C628795FF83FF8AFF1B1671FCB3DE4A" httpUri="http://treatment.plazi.org/id/0C628795FF83FF8AFF1B1671FCB3DE4A" lastPageId="5" lastPageNumber="368" pageId="4" pageNumber="367">
<subSubSection id="CCD16508FF83FF8BFF1B1671FD34D930" box="[151,645,1065,1092]" pageId="4" pageNumber="367" type="nomenclature">
<paragraph id="84743683FF83FF8BFF1B1671FD34D930" blockId="4.[151,645,1065,1092]" box="[151,645,1065,1092]" pageId="4" pageNumber="367">
<heading id="DF3C81EFFF83FF8BFF1B1671FD34D930" bold="true" box="[151,645,1065,1092]" fontSize="11" level="1" pageId="4" pageNumber="367" reason="1">
<taxonomicName id="43CB4D00FF83FF8BFF1B1671FD34D930" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[151,645,1065,1092]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B6BFEA91FF83FF8BFF1B1671FD34D930" bold="true" box="[151,645,1065,1092]" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF1B1671FE2FD937" bold="true" box="[151,414,1065,1092]" italics="true" pageId="4" pageNumber="367">Mesystoechus ciliatus</emphasis>
<bibRefCitation id="E05A4B72FF83FF8BFE291671FD34D930" author="Waterhouse, C. O." box="[421,645,1065,1091]" pageId="4" pageNumber="367" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCD16508FF83FF8BFF1B1636FDDAD9F5" box="[151,619,1134,1158]" pageId="4" pageNumber="367" type="reference_group">
<paragraph id="84743683FF83FF8BFF1B1636FDDAD9F5" blockId="4.[151,619,1134,1158]" box="[151,619,1134,1158]" pageId="4" pageNumber="367">
<treatmentCitationGroup id="A4DB11ADFF83FF8BFF1B1636FDDAD9F5" box="[151,619,1134,1158]" pageId="4" pageNumber="367">
<treatmentCitation id="056A1092FF83FF8BFF1B1636FDD6D9F5" author="Waterhouse, C. O." box="[151,615,1134,1158]" page="228" pageId="4" pageNumber="367" year="1878">
<taxonomicName id="43CB4D00FF83FF8BFF1B1636FDD6D9F5" authority="Waterhouse, 1878: 228" authorityName="Waterhouse" authorityPageNumber="228" authorityYear="1878" box="[151,615,1134,1158]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B6BFEA91FF83FF8BFF1B1636FEC4D9F5" box="[151,373,1134,1158]" italics="true" pageId="4" pageNumber="367">Mesystoechus ciliatus</emphasis>
<bibRefCitation id="E05A4B72FF83FF8BFEF71636FDD6D9F5" author="Waterhouse, C. O." box="[379,615,1134,1158]" pageId="4" pageNumber="367" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse, 1878: 228</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CCD16508FF83FF8BFF1B16ECFF54DB45" pageId="4" pageNumber="367" type="materials_examined">
<paragraph id="84743683FF83FF8BFF1B16ECFA38D865" blockId="4.[151,1437,1203,2023]" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF1B16ECFE93D9BD" bold="true" box="[151,290,1204,1230]" pageId="4" pageNumber="367">Type series.</emphasis>
<materialsCitation id="34A33CDEFF83FF8BFEA416ECFA38D865" ID-GBIF-Occurrence="3110759334" collectionCode="NHML" collectorName="Type C. Waterh. &amp; P. Allsopp" country="Australia" location="Moreton Bay" pageId="4" pageNumber="367" specimenCount="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="lectotype">
<emphasis id="B6BFEA91FF83FF8BFEA416ECFD3AD9BD" bold="true" box="[296,651,1204,1232]" pageId="4" pageNumber="367">
<typeStatus id="5B708821FF83FF8BFEA416ECFE2FD9BD" box="[296,414,1204,1230]" pageId="4" pageNumber="367" type="lectotype">Lectotype</typeStatus>
<specimenCount id="92CDFD0AFF83FF8BFE2816EDFE08D9A3" box="[420,441,1205,1232]" pageId="4" pageNumber="367" type="male">♂</specimenCount>
(here designated)
</emphasis>
:
<collectingCountry id="FCDC7613FF83FF8BFD1416ECFC8AD9BD" box="[664,827,1204,1230]" name="Australia" pageId="4" pageNumber="367">AUSTRALIA</collectingCountry>
:
<collectingRegion id="460FF861FF83FF8BFCC816ECFBB4D9BD" box="[836,1029,1204,1230]" country="Australia" name="Queensland" pageId="4" pageNumber="367">QUEENSLAND</collectingRegion>
: Type (red-ringed, circular, printed] |
<location id="81146058FF83FF8BFF1B1681FE9DD980" LSID="urn:lsid:plazi:treatment:0C628795FF83FF8AFF1B1671FCB3DE4A:81146058FF83FF8BFF1B1681FE9DD980" box="[151,300,1241,1267]" country="Australia" name="Moreton Bay" pageId="4" pageNumber="367" stateProvince="Queensland">Moreton Bay</location>
, 61 53 [violet, printed] | dissected mouthparts |
<taxonomicName id="43CB4D00FF83FF8BFCB21680FB84D981" authorityName="Waterhouse" authorityYear="1878" box="[830,1077,1240,1266]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="ciliatus">Mesystoechus ciliatus</taxonomicName>
<collectorName id="293E5355FF83FF8BFBB71681FB40D981" box="[1083,1265,1240,1267]" pageId="4" pageNumber="367">Type C. Waterh.</collectorName>
[handwritten] |
<typeStatus id="5B708821FF83FF8BFF1B16A4FE86D865" box="[151,311,1276,1302]" pageId="4" pageNumber="367" type="lectotype">LECTOTYPE</typeStatus>
<specimenCount id="92CDFD0AFF83FF8BFEB216A5FEE4D864" box="[318,341,1277,1303]" pageId="4" pageNumber="367" type="male">♂</specimenCount>
,
<taxonomicName id="43CB4D00FF83FF8BFEED16A4FD51D865" authority="Waterhouse" authorityName="Waterhouse" authorityYear="1878" box="[353,736,1276,1302]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B6BFEA91FF83FF8BFEED16A4FDE4D865" box="[353,597,1276,1302]" italics="true" pageId="4" pageNumber="367">Mesystoechus ciliatus</emphasis>
Waterhouse
</taxonomicName>
des.
<collectorName id="293E5355FF83FF8BFC9716A5FC3CD865" box="[795,909,1276,1303]" pageId="4" pageNumber="367">P. Allsopp</collectorName>
2021 [yellow highlighted, printed] [
<collectionCode id="E2DAAE46FF83FF8BFAAF16A3FACFD865" box="[1315,1406,1275,1302]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="367">NHML</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="84743683FF83FF8BFF4B1778FF54DB45" blockId="4.[151,1437,1203,2023]" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF4B1778FDB6D849" bold="true" box="[199,519,1312,1338]" pageId="4" pageNumber="367">Other material examined.</emphasis>
<materialsCitation id="34A33CDEFF83FF8BFD9F1778FA91D849" ID-GBIF-Occurrence="3110759337" box="[531,1312,1311,1339]" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="R. Illidge" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<collectingCountry id="FCDC7613FF83FF8BFD9F1778FD04D849" box="[531,693,1312,1338]" name="Australia" pageId="4" pageNumber="367">AUSTRALIA</collectingCountry>
.
<collectingRegion id="460FF861FF83FF8BFD491778FC37D849" box="[709,902,1312,1338]" country="Australia" name="Queensland" pageId="4" pageNumber="367">QUEENSLAND</collectingRegion>
:
<specimenCount id="92CDFD0AFF83FF8BFC1A1778FC0DD849" box="[918,956,1312,1338]" pageId="4" pageNumber="367" type="male">2♂</specimenCount>
,
<collectorName id="293E5355FF83FF8BFC411778FBEAD848" box="[973,1115,1312,1339]" pageId="4" pageNumber="367">Brisbane, R</collectorName>
. Illidge [
<collectionCode id="E2DAAE46FF83FF8BFB441747FAA4D849" box="[1224,1301,1311,1338]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="4" pageNumber="367">UQIC</collectionCode>
];
</materialsCitation>
<materialsCitation id="34A33CDEFF83FF8BFAA11778FD6BD82D" ID-GBIF-Occurrence="3110759333" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="M. Permiakoff " country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFAA11778FAE3D849" box="[1325,1362,1312,1338]" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
, Brisbane,
<date id="F0751043FF83FF8BFF57171CFEF9D82C" box="[219,328,1348,1375]" pageId="4" pageNumber="367" value="1957-11-03">
<collectingDate id="E031E9ABFF83FF8BFF57171CFEF9D82C" box="[219,328,1348,1375]" pageId="4" pageNumber="367" value="1957-11-03">3.xi.1957</collectingDate>
</date>
,
<collectorName id="293E5355FF83FF8BFED9171DFE4FD82D" box="[341,510,1348,1375]" pageId="4" pageNumber="367">M. Permiakoff</collectorName>
, sweeping [
<collectionCode id="E2DAAE46FF83FF8BFD0E171BFD7ED82D" box="[642,719,1347,1374]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="4" pageNumber="367">UQIC</collectionCode>
];
</materialsCitation>
<materialsCitation id="34A33CDEFF83FF8BFD6F171CFA2CD8F1" ID-GBIF-Occurrence="3110759338" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="M. De Baar " country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFD6F171CFCA7D82D" box="[739,790,1348,1374]" pageId="4" pageNumber="367" type="male">13♂</specimenCount>
, Corinda, Brisbane,
<collectingDate id="E031E9ABFF83FF8BFB8C171CFBB9D8F1" isEnumeration="true" pageId="4" pageNumber="367">
<date id="F0751043FF83FF8BFB8C171CFBCBD82D" box="[1024,1146,1348,1374]" pageId="4" pageNumber="367" value="1991-11-29">29.xi.1991</date>
,
<date id="F0751043FF83FF8BFB0B171CFAB9D82C" box="[1159,1288,1348,1375]" pageId="4" pageNumber="367" value="1991-12-15">15.xii.1991</date>
,
<date id="F0751043FF83FF8BFA9A171CFA26D82D" box="[1302,1431,1348,1374]" pageId="4" pageNumber="367" value="1991-12-17">17.xii.1991</date>
,
<date id="F0751043FF83FF8BFF1B1730FEB2D8F1" box="[151,259,1384,1410]" pageId="4" pageNumber="367" value="1992-11-06">6.xi.1992</date>
,
<date id="F0751043FF83FF8BFE9D1730FECCD8F1" box="[273,381,1384,1410]" pageId="4" pageNumber="367" value="1992-11-08">8.xi.1992</date>
,
<date id="F0751043FF83FF8BFE071730FDB4D8F1" box="[395,517,1384,1410]" pageId="4" pageNumber="367" value="1992-11-23">23.xi.1992</date>
,
<date id="F0751043FF83FF8BFD9E1730FDCFD8F1" box="[530,638,1384,1410]" pageId="4" pageNumber="367" value="1993-11-02">2.xi.1993</date>
,
<date id="F0751043FF83FF8BFD001730FD49D8F1" box="[652,760,1384,1410]" pageId="4" pageNumber="367" value="1993-11-03">3.xi.1993</date>
,
<date id="F0751043FF83FF8BFC8A1730FC31D8F1" box="[774,896,1384,1410]" pageId="4" pageNumber="367" value="1993-11-26">26.xi.1993</date>
,
<date id="F0751043FF83FF8BFC021730FBB9D8F1" box="[910,1032,1384,1410]" pageId="4" pageNumber="367" value="1994-11-24">24.xi.1994</date>
</collectingDate>
,
<collectorName id="293E5355FF83FF8BFB991731FB2CD8F0" box="[1045,1181,1385,1411]" pageId="4" pageNumber="367">M. De Baar</collectorName>
[
<collectionCode id="E2DAAE46FF83FF8BFB201730FB47D8F1" box="[1196,1270,1384,1410]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="367">ANIC</collectionCode>
,
<collectionCode id="E2DAAE46FF83FF8BFA8F1730FAE0D8F1" box="[1283,1361,1384,1410]" country="Australia" lsid="urn:lsid:biocol.org:col:34157" name="Queensland Primary Industries Insect Collection" pageId="4" pageNumber="367" type="Other type of research institution/biorepository">QDPC</collectionCode>
,
<collectionCode id="E2DAAE46FF83FF8BFAD2173FFA20D8F1" box="[1374,1425,1383,1410]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="4" pageNumber="367" type="Museum">QM</collectionCode>
];
</materialsCitation>
<materialsCitation id="34A33CDEFF83FF8BFF1B17D4FDD3D8D5" ID-GBIF-Occurrence="3110759321" box="[151,610,1419,1447]" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="R. Illidge" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFF1B17D4FF0DD8D5" box="[151,188,1420,1446]" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
, Darra,
<collectorName id="293E5355FF83FF8BFE9617D4FE15D8D4" box="[282,420,1420,1447]" pageId="4" pageNumber="367">Brisbane, R</collectorName>
. Illidge [
<collectionCode id="E2DAAE46FF83FF8BFD8617D3FDE6D8D5" box="[522,599,1419,1446]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="4" pageNumber="367">UQIC</collectionCode>
];
</materialsCitation>
<materialsCitation id="34A33CDEFF83FF8BFDE717D4FBBBD8D5" ID-GBIF-Occurrence="3110759304" box="[619,1034,1419,1447]" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFDE717D4FD20D8D5" box="[619,657,1420,1446]" pageId="4" pageNumber="367" type="male">6♂</specimenCount>
, no further data [
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,
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<collectingRegion id="460FF861FF83FF8BFB9F17D5FA95D8D5" box="[1043,1316,1420,1447]" country="Australia" name="New South Wales" pageId="4" pageNumber="367">NEW SOUTH WALES</collectingRegion>
:
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, Clarence River [
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];
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,
<collectorName id="293E5355FF83FF8BFE2717E8FDECD8B9" box="[427,605,1456,1482]" pageId="4" pageNumber="367">Ulmarra, S.L.S.</collectorName>
[
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];
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<specimenCount id="92CDFD0AFF83FF8BFD5617E8FD4ED8B9" box="[730,767,1456,1482]" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
, Grafton, xii.94, N.H. [
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];
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<materialsCitation id="34A33CDEFF83FF8BFBED17E8FE87D89D" ID-GBIF-Occurrence="3110759320" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="M. Watt" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFBED17E8FB37D8B9" box="[1121,1158,1456,1482]" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
, Maclean,
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<collectingDate id="E031E9ABFF83FF8BFA8F17E8FADED8B8" box="[1283,1391,1456,1483]" pageId="4" pageNumber="367" value="1959-01-25">25.i.1959</collectingDate>
</date>
,
<collectorName id="293E5355FF83FF8BFAF017E9FF7AD89C" pageId="4" pageNumber="367">M. Watt</collectorName>
[
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];
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<materialsCitation id="34A33CDEFF83FF8BFEB3178CFB0AD89D" ID-GBIF-Occurrence="3110759306" box="[319,1211,1491,1519]" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" collectorName="R. H. Eldridge" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFEB3178CFED4D89D" box="[319,357,1492,1518]" pageId="4" pageNumber="367" type="male">2♂</specimenCount>
,
<collectorName id="293E5355FF83FF8BFEFE178CFDBBD89D" box="[370,522,1492,1518]" pageId="4" pageNumber="367">Southgate SF</collectorName>
[
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], Grafton,
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<collectingDate id="E031E9ABFF83FF8BFCAF178CFC14D89D" box="[803,933,1492,1518]" pageId="4" pageNumber="367" value="1984-12-18">18.xii.1984</collectingDate>
</date>
,
<collectorName id="293E5355FF83FF8BFC3E178DFBE1D89D" box="[946,1104,1492,1519]" pageId="4" pageNumber="367">R.H. Eldridge</collectorName>
[
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];
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<materialsCitation id="34A33CDEFF83FF8BFB48178CFE43DB61" ID-GBIF-Occurrence="3110759318" collectingDate="1957-11-03" collectingDateMax="1984-12-18" collectingDateMin="1957-11-03" collectionCode="UQIC, ANIC, QDPC, QM, NHML, SAM, ASCT" country="Australia" location="State Forest" pageId="4" pageNumber="367" specimenCount="31" specimenCount-male="31" stateProvince="Queensland">
<specimenCount id="92CDFD0AFF83FF8BFB48178CFB58D89D" box="[1220,1257,1492,1518]" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
, Tabbimoble nr Woodburn,
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<collectingDate id="E031E9ABFF83FF8BFE9617A0FE23DB60" box="[282,402,1528,1555]" pageId="4" pageNumber="367" value="1957-11-15">15.xi.1957</collectingDate>
</date>
[
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]
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.
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<collectingCountry id="FCDC7613FF83FF8BFE7617A0FD2CDB61" box="[506,669,1528,1554]" name="Australia" pageId="4" pageNumber="367">AUSTRALIA</collectingCountry>
:
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, no further data [
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].
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:
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</quantity>
.
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[
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,
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,
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,
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.
</paragraph>
</subSubSection>
<subSubSection id="CCD16508FF83FF8BFF4B1418FE13DAB1" pageId="4" pageNumber="367" type="diagnosis">
<paragraph id="84743683FF83FF8BFF4B1418FE73DAEC" blockId="4.[151,1437,1203,2023]" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF4B1418FDA5DB29" bold="true" box="[199,532,1600,1626]" pageId="4" pageNumber="367">
Diagnosis. Male (
<figureCitation id="1CF02A06FF83FF8BFE1B1418FDB6DB29" box="[407,519,1600,1626]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Figs. 7–8</figureCitation>
).
</emphasis>
Body
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long; dorsum generally yellowish brown; clypeus and V- to Tshaped central section of frons yellowish brown, lateral frons dark brown to black; scutellum dark brownish black; pronotum with a variable pattern of dark yellowish black, fully developed it consists of a median longitudinal zone broadening posteriorly, a pair of symmetrical lateral curved bands and a pair of indistinct lateral patches, but darker colour may cover almost the entire pronotum or be reduced to a few darker spots on either side; abdominal tergites, sides of the coxae and femora light yellowish brown; pygidium, abdominal ventrites, and legs dark brown. Mentum with anterior margin evenly convex. Anterior margin of upper section of clypeus weakly recurved. Anterior margin of pronotum glabrous. Elytral intervals weakly costate. Larger claws with a minute tooth at about half length. Description by
<bibRefCitation id="E05A4B72FF83FF8BFEC81538FE62DA08" author="Carne, P. B." box="[324,467,1888,1915]" pageId="4" pageNumber="367" pagination="36 - 40" refId="ref7682" refString="Carne, P. B. (1954) Notes on the Australian Rutelinae (Coleoptera) and description of a new genus. Proceedings of the Royal Entomological Society of London B, 23, 36 - 40. https: // doi. org / 10.1111 / j. 1365 - 3113.1954. tb 00089. x" type="journal article" year="1954">Carne (1954</bibRefCitation>
: metatibia of male
<figureCitation id="1CF02A06FF83FF8BFD251538FD5CDA09" box="[681,749,1888,1914]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1918,1942]" captionTargetBox="[212,1364,181,1894]" captionTargetId="figure-15@3.[212,1375,181,1894]" captionTargetPageId="3" captionText="FIGURES 1–6. Amblochilus bicolor Blanchard, 1851. 1–2, male lectotype: 1, lateral view; 2, labels; 3–5, male: 3, dorsal view; 4, lateral view; 5, ventral view; 6, known distribution of Amblochilus bicolor in southeastern Queensland and northeastern New South Wales. Figs. 1–2 photographed by Muséum national d’Histoire naturelle. Figs. 3–5 photographed by Matthias Seidel." figureDoi="http://doi.org/10.5281/zenodo.4751853" httpUri="https://zenodo.org/record/4751853/files/figure.png" pageId="4" pageNumber="367">Fig. 6</figureCitation>
; parameres of male
<figureCitation id="1CF02A06FF83FF8BFC5C1538FB87DA09" box="[976,1078,1888,1914]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="4" pageNumber="367">Figs. 7-8</figureCitation>
);
<bibRefCitation id="E05A4B72FF83FF8BFBC71538FAADDA09" author="Carne, P. B." box="[1099,1308,1888,1915]" pageId="4" pageNumber="367" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne (1958: 217)</bibRefCitation>
;
<bibRefCitation id="E05A4B72FF83FF8BFAAB1538FF6DDAED" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." pageId="4" pageNumber="367" pagination="1919" refId="ref8231" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Rutelinae Macleay 1919 [sic]. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria, pp. 508 - 515." type="book chapter" year="2019">
Weir
<emphasis id="B6BFEA91FF83FF8BFAE81539FA2CDA09" box="[1380,1437,1888,1914]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
(2019
</bibRefCitation>
: dorsum Plate 68A).
</paragraph>
<paragraph id="84743683FF83FF8BFF4B15F0FE13DAB1" blockId="4.[151,1437,1203,2023]" box="[199,418,1960,1986]" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF4B15F0FE94DAB1" bold="true" box="[199,293,1960,1986]" pageId="4" pageNumber="367">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="CCD16508FF83FF8AFF4B1594FB1FDC32" lastPageId="5" lastPageNumber="368" pageId="4" pageNumber="367" type="discussion">
<paragraph id="84743683FF83FF8AFF4B1594FB1FDC32" blockId="4.[151,1437,1203,2023]" lastBlockId="5.[151,1437,150,825]" lastPageId="5" lastPageNumber="368" pageId="4" pageNumber="367">
<emphasis id="B6BFEA91FF83FF8BFF4B1594FE8ADA95" bold="true" box="[199,315,1996,2022]" pageId="4" pageNumber="367">Remarks.</emphasis>
The sex of the
<typeStatus id="5B708821FF83FF8BFE7D1594FDE2DA95" box="[497,595,1996,2022]" pageId="4" pageNumber="367" type="holotype">holotype</typeStatus>
was not stated explicitly in the original description,
<bibRefCitation id="E05A4B72FF83FF8BFB231594FAFDDA95" author="Ohaus, F." box="[1199,1356,1996,2022]" pageId="4" pageNumber="367" pagination="57 - 175" refId="ref8042" refString="Ohaus, F. (1904) Revision der Anoplognathiden. (Coleoptera Lamellicornia). Stettiner entomologische Zeitung, 65, 57 - 175." type="journal article" year="1904">Ohaus (1904)</bibRefCitation>
,
<bibRefCitation id="E05A4B72FF83FF8AFAD51594FF55DDC2" author="Carne, P. B." lastPageId="5" lastPageNumber="368" pageId="4" pageNumber="367" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne (1958)</bibRefCitation>
, or
<bibRefCitation id="E05A4B72FF82FF8AFE9E12CFFDBDDDC2" author="Cassis, G. &amp; Weir, T. A." box="[274,524,151,177]" pageId="5" pageNumber="368" refId="ref7784" refString="Cassis, G. &amp; Weir, T. A. (1992) Rutelinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Vol. 9. Australian Government Publishing Service, Canberra, pp. 359 - 382." type="book" year="1992">Cassis &amp; Weir (1992)</bibRefCitation>
;
<bibRefCitation id="E05A4B72FF82FF8AFD9612CFFD01DDC2" author="Carne, P. B." box="[538,688,150,177]" pageId="5" pageNumber="368" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne (1958)</bibRefCitation>
did not state whether he had examined the type, but he saw only males of this species.
<bibRefCitation id="E05A4B72FF82FF8AFE0412E3FDEDDDA6" author="Waterhouse, C. O." box="[392,604,187,213]" pageId="5" pageNumber="368" pagination="225 - 237" refId="ref8174" refString="Waterhouse, C. O. (1878) New Coleoptera from Australia and Tasmania, in the collection of the British Museum. Transactions of the Entomological Society of London, 1878, 225 - 237. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01527. x" type="journal article" year="1878">Waterhouse (1878)</bibRefCitation>
also did not state how many specimens he saw, and he gave no range in the length measurement. He did, however, say that “I have only dissected
<specimenCount id="92CDFD0AFF82FF8AFC121287FB87DD8A" box="[926,1078,223,249]" pageId="5" pageNumber="368" type="generic">one specimen</specimenCount>
...”, which can be taken to mean he had more than
<specimenCount id="92CDFD0AFF82FF8AFEED135BFE5DDC6E" box="[353,492,259,285]" pageId="5" pageNumber="368" type="generic" typeStatus="syntype">one syntype</specimenCount>
. There is only a specimen in NHML with dissected mouthparts and marked “Type”, and I designate it the
<typeStatus id="5B708821FF82FF8AFE0B137FFE5FDC32" box="[391,494,295,321]" pageId="5" pageNumber="368" type="lectotype">lectotype</typeStatus>
in order to stabilise nomenclature and have labelled it as such.
</paragraph>
</subSubSection>
<subSubSection id="CCD16508FF82FF8AFF4B1312FCB3DE4A" pageId="5" pageNumber="368" type="distribution">
<paragraph id="84743683FF82FF8AFF4B1312FE04DC86" blockId="5.[151,1437,150,825]" pageId="5" pageNumber="368">
<emphasis id="B6BFEA91FF82FF8AFF4B1312FE60DC17" bold="true" box="[199,465,330,356]" pageId="5" pageNumber="368">
Distribution (
<figureCitation id="1CF02A06FF82FF8AFEE71312FE73DC17" box="[363,450,330,356]" captionStart="FIGURES 7–15" captionStartId="6.[151,264,1869,1893]" captionTargetBox="[198,1389,181,1845]" captionTargetId="figure-20@6.[198,1389,181,1845]" captionTargetPageId="6" captionText="FIGURES 7–15. Mesystoechus species. 7–8, Mesystoechus ciliatus Waterhouse, 1878, male: 7, dorsal view; 8, lateral view; 9–10, Mesystoechus costatus Carne, 1958, male: 9, dorsal view; 10, lateral view; 11–14, Mesystoechus lithgowae, new species, male holotype: 11, dorsal view; 12, lateral view; 13, parameres, frontal view; 14, aedeagus, lateral view; 15, known distribution of Mesystoechus spp. in southeastern Queensland and northeastern New South Wales: yellow squares—M. ciliatus; blue triangles—M. costatus; red star—M. lithgowae. Figs. 7–14 photographed by Greg Daniels." figureDoi="http://doi.org/10.5281/zenodo.4751863" httpUri="https://zenodo.org/record/4751863/files/figure.png" pageId="5" pageNumber="368">Fig. 15</figureCitation>
).
</emphasis>
Known from coastal areas from the Brisbane area and south to the Clarence River/ Grafton area,
<collectingRegion id="460FF861FF82FF8AFEBF1337FE4DDCFA" box="[307,508,367,393]" country="Australia" name="New South Wales" pageId="5" pageNumber="368">New South Wales</collectingRegion>
(
<bibRefCitation id="E05A4B72FF82FF8AFD871337FD23DCFA" author="Carne, P. B." box="[523,658,366,393]" pageId="5" pageNumber="368" pagination="162 - 240" refId="ref7741" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. https: // doi. org / 10.1071 / ZO 9580162" type="journal article" year="1958">Carne 1958</bibRefCitation>
;
<bibRefCitation id="E05A4B72FF82FF8AFD121337FC8EDCFA" author="De Baar, M." box="[670,831,367,393]" pageId="5" pageNumber="368" pagination="20 - 21" refId="ref7841" refString="De Baar, M. (1992) Some observations on the ruteline beetle Mesystoechus ciliatus Waterhouse, 1878 (Scarabaeidae). News Bulletin, Entomological Society of Queensland, 20, 20 - 21." type="journal article" year="1992">De Baar 1992</bibRefCitation>
).
<bibRefCitation id="E05A4B72FF82FF8AFCDE1337FB0CDCFA" author="Atlas of Living Australia" box="[850,1213,367,393]" pageId="5" pageNumber="368" refId="ref7470" refString="Atlas of Living Australia (2019) Occurrence record: Entomology: K. 31398. Available from: https: // biocache. ala. org. au / occurrences / b 687 bafa-d 0 e 0 - 4 c 83 - 8 a 99 - b 2 e 852 b 741 fd (accessed 30 November 2019)" type="url" year="2019">
Atlas of Living
<collectingCountry id="FCDC7613FF82FF8AFB881337FBDADCFA" box="[1028,1131,367,393]" name="Australia" pageId="5" pageNumber="368">Australia</collectingCountry>
(2019)
</bibRefCitation>
records a specimen from ‘Mitchell River, New South Wales’—Mitchell River is an old name for the Mann River (
<geoCoordinate id="E1FF5044FF82FF8AFB3513CBFAA7DCDE" box="[1209,1302,403,429]" degrees="29.67" direction="south" orientation="latitude" pageId="5" pageNumber="368" precision="555" value="-29.67">29.67°S</geoCoordinate>
,
<geoCoordinate id="E1FF5044FF82FF8AFAAF13CAFA21DCDE" box="[1315,1424,402,429]" degrees="152.15" direction="east" orientation="longitude" pageId="5" pageNumber="368" precision="555" value="152.15">152.15°E</geoCoordinate>
), a tributary of the Clarence River (https://www.gnb.nsw.gov.au/__data/assets/pdf_file/0019/180532/7_LocalitiesNamesNSW1954_M-P.pdf).
</paragraph>
<paragraph id="84743683FF82FF8AFF4B13A7FCB3DE4A" blockId="5.[151,1437,150,825]" pageId="5" pageNumber="368">
<emphasis id="B6BFEA91FF82FF8AFF4B13A7FE37DF6B" bold="true" box="[199,390,510,537]" pageId="5" pageNumber="368">Natural history.</emphasis>
Adult males of
<taxonomicName id="43CB4D00FF82FF8AFDB313A7FD09DF6A" authorityName="Waterhouse" authorityYear="1878" box="[575,696,511,537]" class="Insecta" family="Rutelidae" genus="Mesystoechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B6BFEA91FF82FF8AFDB313A7FD09DF6A" box="[575,696,511,537]" italics="true" pageId="5" pageNumber="368">M. ciliatus</emphasis>
</taxonomicName>
are diurnal, actively flying during hot, sunny periods and resting during cloud cover (
<bibRefCitation id="E05A4B72FF82FF8AFEF7107BFDACDF4E" author="De Baar, M." box="[379,541,547,573]" pageId="5" pageNumber="368" pagination="20 - 21" refId="ref7841" refString="De Baar, M. (1992) Some observations on the ruteline beetle Mesystoechus ciliatus Waterhouse, 1878 (Scarabaeidae). News Bulletin, Entomological Society of Queensland, 20, 20 - 21." type="journal article" year="1992">De Baar 1992</bibRefCitation>
). Flight is rapid and maintained up to a height of
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and (rarely higher), but more usually lower above grass and in and out of thick hedges with leaf litter. Flight patterns are in straight lines or circular. De Baar’s observations during 1984–1991 were that all flew between 1000 and 1400 hr between 26 October and 14 November. Following a long dry period, emergences during 1991 were delayed until 29 November to 17 December and followed heavy storm rains. During 15–17 December there were very hot and steamy mornings and adults were active as early as 0900 hr but not in the afternoon. No females have been collected, and, based on De Baar’s observations of males dropping into leaf litter, females probably remain in the soil, emit pheromones, copulate on the surface of the soil, and oviposit nearby.
</paragraph>
</subSubSection>
</treatment>
</document>
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