492 lines
66 KiB
XML
492 lines
66 KiB
XML
<document id="CCF99A76C49AE01A5A5B0F736E4985E0" ID-CLB-Dataset="45532" ID-DOI="10.5281/zenodo.279945" ID-GBIF-Dataset="29ff95c8-b1d7-4cfa-966c-ca6bd1627974" ID-ISSN="1175-5326" ID-Zenodo-Dep="279945" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434813551" checkinUser="plazi" docAuthor="Pilato, Giovanni, Kiosya, Yevgen, Lisi, Oscar & Sabella, Giorgio" docDate="2012" docId="03F03A099B50B467E5F967FEFB9024F3" docLanguage="en" docName="zt03179p060.pdf" docOrigin="Zootaxa 3179" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Macrobiotus sottilei Pilato, Kiosya, Lisi & Sabella, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="FFC942719B55B46EE56E6649FFE8234B" masterDocTitle="New records of Eutardigrada from Belarus with the description of three new species" masterLastPageNumber="60" masterPageNumber="39" pageNumber="44" updateTime="1698331938802" updateUser="ExternalLinkService">
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<mods:titleInfo id="040D3094CBD8EA35399C3D7CE6DB2F46">
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<mods:title id="4EAFB2083DCAD3D8CC432F17D2BF7A0A">New records of Eutardigrada from Belarus with the description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="1A49B14AD29FB893DEF0A96143F14B31" type="personal">
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<mods:roleTerm id="E9314EBC82CC0CB4C6407A9614B4873C">Author</mods:roleTerm>
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<mods:namePart id="9B98CC8262856DB360F12A16E9DAD6CC">Pilato, Giovanni</mods:namePart>
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<mods:roleTerm id="BCC86761B340B28D479814A1EAB3CEFB">Author</mods:roleTerm>
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<mods:namePart id="2B2014E0DF0E5BFF2BF4369758425829">Kiosya, Yevgen</mods:namePart>
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</mods:name>
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<mods:name id="F5081BC3D60E34EF325C3F5558B180C0" type="personal">
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<mods:roleTerm id="55915A7B4B120DBAAF4B7103E16EFD1B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FCA1E939436331BCE4FB1BF91A1039FE">Lisi, Oscar</mods:namePart>
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</mods:name>
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<mods:name id="CBE80E4B40493C0FB95C8FE48934B97D" type="personal">
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<mods:roleTerm id="36336BDB651C3E5FD67DCF09E32EDBE5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4E9DE41A2225654523DEF1671A5BD38F">Sabella, Giorgio</mods:namePart>
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<mods:typeOfResource id="01601A4B379831BA0132492536E76D85">text</mods:typeOfResource>
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<mods:title id="ECDA79991D4070B136523A0520831C0C">Zootaxa</mods:title>
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<mods:part id="CD9F34B1E20954A4CBB810710B080B33">
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<mods:date id="2DC3C5CBBFAF97A51739FB2347024B18">2012</mods:date>
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<mods:detail id="775239D8A1B47547154F8F567AB1B593" type="volume">
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<mods:number id="937985658F94CB375AA802262A809811">3179</mods:number>
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<mods:identifier id="99A75EA05CFE77899787FF2DF80042C9" type="CLB-Dataset">45532</mods:identifier>
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<mods:identifier id="2C3FC5B1DE7CBAEA5779161FC48951EC" type="DOI">10.5281/zenodo.279945</mods:identifier>
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<treatment id="03F03A099B50B467E5F967FEFB9024F3" ID-DOI="http://doi.org/10.5281/zenodo.6180889" ID-GBIF-Taxon="119431495" ID-Zenodo-Dep="6180889" LSID="urn:lsid:plazi:treatment:03F03A099B50B467E5F967FEFB9024F3" httpUri="http://treatment.plazi.org/id/03F03A099B50B467E5F967FEFB9024F3" lastPageId="9" lastPageNumber="48" pageId="5" pageNumber="44">
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<subSubSection id="C343D8949B50B46BE5F967FEFF0322BF" pageId="5" pageNumber="44" type="nomenclature">
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<paragraph id="8BE68B1F9B50B46BE5F967FEFE35229B" blockId="5.[151,477,439,500]" box="[151,477,439,464]" pageId="5" pageNumber="44">
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<heading id="D0AE3C739B50B46BE5F967FEFE35229B" bold="true" box="[151,477,439,464]" fontSize="10" level="2" pageId="5" pageNumber="44" reason="2">
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||
<emphasis id="B92D570D9B50B46BE5F967FEFE35229B" bold="true" box="[151,477,439,464]" pageId="5" pageNumber="44">
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||
<taxonomicName id="4C59F09C9B50B46BE5F967FEFE94229B" ID-CoL="4a56c760-d1a8-4108-95c4-c07f0fa145c1" box="[151,380,439,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="44" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
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<emphasis id="B92D570D9B50B46BE5F967FEFE94229B" bold="true" box="[151,380,439,464]" italics="true" pageId="5" pageNumber="44">Macrobiotus sottilei</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A21EEA769B50B46BE4ED67F1FE35229B" box="[387,477,440,464]" pageId="5" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BE68B1F9B50B46BE5F96795FF0322BF" blockId="5.[151,477,439,500]" box="[151,235,476,500]" pageId="5" pageNumber="44">
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||
(
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||
<figureCitation id="1362979A9B50B46BE5F16795FF0B22BF" box="[159,227,476,500]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C343D8949B50B46BE5F9646DFBAD208C" pageId="5" pageNumber="44" type="materials_examined">
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<paragraph id="8BE68B1F9B50B46BE5F9646DFF1C21CF" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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<emphasis id="B92D570D9B50B46BE5F9646DFEAE2177" bold="true" box="[151,326,547,572]" pageId="5" pageNumber="44">
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<typeStatus id="54E235BD9B50B46BE5F9646DFF3A2177" box="[151,210,548,572]" pageId="5" pageNumber="44">Type</typeStatus>
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material.
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</emphasis>
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Minsk Oblast, vicinity of Zhodino, Pine forest (approximately 54°6ʹ N, 28°21ʹ E), moss on stone collected (
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<date id="FFE7ADDF9B50B46BE465640EFE2D2114" box="[267,453,583,607]" pageId="5" pageNumber="44" value="2008-11">November 2008</date>
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) by S.A. Dobrovolsky:
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<typeStatus id="54E235BD9B50B46BE7BB640EFCD22114" box="[725,826,583,607]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
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, 39
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<typeStatus id="54E235BD9B50B46BE6046401FC30212B" box="[874,984,584,608]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
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, and
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<specimenCount id="9D5F40969B50B46BE17A6401FB88212B" box="[1044,1120,584,608]" pageId="5" pageNumber="44" type="egg">7 eggs</specimenCount>
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; 2 of which with developed embryo.
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</paragraph>
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<paragraph id="8BE68B1F9B50B46BE5A964D9FAD5205F" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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<emphasis id="B92D570D9B50B46BE5A964D9FE6421E3" bold="true" box="[199,396,655,680]" pageId="5" pageNumber="44">
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<typeStatus id="54E235BD9B50B46BE5A964D9FEEA21E3" box="[199,258,656,680]" pageId="5" pageNumber="44">Type</typeStatus>
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repository:
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</emphasis>
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<typeStatus id="54E235BD9B50B46BE4FC64C6FE1221EC" box="[402,506,655,679]" pageId="5" pageNumber="44" type="holotype">Holotype</typeStatus>
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(slide No. 5428), 16
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<typeStatus id="54E235BD9B50B46BE78964D9FCBB21E3" box="[743,851,656,680]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
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and
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<specimenCount id="9D5F40969B50B46BE6E664C6FC3821E3" box="[904,976,655,680]" pageId="5" pageNumber="44" type="egg">3 eggs</specimenCount>
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(slides No. 5427, 5428 and 5438) deposited in the collection of Binda & Pilato (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania,
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<collectingCountry id="F34ECB8F9B50B46BE403649EFE4D21A4" box="[365,421,727,751]" name="Italy" pageId="5" pageNumber="44">Italy</collectingCountry>
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). 23
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<typeStatus id="54E235BD9B50B46BE4B56491FDAE21BB" box="[475,582,728,752]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
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and
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<specimenCount id="9D5F40969B50B46BE713649EFD2E21BB" box="[637,710,727,752]" pageId="5" pageNumber="44" type="egg">4 eggs</specimenCount>
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from the same moss sample as well as numerous specimens and eggs from other samples are deposited in the collection of Kiosya (Kharkiv National University,
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<collectingCountry id="F34ECB8F9B50B46BE1BC64B5FAD9205F" box="[1234,1329,764,788]" name="Ukraine" pageId="5" pageNumber="44">Ukraine</collectingCountry>
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).
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</paragraph>
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<paragraph id="8BE68B1F9B50B46BE5A96556FBAD208C" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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<emphasis id="B92D570D9B50B46BE5A96556FE4D2073" bold="true" box="[199,421,799,824]" pageId="5" pageNumber="44">Specific diagnosis:</emphasis>
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Cuticle smooth with small pores, eye spots present, dorsal ridges of the buccal armature joined forming a continuous arc, pharyngeal bulb with two macroplacoids and a microplacoid; claws, well developed, of
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<taxonomicName id="4C59F09C9B50B46BE5946521FE8C20CB" box="[250,356,872,896]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="44" phylum="Tardigrada" rank="species" species="hufelandi">
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<emphasis id="B92D570D9B50B46BE5946521FE8C20CB" box="[250,356,872,896]" italics="true" pageId="5" pageNumber="44">hufelandi</emphasis>
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</taxonomicName>
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-
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<typeStatus id="54E235BD9B50B46BE4026521FE4920CB" box="[364,417,872,896]" pageId="5" pageNumber="44">type</typeStatus>
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; accessory points and lunules present; eggs, freely laid, spherical, with trunco-conical processes with indented distal disc. Egg shell with a very dense and regular reticular design with extremely small mesh; that around the base of the processes is not larger than elsewhere on the shell.
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</paragraph>
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</subSubSection>
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<subSubSection id="C343D8949B50B46BE5A9659AFBBB2594" pageId="5" pageNumber="44" type="description">
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<paragraph id="8BE68B1F9B50B46BE5A9659AFDD42744" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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<emphasis id="B92D570D9B50B46BE5A9659AFDFE20A7" bold="true" box="[199,534,979,1004]" pageId="5" pageNumber="44">
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Description of the
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<typeStatus id="54E235BD9B50B46BE4C6659DFDF920A7" box="[424,529,980,1004]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body length 337 µm, colourless; eye spots present. Cuticle smooth with small pores and very small dots on the legs.
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</paragraph>
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<paragraph id="8BE68B1F9B50B46BE5A76255FD6D2647" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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Bucco-pharyngeal apparatus of the
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<taxonomicName id="4C59F09C9B50B46BE7066255FD1E277F" box="[616,758,1052,1076]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="44" phylum="Tardigrada" rank="genus">
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<emphasis id="B92D570D9B50B46BE7066255FD1E277F" box="[616,758,1052,1076]" italics="true" pageId="5" pageNumber="44">Macrobiotus</emphasis>
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</taxonomicName>
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-
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<typeStatus id="54E235BD9B50B46BE7936254FCC7277E" box="[765,815,1053,1077]" pageId="5" pageNumber="44">type</typeStatus>
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(
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<figureCitation id="1362979A9B50B46BE62D6255FC49277F" box="[835,929,1052,1076]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2A</figureCitation>
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); mouth terminal with ten small peribuccal lamellae, a posterior ring of small teeth and, more caudally, a system of three ventral thin transverse ridges and a continuous dorsal transverse ridge (
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<figureCitation id="1362979A9B50B46BE740622DFD612737" box="[558,649,1124,1148]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2B</figureCitation>
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). Buccal tube 34.1 µm long and 4.4 µm wide externally (
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<emphasis id="B92D570D9B50B46BE04D622CFAD12737" box="[1315,1337,1125,1148]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 12.9). Stylet supports inserted on the buccal tube at 76.6 % of its length (
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<emphasis id="B92D570D9B50B46BE61462C0FC6727EB" box="[890,911,1161,1184]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
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||
= 76.6). Pharyngeal bulb (30.6 µm × 25.4 µm) with apophyses, two macroplacoids, and a microplacoid (
|
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<figureCitation id="1362979A9B50B46BE67962E5FC99278F" box="[791,881,1196,1220]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2A</figureCitation>
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). Length of first macroplacoid 9.8 µm (
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||
<emphasis id="B92D570D9B50B46BE04662E4FAD6278F" box="[1320,1342,1197,1220]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 28.7), second 6.3 µm (
|
||
<emphasis id="B92D570D9B50B46BE4386298FE8327A3" box="[342,363,1233,1256]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 18.5), and microplacoid 2.1 µm (
|
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<emphasis id="B92D570D9B50B46BE6786298FCC427A3" box="[790,812,1233,1256]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 6.2); entire placoid row 19.3 µm long (
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||
<emphasis id="B92D570D9B50B46BE0716298FADD27A3" box="[1311,1333,1233,1256]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 56.6); macroplacoid row 17.1 µm long (
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<emphasis id="B92D570D9B50B46BE77E62BCFDCE2647" box="[528,550,1269,1292]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 50.1).
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||
</paragraph>
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<paragraph id="8BE68B1F9B50B46BE5A7635EFD492543" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
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Claws, well developed, of
|
||
<taxonomicName id="4C59F09C9B50B46BE76C6351FD84267B" box="[514,620,1304,1328]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="44" phylum="Tardigrada" rank="species" species="hufelandi">
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||
<emphasis id="B92D570D9B50B46BE76C6351FD84267B" box="[514,620,1304,1328]" italics="true" pageId="5" pageNumber="44">hufelandi</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54E235BD9B50B46BE71D6350FD4D267A" box="[627,677,1305,1329]" pageId="5" pageNumber="44">type</typeStatus>
|
||
(
|
||
<figureCitation id="1362979A9B50B46BE7D6635EFD152664" box="[696,765,1303,1327]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2</figureCitation>
|
||
C-E) with main and secondary branches diverging after a short common portion; accessory points on the main branches present. External and internal claws of the first pair of legs 11.2 µm long (
|
||
<emphasis id="B92D570D9B50B46BE4E36328FE4B2633" box="[397,419,1377,1400]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 32.8) and 10.0 µm long (
|
||
<emphasis id="B92D570D9B50B46BE7BB6328FD032633" box="[725,747,1377,1400]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 29.3) respectively; those of the second pair of legs 12.4 µm long (
|
||
<emphasis id="B92D570D9B50B46BE5B463CCFF1826D7" box="[218,240,1413,1436]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 36.4) and 10.9 µm long (
|
||
<emphasis id="B92D570D9B50B46BE74263CCFDAA26D7" box="[556,578,1413,1436]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 32.0) respectively; length of posterior and anterior claws of the hind legs 12.5 µm (
|
||
<emphasis id="B92D570D9B50B46BE46A63E0FEF2268B" box="[260,282,1449,1472]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 36.7) and 11.3 µm (
|
||
<emphasis id="B92D570D9B50B46BE77A63E0FDC2268B" box="[532,554,1449,1472]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
= 33.1) respectively. Lunules present, small and smooth on the first three pairs of legs, slightly larger and with a delicately indented margin on the hind legs. Two cuticular bars present below the claws on each leg of the first two pairs of legs.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B50B46BE5A9605AFBBB2594" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
|
||
<emphasis id="B92D570D9B50B46BE5A9605AFE352567" bold="true" box="[199,477,1555,1580]" pageId="5" pageNumber="44">Description of the eggs.</emphasis>
|
||
The eggs, freely laid, were spherical, with trunco-conical processes and indented distal disc (
|
||
<figureCitation id="1362979A9B50B46BE594607EFED72504" box="[250,319,1591,1615]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2</figureCitation>
|
||
F, arrows); we counted 29 processes on the circumference and about 120 on the hemisphere. The diameter was about 72 µm excluding the processes and about 83 µm including. The processes were 5.4–6.2 µm high with a basal diameter of 4.8–5.2 µm; the distal discs have a diameter of 3.6–4.4 µm. The egg shell had a uniform, very dense and regular reticular design with extremely small mesh (
|
||
<figureCitation id="1362979A9B50B46BE6BD60EDFBFC25F7" box="[979,1044,1700,1724]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="5" pageNumber="44">Fig. 2</figureCitation>
|
||
G, arrows), that showed no distinction between that around the base of the processes and the between process egg shell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343D8949B50B46BE5A960A5FDF32407" pageId="5" pageNumber="44" type="discussion">
|
||
<paragraph id="8BE68B1F9B50B46BE5A960A5FDF32407" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
|
||
<emphasis id="B92D570D9B50B46BE5A960A5FED3244F" bold="true" box="[199,315,1772,1796]" pageId="5" pageNumber="44">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54E235BD9B50B46BE41B60A4FE08244E" box="[373,480,1773,1797]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
|
||
were similar to the
|
||
<typeStatus id="54E235BD9B50B46BE7AF60A5FCCB244F" box="[705,803,1772,1796]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
in both qualitative and metric characters (we referred to the
|
||
<emphasis id="B92D570D9B50B46BE5AD6158FF312463" box="[195,217,1809,1832]" italics="true" pageId="5" pageNumber="44">pt</emphasis>
|
||
index values when the specimens had different body length). The measurements of the
|
||
<typeStatus id="54E235BD9B50B46BE1BA6146FADE246C" box="[1236,1334,1807,1831]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="54E235BD9B50B46BE5F9617CFEEA2406" box="[151,258,1845,1869]" pageId="5" pageNumber="44" type="paratype">paratypes</typeStatus>
|
||
are presented in Table 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343D8949B50B467E5A96111FB9024F3" lastPageId="9" lastPageNumber="48" pageId="5" pageNumber="44" type="etymology">
|
||
<paragraph id="8BE68B1F9B50B46BE5A96111FD0024DF" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
|
||
<emphasis id="B92D570D9B50B46BE5A96111FEA6243B" bold="true" box="[199,334,1880,1904]" pageId="5" pageNumber="44">Etymology.</emphasis>
|
||
The specific name is in honour of the histologist Lorenzo Sottile, who has recently died and who had been a former scholar and colleague of G. Pilato.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B50B46BE5A961D6FA742497" blockId="5.[151,1436,547,2012]" pageId="5" pageNumber="44">
|
||
<emphasis id="B92D570D9B50B46BE5A961D6FE2624F3" bold="true" box="[199,462,1951,1976]" pageId="5" pageNumber="44">Differential diagnosis.</emphasis>
|
||
Within the genus
|
||
<taxonomicName id="4C59F09C9B50B46BE7F261E9FCC224F3" box="[668,810,1952,1976]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="44" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B92D570D9B50B46BE7F261E9FCC224F3" box="[668,810,1952,1976]" italics="true" pageId="5" pageNumber="44">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
11 species were known to have two macroplacoids and a microplacoid and, in the buccal armature, the dorsal transverse ridges joined to form a continuous arc. Three,
|
||
<emphasis id="B92D570D9B50B46BE013618CFA742497" box="[1405,1436,1989,2012]" italics="true" pageId="5" pageNumber="44">M.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B53B468E5F966D1FDC4222F" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="45">
|
||
<taxonomicName id="4C59F09C9B53B468E5F966D1FEDA23FB" box="[151,306,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="echinogenitus">
|
||
<emphasis id="B92D570D9B53B468E5F966D1FEDA23FB" box="[151,306,152,176]" italics="true" pageId="6" pageNumber="45">echinogenitus</emphasis>
|
||
</taxonomicName>
|
||
(according to
|
||
<bibRefCitation id="EFC8F6EE9B53B468E4B866DEFD8B23E4" author="Binda" box="[470,611,151,175]" pageId="6" pageNumber="45" refString="Binda, M. G. (1988) Ridescrizione di Macrobiotus echinogenitus Richters, 1904 e sul valore di buona specie di Macrobiotus crenulatus Richters, 1904 (Eutardigrada). Animalia, 15, 201 - 210." type="journal article" year="1988">Binda, 1988</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C59F09C9B53B468E71966D0FCC323FB" box="[631,811,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="semmelweisi">
|
||
<emphasis id="B92D570D9B53B468E71966D0FCC323FB" box="[631,811,152,176]" italics="true" pageId="6" pageNumber="45">M. semmelweisi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B92D570D9B53B468E60C66D0FBCA23FB" box="[866,1058,152,176]" italics="true" pageId="6" pageNumber="45">
|
||
<taxonomicName id="4C59F09C9B53B468E60C66D0FBF623FB" box="[866,1054,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="drakensbergi">M. drakensbergi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
are clearly different from
|
||
<taxonomicName id="4C59F09C9B53B468E02266D0FF2B239F" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B53B468E02266D0FA8B23FB" box="[1356,1379,153,176]" italics="true" pageId="6" pageNumber="45">M</emphasis>
|
||
.
|
||
<emphasis id="B92D570D9B53B468E01C66D0FF2B239F" italics="true" pageId="6" pageNumber="45">sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B53B468E5A566F5FECF239F" box="[203,295,188,212]" pageId="6" pageNumber="45" rank="species">
|
||
<emphasis id="B92D570D9B53B468E5A566F5FECF239F" bold="true" box="[203,295,188,212]" pageId="6" pageNumber="45">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in characters of both the specimens and eggs:
|
||
<taxonomicName id="4C59F09C9B53B468E62C66F4FC15239F" box="[834,1021,188,212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="echinogentus">
|
||
<emphasis id="B92D570D9B53B468E62C66F4FC15239F" box="[834,1021,188,212]" italics="true" pageId="6" pageNumber="45">M. echinogentus</emphasis>
|
||
</taxonomicName>
|
||
has more visible teeth in the buccal cavity wall and eggs with large conical processes.
|
||
<taxonomicName id="4C59F09C9B53B468E7BD66A8FC6F23B3" box="[723,903,224,248]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="semmelweisi">
|
||
<emphasis id="B92D570D9B53B468E7BD66A8FC6F23B3" box="[723,903,224,248]" italics="true" pageId="6" pageNumber="45">M. semmelweisi</emphasis>
|
||
</taxonomicName>
|
||
has bands of dorsal gibbosities, stylet supports inserted on the buccal tube in a more cephalic position (
|
||
<emphasis id="B92D570D9B53B468E662674CFCCA2257" box="[780,802,261,284]" italics="true" pageId="6" pageNumber="45">pt</emphasis>
|
||
=
|
||
<quantity id="4CA126FA9B53B468E62E674DFC4B2257" box="[832,931,260,284]" metricMagnitude="0" metricUnit="m" metricValue="1.8415" metricValueMax="1.8542" metricValueMin="1.8288" pageId="6" pageNumber="45" unit="in" value="72.5" valueMax="73.0" valueMin="72.0">72–73 in</quantity>
|
||
<taxonomicName id="4C59F09C9B53B468E6C4674CFBB62257" box="[938,1118,260,284]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="semmelweisi">
|
||
<emphasis id="B92D570D9B53B468E6C4674CFBB62257" box="[938,1118,260,284]" italics="true" pageId="6" pageNumber="45">M. semmelweisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CA126FA9B53B468E105674DFB112257" box="[1131,1273,260,284]" metricMagnitude="0" metricUnit="m" metricValue="1.92913" metricValueMax="1.9456399999999998" metricValueMin="1.91262" pageId="6" pageNumber="45" unit="in" value="75.94999999999999" valueMax="76.6" valueMin="75.3">75.3–76.6 in</quantity>
|
||
M.
|
||
<taxonomicName id="4C59F09C9B53B468E049674DFA9B2257" box="[1319,1395,260,284]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B53B468E049674DFA9B2257" box="[1319,1395,260,284]" italics="true" pageId="6" pageNumber="45">sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B53B468E014674DFF20220B" pageId="6" pageNumber="45" rank="species">
|
||
<emphasis id="B92D570D9B53B468E014674DFF20220B" bold="true" pageId="6" pageNumber="45">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
), claws stout and eggs with slender conical processes.
|
||
<taxonomicName id="4C59F09C9B53B468E6596761FB88220B" box="[823,1120,296,320]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="drakensbergi">
|
||
<emphasis id="B92D570D9B53B468E6596761FB88220B" box="[823,1120,296,320]" italics="true" pageId="6" pageNumber="45">Macrobiotus drakensbergi</emphasis>
|
||
</taxonomicName>
|
||
has strongly indented lunulae and eggs with conical processes.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B53B468E5A96726FEF922E7" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="45">Comparing adults of, where possible, comparable size the new species can be from the other species (Table 2) as follows:</paragraph>
|
||
<paragraph id="8BE68B1F9B53B468E5A967FEFF2022BF" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="45">
|
||
Characters of the eggs easily distinguish two species,
|
||
<taxonomicName id="4C59F09C9B53B468E65A67F0FC51229B" box="[820,953,440,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="diversus">
|
||
<emphasis id="B92D570D9B53B468E65A67F0FC51229B" box="[820,953,440,464]" italics="true" pageId="6" pageNumber="45">M. diversus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C59F09C9B53B468E69A67F0FB45229B" box="[1012,1197,440,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="seychellensis">
|
||
<emphasis id="B92D570D9B53B468E69A67F0FB45229B" box="[1012,1197,440,464]" italics="true" pageId="6" pageNumber="45">M. seychellensis</emphasis>
|
||
</taxonomicName>
|
||
, from
|
||
<taxonomicName id="4C59F09C9B53B468E19367F0FA99229B" box="[1277,1393,440,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B53B468E19367F0FAFC229B" box="[1277,1300,441,464]" italics="true" pageId="6" pageNumber="45">M</emphasis>
|
||
.
|
||
<emphasis id="B92D570D9B53B468E04B67F1FA99229B" box="[1317,1393,440,464]" italics="true" pageId="6" pageNumber="45">sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B53B468E01467F1FF2022BF" pageId="6" pageNumber="45" rank="species">
|
||
<emphasis id="B92D570D9B53B468E01467F1FF2022BF" bold="true" pageId="6" pageNumber="45">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B53B468E5A967B6FD7521CF" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="45">
|
||
The eggs of
|
||
<taxonomicName id="4C59F09C9B53B468E43C6448FE3E2153" box="[338,470,512,536]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="diversus">
|
||
<emphasis id="B92D570D9B53B468E43C6448FE3E2153" box="[338,470,512,536]" italics="true" pageId="6" pageNumber="45">M. diversus</emphasis>
|
||
</taxonomicName>
|
||
had egg processes shaped as a bowling-pin with cup-shaped extremities (the diameter of which was clearly shorter than the basal diameter of the processes); the egg shell was rough but without a true reticular design. In
|
||
<taxonomicName id="4C59F09C9B53B468E4016400FE37212B" box="[367,479,584,608]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B53B468E4016400FE37212B" box="[367,479,584,608]" italics="true" pageId="6" pageNumber="45">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B53B468E4886401FDD7212B" box="[486,575,584,608]" pageId="6" pageNumber="45" rank="species">
|
||
<emphasis id="B92D570D9B53B468E4886401FDD7212B" bold="true" box="[486,575,584,608]" pageId="6" pageNumber="45">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the distal disc of each egg process was larger and had a clearly indented margin; the egg shell had a dense reticular design.
|
||
</paragraph>
|
||
<caption id="DF26DB979B53B468E5F96116FC4524BB" httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="6" pageNumber="45" targetBox="[181,1386,723,1826]" targetPageId="6">
|
||
<paragraph id="8BE68B1F9B53B468E5F96116FC4524BB" blockId="6.[151,1436,1887,2032]" pageId="6" pageNumber="45">
|
||
<emphasis id="B92D570D9B53B468E5F96116FEE5243E" bold="true" box="[151,269,1887,1910]" pageId="6" pageNumber="45">FIGURE 2</emphasis>
|
||
. A–D,
|
||
<taxonomicName id="4C59F09C9B53B468E4326129FDC1243D" box="[348,553,1888,1910]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="45" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B53B468E4326129FDC1243D" box="[348,553,1888,1910]" italics="true" pageId="6" pageNumber="45">Macrobiotus sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B53B468E75C6129FD6B243D" box="[562,643,1888,1910]" pageId="6" pageNumber="45" rank="species">
|
||
<emphasis id="B92D570D9B53B468E75C6129FD95243E" bold="true" box="[562,637,1888,1909]" pageId="6" pageNumber="45">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF26DB979B5DB466E5F96050FBB72526" httpUri="https://zenodo.org/record/279948/files/figure.png" pageId="8" pageNumber="47" targetBox="[195,1383,198,1484]" targetPageId="8">
|
||
<paragraph id="8BE68B1F9B5DB466E5F96050FBB72526" blockId="8.[151,1436,1561,1646]" pageId="8" pageNumber="47">
|
||
<emphasis id="B92D570D9B5DB466E5F96050FEFC2564" bold="true" box="[151,276,1561,1584]" pageId="8" pageNumber="47">FIGURE 3.</emphasis>
|
||
A,B,
|
||
<taxonomicName id="4C59F09C9B5DB466E43A6053FD88257B" box="[340,608,1562,1584]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="seychellensis">
|
||
<emphasis id="B92D570D9B5DB466E43A6053FD88257B" box="[340,608,1562,1584]" italics="true" pageId="8" pageNumber="47">Macrobiotus seychellensis</emphasis>
|
||
</taxonomicName>
|
||
: A, Bucco-pharyngeal apparatus of a paratype (slide No. 4227). B, Detail of an egg. The arrows indicate terminal discs of the processes with margins prolonged in well visible lobes. C, D,
|
||
<taxonomicName id="4C59F09C9B5DB466E18C6071FF512526" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="sapiens">
|
||
<emphasis id="B92D570D9B5DB466E18C6071FF512526" italics="true" pageId="8" pageNumber="47">Macrobiotus sapiens</emphasis>
|
||
</taxonomicName>
|
||
(slide No.3309). C, Bucco-pharyngeal apparatus. D, Detail of an egg. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE68B1F9B5DB466E5A960DDFEBD24CF" blockId="8.[151,1436,1684,2033]" pageId="8" pageNumber="47">
|
||
The egg processes of
|
||
<taxonomicName id="4C59F09C9B5DB466E4D260DFFD9C25E6" box="[444,628,1685,1709]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="seychellensis">
|
||
<emphasis id="B92D570D9B5DB466E4D260DFFD9C25E6" box="[444,628,1685,1709]" italics="true" pageId="8" pageNumber="47">M. seychellensis</emphasis>
|
||
</taxonomicName>
|
||
had a terminal disc with a diameter greater than the basal diameter and the terminal disc margin prolonged into obvious lobes, very different from
|
||
<taxonomicName id="4C59F09C9B5DB466E6B060F3FBA7259A" box="[990,1103,1721,1745]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5DB466E6B060F3FBA7259A" box="[990,1103,1721,1745]" italics="true" pageId="8" pageNumber="47">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5DB466E13860F0FB58259A" box="[1110,1200,1721,1745]" pageId="8" pageNumber="47" rank="species">
|
||
<emphasis id="B92D570D9B5DB466E13860F0FB58259A" bold="true" box="[1110,1200,1721,1745]" pageId="8" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
where the distal disc of each egg process was shorter than the basal diameter and the margin indented (
|
||
<figureCitation id="1362979A9B5DB466E1296095FB6625BF" box="[1095,1166,1756,1780]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="8" pageNumber="47">Figs 2</figureCitation>
|
||
F and 3B, arrows). The reticular design of the egg shell of
|
||
<taxonomicName id="4C59F09C9B5DB466E74F614BFD7B2452" box="[545,659,1793,1817]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5DB466E74F614BFDD02452" box="[545,568,1794,1817]" italics="true" pageId="8" pageNumber="47">M</emphasis>
|
||
.
|
||
<emphasis id="B92D570D9B5DB466E7296148FD7B2452" box="[583,659,1793,1817]" italics="true" pageId="8" pageNumber="47">sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5DB466E7F56148FD1D2453" box="[667,757,1793,1817]" pageId="8" pageNumber="47" rank="species">
|
||
<emphasis id="B92D570D9B5DB466E7F56148FD062452" bold="true" box="[667,750,1793,1817]" pageId="8" pageNumber="47">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was more dense than the eggs of
|
||
<taxonomicName id="4C59F09C9B5DB466E11A614BFAC42452" box="[1140,1324,1793,1817]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="seychellensis">
|
||
<emphasis id="B92D570D9B5DB466E11A614BFAC42452" box="[1140,1324,1793,1817]" italics="true" pageId="8" pageNumber="47">M. seychellensis</emphasis>
|
||
</taxonomicName>
|
||
, having a clearly smaller mesh (
|
||
<figureCitation id="1362979A9B5DB466E4E0616DFE342477" box="[398,476,1828,1852]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="8" pageNumber="47">Figs. 2</figureCitation>
|
||
G and 3B). In addition, with respect to adult animals,
|
||
<taxonomicName id="4C59F09C9B5DB466E156616FFB412476" box="[1080,1193,1829,1853]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5DB466E156616FFB412476" box="[1080,1193,1829,1853]" italics="true" pageId="8" pageNumber="47">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5DB466E1DE616CFAE22476" box="[1200,1290,1829,1853]" pageId="8" pageNumber="47" rank="species">
|
||
<emphasis id="B92D570D9B5DB466E1DE616CFAE22476" bold="true" box="[1200,1290,1829,1853]" pageId="8" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
had a longer second placoid, and claws with a shorter common portion with respect to the total length of the claws than seen in
|
||
<taxonomicName id="4C59F09C9B5DB466E5F96127FEA624CE" box="[151,334,1901,1925]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="seychellensis">
|
||
<emphasis id="B92D570D9B5DB466E5F96127FEA624CE" box="[151,334,1901,1925]" italics="true" pageId="8" pageNumber="47">M. seychellensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5DB466E5A961DBFC2124BB" blockId="8.[151,1436,1684,2033]" pageId="8" pageNumber="47">
|
||
<taxonomicName id="4C59F09C9B5DB466E5A961DBFED024E2" box="[199,312,1937,1961]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5DB466E5A961DBFED024E2" box="[199,312,1937,1961]" italics="true" pageId="8" pageNumber="47">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5DB466E42E61D8FE7224E2" box="[320,410,1937,1961]" pageId="8" pageNumber="47" rank="species">
|
||
<emphasis id="B92D570D9B5DB466E42E61D8FE7224E2" bold="true" box="[320,410,1937,1961]" pageId="8" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<taxonomicName id="4C59F09C9B5DB466E75061DBFD5424E2" box="[574,700,1937,1961]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="47" phylum="Tardigrada" rank="species" species="biserovi">
|
||
<emphasis id="B92D570D9B5DB466E75061DBFD5424E2" box="[574,700,1937,1961]" italics="true" pageId="8" pageNumber="47">M. biserovi</emphasis>
|
||
</taxonomicName>
|
||
in having a longer second macroplacoid, more slender claws and clearly different eggs, with smaller processes, distal disc diameter shorter than the basal diameter, margins of distal discs clearly indented, and a more dense reticular design of the egg shell.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5CB467E5A966D0FDB323BC" blockId="9.[151,1436,151,356]" pageId="9" pageNumber="48">
|
||
<taxonomicName id="4C59F09C9B5CB467E5A966D0FED123FB" box="[199,313,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5CB467E5A966D0FED123FB" box="[199,313,152,176]" italics="true" pageId="9" pageNumber="48">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5CB467E42C66D1FE7423E4" box="[322,412,152,176]" pageId="9" pageNumber="48" rank="species">
|
||
<emphasis id="B92D570D9B5CB467E42C66D1FE7D23FB" bold="true" box="[322,405,152,176]" pageId="9" pageNumber="48">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<taxonomicName id="4C59F09C9B5CB467E72D66D0FD5523FB" box="[579,701,153,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="sapiens">
|
||
<emphasis id="B92D570D9B5CB467E72D66D0FD5523FB" box="[579,701,153,176]" italics="true" pageId="9" pageNumber="48">M. sapiens</emphasis>
|
||
</taxonomicName>
|
||
in having a narrower buccal tube, stylet supports inserted on the buccal tube in a more cephalic position (Table 2;
|
||
<figureCitation id="1362979A9B5CB467E7B266F5FCA8239F" box="[732,832,188,212]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs. 2A</figureCitation>
|
||
and
|
||
<figureCitation id="1362979A9B5CB467E61266F5FC62239F" box="[892,906,188,212]" captionStart="FIGURE 3. A, B" captionStartId="8.[151,250,1561,1584]" captionTargetBox="[195,1383,198,1484]" captionTargetId="figure@8.[151,1436,155,1540]" captionTargetPageId="8" captionText="FIGURE 3. A, B, Macrobiotus seychellensis: A, Bucco-pharyngeal apparatus of a paratype (slide No. 4227). B, Detail of an egg. The arrows indicate terminal discs of the processes with margins prolonged in well visible lobes. C, D, Macrobiotus sapiens (slide No. 3309). C, Bucco-pharyngeal apparatus. D, Detail of an egg. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279948/files/figure.png" pageId="9" pageNumber="48">3</figureCitation>
|
||
C), and a more dense, smaller mesh, reticular design of the egg shell (
|
||
<figureCitation id="1362979A9B5CB467E4CF6696FE3523BC" box="[417,477,223,247]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Fig 2</figureCitation>
|
||
G and 3D).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5CB467E5A9674DFCCF222F" blockId="9.[151,1436,151,356]" pageId="9" pageNumber="48">
|
||
The new species differed from
|
||
<taxonomicName id="4C59F09C9B5CB467E74A674CFD5A2257" box="[548,690,260,284]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="modestus">
|
||
<emphasis id="B92D570D9B5CB467E74A674CFD5A2257" box="[548,690,260,284]" italics="true" pageId="9" pageNumber="48">M. modestus</emphasis>
|
||
</taxonomicName>
|
||
in having a wider buccal tube; stylet supports inserted on the buccal tube in a more caudal position; longer placoids (Table 2;
|
||
<figureCitation id="1362979A9B5CB467E624676EFC4D2274" box="[842,933,295,319]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs 2A</figureCitation>
|
||
and
|
||
<figureCitation id="1362979A9B5CB467E6B3676EFBEB2274" box="[989,1027,295,319]" captionStart="FIGURE 4. A, B" captionStartId="9.[151,250,1536,1559]" captionTargetBox="[217,1355,435,1477]" captionTargetId="figure@9.[191,1395,409,1515]" captionTargetPageId="9" captionText="FIGURE 4. A, B, Macrobiotus modestus: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5250). B, Detail of an egg. C, D, Macrobiotus humilis: C, Bucco-pharyngeal apparatus of the holotype (slide No. 4810). D, Detail of an egg. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279949/files/figure.png" pageId="9" pageNumber="48">4A</figureCitation>
|
||
), and the reticular design of the egg shell was more delicate and less visible (
|
||
<figureCitation id="1362979A9B5CB467E7306705FD43222F" box="[606,683,332,356]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs. 2</figureCitation>
|
||
G and 4B).
|
||
</paragraph>
|
||
<caption id="DF26DB979B5CB467E5F96049FF002518" httpUri="https://zenodo.org/record/279949/files/figure.png" pageId="9" pageNumber="48" targetBox="[217,1355,435,1477]" targetPageId="9">
|
||
<paragraph id="8BE68B1F9B5CB467E5F96049FF002518" blockId="9.[151,1437,1536,1619]" pageId="9" pageNumber="48">
|
||
<emphasis id="B92D570D9B5CB467E5F96049FEFA255D" bold="true" box="[151,274,1536,1559]" pageId="9" pageNumber="48">FIGURE 4.</emphasis>
|
||
A, B,
|
||
<taxonomicName id="4C59F09C9B5CB467E43D6049FDD0255D" box="[339,568,1536,1558]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="modestus">
|
||
<emphasis id="B92D570D9B5CB467E43D6049FDD0255D" box="[339,568,1536,1558]" italics="true" pageId="9" pageNumber="48">Macrobiotus modestus</emphasis>
|
||
</taxonomicName>
|
||
: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5250). B, Detail of an egg. C, D,
|
||
<taxonomicName id="4C59F09C9B5CB467E5BD6056FE4D257E" box="[211,421,1567,1589]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="humilis">
|
||
<emphasis id="B92D570D9B5CB467E5BD6056FE4D257E" box="[211,421,1567,1589]" italics="true" pageId="9" pageNumber="48">Macrobiotus humilis</emphasis>
|
||
</taxonomicName>
|
||
: C, Bucco-pharyngeal apparatus of the holotype (slide No. 4810). D, Detail of an egg. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE68B1F9B5CB467E5A960C8FBEC25AB" blockId="9.[151,1436,1664,1977]" pageId="9" pageNumber="48">
|
||
<taxonomicName id="4C59F09C9B5CB467E5A960C8FED025D3" box="[199,312,1664,1688]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5CB467E5A960C8FED025D3" box="[199,312,1664,1688]" italics="true" pageId="9" pageNumber="48">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5CB467E45160C8FE7125D3" box="[319,409,1665,1689]" pageId="9" pageNumber="48" rank="species">
|
||
<emphasis id="B92D570D9B5CB467E45160C8FE7A25D2" bold="true" box="[319,402,1665,1689]" pageId="9" pageNumber="48">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<taxonomicName id="4C59F09C9B5CB467E75260C8FD5D25D3" box="[572,693,1664,1688]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="humilis">
|
||
<emphasis id="B92D570D9B5CB467E75260C8FD5D25D3" box="[572,693,1664,1688]" italics="true" pageId="9" pageNumber="48">M. humilis</emphasis>
|
||
</taxonomicName>
|
||
in having the stylet supports inserted on the buccal tube in a more caudal position; longer placoids (Table 2;
|
||
<figureCitation id="1362979A9B5CB467E71B60EDFD2725F7" box="[629,719,1700,1724]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs 2A</figureCitation>
|
||
and
|
||
<figureCitation id="1362979A9B5CB467E66660EDFCF025F7" box="[776,792,1700,1724]" captionStart="FIGURE 4. A, B" captionStartId="9.[151,250,1536,1559]" captionTargetBox="[217,1355,435,1477]" captionTargetId="figure@9.[191,1395,409,1515]" captionTargetPageId="9" captionText="FIGURE 4. A, B, Macrobiotus modestus: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5250). B, Detail of an egg. C, D, Macrobiotus humilis: C, Bucco-pharyngeal apparatus of the holotype (slide No. 4810). D, Detail of an egg. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279949/files/figure.png" pageId="9" pageNumber="48">4</figureCitation>
|
||
C); longer and more slender claws (Table 2) with shorter common tract; and less visible reticular design of egg shell (
|
||
<figureCitation id="1362979A9B5CB467E6576081FC6E25AB" box="[825,902,1736,1760]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs. 2</figureCitation>
|
||
G and 4D).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5CB467E5A960A5FBD0243B" blockId="9.[151,1436,1664,1977]" pageId="9" pageNumber="48">
|
||
The new species differed from
|
||
<taxonomicName id="4C59F09C9B5CB467E74960A7FD3C244E" box="[551,724,1773,1797]" genus="Macrobotus" pageId="9" pageNumber="48" rank="species" species="madegassus">
|
||
<emphasis id="B92D570D9B5CB467E74960A7FD3C244E" box="[551,724,1773,1797]" italics="true" pageId="9" pageNumber="48">M. madegassus</emphasis>
|
||
</taxonomicName>
|
||
in having eye spots; slightly wider buccal tube; stylet supports inserted on the buccal tube in a more caudal position, longer placoids (Table 2;
|
||
<figureCitation id="1362979A9B5CB467E1496159FB6A2463" box="[1063,1154,1808,1832]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs 2A</figureCitation>
|
||
and
|
||
<figureCitation id="1362979A9B5CB467E1D26159FB0A2463" box="[1212,1250,1808,1832]" captionStart="FIGURE 5. A, B" captionStartId="10.[151,250,1427,1450]" captionTargetBox="[170,1405,206,1380]" captionTargetId="figure@10.[151,1436,195,1406]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Macrobiotus madegassus: A, Bucco-pharyngeal apparatus of a paratype from the Maucci collection. B, Detail of an egg. C, D, Macrobiotus iharosi: C, Bucco-pharyngeal apparatus of the holotype (slide No. 3706). D, Detail of an egg. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279950/files/figure.png" pageId="9" pageNumber="48">5A</figureCitation>
|
||
); slightly longer and more slender claws with shorter common tract; margins of the distal discs of the egg processes more clearly indented; and a smaller mesh reticular design of the egg shell (
|
||
<figureCitation id="1362979A9B5CB467E63A6111FC49243B" box="[852,929,1880,1904]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs. 2</figureCitation>
|
||
F, G and 5B).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5CB467E5A96137FB9024F3" blockId="9.[151,1436,1664,1977]" pageId="9" pageNumber="48">
|
||
<taxonomicName id="4C59F09C9B5CB467E5A96137FED124DE" box="[199,313,1917,1941]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="sottilei" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5CB467E5A96137FED124DE" box="[199,313,1917,1941]" italics="true" pageId="9" pageNumber="48">M. sottilei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5CB467E42C6134FE7524DF" box="[322,413,1917,1941]" pageId="9" pageNumber="48" rank="species">
|
||
<emphasis id="B92D570D9B5CB467E42C6134FE7E24DE" bold="true" box="[322,406,1917,1941]" pageId="9" pageNumber="48">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<taxonomicName id="4C59F09C9B5CB467E72B6137FD5324DE" box="[581,699,1917,1941]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="48" phylum="Tardigrada" rank="species" species="iharosi">
|
||
<emphasis id="B92D570D9B5CB467E72B6137FD5324DE" box="[581,699,1917,1941]" italics="true" pageId="9" pageNumber="48">M. iharosi</emphasis>
|
||
</taxonomicName>
|
||
in having a slightly wider buccal tube; longer placoids (Table 2;
|
||
<figureCitation id="1362979A9B5CB467E5F961E9FF1924F3" box="[151,241,1952,1976]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs 2A</figureCitation>
|
||
and
|
||
<figureCitation id="1362979A9B5CB467E44961E9FEDD24F3" box="[295,309,1952,1976]" captionStart="FIGURE 5. A, B" captionStartId="10.[151,250,1427,1450]" captionTargetBox="[170,1405,206,1380]" captionTargetId="figure@10.[151,1436,195,1406]" captionTargetPageId="10" captionText="FIGURE 5. A, B, Macrobiotus madegassus: A, Bucco-pharyngeal apparatus of a paratype from the Maucci collection. B, Detail of an egg. C, D, Macrobiotus iharosi: C, Bucco-pharyngeal apparatus of the holotype (slide No. 3706). D, Detail of an egg. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279950/files/figure.png" pageId="9" pageNumber="48">5</figureCitation>
|
||
C); and a smaller mesh reticular design of the egg shell (
|
||
<figureCitation id="1362979A9B5CB467E6C361E9FC1224F3" box="[941,1018,1952,1976]" captionStart="FIGURE 2. A – D" captionStartId="6.[151,250,1887,1910]" captionTargetBox="[181,1386,723,1826]" captionTargetId="figure@6.[151,1436,697,1866]" captionTargetPageId="6" captionText="FIGURE 2. A – D, Macrobiotus sottilei sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5428). B, Buccopharyngeal apparatus the holotype. The arrow indicates the dorsal transverse ridges forming a continuous arc. C, Claws of the first pair of legs of the holotype. D, Claws of the second pair of legs of the holotype. E, Claws of the third pair of legs of the holotype. F, Detail of an egg. The arrows indicate indented terminal discs of the egg processes. G, Detail of an egg. The arrows indicate the very dense reticular design of the egg shell. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279947/files/figure.png" pageId="9" pageNumber="48">Figs. 2</figureCitation>
|
||
G and 5D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |