treatments-xml/data/03/D2/1F/03D21F54FFD4FFBE77F3FDA8FC2BF88F.xml
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<mods:title id="A9B4E4BCE30719709A21A976DB84F1B4">Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages</mods:title>
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<mods:namePart id="F79AC477DE0BFE9E232EF10D2938F86B">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="1B2F79010A98006E2A9F142D30611A0F">Kodada, Ján</mods:namePart>
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<treatment id="03D21F54FFD4FFBE77F3FDA8FC2BF88F" ID-DOI="http://doi.org/10.5281/zenodo.6031711" ID-GBIF-Taxon="132585009" ID-Zenodo-Dep="6031711" LSID="urn:lsid:plazi:treatment:03D21F54FFD4FFBE77F3FDA8FC2BF88F" httpUri="http://treatment.plazi.org/id/03D21F54FFD4FFBE77F3FDA8FC2BF88F" lastPageNumber="126" pageId="5" pageNumber="126">
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<paragraph id="8BC4AE42FFD4FFBE77F3FDA8FFAAFB65" blockId="5.[151,681,1158,1218]" box="[151,681,1158,1184]" pageId="5" pageNumber="126">
<heading id="D08C192EFFD4FFBE77F3FDA8FFAAFB65" bold="true" box="[151,681,1158,1184]" fontSize="11" level="1" pageId="5" pageNumber="126" reason="1">
<taxonomicName id="4C7BD5C1FFD4FFBE77F3FDA8FFAAFB65" ID-CoL="66NG7" authority="Freitag &amp; Kodada, 2017" authorityName="Freitag &amp; Kodada" authorityYear="2017" box="[151,681,1158,1184]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="tobada">
<emphasis id="B90F7250FFD4FFBE77F3FDA8FFAAFB65" bold="true" box="[151,681,1158,1184]" pageId="5" pageNumber="126">
<emphasis id="B90F7250FFD4FFBE77F3FDA8FC70FB65" bold="true" box="[151,371,1158,1184]" italics="true" pageId="5" pageNumber="126">Ancyronyx tobada</emphasis>
<bibRefCitation id="EFEAD3B3FFD4FFBE761FFDA8FFAAFB65" author="Freitag" box="[379,681,1158,1184]" pageId="5" pageNumber="126" refString="Freitag, H. &amp; Kodada, J. (2017) A taxonomic review of the genus Ancyronyx Erichson, 1847 from Sulawesi (Insecta: Coleoptera: Elmidae). Journal Natural History, 51, 561 - 606." type="journal article" year="2017">Freitag &amp; Kodada, 2017</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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(
<figureCitation id="1340B2C7FFD4FFBE77FBFD87FDE5FB07" box="[159,230,1193,1218]" captionStart="FIGURE 2" captionStartId="3.[151,250,1822,1844]" captionTargetBox="[170,1430,878,1783]" captionTargetId="figure@3.[151,1436,873,1800]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Ancyronyx larvae from Sulawesi, A: A. schoedli (body length 2.90 mm), B: A. skalei (body length 2.65 mm), C: A. tobada (body length 2.95 mm), D: A. longiparamerus (body length 2.95 mm), E: A. toraja (body length 3.30 mm), F: A. henningi (body length 2.35 mm)." httpUri="https://zenodo.org/record/835154/files/figure.png" pageId="5" pageNumber="126">Figs 2</figureCitation>
C; 4A, C, E)
</paragraph>
</subSubSection>
<subSubSection id="C361FDC9FFD4FFBE77F3FDDFF9CDFA7B" pageId="5" pageNumber="126" type="materials_examined">
<paragraph id="8BC4AE42FFD4FFBE77F3FDDFF9CDFA7B" blockId="5.[151,1437,1265,1866]" pageId="5" pageNumber="126">
<materialsCitation id="3B13A41FFFD4FFBE77F3FDDFFFFDFAEB" ID-GBIF-Occurrence="1580520945" collectionCode="CKB, CFM" collectorName="Material" country="Indonesia" elevation="1680" location="Tentena-Gintu" municipality="Rd. Km" pageId="5" pageNumber="126" specimenCount="1" stateProvince="Sulawesi">
<emphasis id="B90F7250FFD4FFBE77F3FDDFFC83FACF" bold="true" box="[151,384,1265,1290]" pageId="5" pageNumber="126">
<collectorName id="268ECB94FFD4FFBE77F3FDDFFDFCFACF" box="[151,255,1265,1290]" pageId="5" pageNumber="126">Material</collectorName>
examined.
</emphasis>
3LL (
<collectionCode id="ED6A3687FFD4FFBE76A1FDDCFF01FACF" box="[453,514,1266,1290]" pageId="5" pageNumber="126">CKB</collectionCode>
,
<collectionCode id="ED6A3687FFD4FFBE7568FDDCFF44FACF" box="[524,583,1266,1290]" pageId="5" pageNumber="126">CFM</collectionCode>
[FR114]): “
<collectingCountry id="F36CEED2FFD4FFBE75AAFDDCFE6EFACF" box="[718,877,1266,1290]" name="Indonesia" pageId="5" pageNumber="126">INDONESIA</collectingCountry>
: C
<collectingRegion id="49BF60A0FFD4FFBE74EBFDDFFEF4FACF" box="[911,1015,1265,1290]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="126">Sulawesi</collectingRegion>
,
<collectingMunicipality id="6BA03438FFD4FFBE7366FDDFF959FACC" box="[1026,1114,1265,1290]" pageId="5" pageNumber="126">Rd. Km</collectingMunicipality>
43
<location id="8EA4F899FFD4FFBE73E7FDDFF82BFACF" LSID="urn:lsid:plazi:treatment:03D21F54FFD4FFBE77F3FDA8FC2BF88F:8EA4F899FFD4FFBE73E7FDDFF82BFACF" box="[1155,1320,1265,1290]" country="Indonesia" municipality="Rd. Km" name="Tentena-Gintu" pageId="5" pageNumber="126" stateProvince="Sulawesi">Tentena-Gintu</location>
; mountain creek; cloud forest; submerged wood, c.
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<quantity id="4C8303A7FFD4FFBE751EFC3BFFCFFAEB" box="[634,716,1301,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="5" pageNumber="126" unit="m" value="1680.0">1680m</quantity>
asl
</elevation>
</materialsCitation>
.,
<materialsCitation id="3B13A41FFFD4FFBE7473FC38FE2AFAB3" ID-GBIF-Occurrence="1580520924" collectingDate="1998-01-04" collectionCode="CKB, NMW, CFM" collectorName="Freitag &amp; Sul" country="Indonesia" elevation="860" latitude="-1.8108333" location="Tentena" longLatPrecision="925" longitude="0.06666667" municipality="Rd. Tentena-Pendola" pageId="5" pageNumber="126" specimenCount="1" stateProvince="Sulawesi">
c.
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120°2317
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</geoCoordinate>
0
<collectingDate id="EF81716AFFD4FFBE73E1FC38F805FAEB" box="[1157,1286,1302,1326]" pageId="5" pageNumber="126" value="1998-01-04">
<date id="FFC58882FFD4FFBE73E1FC38F805FAEB" box="[1157,1286,1302,1326]" pageId="5" pageNumber="126" value="1998-01-04">4 Jan.1998</date>
</collectingDate>
leg.
<collectorName id="268ECB94FFD4FFBE7229FC3BF89FFAEB" box="[1357,1436,1301,1326]" pageId="5" pageNumber="126">Freitag</collectorName>
(
<collectorName id="268ECB94FFD4FFBE77FBFC17FDCBFA97" box="[159,200,1337,1362]" pageId="5" pageNumber="126">Sul</collectorName>
14f)”; 6LL (
<collectionCode id="ED6A3687FFD4FFBE7636FC14FC8EFA97" box="[338,397,1338,1362]" pageId="5" pageNumber="126">CKB</collectionCode>
[SEM],
<collectionCode id="ED6A3687FFD4FFBE7697FC14FF38FA94" box="[499,571,1338,1361]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="126">NMW</collectionCode>
,
<collectionCode id="ED6A3687FFD4FFBE752CFC14FF80FA97" box="[584,643,1338,1362]" pageId="5" pageNumber="126">CFM</collectionCode>
[FR 119]): “
<collectingCountry id="F36CEED2FFD4FFBE7473FC14FEB5FA97" box="[791,950,1338,1362]" name="Indonesia" pageId="5" pageNumber="126">INDONESIA</collectingCountry>
: C
<collectingRegion id="49BF60A0FFD4FFBE74BAFC17F945FA97" box="[990,1094,1337,1362]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="126">Sulawesi</collectingRegion>
,
<collectingMunicipality id="6BA03438FFD4FFBE7330FC17F840FA97" box="[1108,1347,1337,1362]" pageId="5" pageNumber="126">Rd. Tentena-Pendola</collectingMunicipality>
,
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E
<location id="8EA4F899FFD4FFBE77F3FC70FDF1FAB0" LSID="urn:lsid:plazi:treatment:03D21F54FFD4FFBE77F3FDA8FC2BF88F:8EA4F899FFD4FFBE77F3FC70FDF1FAB0" box="[151,242,1374,1397]" country="Indonesia" latitude="-1.8108333" longLatPrecision="925" longitude="0.06666667" municipality="Rd. Tentena-Pendola" name="Tentena" pageId="5" pageNumber="126" stateProvince="Sulawesi">Tentena</location>
, secondary forest; submerged wood, c.
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<quantity id="4C8303A7FFD4FFBE75DFFC73FFF8FAB3" box="[699,763,1373,1398]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="5" pageNumber="126" unit="m" value="860.0">860m</quantity>
asl
</elevation>
</materialsCitation>
.,
<materialsCitation id="3B13A41FFFD4FFBE745FFC70FCF4FA7B" ID-GBIF-Occurrence="1580520930" collectingDate="1998-01-03" collectionCode="CFM" collectorName="Freitag &amp; Sul" country="Indonesia" elevation="1190" latitude="-1.7752777" location="Tentena-Gintu" longLatPrecision="925" longitude="0.05" municipality="Rd. Km" pageId="5" pageNumber="126" specimenCount="1" stateProvince="Sulawesi">
c.
<geoCoordinate id="EE4FC885FFD4FFBE743CFC73FEE0FAB3" box="[856,995,1373,1398]" direction="south" orientation="latitude" pageId="5" pageNumber="126" precision="15" value="-1.7752777">01°4631 S</geoCoordinate>
<geoCoordinate id="EE4FC885FFD4FFBE7488FC73F97EFAB3" box="[1004,1149,1373,1398]" direction="east" orientation="longitude" pageId="5" pageNumber="126" precision="15" value="120.718056">
120°4305
<geoCoordinate id="EE4FC885FFD4FFBE730FFC73F97EFAB3" box="[1131,1149,1373,1398]" direction="east" orientation="longitude" pageId="5" pageNumber="126" precision="925" value="0.05">E</geoCoordinate>
</geoCoordinate>
0
<collectingDate id="EF81716AFFD4FFBE73F2FC70F812FAB3" box="[1174,1297,1374,1398]" pageId="5" pageNumber="126" value="1998-01-03">
<date id="FFC58882FFD4FFBE73F2FC70F812FAB3" box="[1174,1297,1374,1398]" pageId="5" pageNumber="126" value="1998-01-03">3 Jan.1998</date>
</collectingDate>
leg.
<collectorName id="268ECB94FFD4FFBE7229FC73F89FFAB3" box="[1357,1436,1373,1398]" pageId="5" pageNumber="126">Freitag</collectorName>
(
<collectorName id="268ECB94FFD4FFBE77FBFCAFFDCBFA5F" box="[159,200,1409,1434]" pageId="5" pageNumber="126">Sul</collectorName>
10f)”; 4LL (
<collectionCode id="ED6A3687FFD4FFBE7639FCACFC99FA5F" box="[349,410,1410,1434]" pageId="5" pageNumber="126">CFM</collectionCode>
[FR121]): “
<collectingCountry id="F36CEED2FFD4FFBE7555FCACFFD3FA5F" box="[561,720,1410,1434]" name="Indonesia" pageId="5" pageNumber="126">INDONESIA</collectingCountry>
: S
<collectingRegion id="49BF60A0FFD4FFBE7465FCAFFE6AFA5F" box="[769,873,1409,1434]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="126">Sulawesi</collectingRegion>
,
<collectingMunicipality id="6BA03438FFD4FFBE7418FCAFFEDFFA5C" box="[892,988,1409,1434]" pageId="5" pageNumber="126">Rd. Km</collectingMunicipality>
56
<location id="8EA4F899FFD4FFBE7371FCAFF9B9FA5F" LSID="urn:lsid:plazi:treatment:03D21F54FFD4FFBE77F3FDA8FC2BF88F:8EA4F899FFD4FFBE7371FCAFF9B9FA5F" box="[1045,1210,1409,1434]" country="Indonesia" latitude="-1.7752777" longLatPrecision="925" longitude="0.05" municipality="Rd. Km" name="Tentena-Gintu" pageId="5" pageNumber="126" stateProvince="Sulawesi">Tentena-Gintu</location>
; black-water river; submerged wood, c.
<elevation id="00564971FFD4FFBE761BFC88FCF4FA7B" box="[383,503,1445,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="5" pageNumber="126" unit="m" value="1190.0">
<quantity id="4C8303A7FFD4FFBE761BFC88FCC8FA7B" box="[383,459,1446,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="5" pageNumber="126" unit="m" value="1190.0">1190m</quantity>
asl
</elevation>
</materialsCitation>
., c. 01°4928 S 120°2143E 0
<date id="FFC58882FFD4FFBE7432FC88FECCFA7B" box="[854,975,1446,1470]" pageId="5" pageNumber="126" value="1998-01-04">4 Jan.1998</date>
leg. Freitag (Sul16f)”.
</paragraph>
</subSubSection>
<subSubSection id="C361FDC9FFD4FFBE77A3FCE7FC2BF88F" pageId="5" pageNumber="126" type="diagnosis">
<paragraph id="8BC4AE42FFD4FFBE77A3FCE7FC2BF88F" blockId="5.[151,1437,1265,1866]" pageId="5" pageNumber="126">
<emphasis id="B90F7250FFD4FFBE77A3FCE7FCC9FA27" bold="true" box="[199,458,1481,1506]" pageId="5" pageNumber="126">Differential diagnosis</emphasis>
(based on presumably sixth instar, HW:
<quantity id="4C8303A7FFD4FFBE74CBFCE4F918FA27" box="[943,1051,1482,1506]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="126" unit="mm" value="0.32">0.32 mm</quantity>
). Generally similar to
<taxonomicName id="4C7BD5C1FFD4FFBE7241FCE4FDF7F9C3" authorityName="Freitag &amp; Kodada" authorityYear="2017" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="schoedli">
<emphasis id="B90F7250FFD4FFBE7241FCE4FDF7F9C3" italics="true" pageId="5" pageNumber="126">Ancyronyx schoedli</emphasis>
</taxonomicName>
except for the following characters: pronotum yellowish in anterior third, yellowish lateral portions of head extending posteriad along the frontal suture; posterior margins of thoracic and abdominal terga slightly paler, somewhat dark yellow-brown; terminal abdominal segment dark along anterior 0.1, yellow-brown along anterior 0.10.6, dark brown along posterior 0.60.9 and pale yellow-brown at the tip. Entire ventral side pale yellowish, only abdominal segment IX occasionally slightly darker, but still somewhat pale. Dorsal tubercles moderately elevated (
<figureCitation id="1340B2C7FFD4FFBE7666FF8FFC49F97F" box="[258,330,1697,1722]" captionStart="FIGURE 4" captionStartId="6.[151,250,1570,1592]" captionTargetBox="[313,1273,187,1548]" captionTargetId="figure@6.[313,1273,187,1548]" captionTargetPageId="6" captionText="FIGURE 4. Ancyronyx larvae, dorsal (SEM photographs), A: head and pronotum of A. tobada; B: head and pronotum of A. longiparamerus; C: details of abdominal segments I III of A. tobada with tubercles, lateral abdominal projections, and setae; D: same aspect of A. longiparamerus; E: abdominal segments IX of A. tobada; F: abdominal segments IX of A. longiparamerus." httpUri="https://zenodo.org/record/835158/files/figure.png" pageId="5" pageNumber="126">Figs 4</figureCitation>
A, C). Head (
<figureCitation id="1340B2C7FFD4FFBE76B9FF8FFF26F97F" box="[477,549,1697,1722]" captionStart="FIGURE 2" captionStartId="3.[151,250,1822,1844]" captionTargetBox="[170,1430,878,1783]" captionTargetId="figure@3.[151,1436,873,1800]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Ancyronyx larvae from Sulawesi, A: A. schoedli (body length 2.90 mm), B: A. skalei (body length 2.65 mm), C: A. tobada (body length 2.95 mm), D: A. longiparamerus (body length 2.95 mm), E: A. toraja (body length 3.30 mm), F: A. henningi (body length 2.35 mm)." httpUri="https://zenodo.org/record/835154/files/figure.png" pageId="5" pageNumber="126">Figs 2</figureCitation>
C, 4A) slightly tapering anteriad, widest at posterior 0.2; antennae overreaching anterior margin of labrum. Abdominal segments gently elevated near entire posterior portion; dorsum with scattered short setae (
<figureCitation id="1340B2C7FFD4FFBE76F4FFC7FCD8F8C7" box="[400,475,1769,1794]" captionStart="FIGURE 4" captionStartId="6.[151,250,1570,1592]" captionTargetBox="[313,1273,187,1548]" captionTargetId="figure@6.[313,1273,187,1548]" captionTargetPageId="6" captionText="FIGURE 4. Ancyronyx larvae, dorsal (SEM photographs), A: head and pronotum of A. tobada; B: head and pronotum of A. longiparamerus; C: details of abdominal segments I III of A. tobada with tubercles, lateral abdominal projections, and setae; D: same aspect of A. longiparamerus; E: abdominal segments IX of A. tobada; F: abdominal segments IX of A. longiparamerus." httpUri="https://zenodo.org/record/835158/files/figure.png" pageId="5" pageNumber="126">Figs 4</figureCitation>
C, E); lateral protuberances with tubercles almost as large as spiracles; segment IX (
<figureCitation id="1340B2C7FFD4FFBE77FBFE23FDE5F8E3" box="[159,230,1805,1830]" captionStart="FIGURE 2" captionStartId="3.[151,250,1822,1844]" captionTargetBox="[170,1430,878,1783]" captionTargetId="figure@3.[151,1436,873,1800]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Ancyronyx larvae from Sulawesi, A: A. schoedli (body length 2.90 mm), B: A. skalei (body length 2.65 mm), C: A. tobada (body length 2.95 mm), D: A. longiparamerus (body length 2.95 mm), E: A. toraja (body length 3.30 mm), F: A. henningi (body length 2.35 mm)." httpUri="https://zenodo.org/record/835154/files/figure.png" pageId="5" pageNumber="126">Figs 2</figureCitation>
C, 4E) very long, ca. 2.4 times as long as wide; basomedian portion glabrous, without tubercles; apex deeply emarginated.
</paragraph>
</subSubSection>
</treatment>
</document>