treatments-xml/data/08/26/87/082687BEFFE0C57873CAFB7DFA3CA942.xml
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<document id="823D91FCBB6CB49CD35942CF166F9B3B" ID-CLB-Dataset="21785" ID-DOI="10.11646/zootaxa.4786.1.6" ID-GBIF-Dataset="3ad91afc-d278-46b7-acf3-2df2f9f71f1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3865007" ID-ZooBank="EE141AE8-282F-4035-B332-7F3D6E352EC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590737493716" checkinUser="plazi" docAuthor="Dantas, Galileu P. S., Pinheiro, Marcos P. G. &amp; Hamada, Neusa" docDate="2020" docId="082687BEFFE0C57873CAFB7DFA3CA942" docLanguage="en" docName="zootaxa.4786.1.6.pdf" docOrigin="Zootaxa 4786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pentaneura herbeti Dantas &amp; Pinheiro &amp; Hamada 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="89" masterDocId="F41FFFC6FFE1C570735DFFD3FFBEAF11" masterDocTitle="An unusual new species of Pentaneura Philippi (Diptera: Chironomidae) from northeastern Brazil, with an emended diagnosis to the genus" masterLastPageNumber="92" masterPageNumber="81" pageNumber="82" updateTime="1698841875954" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CC509F492A5C6ABA378D31A6412F0E64">An unusual new species of Pentaneura Philippi (Diptera: Chironomidae) from northeastern Brazil, with an emended diagnosis to the genus</mods:title>
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<mods:namePart id="DB3B21566B45842D86A7B3E002A4B20D">Dantas, Galileu P. S.</mods:namePart>
<mods:affiliation id="F634FDA8F72C40076DB61172BBD6283D">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt), Caixa Postal 478, 69067 - 375, Manaus, AM, Brazil</mods:affiliation>
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<mods:namePart id="18FD7203CB8BC0CE1CD72AD349E48736">Pinheiro, Marcos P. G.</mods:namePart>
<mods:affiliation id="58EA3AE567E1DD737F69965E667B2867">Universidade Federal do Rio Grande do Norte, Avenida Senador Salgado Filho, 3000, Natal, Rio Grande do Norte, Brazil mpgpinheiro @ gmail. com; https: // orcid. org / 0000 - 0002 - 8465 - 6570</mods:affiliation>
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<mods:namePart id="C0D93779574DF01F4E288EB929C0142A">Hamada, Neusa</mods:namePart>
<mods:affiliation id="D518E824B98411513AA7B6493DC02F92">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt), Caixa Postal 478, 69067 - 375, Manaus, AM, Brazil &amp; neusaha @ gmail. com; https: // orcid. org / 0000 - 0002 - 3526 - 5426</mods:affiliation>
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<mods:title id="5550A8DB8DC07BFC0766C1C1ECD8019D">Zootaxa</mods:title>
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<mods:date id="1734F9019865DF4A20872388F877991E">2020</mods:date>
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<treatment id="082687BEFFE0C57873CAFB7DFA3CA942" ID-DOI="http://doi.org/10.5281/zenodo.3866582" ID-GBIF-Taxon="164392532" ID-Zenodo-Dep="3866582" LSID="urn:lsid:plazi:treatment:082687BEFFE0C57873CAFB7DFA3CA942" httpUri="http://treatment.plazi.org/id/082687BEFFE0C57873CAFB7DFA3CA942" lastPageId="8" lastPageNumber="89" pageId="1" pageNumber="82">
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<heading id="DB7881C4FFE0C57173CAFB7DFE58ABD9" bold="true" box="[151,486,1197,1224]" fontSize="11" level="1" pageId="1" pageNumber="82" reason="1">
<emphasis id="B2FBEABAFFE0C57173CAFB7DFE58ABD9" bold="true" box="[151,486,1197,1224]" pageId="1" pageNumber="82">
<taxonomicName id="478F4D2BFFE0C57173CAFB7DFE3FABD9" ID-CoL="9HX65" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[151,385,1197,1224]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE0C57173CAFB7DFE3FABD9" bold="true" box="[151,385,1197,1224]" italics="true" pageId="1" pageNumber="82">Pentaneura herbeti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9C857C1FFE0C57172D5FB7DFE58ABD9" box="[392,486,1198,1224]" pageId="1" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<emphasis id="B2FBEABAFFE0C57173CAFB20FEFDAA1C" bold="true" box="[151,323,1266,1293]" pageId="1" pageNumber="82">Type material</emphasis>
.
<materialsCitation id="30E73CF5FFE0C571720BFB20FBC2AA3E" ID-GBIF-Occurrence="2628394301" collectingDate="2015-09-01" collectionCode="INPA" collectorName="G. P. S. Dantas &amp; M. P. G. Pinheiro" country="Brazil" latitude="-5.9584723" location="Pium river" longLatPrecision="1" longitude="-35.167225" municipality="Parnamirim" pageId="1" pageNumber="82" specimenCount="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<typeStatus id="5F34880AFFE0C571720BFB20FE7DAA1C" box="[342,451,1267,1293]" pageId="1" pageNumber="82" type="holotype">Holotype</typeStatus>
: male with pupal exuvia,
<collectingCountry id="F8987638FFE0C57171A1FB20FCDAAA1C" box="[764,868,1267,1293]" name="Brazil" pageId="1" pageNumber="82">BRAZIL</collectingCountry>
,
<collectingRegion id="424BF84AFFE0C5717029FB20FBCDAA1C" box="[884,1139,1267,1293]" country="Brazil" name="Rio Grande do Norte" pageId="1" pageNumber="82">Rio Grande do Norte</collectingRegion>
,
<collectingMunicipality id="6054ACD2FFE0C57177DEFB20FAB4AA1C" box="[1155,1290,1267,1293]" pageId="1" pageNumber="82">Parnamirim</collectingMunicipality>
,
<location id="85506073FFE0C5717647FB20FA26AA1C" LSID="urn:lsid:plazi:treatment:082687BEFFE0C57873CAFB7DFA3CA942:85506073FFE0C5717647FB20FA26AA1C" box="[1306,1432,1267,1293]" country="Brazil" latitude="-5.9584723" longLatPrecision="1" longitude="-35.167225" municipality="Parnamirim" name="Pium river" pageId="1" pageNumber="82" stateProvince="Rio Grande do Norte">Pium river</location>
,
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/
<geoCoordinate id="E5BB506FFFE0C571721DFAC6FE5CAA21" box="[320,482,1301,1328]" degrees="35" direction="west" minutes="10" orientation="longitude" pageId="1" pageNumber="82" precision="1" seconds="02.0" value="-35.167225">35º1002.0”W</geoCoordinate>
,
<date id="F4311068FFE0C57172ADFAC6FDD4AA21" box="[496,618,1301,1328]" pageId="1" pageNumber="82" value="2015-09-01">
<collectingDate id="E475E980FFE0C57172ADFAC6FDD4AA21" box="[496,618,1301,1328]" pageId="1" pageNumber="82" value="2015-09-01">01.ix.2015</collectingDate>
</date>
,
<collectorName id="2D7A537EFFE0C5717124FAC6FCA8AA21" box="[633,790,1301,1328]" pageId="1" pageNumber="82">G.P.S. Dantas</collectorName>
and
<collectorName id="2D7A537EFFE0C571700FFAC5FBA3AA3E" box="[850,1053,1301,1328]" pageId="1" pageNumber="82">M. P. G. Pinheiro</collectorName>
(
<collectionCode id="E69EAE6DFFE0C5717773FAC6FBCAAA3E" box="[1070,1140,1301,1327]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="82">INPA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="5F34880AFFE0C57177D4FAC5FB44AA21" box="[1161,1274,1302,1328]" pageId="1" pageNumber="82" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9689FD21FFE0C571765AFAC6FADEAA3E" box="[1287,1376,1301,1327]" pageId="1" pageNumber="82" type="male">2 males</specimenCount>
with pupal exuviae, as
<typeStatus id="5F34880AFFE0C5717239FAEBFE74AA43" box="[356,458,1336,1362]" pageId="1" pageNumber="82" type="holotype">holotype</typeStatus>
;
<materialsCitation id="30E73CF5FFE0C571728BFAEBFC44AA43" ID-GBIF-Occurrence="2628394303" box="[470,1018,1336,1363]" collectionCode="INPA" country="Brazil" pageId="1" pageNumber="82" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<specimenCount id="9689FD21FFE0C571728BFAEBFD86AA43" box="[470,568,1336,1362]" pageId="1" pageNumber="82" type="female">1 female</specimenCount>
with pupal exuvia, as
<typeStatus id="5F34880AFFE0C5717067FAEBFC22AA43" box="[826,924,1336,1362]" pageId="1" pageNumber="82" type="holotype">holotype</typeStatus>
(
<collectionCode id="E69EAE6DFFE0C57170F1FAEBFC4CAA43" box="[940,1010,1336,1362]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="82">INPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30E73CF5FFE0C571775BFAEBFD09AA64" ID-GBIF-Occurrence="2628394302" collectionCode="INPA" country="Brazil" pageId="1" pageNumber="82" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Norte" typeStatus="paratype">
<specimenCount id="9689FD21FFE0C571775BFAEBFBD6AA43" box="[1030,1128,1336,1362]" pageId="1" pageNumber="82" type="female">1 female</specimenCount>
gynandromoph (with male genitalia), with pupal and larval exuviae (
<collectionCode id="E69EAE6DFFE0C5717134FA88FD11AA64" box="[617,687,1371,1397]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="82">INPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<emphasis id="B2FBEABAFFE0C571739AFAAEFD9CAA89" bold="true" box="[199,546,1405,1432]" pageId="1" pageNumber="82">Diagnostic characters. Male:</emphasis>
spur of front tibia with 8 teeth; gonostylus with a dorsal lamella at the apex.
</paragraph>
<paragraph id="803036A8FFE0C57173CAFA73FA92AAEE" blockId="1.[151,1437,1266,2024]" pageId="1" pageNumber="82">
<emphasis id="B2FBEABAFFE0C57173CAFA73FF61AAAB" bold="true" box="[151,223,1440,1466]" pageId="1" pageNumber="82">Pupa:</emphasis>
respiratory atrium (
<collectionCode id="E69EAE6DFFE0C57172E6FA73FE5BAAAB" box="[443,485,1440,1466]" pageId="1" pageNumber="82">RA</collectionCode>
) with two diverticula, one apical and one in the mid portion; RA connected to the subbasal portion of the small plastron plate; internal margins of the anal lobe parallel-sided.
<emphasis id="B2FBEABAFFE0C5717733FA10FB7EAACC" bold="true" box="[1134,1216,1475,1501]" pageId="1" pageNumber="82">Larva:</emphasis>
S7 multi-branched, S9 bifid; S5 markedly posteromedial to DP; inner teeth of the ligula with tips strongly curved outwardly.
</paragraph>
</subSubSection>
<subSubSection id="C8956523FFE0C578739AF9DBFA3CA942" lastPageId="8" lastPageNumber="89" pageId="1" pageNumber="82" type="etymology">
<paragraph id="803036A8FFE0C571739AF9DBFA8EA954" blockId="1.[151,1437,1266,2024]" pageId="1" pageNumber="82">
<emphasis id="B2FBEABAFFE0C571739AF9DBFEF6A933" bold="true" box="[199,328,1544,1570]" pageId="1" pageNumber="82">Etymology</emphasis>
: the name is in honor of Dr. Herbet Tadeu de Almeida Andrade, for his constant support and in recognition of his initiative in study on aquatic macroinvertebrates in the
<collectingRegion id="424BF84AFFE0C57170F6F9F8FB67A954" box="[939,1241,1579,1605]" country="Brazil" name="Rio Grande do Norte" pageId="1" pageNumber="82">Rio Grande do Norte State</collectingRegion>
,
<collectingCountry id="F8987638FFE0C57177B9F9F8FA92A954" box="[1252,1324,1579,1605]" name="Brazil" pageId="1" pageNumber="82">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="803036A8FFE0C571739AF99EFEC0A99B" blockId="1.[151,1437,1266,2024]" pageId="1" pageNumber="82">
<emphasis id="B2FBEABAFFE0C571739AF99EFEF3A979" bold="true" box="[199,333,1613,1640]" pageId="1" pageNumber="82">Adult male</emphasis>
(n= 13). Total length
<quantity id="47779B4DFFE0C571710DF99EFD48A979" box="[592,758,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" metricValueMax="2.21" metricValueMin="1.95" pageId="1" pageNumber="82" unit="mm" value="2.08" valueMax="2.21" valueMin="1.95">1.952.21 mm</quantity>
. Total length/ wing length 1.501.69. Wing length/ length of profemur 2.102.18.
</paragraph>
<paragraph id="803036A8FFE0C571739AF940FE9FA9E3" blockId="1.[151,1437,1266,2024]" pageId="1" pageNumber="82">
<emphasis id="B2FBEABAFFE0C571739AF940FEF4A9BC" box="[199,330,1683,1709]" italics="true" pageId="1" pageNumber="82">Coloration.</emphasis>
Head, thorax and abdomen brown. Antenna brown, palpomeres light brown. Wings with unmarked membrane, covered with macrotrichia. Front leg brown, mid and hind legs with femurs yellow, tibiae and tarsus light brown.
</paragraph>
<paragraph id="803036A8FFE0C571739AF928FB89A8F9" blockId="1.[151,1437,1266,2024]" pageId="1" pageNumber="82">
<emphasis id="B2FBEABAFFE0C571739AF928FEBAA804" box="[199,260,1787,1813]" italics="true" pageId="1" pageNumber="82">Head</emphasis>
(
<figureCitation id="18B42A2DFFE0C571724EF928FED5A804" box="[275,363,1787,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1435,345,1876]" captionTargetId="figure@2.[151,1435,345,1885]" captionTargetPageId="2" captionText="FIGURE 1. Pentaneura herbeti sp. nov., male. (A) Head, frontal view. (B) Detail of the head showing the setigerous lobes. (C) Thorax, lateral view. (D) Wing." figureDoi="http://doi.org/10.5281/zenodo.3865009" httpUri="https://zenodo.org/record/3865009/files/figure.png" pageId="1" pageNumber="82">Fig. 1A</figureCitation>
). Antenna: AR 1.301.34, flagellum
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long, apical flagellomere not offset, evenly tapering from base to apex, conical,
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long,
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wide, apical seta
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long, pedicel with 4 setae on inferior margin. Eyes bare; dorsomedial extension well developed,
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long,
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wide, medially separated by
<quantity id="47779B4DFFE0C5717271F8B1FE1CA86C" box="[300,418,1890,1917]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="1" pageNumber="82" unit="mm" value="56.5" valueMax="58.0" valueMin="55.0">5558 µm</quantity>
; setigerous lobe dorsolaterally between base of pedicel and eyes (
<figureCitation id="18B42A2DFFE0C57177D8F8B0FB62A86C" box="[1157,1244,1891,1917]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1435,345,1876]" captionTargetId="figure@2.[151,1435,345,1885]" captionTargetPageId="2" captionText="FIGURE 1. Pentaneura herbeti sp. nov., male. (A) Head, frontal view. (B) Detail of the head showing the setigerous lobes. (C) Thorax, lateral view. (D) Wing." figureDoi="http://doi.org/10.5281/zenodo.3865009" httpUri="https://zenodo.org/record/3865009/files/figure.png" pageId="1" pageNumber="82">Fig. 1B</figureCitation>
). Temporal setae 1012, uniserial. Postorbital 1. Clypeus 92105 μm long, 6265 μm maximum width, with 1821 setae. Cibarial pump with anterior margin slightly concave, 208218 μm long, orifice 5258 from apex. Tentorium 145156 μm long. Palpomere lengths (IV in μm): 3540; 6272; 128146; 150174; 220250.
</paragraph>
<paragraph id="803036A8FFE3C572739AFF4AFA35AFC6" blockId="2.[151,1436,153,323]" pageId="2" pageNumber="83">
<emphasis id="B2FBEABAFFE3C572739AFF4AFEA6AFA2" box="[199,280,153,179]" italics="true" pageId="2" pageNumber="83">Thorax</emphasis>
(
<figureCitation id="18B42A2DFFE3C572727AFF4AFEC0AFA2" box="[295,382,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1435,345,1876]" captionTargetId="figure@2.[151,1435,345,1885]" captionTargetPageId="2" captionText="FIGURE 1. Pentaneura herbeti sp. nov., male. (A) Head, frontal view. (B) Detail of the head showing the setigerous lobes. (C) Thorax, lateral view. (D) Wing." figureDoi="http://doi.org/10.5281/zenodo.3865009" httpUri="https://zenodo.org/record/3865009/files/figure.png" pageId="2" pageNumber="83">Fig. 1C</figureCitation>
). Acrostichals 4046, biserial, diverging posteriorly; dorsocentrals 1823; prealars 67; supraalar 1. Antepronotum with 2 lateral setae, small tubercle present. Scutellum with 1417 setae, biserial or triserial.
</paragraph>
<paragraph id="803036A8FFE3C572739AFF31FF6EAE52" blockId="2.[151,1436,153,323]" pageId="2" pageNumber="83">
<emphasis id="B2FBEABAFFE3C572739AFF31FEBFAFEA" box="[199,257,226,251]" italics="true" pageId="2" pageNumber="83">Wing</emphasis>
(
<figureCitation id="18B42A2DFFE3C572724FFF32FED3AFEA" box="[274,365,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1435,345,1876]" captionTargetId="figure@2.[151,1435,345,1885]" captionTargetPageId="2" captionText="FIGURE 1. Pentaneura herbeti sp. nov., male. (A) Head, frontal view. (B) Detail of the head showing the setigerous lobes. (C) Thorax, lateral view. (D) Wing." figureDoi="http://doi.org/10.5281/zenodo.3865009" httpUri="https://zenodo.org/record/3865009/files/figure.png" pageId="2" pageNumber="83">Fig. 1D</figureCitation>
). Length
<quantity id="47779B4DFFE3C5727286FF32FD3CAFEA" box="[475,642,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.295" metricValueMax="1.31" metricValueMin="1.28" pageId="2" pageNumber="83" unit="mm" value="1.295" valueMax="1.31" valueMin="1.28">1.281.31 mm</quantity>
, width
<quantity id="47779B4DFFE3C572718AFF32FCC1AFEA" box="[727,895,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="3.9" metricValueMin="3.8" pageId="2" pageNumber="83" unit="mm" value="0.385" valueMax="0.39" valueMin="0.38">0.380.39 mm</quantity>
. Membrane densely covered with macrotrichia; C slightly produced beyond apex of R
<subScript id="1C0B34EDFFE3C5727113FEC5FDD6AE35" attach="left" box="[590,616,278,292]" fontSize="6" pageId="2" pageNumber="83">4+5</subScript>
. Brachiolum with 3 setae. Squama with 58 setae. VR 0.920.93. WW 0.30.
</paragraph>
<caption id="D4F06620FFE3C57273CAF8A5FE69A8A2" ID-DOI="http://doi.org/10.5281/zenodo.3865009" ID-Zenodo-Dep="3865009" httpUri="https://zenodo.org/record/3865009/files/figure.png" pageId="2" pageNumber="83" startId="2.[151,250,1910,1935]" targetBox="[151,1435,345,1876]" targetPageId="2">
<paragraph id="803036A8FFE3C57273CAF8A5FE69A8A2" blockId="2.[151,1437,1910,1971]" pageId="2" pageNumber="83">
<emphasis id="B2FBEABAFFE3C57273CAF8A5FEB3A89E" bold="true" box="[151,269,1910,1935]" pageId="2" pageNumber="83">FIGURE 1</emphasis>
.
<taxonomicName id="478F4D2BFFE3C5727245F8A4FE62A89F" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[280,476,1910,1934]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE3C5727245F8A4FE62A89F" box="[280,476,1910,1934]" italics="true" pageId="2" pageNumber="83">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE3C57272BFF8A4FD8CA89E" bold="true" box="[482,562,1911,1935]" pageId="2" pageNumber="83">
<taxonomicNameLabel id="A9C857C1FFE3C57272BFF8A4FD8CA89E" box="[482,562,1911,1935]" pageId="2" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. (A) Head, frontal view. (B) Detail of the head showing the setigerous lobes. (C) Thorax, lateral view. (D) Wing.
</paragraph>
</caption>
<paragraph id="803036A8FFE2C573739AFF49FE05AE52" blockId="3.[151,1436,153,323]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFF49FF42AFA2" box="[199,252,154,179]" italics="true" pageId="3" pageNumber="84">Legs</emphasis>
. Foretibia 4042 μm wide at apex, with one lyrate apical spur 23 μm long, bearing 8 teeth. Mid tibia 4243 μm wide at apex, with two apical somewhat lyrate spurs, 24 and 26 μm long, bearing 3 and 5 lateral teeth, respec- tively. Hind tibia 42 μm wide at apex, with two apical somewhat lyrate spurs, 26 and 27 μm long, bearing 3 and 5 teeth, respectively; tibial comb reduced, with 5 bristles. All legs with slender, curved claws. Lengths and proportions of male legs as in
<tableCitation id="CD0D0313FFE2C573723FFEFAFE09AE52" box="[354,439,297,323]" captionStart="TABLE 2" captionStartId="3.[151,239,370,394]" captionText="TABLE 2. Length (in μm) and proportion of male legs of Pentaneura herbeti sp. nov. (Chironomidae: Tanypodinae), male (n= 3):" pageId="3" pageNumber="84">Table 2</tableCitation>
.
</paragraph>
<caption id="D4F06620FFE2C57373CAFEA1FEA5AEBE" pageId="3" pageNumber="84">
<paragraph id="803036A8FFE2C57373CAFEA1FEA5AEBE" blockId="3.[151,1436,370,431]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C57373CAFEA1FEB2AE9A" bold="true" box="[151,268,370,395]" pageId="3" pageNumber="84">TABLE 2.</emphasis>
Length (in μm) and proportion of male legs of
<taxonomicName id="478F4D2BFFE2C5737044FEA0FC54AE9A" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[793,1002,371,395]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE2C5737044FEA0FC54AE9A" box="[793,1002,371,395]" italics="true" pageId="3" pageNumber="84">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE2C57370AEFEA0FBF4AE9A" bold="true" box="[1011,1098,371,395]" pageId="3" pageNumber="84">
<taxonomicNameLabel id="A9C857C1FFE2C57370AEFEA0FBF4AE9A" box="[1011,1098,371,395]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="478F4D2BFFE2C5737707FEA0FB47AE9A" authorityName="Newman" authorityYear="1834" box="[1114,1273,371,395]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
:
<taxonomicName id="478F4D2BFFE2C573765EFEA0FA2FAE9A" authorityName="Kieffer" authorityYear="1906" box="[1283,1425,371,395]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="subFamily" subFamily="Tanypodinae">Tanypodinae</taxonomicName>
), male (n= 3):
</paragraph>
</caption>
<paragraph id="803036A8FFE2C57373BAFE6DFA33AD41" pageId="3" pageNumber="84">
<table id="F28FC408FFE23A8F73FBFE6DFA33AD45" box="[166,1421,446,596]" gridcols="11" gridrows="4" pageId="3" pageNumber="84" rowsContinueIn="3.[166,1378,1580,1731]">
<tr id="3EBF34EAFFE23A8F73FBFE6DFA33AECD" box="[166,1421,446,476]" gridrow="0" pageId="3" pageNumber="84" rowspan-0="1">
<th id="7D6E5D96FFE23A8F73BAFE6DFEFEAECD" box="[231,320,446,476]" gridcol="1" gridrow="0" pageId="3" pageNumber="84">fe</th>
<th id="7D6E5D96FFE23A8F7200FE6DFE08AECD" box="[349,438,446,476]" gridcol="2" gridrow="0" pageId="3" pageNumber="84">ti</th>
<th id="7D6E5D96FFE23A8F728FFE6DFD95AECD" box="[466,555,446,476]" gridcol="3" gridrow="0" pageId="3" pageNumber="84">ta1</th>
<th id="7D6E5D96FFE23A8F7115FE6DFD1FAECD" box="[584,673,446,476]" gridcol="4" gridrow="0" pageId="3" pageNumber="84">ta2</th>
<th id="7D6E5D96FFE23A8F71E3FE6DFCA9AECD" box="[702,791,446,476]" gridcol="5" gridrow="0" pageId="3" pageNumber="84">ta3</th>
<th id="7D6E5D96FFE23A8F7069FE6DFC33AECD" box="[820,909,446,476]" gridcol="6" gridrow="0" pageId="3" pageNumber="84">ta4</th>
<th id="7D6E5D96FFE23A8F70F4FE6DFC57AECD" box="[937,1001,446,476]" gridcol="7" gridrow="0" pageId="3" pageNumber="84">ta5</th>
<th id="7D6E5D96FFE23A8F7746FE6DFB3EAECD" box="[1051,1152,446,476]" gridcol="8" gridrow="0" pageId="3" pageNumber="84">LR</th>
<th id="7D6E5D96FFE23A8F77FCFE6DFAB9AECD" box="[1185,1287,446,476]" gridcol="9" gridrow="0" pageId="3" pageNumber="84">BV</th>
<th id="7D6E5D96FFE23A8F7675FE6DFA33AECD" box="[1320,1421,446,476]" gridcol="10" gridrow="0" pageId="3" pageNumber="84">SV</th>
</tr>
<tr id="3EBF34EAFFE23A8F73FBFE35FA33AD15" box="[166,1421,486,516]" gridrow="1" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBFE35FF05AD15" box="[166,187,486,516]" gridcol="0" gridrow="1" pageId="3" pageNumber="84">P1</th>
<td id="7D6E5D96FFE23A8F73BAFE35FEFEAD15" box="[231,320,486,516]" gridcol="1" gridrow="1" pageId="3" pageNumber="84">585620</td>
<td id="7D6E5D96FFE23A8F7200FE35FE08AD15" box="[349,438,486,516]" gridcol="2" gridrow="1" pageId="3" pageNumber="84">641684</td>
<td id="7D6E5D96FFE23A8F728FFE35FD95AD15" box="[466,555,486,516]" gridcol="3" gridrow="1" pageId="3" pageNumber="84">570630</td>
<td id="7D6E5D96FFE23A8F7115FE35FD1FAD15" box="[584,673,486,516]" gridcol="4" gridrow="1" pageId="3" pageNumber="84">276303</td>
<td id="7D6E5D96FFE23A8F71E3FE35FCA9AD15" box="[702,791,486,516]" gridcol="5" gridrow="1" pageId="3" pageNumber="84">190230</td>
<td id="7D6E5D96FFE23A8F7069FE35FC33AD15" box="[820,909,486,516]" gridcol="6" gridrow="1" pageId="3" pageNumber="84">132150</td>
<td id="7D6E5D96FFE23A8F70F4FE35FC57AD15" box="[937,1001,486,516]" gridcol="7" gridrow="1" pageId="3" pageNumber="84">7585</td>
<td id="7D6E5D96FFE23A8F7746FE35FB3EAD15" box="[1051,1152,486,516]" gridcol="8" gridrow="1" pageId="3" pageNumber="84">0.870.92</td>
<td id="7D6E5D96FFE23A8F77FCFE35FAB9AD15" box="[1185,1287,486,516]" gridcol="9" gridrow="1" pageId="3" pageNumber="84">2.522.66</td>
<td id="7D6E5D96FFE23A8F7675FE35FA33AD15" box="[1320,1421,486,516]" gridcol="10" gridrow="1" pageId="3" pageNumber="84">2.042.21</td>
</tr>
<tr id="3EBF34EAFFE23A8F73FBFDDCFA33AD3D" box="[166,1421,527,556]" gridrow="2" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBFDDCFF05AD3D" box="[166,187,527,556]" gridcol="0" gridrow="2" pageId="3" pageNumber="84">P2</th>
<td id="7D6E5D96FFE23A8F73BAFDDCFEFEAD3D" box="[231,320,527,556]" gridcol="1" gridrow="2" pageId="3" pageNumber="84">683717</td>
<td id="7D6E5D96FFE23A8F7200FDDCFE08AD3D" box="[349,438,527,556]" gridcol="2" gridrow="2" pageId="3" pageNumber="84">771797</td>
<td id="7D6E5D96FFE23A8F728FFDDCFD95AD3D" box="[466,555,527,556]" gridcol="3" gridrow="2" pageId="3" pageNumber="84">626720</td>
<td id="7D6E5D96FFE23A8F7115FDDCFD1FAD3D" box="[584,673,527,556]" gridcol="4" gridrow="2" pageId="3" pageNumber="84">270309</td>
<td id="7D6E5D96FFE23A8F71E3FDDCFCA9AD3D" box="[702,791,527,556]" gridcol="5" gridrow="2" pageId="3" pageNumber="84">170192</td>
<td id="7D6E5D96FFE23A8F7069FDDCFC33AD3D" box="[820,909,527,556]" gridcol="6" gridrow="2" pageId="3" pageNumber="84">125131</td>
<td id="7D6E5D96FFE23A8F70F4FDDCFC57AD3D" box="[937,1001,527,556]" gridcol="7" gridrow="2" pageId="3" pageNumber="84">7275</td>
<td id="7D6E5D96FFE23A8F7746FDDCFB3EAD3D" box="[1051,1152,527,556]" gridcol="8" gridrow="2" pageId="3" pageNumber="84">0.810.91</td>
<td id="7D6E5D96FFE23A8F77FCFDDCFAB9AD3D" box="[1185,1287,527,556]" gridcol="9" gridrow="2" pageId="3" pageNumber="84">3.173.20</td>
<td id="7D6E5D96FFE23A8F7675FDDCFA33AD3D" box="[1320,1421,527,556]" gridcol="10" gridrow="2" pageId="3" pageNumber="84">2.102.25</td>
</tr>
<tr id="3EBF34EAFFE23A8F73FBFDE4FA33AD45" box="[166,1421,567,596]" gridrow="3" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBFDE4FF05AD45" box="[166,187,567,596]" gridcol="0" gridrow="3" pageId="3" pageNumber="84">P3</th>
<td id="7D6E5D96FFE23A8F73BAFDE4FEFEAD45" box="[231,320,567,596]" gridcol="1" gridrow="3" pageId="3" pageNumber="84">581594</td>
<td id="7D6E5D96FFE23A8F7200FDE4FE08AD45" box="[349,438,567,596]" gridcol="2" gridrow="3" pageId="3" pageNumber="84">824898</td>
<td id="7D6E5D96FFE23A8F728FFDE4FD95AD45" box="[466,555,567,596]" gridcol="3" gridrow="3" pageId="3" pageNumber="84">596666</td>
<td id="7D6E5D96FFE23A8F7115FDE4FD1FAD45" box="[584,673,567,596]" gridcol="4" gridrow="3" pageId="3" pageNumber="84">317350</td>
<td id="7D6E5D96FFE23A8F71E3FDE4FCA9AD45" box="[702,791,567,596]" gridcol="5" gridrow="3" pageId="3" pageNumber="84">234253</td>
<td id="7D6E5D96FFE23A8F7069FDE4FC33AD45" box="[820,909,567,596]" gridcol="6" gridrow="3" pageId="3" pageNumber="84">156170</td>
<td id="7D6E5D96FFE23A8F70F4FDE4FC57AD45" box="[937,1001,567,596]" gridcol="7" gridrow="3" pageId="3" pageNumber="84">8084</td>
<td id="7D6E5D96FFE23A8F7746FDE4FB3EAD45" box="[1051,1152,567,596]" gridcol="8" gridrow="3" pageId="3" pageNumber="84">0.720.76</td>
<td id="7D6E5D96FFE23A8F77FCFDE4FAB9AD45" box="[1185,1287,567,596]" gridcol="9" gridrow="3" pageId="3" pageNumber="84">2.502.55</td>
<td id="7D6E5D96FFE23A8F7675FDE4FA33AD45" box="[1320,1421,567,596]" gridcol="10" gridrow="3" pageId="3" pageNumber="84">2.202.31</td>
</tr>
</table>
</paragraph>
<paragraph id="803036A8FFE2C573739AFD53FC6FAC14" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFD53FEE8AD88" box="[199,342,640,665]" italics="true" pageId="3" pageNumber="84">Hypopygium</emphasis>
(
<figureCitation id="18B42A2DFFE2C573723BFDACFE0AAD88" box="[358,436,639,665]" captionStart="FIGURE 2" captionStartId="4.[151,250,1821,1846]" captionTargetBox="[180,1413,193,1789]" captionTargetId="figure@4.[165,1423,181,1795]" captionTargetPageId="4" captionText="FIGURE 2. Pentaneura herbeti sp. nov., male. (A) Hypopygium, dorsal view. (B) Gonostylus. (C) Hypopygium, the arrow indicates the dorsal lamella, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865011" httpUri="https://zenodo.org/record/3865011/files/figure.png" pageId="3" pageNumber="84">Figs. 2</figureCitation>
AC). Tergite IX with 47 setae. Anal point broad. Phallapodeme markedly elongate and curved, 92 μm long; sternapodeme with a weak anterior process. Gonocoxite robust, 8085 μm long, with a dor- somedial setigerous basal area. Inferior volsella absent. Gonostylus slightly curved, 4546 μm long, with a dorsal lamella at the apex; megaseta robust, 9 μm long. HR 1.781.85; HV 4.80.
</paragraph>
<paragraph id="803036A8FFE2C573739AFCDCFA3CAC38" blockId="3.[151,1437,639,1458]" box="[199,1410,783,810]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFCDCFEA0AC38" bold="true" box="[199,286,783,809]" pageId="3" pageNumber="84">Female</emphasis>
(n= 1). Total length
<quantity id="47779B4DFFE2C5737158FCDCFDD2AC3B" box="[517,620,783,810]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="3" pageNumber="84" unit="mm" value="1.66">1.66 mm</quantity>
. Total length/ wing length 1.42. Wing length/ length of profemur 2.04.
</paragraph>
<paragraph id="803036A8FFE2C573739AFCE0FF58AC84" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFCE0FEF4AC5C" box="[199,330,819,845]" italics="true" pageId="3" pageNumber="84">Coloration.</emphasis>
Head, thorax and abdomen brown. Antenna brown, palpomeres light brown. Wings with unmarked membrane, covered with macrotrichia. Legs with femurs light brown, slightly darker at apex; tibiae and tarsus brown.
</paragraph>
<paragraph id="803036A8FFE2C573739AFC4CFA3CAB58" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFC4CFEBAACA8" box="[199,260,927,953]" italics="true" pageId="3" pageNumber="84">Head</emphasis>
(
<figureCitation id="18B42A2DFFE2C573724FFC4CFED2ACA8" box="[274,364,927,953]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[175,1413,181,1942]" captionTargetId="figure@5.[175,1413,181,1944]" captionTargetPageId="5" captionText="FIGURE 3. Pentaneura herbeti sp. nov., female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus)." figureDoi="http://doi.org/10.5281/zenodo.3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="3" pageNumber="84">Fig. 3A</figureCitation>
). Antenna: AR 0.30, flagellum
<quantity id="47779B4DFFE2C5737198FC4CFCA2ACAB" box="[709,796,927,954]" metricMagnitude="-1" metricUnit="m" metricValue="9.62" pageId="3" pageNumber="84" unit="mm" value="962.0">962 µm</quantity>
long, apical flagellomere
<quantity id="47779B4DFFE2C5737763FC4CFB2AACAB" box="[1086,1172,927,954]" metricMagnitude="-1" metricUnit="m" metricValue="2.2199999999999998" pageId="3" pageNumber="84" unit="mm" value="222.0">222 µm</quantity>
long, apical seta
<quantity id="47779B4DFFE2C5737609FC4CFA22ACAB" box="[1364,1436,927,954]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="84" unit="mm" value="66.0">66 µm</quantity>
long, pedicel with a row of 7 setae, scape with 5 ventral setae. Eyes bare; dorsomedial extension well developed,
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long,
<quantity id="47779B4DFFE2C573727AFC35FED1AB13" box="[295,367,998,1026]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="84" unit="mm" value="75.0">75 µm</quantity>
wide, medially separated by
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. Temporal setae 9, uniserial. Postorbital 1. Clypeus 102 μm long, 72 μm maximum width, with 26 setae. Cibarial pump with anterior margin slightly concave, 215 μm long, orifice placed 68 μm from apex. Tentorium 133 μm long. Palpomere lengths (IV in μm): 37; 70; 133; 171; 287.
</paragraph>
<paragraph id="803036A8FFE2C573739AFB80FDD0ABA4" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFB80FEA6AB7C" box="[199,280,1107,1133]" italics="true" pageId="3" pageNumber="84">Thorax</emphasis>
(
<figureCitation id="18B42A2DFFE2C5737276FB80FE38AB7C" box="[299,390,1107,1133]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[175,1413,181,1942]" captionTargetId="figure@5.[175,1413,181,1944]" captionTargetPageId="5" captionText="FIGURE 3. Pentaneura herbeti sp. nov., female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus)." figureDoi="http://doi.org/10.5281/zenodo.3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="3" pageNumber="84">Fig. 3B</figureCitation>
). Acrostichals about 44, biserial, diverging posteriorly; dorsocentrals about 35, multiserial anteriorly, uniserial and biserial posteriorly; prealars 9; supraalar 1. Antepronotum with 2 lateral setae, tubercle present. Scutellum with 18 setae, triserial.
</paragraph>
<paragraph id="803036A8FFE2C573739AFB13FB0FABEC" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFB13FEBFABC8" box="[199,257,1216,1241]" italics="true" pageId="3" pageNumber="84">Wing</emphasis>
(
<figureCitation id="18B42A2DFFE2C573724CFB6CFED5ABC8" box="[273,363,1215,1241]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[175,1413,181,1942]" captionTargetId="figure@5.[175,1413,181,1944]" captionTargetPageId="5" captionText="FIGURE 3. Pentaneura herbeti sp. nov., female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus)." figureDoi="http://doi.org/10.5281/zenodo.3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="3" pageNumber="84">Fig. 3C</figureCitation>
). Length
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, width
<quantity id="47779B4DFFE2C57371C9FB6CFD42ABCB" box="[660,764,1215,1242]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="84" unit="mm" value="0.43">0.43 mm</quantity>
. Membrane covered with macrotrichia; C not produced beyond apex of R
<subScript id="1C0B34EDFFE2C573721EFB26FEE3AA12" attach="left" box="[323,349,1269,1283]" fontSize="6" pageId="3" pageNumber="84">4+5</subScript>
. Brachiolum with 2 setae. Squama with about 10 setae. VR 0.92. WW 0.36.
</paragraph>
<paragraph id="803036A8FFE2C573739AFADBFE7AAAA0" blockId="3.[151,1437,639,1458]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AFADBFF42AA30" box="[199,252,1288,1313]" italics="true" pageId="3" pageNumber="84">Legs</emphasis>
. Foretibia 45 μm wide at apex, with one lyrate apical spur 21 μm long, bearing 8 teeth (
<figureCitation id="18B42A2DFFE2C5737794FAD4FA9CAA30" box="[1225,1314,1287,1313]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[175,1413,181,1942]" captionTargetId="figure@5.[175,1413,181,1944]" captionTargetPageId="5" captionText="FIGURE 3. Pentaneura herbeti sp. nov., female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus)." figureDoi="http://doi.org/10.5281/zenodo.3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="3" pageNumber="84">Fig. 3D</figureCitation>
). Mid tibia 45 μm wide at apex, with two apical somewhat lyrate spurs, 23 and 25 μm long, bearing 3 and 5 teeth, respectively. Hind tibia 45 μm wide at apex, with two apical somewhat lyrate spurs, 25 and 27 μm long, bearing 3 and 5 teeth, respectively; tibial comb indistinct, with 5 bristles. All legs with slender, curved claws. Lengths and proportions of leg segments as in
<tableCitation id="CD0D0313FFE2C5737237FA44FE01AAA0" box="[362,447,1431,1457]" captionStart="TABLE 3" captionStartId="3.[151,239,1505,1529]" captionText="TABLE 3. Length (in μm) and proportion of female legs of Pentaneura herbeti sp. nov. (Chironomidae: Tanypodinae), female (n= 1):" pageId="3" pageNumber="84">Table 3</tableCitation>
.
</paragraph>
<caption id="D4F06620FFE2C57373CAFA32FE8EA90C" pageId="3" pageNumber="84">
<paragraph id="803036A8FFE2C57373CAFA32FE8EA90C" blockId="3.[151,1436,1505,1565]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C57373CAFA32FEB4AAE8" bold="true" box="[151,266,1505,1529]" pageId="3" pageNumber="84">TABLE 3.</emphasis>
Length (in μm) and proportion of female legs of
<taxonomicName id="478F4D2BFFE2C573707DFA31FC4EAAE8" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[800,1008,1505,1529]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE2C573707DFA31FC4EAAE8" box="[800,1008,1505,1529]" italics="true" pageId="3" pageNumber="84">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE2C57370ABFA32FBF3AAE8" bold="true" box="[1014,1101,1505,1529]" pageId="3" pageNumber="84">
<taxonomicNameLabel id="A9C857C1FFE2C57370ABFA32FBF3AAE8" box="[1014,1101,1505,1529]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="478F4D2BFFE2C5737706FA32FB44AAE8" authorityName="Newman" authorityYear="1834" box="[1115,1274,1505,1529]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
:
<taxonomicName id="478F4D2BFFE2C573765EFA32FA2FAAE8" authorityName="Kieffer" authorityYear="1906" box="[1283,1425,1505,1529]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="subFamily" subFamily="Tanypodinae">Tanypodinae</taxonomicName>
), female (n= 1):
</paragraph>
</caption>
<paragraph id="803036A8FFE2C5737247F9FFFADCA9AF" pageId="3" pageNumber="84">
<table id="F28FC408FFE23A8F73FBF9FFFADCA9D2" box="[166,1378,1580,1731]" gridcols="11" gridrows="4" pageId="3" pageNumber="84" rowsContinueFrom="3.[166,1421,446,596]">
<tr id="3EBF34EAFFE23A8F73FBF9FFFADCA95B" box="[166,1378,1580,1610]" gridrow="0" pageId="3" pageNumber="84" rowspan-0="1">
<th id="7D6E5D96FFE23A8F7247F9FFFEFEA95B" box="[282,320,1580,1610]" gridcol="1" gridrow="0" pageId="3" pageNumber="84">fe</th>
<th id="7D6E5D96FFE23A8F72D2F9FFFE0BA95B" box="[399,437,1580,1610]" gridcol="2" gridrow="0" pageId="3" pageNumber="84">ti</th>
<th id="7D6E5D96FFE23A8F7159F9FFFD94A95B" box="[516,554,1580,1610]" gridcol="3" gridrow="0" pageId="3" pageNumber="84">ta1</th>
<th id="7D6E5D96FFE23A8F7124F9FFFD21A95B" box="[633,671,1580,1610]" gridcol="4" gridrow="0" pageId="3" pageNumber="84">ta2</th>
<th id="7D6E5D96FFE23A8F71B3F9FFFCAAA95B" box="[750,788,1580,1610]" gridcol="5" gridrow="0" pageId="3" pageNumber="84">ta3</th>
<th id="7D6E5D96FFE23A8F703FF9FFFC36A95B" box="[866,904,1580,1610]" gridcol="6" gridrow="0" pageId="3" pageNumber="84">ta4</th>
<th id="7D6E5D96FFE23A8F708AF9FFFC4FA95B" box="[983,1009,1580,1610]" gridcol="7" gridrow="0" pageId="3" pageNumber="84">ta5</th>
<th id="7D6E5D96FFE23A8F7711F9FFFBC6A95B" box="[1100,1144,1580,1610]" gridcol="8" gridrow="0" pageId="3" pageNumber="84">LR</th>
<th id="7D6E5D96FFE23A8F779CF9FFFB53A95B" box="[1217,1261,1580,1610]" gridcol="9" gridrow="0" pageId="3" pageNumber="84">BV</th>
<th id="7D6E5D96FFE23A8F766BF9FFFADCA95B" box="[1334,1378,1580,1610]" gridcol="10" gridrow="0" pageId="3" pageNumber="84">SV</th>
</tr>
<tr id="3EBF34EAFFE23A8F73FBF986FADCA963" box="[166,1378,1621,1650]" gridrow="1" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBF986FF05A963" box="[166,187,1621,1650]" gridcol="0" gridrow="1" pageId="3" pageNumber="84">P1</th>
<td id="7D6E5D96FFE23A8F7247F986FEFEA963" box="[282,320,1621,1650]" gridcol="1" gridrow="1" pageId="3" pageNumber="84">575</td>
<td id="7D6E5D96FFE23A8F72D2F986FE0BA963" box="[399,437,1621,1650]" gridcol="2" gridrow="1" pageId="3" pageNumber="84">650</td>
<td id="7D6E5D96FFE23A8F7159F986FD94A963" box="[516,554,1621,1650]" gridcol="3" gridrow="1" pageId="3" pageNumber="84">581</td>
<td id="7D6E5D96FFE23A8F7124F986FD21A963" box="[633,671,1621,1650]" gridcol="4" gridrow="1" pageId="3" pageNumber="84">286</td>
<td id="7D6E5D96FFE23A8F71B3F986FCAAA963" box="[750,788,1621,1650]" gridcol="5" gridrow="1" pageId="3" pageNumber="84">210</td>
<td id="7D6E5D96FFE23A8F703FF986FC36A963" box="[866,904,1621,1650]" gridcol="6" gridrow="1" pageId="3" pageNumber="84">147</td>
<td id="7D6E5D96FFE23A8F708AF986FC4FA963" box="[983,1009,1621,1650]" gridcol="7" gridrow="1" pageId="3" pageNumber="84">75</td>
<td id="7D6E5D96FFE23A8F7711F986FBC6A963" box="[1100,1144,1621,1650]" gridcol="8" gridrow="1" pageId="3" pageNumber="84">0.89</td>
<td id="7D6E5D96FFE23A8F779CF986FB53A963" box="[1217,1261,1621,1650]" gridcol="9" gridrow="1" pageId="3" pageNumber="84">2.52</td>
<td id="7D6E5D96FFE23A8F766BF986FADCA963" box="[1334,1378,1621,1650]" gridcol="10" gridrow="1" pageId="3" pageNumber="84">2.11</td>
</tr>
<tr id="3EBF34EAFFE23A8F73FBF9AEFADCA98B" box="[166,1378,1661,1690]" gridrow="2" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBF9AEFF05A98B" box="[166,187,1661,1690]" gridcol="0" gridrow="2" pageId="3" pageNumber="84">P2</th>
<td id="7D6E5D96FFE23A8F7247F9AEFEFEA98B" box="[282,320,1661,1690]" gridcol="1" gridrow="2" pageId="3" pageNumber="84">670</td>
<td id="7D6E5D96FFE23A8F72D2F9AEFE0BA98B" box="[399,437,1661,1690]" gridcol="2" gridrow="2" pageId="3" pageNumber="84">775</td>
<td id="7D6E5D96FFE23A8F7159F9AEFD94A98B" box="[516,554,1661,1690]" gridcol="3" gridrow="2" pageId="3" pageNumber="84">656</td>
<td id="7D6E5D96FFE23A8F7124F9AEFD21A98B" box="[633,671,1661,1690]" gridcol="4" gridrow="2" pageId="3" pageNumber="84">280</td>
<td id="7D6E5D96FFE23A8F71B3F9AEFCAAA98B" box="[750,788,1661,1690]" gridcol="5" gridrow="2" pageId="3" pageNumber="84">175</td>
<td id="7D6E5D96FFE23A8F703FF9AEFC36A98B" box="[866,904,1661,1690]" gridcol="6" gridrow="2" pageId="3" pageNumber="84">133</td>
<td id="7D6E5D96FFE23A8F708AF9AEFC4FA98B" box="[983,1009,1661,1690]" gridcol="7" gridrow="2" pageId="3" pageNumber="84">77</td>
<td id="7D6E5D96FFE23A8F7711F9AEFBC6A98B" box="[1100,1144,1661,1690]" gridcol="8" gridrow="2" pageId="3" pageNumber="84">0.85</td>
<td id="7D6E5D96FFE23A8F779CF9AEFB53A98B" box="[1217,1261,1661,1690]" gridcol="9" gridrow="2" pageId="3" pageNumber="84">3.16</td>
<td id="7D6E5D96FFE23A8F766BF9AEFADCA98B" box="[1334,1378,1661,1690]" gridcol="10" gridrow="2" pageId="3" pageNumber="84">2.20</td>
</tr>
<tr id="3EBF34EAFFE23A8F73FBF976FADCA9D2" box="[166,1378,1701,1731]" gridrow="3" pageId="3" pageNumber="84">
<th id="7D6E5D96FFE23A8F73FBF976FF05A9D2" box="[166,187,1701,1731]" gridcol="0" gridrow="3" pageId="3" pageNumber="84">P3</th>
<td id="7D6E5D96FFE23A8F7247F976FEFEA9D2" box="[282,320,1701,1731]" gridcol="1" gridrow="3" pageId="3" pageNumber="84">582</td>
<td id="7D6E5D96FFE23A8F72D2F976FE0BA9D2" box="[399,437,1701,1731]" gridcol="2" gridrow="3" pageId="3" pageNumber="84">870</td>
<td id="7D6E5D96FFE23A8F7159F976FD94A9D2" box="[516,554,1701,1731]" gridcol="3" gridrow="3" pageId="3" pageNumber="84">628</td>
<td id="7D6E5D96FFE23A8F7124F976FD21A9D2" box="[633,671,1701,1731]" gridcol="4" gridrow="3" pageId="3" pageNumber="84">328</td>
<td id="7D6E5D96FFE23A8F71B3F976FCAAA9D2" box="[750,788,1701,1731]" gridcol="5" gridrow="3" pageId="3" pageNumber="84">255</td>
<td id="7D6E5D96FFE23A8F703FF976FC36A9D2" box="[866,904,1701,1731]" gridcol="6" gridrow="3" pageId="3" pageNumber="84">174</td>
<td id="7D6E5D96FFE23A8F708AF976FC4FA9D2" box="[983,1009,1701,1731]" gridcol="7" gridrow="3" pageId="3" pageNumber="84">86</td>
<td id="7D6E5D96FFE23A8F7711F976FBC6A9D2" box="[1100,1144,1701,1731]" gridcol="8" gridrow="3" pageId="3" pageNumber="84">0.72</td>
<td id="7D6E5D96FFE23A8F779CF976FB53A9D2" box="[1217,1261,1701,1731]" gridcol="9" gridrow="3" pageId="3" pageNumber="84">2.47</td>
<td id="7D6E5D96FFE23A8F766BF976FADCA9D2" box="[1334,1378,1701,1731]" gridcol="10" gridrow="3" pageId="3" pageNumber="84">2.31</td>
</tr>
</table>
</paragraph>
<paragraph id="803036A8FFE2C573739AF93DFDE4A85E" blockId="3.[151,1437,1773,2016]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AF93DFE8FA819" box="[199,305,1774,1800]" italics="true" pageId="3" pageNumber="84">Genitalia</emphasis>
(
<figureCitation id="18B42A2DFFE2C573721DF93EFE29A819" box="[320,407,1773,1800]" captionStart="FIGURE 3" captionStartId="5.[151,250,1970,1995]" captionTargetBox="[175,1413,181,1942]" captionTargetId="figure@5.[175,1413,181,1944]" captionTargetPageId="5" captionText="FIGURE 3. Pentaneura herbeti sp. nov., female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus)." figureDoi="http://doi.org/10.5281/zenodo.3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="3" pageNumber="84">Fig. 3E</figureCitation>
). Three spherical seminal capsules, with
<quantity id="47779B4DFFE2C573703DF93EFC17A819" box="[864,937,1773,1800]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="84" unit="mm" value="60.0">60 µm</quantity>
in diameter; spermathecal ducts separate for all their length. Coxosternapodeme curved,
<quantity id="47779B4DFFE2C57371D9F8C2FD73A83D" box="[644,717,1809,1836]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="84" unit="mm" value="85.0">85 µm</quantity>
long. Tergite X
<quantity id="47779B4DFFE2C57370D8F8C1FC63A83D" box="[901,989,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="84" unit="mm" value="120.0">120 µm</quantity>
wide, with about 12 setae on each side. Notum
<quantity id="47779B4DFFE2C57373B1F8E5FE8BA841" box="[236,309,1846,1872]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="84" unit="mm" value="80.0">80 µm</quantity>
long. Cercus
<quantity id="47779B4DFFE2C573728DF8E5FDA7A841" box="[464,537,1846,1872]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="84" unit="mm" value="21.0">21 µm</quantity>
long.
</paragraph>
<paragraph id="803036A8FFE2C573739AF889FC4FA865" blockId="3.[151,1437,1773,2016]" box="[199,1009,1881,1908]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AF889FEBBA865" bold="true" box="[199,261,1882,1908]" pageId="3" pageNumber="84">Pupa</emphasis>
(n= 3). Total length
<quantity id="47779B4DFFE2C57372B0F889FD2EA865" box="[493,656,1882,1908]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="2.97" metricValueMin="2.83" pageId="3" pageNumber="84" unit="mm" value="2.9" valueMax="2.97" valueMin="2.83">2.832.97 mm</quantity>
(abdomen and thorax of male).
</paragraph>
<paragraph id="803036A8FFE2C573739AF8ADFCC4A8CE" blockId="3.[151,1437,1773,2016]" pageId="3" pageNumber="84">
<emphasis id="B2FBEABAFFE2C573739AF8ADFEFCA889" box="[199,322,1918,1944]" italics="true" pageId="3" pageNumber="84">Coloration</emphasis>
. Cephalothorax and wing sheath light brown, without light spots (
<figureCitation id="18B42A2DFFE2C573776FF8AEFB32A889" box="[1074,1164,1917,1944]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="3" pageNumber="84">Fig. 4B</figureCitation>
); thoracic horn with external membrane and plastron plate light-brown, respiratory atrium dark brown. Abdomen as in
<figureCitation id="18B42A2DFFE2C5737796F872FA81A8AD" box="[1227,1343,1953,1980]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="3" pageNumber="84">Figure 4C</figureCitation>
, all segments brown, IVII with lighter longitudinal patches on the sides.
</paragraph>
<caption id="D4F06620FFE5C57473CAF8CEFD8DA84B" ID-DOI="http://doi.org/10.5281/zenodo.3865011" ID-Zenodo-Dep="3865011" httpUri="https://zenodo.org/record/3865011/files/figure.png" pageId="4" pageNumber="85" startId="4.[151,250,1821,1846]" targetBox="[180,1413,193,1789]" targetPageId="4">
<paragraph id="803036A8FFE5C57473CAF8CEFD8DA84B" blockId="4.[151,1436,1821,1882]" pageId="4" pageNumber="85">
<emphasis id="B2FBEABAFFE5C57473CAF8CEFEB1A827" bold="true" box="[151,271,1821,1846]" pageId="4" pageNumber="85">FIGURE 2</emphasis>
.
<taxonomicName id="478F4D2BFFE5C5747240F8CDFE5DA824" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[285,483,1821,1845]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE5C5747240F8CDFE5DA824" box="[285,483,1821,1845]" italics="true" pageId="4" pageNumber="85">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE5C57472B6F8CDFD80A827" bold="true" box="[491,574,1822,1846]" pageId="4" pageNumber="85">
<taxonomicNameLabel id="A9C857C1FFE5C57472B6F8CDFD80A827" box="[491,574,1822,1846]" pageId="4" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. (A) Hypopygium, dorsal view. (B) Gonostylus. (C) Hypopygium, the arrow indicates the dorsal lamella, dorsal view.
</paragraph>
</caption>
<caption id="D4F06620FFE4C57573CAF861FB5AA8FE" ID-DOI="http://doi.org/10.5281/zenodo.3865013" ID-Zenodo-Dep="3865013" httpUri="https://zenodo.org/record/3865013/files/figure.png" pageId="5" pageNumber="86" startId="5.[151,250,1970,1995]" targetBox="[175,1413,181,1942]" targetPageId="5">
<paragraph id="803036A8FFE4C57573CAF861FB5AA8FE" blockId="5.[151,1436,1970,2031]" pageId="5" pageNumber="86">
<emphasis id="B2FBEABAFFE4C57573CAF861FEB0A8DA" bold="true" box="[151,270,1970,1995]" pageId="5" pageNumber="86">FIGURE 3</emphasis>
.
<taxonomicName id="478F4D2BFFE4C5757246F860FE5EA8DB" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[283,480,1970,1994]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE4C5757246F860FE5EA8DB" box="[283,480,1970,1994]" italics="true" pageId="5" pageNumber="86">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE4C57572BAF860FD86A8DA" bold="true" box="[487,568,1971,1995]" pageId="5" pageNumber="86">
<taxonomicNameLabel id="A9C857C1FFE4C57572BAF860FD86A8DA" box="[487,568,1971,1995]" pageId="5" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. (A) head, frontal view. (B) Thorax, lateral view. (C) Wing. (D) Spur of fore tibia. (E) Genitalia, ventral view (No= notum; SCa= seminal capsule; Csa= coxosternapodeme; Ce= cercus).
</paragraph>
</caption>
<caption id="D4F06620FFE7C57673CAF87BFE14A8F4" ID-DOI="http://doi.org/10.5281/zenodo.3865017" ID-Zenodo-Dep="3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="6" pageNumber="87" startId="6.[151,250,1960,1985]" targetBox="[160,1427,181,1931]" targetPageId="6">
<paragraph id="803036A8FFE7C57673CAF87BFE14A8F4" blockId="6.[151,1437,1960,2021]" pageId="6" pageNumber="87">
<emphasis id="B2FBEABAFFE7C57673CAF87BFEB0A8D0" bold="true" box="[151,270,1960,1985]" pageId="6" pageNumber="87">FIGURE 4</emphasis>
.
<taxonomicName id="478F4D2BFFE7C5767246F879FE5FA8D0" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[283,481,1961,1985]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE7C5767246F879FE5FA8D0" box="[283,481,1961,1985]" italics="true" pageId="6" pageNumber="87">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE7C57672B5F87AFD84A8D0" bold="true" box="[488,570,1961,1985]" pageId="6" pageNumber="87">
<taxonomicNameLabel id="A9C857C1FFE7C57672B5F87AFD84A8D0" box="[488,570,1961,1985]" pageId="6" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view.
</paragraph>
</caption>
<caption id="D4F06620FFE6C57773CAF875FAF6A8F2" ID-DOI="http://doi.org/10.5281/zenodo.3865019" ID-Zenodo-Dep="3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="7" pageNumber="88" startId="7.[151,250,1958,1983]" targetBox="[224,1369,189,1919]" targetPageId="7">
<paragraph id="803036A8FFE6C57773CAF875FAF6A8F2" blockId="7.[151,1436,1958,2019]" pageId="7" pageNumber="88">
<emphasis id="B2FBEABAFFE6C57773CAF875FEB3A8AE" bold="true" box="[151,269,1958,1983]" pageId="7" pageNumber="88">FIGURE 5</emphasis>
.
<taxonomicName id="478F4D2BFFE6C5777244F874FE63A8AF" authority="Dantas &amp; Pinheiro &amp; Hamada, 2020" authorityName="Dantas &amp; Pinheiro &amp; Hamada" authorityYear="2020" box="[281,477,1958,1982]" class="Insecta" family="Chironomidae" genus="Pentaneura" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="herbeti" status="sp. nov.">
<emphasis id="B2FBEABAFFE6C5777244F874FE63A8AF" box="[281,477,1958,1982]" italics="true" pageId="7" pageNumber="88">Pentaneura herbeti</emphasis>
</taxonomicName>
<emphasis id="B2FBEABAFFE6C57772BFF874FD8CA8AE" bold="true" box="[482,562,1959,1983]" pageId="7" pageNumber="88">
<taxonomicNameLabel id="A9C857C1FFE6C57772BFF874FD8CA8AE" box="[482,562,1959,1983]" pageId="7" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end.
</paragraph>
</caption>
<paragraph id="803036A8FFE9C578739AFF4AFECDAEBE" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFF4AFEC9AFA2" box="[199,375,153,179]" italics="true" pageId="8" pageNumber="89">Cephalothorax.</emphasis>
Dorsal surface slightly roughened. Frontal apotome somewhat triangular, 160150 μm wide at base. Wing sheath smooth, 808820 μm long, 310318 μm maximum width (
<figureCitation id="18B42A2DFFE9C578776CFF6EFB32AFC6" box="[1073,1164,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="8" pageNumber="89">Fig. 4B</figureCitation>
). Thoracic horn arched, rounded at apex, external membrane covered with scattered spines (
<figureCitation id="18B42A2DFFE9C57870C5FF32FC4AAFEA" box="[920,1012,225,251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="8" pageNumber="89">Fig. 4A</figureCitation>
), 196203 μm long, 5660 μm maxi- mum width; plastron plate small, oval-shaped, 5255 μm long. Corona well developed, 122126 μm long. Respira- tory atrium occupying about 50% of the thoracic horn volume, directly connected to the subbasal portion of plastron plate by a small neck; one apical and one medial diverticulum present. Basal lobe well developed, 4648 μm long; thoracic comb composed by 1012 tubercles. Dc
<subScript id="1C0B34EDFFE9C57871E0FE51FD7BAE81" attach="left" box="[701,709,386,400]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
42 μm long; Dc
<subScript id="1C0B34EDFFE9C5787022FE51FC39AE81" attach="left" box="[895,903,386,400]" fontSize="6" pageId="8" pageNumber="89">2</subScript>
21 μm long; Dc
<subScript id="1C0B34EDFFE9C578771DFE51FBF6AE81" attach="left" box="[1088,1096,386,400]" fontSize="6" pageId="8" pageNumber="89">3</subScript>
not observed (absent?) in any analyzed specimen.
</paragraph>
<paragraph id="803036A8FFE9C578739AFE6AFC77ADE2" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFE6AFE8FAEC2" box="[199,305,441,467]" italics="true" pageId="8" pageNumber="89">Abdomen</emphasis>
(
<figureCitation id="18B42A2DFFE9C578721CFE6AFE25AEC2" box="[321,411,441,467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="8" pageNumber="89">Fig. 4C</figureCitation>
). Tergite I with large field of shagreen, formed by small scattered spines, scar 130140 μm long; TIITVIII and anal lobe with large field of shagreen formed by small spines arranged in small transverse lines, less defined in TVIII and anal lobe. Sternite I apparently without shagreen, SIIIV with a large field of shagreen composed by spinules arranged in relatively long transversal lines, SVVIII with large field of shagreen composed by spinules arranged in short transversal lines. TVII with 4 LS setae, all in the posterior half. T VIII with 5 lateral filaments, distal margin straight (
<figureCitation id="18B42A2DFFE9C5787150FDBEFDD7AD96" box="[525,617,621,647]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="8" pageNumber="89">Fig. 4D</figureCitation>
). Anal lobe elongated (
<figureCitation id="18B42A2DFFE9C5787033FDBEFC76AD96" box="[878,968,621,647]" captionStart="FIGURE 4" captionStartId="6.[151,250,1960,1985]" captionTargetBox="[160,1427,181,1931]" captionTargetId="figure@6.[160,1427,181,1935]" captionTargetPageId="6" captionText="FIGURE 4. Pentaneura herbeti sp. nov., pupa. (A) Thoracic horn, lateral view. (B) Wing sheath. (C) Abdomen, dorsal view. (D) Anal lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3865017" httpUri="https://zenodo.org/record/3865017/files/figure.png" pageId="8" pageNumber="89">Fig. 4D</figureCitation>
), 374381 μm long, 217222 μm wide at base, with 2 lateral macrosetae with adhesive sheath; outer margins with about 1416 small spinules; inner margin membranous, parallel-sided, without spinules. Genital sac wedge-shaped, 137142 μm long, 178180 μm wide at base, not reaching insertion of the second macrosetae. GS/AL 0.370.38.
</paragraph>
<paragraph id="803036A8FFE9C578739AFD2EFEDAAC06" blockId="8.[151,1437,152,1619]" box="[199,356,765,791]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFD2EFEAEAC06" bold="true" box="[199,272,765,791]" pageId="8" pageNumber="89">Larva</emphasis>
(n= 1).
</paragraph>
<paragraph id="803036A8FFE9C578739AFCF2FEC6AC92" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFCF2FEFCAC2A" box="[199,322,801,827]" italics="true" pageId="8" pageNumber="89">Coloration</emphasis>
. Head yellowish, postoccipital margin brown; apex of mandible dark brown; ligula brownish, darker at apex. Abdomen yellowish; procercus mostly pale, inner margin brown, anal setae light-brown. Claws of posterior parapods yellowish.
</paragraph>
<paragraph id="803036A8FFE9C578739AFC5EFCB1AB02" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFC5EFEBAACB6" box="[199,260,909,935]" italics="true" pageId="8" pageNumber="89">Head</emphasis>
. Head capsule elongate; cephalic index 0.74. Chaetotaxy as in
<figureCitation id="18B42A2DFFE9C5787089FC5EFB80ACB6" box="[980,1086,908,935]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Figures 5</figureCitation>
AB. Dorsally, S5, DP, and S8 nearly aligned diagonally; S8 and S7 somewhat close from each other, aligned laterally; S7 pit large. Ventrally, S10 posterolateral to S9; VP larger than setal pits, closer to S10 than S9; SSm postero-median to S9. SSm, S7 multibranched, S9 bifid, S5 not observed, S8 and S10 simple.
</paragraph>
<paragraph id="803036A8FFE9C578739AFBCEFE10AB4A" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFBCEFE9AAB26" box="[199,292,1053,1079]" italics="true" pageId="8" pageNumber="89">Antenna</emphasis>
. Total length 316 μm, A
<subScript id="1C0B34EDFFE9C5787165FBFDFDFEAB2D" attach="left" box="[568,576,1070,1084]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
255 μm long, 18 μm maximum width, ring organ placed 155 μm from base, A
<subScript id="1C0B34EDFFE9C57873F6FB81FF0DAB71" attach="left" box="[171,179,1106,1120]" fontSize="6" pageId="8" pageNumber="89">2</subScript>
57 μm long. AR 4.05.
</paragraph>
<paragraph id="803036A8FFE9C578739AFBB6FCD3ABB2" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFBB6FEA0AB6E" box="[199,286,1125,1151]" italics="true" pageId="8" pageNumber="89">Maxilla</emphasis>
(
<figureCitation id="18B42A2DFFE9C5787270FBB6FE3BAB6E" box="[301,389,1124,1151]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Fig. 5C</figureCitation>
). Maxillary palp arising from a prominent palpiger. Basal palp segment 38 μm long and 10 μm wide, ring organ placed 21 μm from base. A
<subScript id="1C0B34EDFFE9C57871D8FB49FD33ABB9" attach="right" box="[645,653,1178,1192]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
/P
<subScript id="1C0B34EDFFE9C57871F9FB49FD12ABB9" attach="left" box="[676,684,1178,1192]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
6.71; A
<subScript id="1C0B34EDFFE9C578705BFB49FCB0ABB9" attach="left" box="[774,782,1178,1192]" fontSize="6" pageId="8" pageNumber="89">2</subScript>
/P
<subScript id="1C0B34EDFFE9C5787078FB49FC93ABB9" attach="left" box="[805,813,1178,1192]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
1.50.
</paragraph>
<paragraph id="803036A8FFE9C578739AFB7EFD82ABFA" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFB7EFE8CABD6" box="[199,306,1197,1223]" italics="true" pageId="8" pageNumber="89">Mandible</emphasis>
(
<figureCitation id="18B42A2DFFE9C578721FFB7EFE25ABD6" box="[322,411,1196,1223]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Fig. 5D</figureCitation>
). Curved and gradually narrowed towards apex, 85 μm long, with 3 lateral setae, ring organ placed 20 μm from base. A
<subScript id="1C0B34EDFFE9C578729BFB31FE70ABE1" attach="left" box="[454,462,1250,1264]" fontSize="6" pageId="8" pageNumber="89">1</subScript>
/Md 3.00.
</paragraph>
<paragraph id="803036A8FFE9C578739AFB25FA38AA22" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFB25FE47AA1E" box="[199,505,1269,1295]" italics="true" pageId="8" pageNumber="89">Mentum and M appendage</emphasis>
. Dorsomental teeth reduced, 3 on each side; labial vesicles rounded, 22 μm long. Pseudoradula well-developed, 85 μm long, uniformly granulated. Submentum with a wide rough area (
<figureCitation id="18B42A2DFFE9C5787645FACAFAC7AA22" box="[1304,1401,1304,1331]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Figs. 5E</figureCitation>
).
</paragraph>
<paragraph id="803036A8FFE9C578739AFAEEFDB9AAD2" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFAEEFE57AA46" box="[199,489,1341,1367]" italics="true" pageId="8" pageNumber="89">Hypopharyngeal complex</emphasis>
(
<figureCitation id="18B42A2DFFE9C57872A7FAEEFDEFAA46" box="[506,593,1340,1367]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Fig. 5F</figureCitation>
). Ligula 65 μm long, 40, 22 and 20 μm wide at apex, middle and base re- spectively, with five teeth, anterior row of teeth somewhat straight; tip of inner teeth curved outwardly; granulated at base not expanding laterally. Paraligula bifid and slender, 35 μm long, with a basal spur. Pecten hypopharyngis with about 14 teeth on each side.
</paragraph>
<paragraph id="803036A8FFE9C578739AFA1EFA3CA942" blockId="8.[151,1437,152,1619]" pageId="8" pageNumber="89">
<emphasis id="B2FBEABAFFE9C578739AFA1EFEBFAAF6" box="[199,257,1485,1511]" italics="true" pageId="8" pageNumber="89">Body</emphasis>
(
<figureCitation id="18B42A2DFFE9C578724DFA1EFED4AAF6" box="[272,362,1484,1511]" captionStart="FIGURE 5" captionStartId="7.[151,250,1958,1983]" captionTargetBox="[224,1369,189,1919]" captionTargetId="figure@7.[207,1381,181,1932]" captionTargetPageId="7" captionText="FIGURE 5. Pentaneura herbeti sp. nov., larva. (A) Chaetotaxy of the head, ventrally. (B) Chaetotaxy of the head, dorsally. (C) Basal segment of the maxillary palp. (D) Mandible apex. (E) Submentum (F) Ligula and paraligula. (G) Posterior end." figureDoi="http://doi.org/10.5281/zenodo.3865019" httpUri="https://zenodo.org/record/3865019/files/figure.png" pageId="8" pageNumber="89">Fig. 5G</figureCitation>
). Abdomen without fringe of swim-setae; with scattered lateral and ventral setae, some of which are multi-branched. Anterior parapods with simple claws, curved at apex. Posterior parapods with hooklets and simple claws of varied format. Subbasal seta of posterior parapod simple, 225 μm long. Procercus 175 μm long, 27 μm wide at base, with 7 anal setae, 440 μm long; L/W 6.49. Supraanal seta 560 μm long. Anal tubules damaged.
</paragraph>
</subSubSection>
</treatment>
</document>