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<document id="01FBDD26CF0F93BD0E6002818288B484" ID-CLB-Dataset="7126" ID-DOI="10.12782/sd.19.1.043" ID-GBIF-Dataset="238dd2d3-f38f-4e11-a2be-87c16ff21dd8" ID-ISSN="2189-7301" ID-Zenodo-Dep="4584938" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614946654621" checkinUser="felipe" docAuthor="Dwibadra, Dhian, Takaku, Gen, Ôhara, Masahiro &amp; Ueda, Akira" docDate="2014" docId="0C62878DFFEB215BFC90FD6CFEC2FD3A" docLanguage="en" docName="SpeciesDiversity.19.1.43-57.pdf" docOrigin="Species Diversity 19 (1)" docSource="http://dx.doi.org/10.12782/sd.19.1.043" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Macrocheles dayaci Dwibadra and Takaku 2014, sp. nov." docType="treatment" docUuid="13DA0BC7-5CB9-40BD-B0B5-5D3748706566" docUuidSource="ZooBank" docVersion="9" lastPageNumber="50" masterDocId="F05BFFF5FFEE215CFFF5FFE1FF89FFDD" masterDocTitle="Mites of the Family Macrochelidae (Acari: Gamasida) from Sungai Wain, East Kalimantan, Indonesia" masterLastPageNumber="57" masterPageNumber="43" pageNumber="48" updateTime="1698932732925" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B45EFD3C678C706408BDDC4D4B49A659">Mites of the Family Macrochelidae (Acari: Gamasida) from Sungai Wain, East Kalimantan, Indonesia</mods:title>
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<mods:namePart id="6E2E85C2F59971E8452767AA7044E60B">Dwibadra, Dhian</mods:namePart>
<mods:affiliation id="4E1CEC974E8728537E30D37757A69362">Systematic Entomology Laboratory, Graduate School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan E-mail: dwibadra _ yk @ yahoo. com &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="91BDD2356BD3AB083553F1A314B50A3B">Takaku, Gen</mods:namePart>
<mods:affiliation id="69060A99E1C9788AFC5493CBF4A47D42">Biological Laboratory, Hokkaido University of Education Sapporo, 5 - 3 - 1 Ainosato, Kita-ku, Sapporo 002 - 8502, Japan</mods:affiliation>
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<mods:namePart id="721B074453A7504183F6838681177EF2">Ôhara, Masahiro</mods:namePart>
<mods:affiliation id="78C4C5E41C0829A2B50EF4FB783DA4A9">The Hokkaido University Museum, N 10 W 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="E269612439292E8DF1FE97AB852BB4A8">Ueda, Akira</mods:namePart>
<mods:affiliation id="9DAF31779AFF3D3CF7A767FEEAB5D454">Kyushu Research Center, Forestry and Forest Products Research Institute (FFPRI), 4 - 11 - 16 Kurokami, Kumamoto 860 - 0862, Japan</mods:affiliation>
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<mods:title id="B1C692031E26297D3CCFA09816F171AB">Species Diversity</mods:title>
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<mods:date id="DF9F39C7C02C482D93F852EEF1198A81">2014</mods:date>
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<treatment id="0C62878DFFEB215BFC90FD6CFEC2FD3A" ID-DOI="http://doi.org/10.5281/zenodo.4730929" ID-GBIF-Taxon="181762127" ID-Zenodo-Dep="4730929" LSID="urn:lsid:zoobank.org:act:13DA0BC7-5CB9-40BD-B0B5-5D3748706566" httpUri="http://treatment.plazi.org/id/0C62878DFFEB215BFC90FD6CFEC2FD3A" lastPageId="7" lastPageNumber="50" pageId="5" pageNumber="48">
<subSubSection id="CCD16510FFEB2159FC90FD6CFB02FD1B" pageId="5" pageNumber="48" type="nomenclature">
<paragraph id="8474369BFFEB2159FC90FD6CFAE1FD78" blockId="5.[869,1384,653,710]" box="[869,1384,653,677]" pageId="5" pageNumber="48">
<heading id="DF3C81F7FFEB2159FC90FD6CFAE1FD78" box="[869,1384,653,677]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="48" reason="2">
<taxonomicName id="43CB4D18FFEB2159FC90FD6CFA9FFD78" authority="Dwibadra and Takaku" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[869,1302,653,677]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="dayaci" status="sp. nov.">
<emphasis id="B6BFEA89FFEB2159FC90FD6CFBA6FD78" bold="true" box="[869,1071,653,677]" italics="true" pageId="5" pageNumber="48">Macrocheles dayaci</emphasis>
Dwibadra and Takaku
</taxonomicName>
<taxonomicNameLabel id="AD8C57F2FFEB2159FAE9FD6FFAE1FD78" box="[1308,1384,654,677]" pageId="5" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8474369BFFEB2159FBB4FD4FFB02FD1B" blockId="5.[869,1384,653,710]" box="[1089,1163,686,710]" pageId="5" pageNumber="48">
<heading id="DF3C81F7FFEB2159FBB4FD4FFB02FD1B" box="[1089,1163,686,710]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="48" reason="2">
(
<figureCitation id="1CF02A1EFFEB2159FBBCFD4FFB08FD1B" box="[1097,1153,686,710]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFEB2159FCBBFD0EFAE0FC54" pageId="5" pageNumber="48" type="materials_examined">
<paragraph id="8474369BFFEB2159FCBBFD0EFAE0FC54" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
<emphasis id="B6BFEA89FFEB2159FCBBFD0EFC5BFCDA" bold="true" box="[846,978,751,775]" pageId="5" pageNumber="48">Type series.</emphasis>
<materialsCitation id="34A33CC6FFEB2159FC1EFD0EFC49FCB4" ID-GBIF-Occurrence="3095730318" collectingDate="2006-12-20" collectionCode="MZB" location="Balikpapan" municipality="Hutan Lindung Sungai Wain" pageId="5" pageNumber="48" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
<typeStatus id="5B708839FFEB2159FC1EFD0EFBC0FCDA" box="[1003,1097,751,775]" pageId="5" pageNumber="48" type="holotype">Holotype</typeStatus>
(
<collectionCode id="E2DAAE5EFFEB2159FB95FD0EFB10FCDA" box="[1120,1177,751,775]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="48" type="Museum">MZB</collectionCode>
. Acar. 7559.2): female,
<collectingMunicipality id="6410ACE1FFEB2159FCC1FCF1FBD7FCF5" box="[820,1118,784,808]" pageId="5" pageNumber="48">Hutan Lindung Sungai Wain</collectingMunicipality>
,
<location id="81146040FFEB2159FB9CFCF1FB54FCF5" LSID="urn:lsid:plazi:treatment:0C62878DFFEB215BFC90FD6CFEC2FD3A:81146040FFEB2159FB9CFCF1FB54FCF5" box="[1129,1245,784,808]" municipality="Hutan Lindung Sungai Wain" name="Balikpapan" pageId="5" pageNumber="48" stateProvince="Kalimantan Timur">Balikpapan</location>
,
<collectingRegion id="460FF879FFEB2159FB1CFCF1FA1DFCF5" box="[1257,1428,784,808]" country="Indonesia" name="Kalimantan Timur" pageId="5" pageNumber="48">East Kalimantan</collectingRegion>
,
<date id="F075105BFFEB2159FCC1FCD1FC70FC95" box="[820,1017,816,840]" pageId="5" pageNumber="48" value="2006-12-20">
<collectingDate id="E031E9B3FFEB2159FCC1FCD1FC70FC95" box="[820,1017,816,840]" pageId="5" pageNumber="48" value="2006-12-20">20 December 2006</collectingDate>
</date>
,
<emphasis id="B6BFEA89FFEB2159FBF0FCD0FB92FC95" box="[1029,1051,817,840]" italics="true" pageId="5" pageNumber="48">ex</emphasis>
<taxonomicName id="43CB4D18FFEB2159FBD1FCD1FC10FCB4" authority=", A. Ueda and D. Dwibadra" authorityName="A. Ueda and D. Dwibadra" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="dayacus">
<emphasis id="B6BFEA89FFEB2159FBD1FCD1FB64FC95" box="[1060,1261,816,840]" italics="true" pageId="5" pageNumber="48">Catharsius dayacus,</emphasis>
A. Ueda and D. Dwibadra
</taxonomicName>
leg
</materialsCitation>
.
<typeStatus id="5B708839FFEB2159FC3CFCB0FBA4FCB4" box="[969,1069,849,873]" pageId="5" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="92CDFD12FFEB2159FBC2FCB0FB07FCB4" box="[1079,1166,849,873]" pageId="5" pageNumber="48" type="female">1 female</specimenCount>
, data same as in
<typeStatus id="5B708839FFEB2159FACCFCB0FA1DFCB4" box="[1337,1428,849,873]" pageId="5" pageNumber="48" type="holotype">holotype</typeStatus>
;
<emphasis id="B6BFEA89FFEB2159FCC1FC90FCF3FC54" bold="true" box="[820,890,881,905]" pageId="5" pageNumber="48">Km 24</emphasis>
:
<specimenCount id="92CDFD12FFEB2159FC73FC90FC6EFC54" box="[902,999,881,905]" pageId="5" pageNumber="48" type="female">2 females</specimenCount>
,
<date id="F075105BFFEB2159FC05FC90FB26FC54" box="[1008,1199,881,905]" pageId="5" pageNumber="48" value="2006-12-26">26 December 2006</date>
,
<emphasis id="B6BFEA89FFEB2159FB4CFC93FB46FC54" box="[1209,1231,882,905]" italics="true" pageId="5" pageNumber="48">ex</emphasis>
residue in vial.
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFEB215BFCBBFC70FD80FE9D" lastPageId="7" lastPageNumber="50" pageId="5" pageNumber="48" type="description">
<paragraph id="8474369BFFEB2159FCBBFC70FB57FC36" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
<emphasis id="B6BFEA89FFEB2159FCBBFC70FC5BFC74" bold="true" box="[846,978,913,937]" pageId="5" pageNumber="48">Description.</emphasis>
<emphasis id="B6BFEA89FFEB2159FC1EFC73FBBBFC77" box="[1003,1074,914,938]" italics="true" pageId="5" pageNumber="48">Female</emphasis>
. Length of dorsal shield 1035 (10201050), width at level of coxae II 627.5 (600660) (
<emphasis id="B6BFEA89FFEB2159FCC8FC32FCC3FC37" box="[829,842,979,1002]" italics="true" pageId="5" pageNumber="48">n</emphasis>
=4). Specimens reddish brown in color.
</paragraph>
<paragraph id="8474369BFFEB2159FCBBFC15FC0BFB73" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
Dorsum (
<figureCitation id="1CF02A1EFFEB2159FC44FC12FB88FBD6" box="[945,1025,1011,1035]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1A</figureCitation>
). Dorsal shield broadly rounded posteriorly with procurved outline, surface of shield covered with strong punctations and reticulation; lateral margin of shield smooth; shield bearing 28 pairs of dorsal setae and 22 pairs of pores; seta j1 pilose distally; z1 simple; other setae pilose distally.
</paragraph>
<paragraph id="8474369BFFEB2159FCBBFB56FA92FA6F" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
Venter (
<figureCitation id="1CF02A1EFFEB2159FC57FB57FC7BFB12" box="[930,1010,1206,1231]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1B</figureCitation>
). Sternal shield wider than long; length 178.8 (175180), width at level of coxae II 207.5 (200210) (
<emphasis id="B6BFEA89FFEB2159FCC8FB19FCC3FAD2" box="[829,842,1272,1295]" italics="true" pageId="5" pageNumber="48">n</emphasis>
=4); shield with 3 pairs of simple setae and 2 pairs of pores; l. ang. and l. m. t. present; l.o.a. with faint punctation; two punctate l. arc. distinct; l. o. p. present as distinct line, with punctations, connected to l. m. t.; a. p. p. with strong and coarse punctations. Metasternal shields oval and free; each shield with 1 simple seta and anterior pore.
</paragraph>
<paragraph id="8474369BFFEB2159FCBBFA5AFB8DF9CE" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
Width of epigynial shield 273.8 (270275) (
<emphasis id="B6BFEA89FFEB2159FAE3FA5AFAAAFA0F" box="[1302,1315,1467,1490]" italics="true" pageId="5" pageNumber="48">n</emphasis>
=4); shield ornamented with punctations, with pair of simple setae on the posterior corner.
</paragraph>
<paragraph id="8474369BFFEB2159FCBBF9FDFC37F90B" blockId="5.[820,1433,751,2076]" pageId="5" pageNumber="48">
Ventrianal shield longer than wide, length 411.3 (400 420), width 317.5 (295340) (
<emphasis id="B6BFEA89FFEB2159FB94F9DFFBE7F988" box="[1121,1134,1598,1621]" italics="true" pageId="5" pageNumber="48">n</emphasis>
=4); shield with strong, dimpled reticulation and punctations; 3 pairs of preanal setae, 1 pair of paranal setae, and 1 postanal seta present; postanal seta pilose, other setae simple; cribrum located posterior to postanal seta.
</paragraph>
<paragraph id="8474369BFFEB215AFCBBF93EFF6BF821" blockId="5.[820,1433,751,2076]" lastBlockId="6.[156,767,1988,2077]" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
Gnathosoma (
<figureCitation id="1CF02A1EFFEB2159FC17F93EFBBAF925" box="[994,1075,1759,1784]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1C</figureCitation>
). Well developed and sclerotized. Deutosternal groove with 6 rows of denticles, anteriormost row divided; 3 pairs of simple hypostomal setae and 1 pair of simple palpcoxal setae present. Epistome (
<figureCitation id="1CF02A1EFFEB2159FAF8F8A0FAD1F884" box="[1293,1368,1857,1881]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1E</figureCitation>
) with median process and pair of lateral elements; median process bifurcate distally, and lacking minute spicules; basal margin serrate. Fixed digit of chelicera (
<figureCitation id="1CF02A1EFFEB2159FB8BF842FB45F866" box="[1150,1228,1955,1979]" captionStart="Fig" captionStartId="6.[156,187,1855,1877]" captionTargetBox="[185,1401,209,1815]" captionTargetId="figure-60@6.[184,1403,207,1817]" captionTargetPageId="6" captionText="Fig. 1. Macrocheles dayaci Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200µm (A, B); 50µm (CE)." figureDoi="http://doi.org/10.5281/zenodo.4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="5" pageNumber="48">Fig. 1D</figureCitation>
) with simple dorsal seta, robust median tooth, small distal tooth,
<emphasis id="B6BFEA89FFEB2159FAE7F822FA1BF806" box="[1298,1426,1987,2011]" italics="true" pageId="5" pageNumber="48">pilus dentilis</emphasis>
, and terminal hook; movable digit with robust bidentate median tooth, small distal tooth, and terminal hook; length of fixed digit 220 (200230) and movable digit 96.9 (95100) (
<emphasis id="B6BFEA89FFE8215AFF51F804FF38F821" box="[164,177,2021,2044]" italics="true" pageId="6" pageNumber="49">n</emphasis>
=4).
</paragraph>
<caption id="D0B46613FFE8215AFF69F8DEFE9CF850" ID-DOI="http://doi.org/10.5281/zenodo.4584940" ID-Zenodo-Dep="4584940" httpUri="https://zenodo.org/record/4584940/files/figure.png" pageId="6" pageNumber="49" startId="6.[156,187,1855,1877]" targetBox="[185,1401,209,1815]" targetPageId="6">
<paragraph id="8474369BFFE8215AFF69F8DEFE9CF850" blockId="6.[156,1432,1855,1933]" pageId="6" pageNumber="49">
Fig. 1.
<taxonomicName id="43CB4D18FFE8215AFF09F8DEFDF1F888" authority="Dwibadra and Takaku" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[252,632,1855,1877]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="dayaci" status="sp. nov.">
<emphasis id="B6BFEA89FFE8215AFF09F8DEFE2EF889" box="[252,423,1855,1876]" italics="true" pageId="6" pageNumber="49">Macrocheles dayaci</emphasis>
Dwibadra and Takaku
</taxonomicName>
,
<taxonomicNameLabel id="AD8C57F2FFE8215AFD74F8DEFD41F889" box="[641,712,1855,1876]" pageId="6" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype (MZB. Acar. 7559.2). A, dorsal shield; B, ventral shield (paratype, MZB. Acar. 7560.2); C, ventral view of gnathosoma; D, chelicera; E, tectum (paratype, MZB. Acar. 7559.1). Scale bars: 200
<emphasis id="B6BFEA89FFE8215AFAC0F8BDFAC8F8AC" box="[1333,1345,1884,1905]" italics="true" pageId="6" pageNumber="49">µ</emphasis>
m (A, B); 50
<emphasis id="B6BFEA89FFE8215AFF41F899FF49F850" box="[180,192,1912,1933]" italics="true" pageId="6" pageNumber="49">µ</emphasis>
m (CE).
</paragraph>
</caption>
<paragraph id="8474369BFFE8215BFF40F7E4FDF1FF22" blockId="6.[156,767,1988,2077]" lastBlockId="7.[156,767,198,743]" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
Legs. Most leg segments with simple and pilose setae except for coxae IIV; trochanter IIII and tarsus I with only simple setae. Leg chaetotaxy typical for this genus. Genu IV with 6 pilose setae. Leg length (excluding ambulacrum,
<emphasis id="B6BFEA89FFE9215BFF69FF26FF20FF03" box="[156,169,199,222]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=4); leg I, 758.8 (720830); leg II, 715.6 (647.5765); leg III, 702.5 (660760); leg IV, 1007.5 (9451045).
</paragraph>
<paragraph id="8474369BFFE9215BFF40FEE6FD9FFEC2" blockId="7.[156,767,198,743]" box="[181,534,263,287]" pageId="7" pageNumber="50">Sacculus foemineus. Not observed.</paragraph>
<paragraph id="8474369BFFE9215BFF40FEC9FD80FE9D" blockId="7.[156,767,198,743]" box="[181,521,296,320]" pageId="7" pageNumber="50">Male and other stages. Unknown.</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFF40FEA8FEEBFE7C" pageId="7" pageNumber="50" type="etymology">
<paragraph id="8474369BFFE9215BFF40FEA8FEEBFE7C" blockId="7.[156,767,198,743]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFF40FEA8FEA5FEBC" bold="true" box="[181,300,329,353]" pageId="7" pageNumber="50">Etymology.</emphasis>
The specific name is derived from the specific epithet of the phoretic host, the scarab beetle
<taxonomicName id="43CB4D18FFE9215BFF69FE68FED5FE7C" box="[156,348,393,417]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="dayacus">
<emphasis id="B6BFEA89FFE9215BFF69FE68FED5FE7C" box="[156,348,393,417]" italics="true" pageId="7" pageNumber="50">Catharsius dayacus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFF40FE4BFEC2FD3A" pageId="7" pageNumber="50" type="discussion">
<paragraph id="8474369BFFE9215BFF40FE4BFEC2FD3A" blockId="7.[156,767,198,743]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFF40FE4BFE90FE1F" bold="true" box="[181,281,426,450]" pageId="7" pageNumber="50">Remarks.</emphasis>
<taxonomicName id="43CB4D18FFE9215BFEC7FE4AFE11FE1F" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[306,408,426,450]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="dayaci">
<emphasis id="B6BFEA89FFE9215BFEC7FE4AFEC1FE1F" box="[306,328,427,450]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFEA2FE4BFE11FE1F" box="[343,408,426,450]" italics="true" pageId="7" pageNumber="50">dayaci</emphasis>
</taxonomicName>
is a member of the
<emphasis id="B6BFEA89FFE9215BFD87FE4BFD27FE1F" box="[626,686,426,450]" italics="true" pageId="7" pageNumber="50">glaber</emphasis>
species group and is similar to
<taxonomicName id="43CB4D18FFE9215BFE7BFE2AFF44FDDE" authority="Walter and Krantz, 1986" authorityName="Walter and Krantz" authorityYear="1986" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="witcoskyanus">
<emphasis id="B6BFEA89FFE9215BFE7BFE2AFE2DFE3F" box="[398,420,459,482]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFE47FE2AFDBCFE3E" box="[434,565,459,483]" italics="true" pageId="7" pageNumber="50">witcoskyanus</emphasis>
Walter and Krantz, 1986
</taxonomicName>
(
<bibRefCitation id="E05A4B6AFFE9215BFF2FFE0AFE57FDDE" author="Walter, D. E. &amp; Krantz, G. W." box="[218,478,491,515]" pageId="7" pageNumber="50" pagination="277 - 294" refId="ref14555" refString="Walter, D. E. and Krantz, G. W. 1986 b. A review of glaber - group (s. str.) species of the genus Macrocheles (Acari: Macrochelidae), and a discussion of species complexes. Acarologia 27: 277 - 294." type="journal article" year="1986">Walter and Krantz 1986b</bibRefCitation>
) in the shape of dorsal setae and shield sternal ornamentation. However,
<taxonomicName id="43CB4D18FFE9215BFD8EFDEDFD6AFDF9" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[635,739,524,548]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="dayaci">
<emphasis id="B6BFEA89FFE9215BFD8EFDEDFD18FDFE" box="[635,657,524,547]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFD57FDEDFD6AFDF9" box="[674,739,524,548]" italics="true" pageId="7" pageNumber="50">dayaci</emphasis>
</taxonomicName>
is distinguishable from
<taxonomicName id="43CB4D18FFE9215BFE81FDCCFD90FD99" authorityName="Walter and Krantz" authorityYear="1986" box="[372,537,556,580]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="witcoskyanus">
<emphasis id="B6BFEA89FFE9215BFE81FDCCFE03FD99" box="[372,394,557,580]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFE63FDCDFD90FD99" box="[406,537,556,580]" italics="true" pageId="7" pageNumber="50">witcoskyanus</emphasis>
</taxonomicName>
by the following characters (corresponding conditions of
<taxonomicName id="43CB4D18FFE9215BFDE7FDACFD30FDB8" authorityName="Walter and Krantz" authorityYear="1986" box="[530,697,589,613]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="witcoskyanus">
<emphasis id="B6BFEA89FFE9215BFDE7FDACFDA1FDB9" box="[530,552,589,612]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFDC3FDACFD30FDB8" box="[566,697,589,613]" italics="true" pageId="7" pageNumber="50">witcoskyanus</emphasis>
</taxonomicName>
in parentheses): 1) j4 and j5 pilose distally (simple); 2) z5 and j
<quantity id="43339B7EFFE9215BFF56FD6FFF44FD7B" box="[163,205,654,678]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="50" unit="in" value="6.0">6 in</quantity>
some cases slightly pilose (simple); and 3) ventrianal shield not expanded, longer than wide (expanded laterally, wider than long).
</paragraph>
</subSubSection>
</treatment>
</document>