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<document id="B352B4E608510439B365C94D1469A931" ID-CLB-Dataset="22494" ID-DOI="10.11646/zootaxa.4765.1.1" ID-GBIF-Dataset="8af5a092-f062-48c7-8d7c-f96ae6282bfa" ID-ISSN="1175-5334" ID-Zenodo-Dep="3774174" ID-ZooBank="urn:lsid:zoobank.org:pub:C442D94C-0EB4-4509-B762-913707214819" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588094957734" checkinUser="carolina" docAuthor="Hodgson, Chris" docDate="2020" docId="03B2EA640A7046322CFCF8F2FC49D433" docLanguage="en" docName="zootaxa.4765.1.1.pdf.imf" docOrigin="Zootaxa 4765 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tanyscelis Hardy &amp; Gullan 2010" docType="treatment" docVersion="5" lastPageNumber="97" masterDocId="FF8B921C0A2F46522C6BFF8EFFCBD747" masterDocTitle="A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males" masterLastPageNumber="264" masterPageNumber="1" pageNumber="96" updateTime="1698837970293" updateUser="ExternalLinkService">
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<mods:title id="70503461F9C5154C64755AAD74CE0B82">A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males</mods:title>
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<mods:namePart id="6F47618A70EF6110668F031AF9E4C2E3">Hodgson, Chris</mods:namePart>
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<mods:date id="E2CE0524F642F57F565E3E87F0C422ED">2020</mods:date>
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<mods:identifier id="1A2033D3BC440878BFB5D5E9767B2D17" type="DOI">10.11646/zootaxa.4765.1.1</mods:identifier>
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<treatment id="03B2EA640A7046322CFCF8F2FC49D433" ID-GBIF-Taxon="163857838" LSID="urn:lsid:plazi:treatment:03B2EA640A7046322CFCF8F2FC49D433" httpUri="http://treatment.plazi.org/id/03B2EA640A7046322CFCF8F2FC49D433" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<subSubSection id="C30108F90A70460D2CFCF8F2FDF8D0D1" box="[151,563,1916,1943]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph id="8BA45B720A70460D2CFCF8F2FDF8D0D1" blockId="95.[151,563,1916,1943]" box="[151,563,1916,1943]" pageId="95" pageNumber="96">
<heading id="D0ECEC1E0A70460D2CFCF8F2FDF8D0D1" bold="true" box="[151,563,1916,1943]" fontSize="10" level="2" pageId="95" pageNumber="96" reason="2">
<emphasis id="B96F87600A70460D2CFCF8F2FDF8D0D1" bold="true" box="[151,563,1916,1943]" pageId="95" pageNumber="96">
<taxonomicName id="4C1B20F10A70460D2CFCF8F2FDE5D0D1" authority="Hardy &amp; Gullan 2010" authorityName="Hardy &amp; Gullan" authorityYear="2010" box="[151,558,1916,1943]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A70460D2CFCF8F2FED8D0D0" bold="true" box="[151,275,1916,1943]" italics="true" pageId="95" pageNumber="96">Tanyscelis</emphasis>
<bibRefCitation id="EF8A26830A70460D2D70F8F3FDE5D0D1" author="Hardy &amp; Gullan" box="[283,558,1916,1943]" firstAuthor="Hardy" pageId="95" pageNumber="96" pagination="1 - 74" refId="ref202999" refString="Hardy, N. B. &amp; Gullan, P. J. (2010) Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species. ZooKeys, 58, 1 - 74. https: // doi. org / 10.3897 / zookeys. 58.507" type="journal article" year="2010">Hardy &amp; Gullan 2010</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30108F90A7046322CFCF834FD9FD68A" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="reference_group">
<paragraph id="8BA45B720A7046322CFCF834FD9FD68A" blockId="95.[151,1345,1977,2002]" lastBlockId="96.[151,1437,152,884]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<taxonomicName id="4C1B20F10A70460D2CFCF834FE2DD095" authority="Hardy &amp; Gullan 2010" authorityName="Hardy &amp; Gullan" authorityYear="2010" box="[151,486,1977,2002]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A70460D2CFCF834FECBD095" box="[151,256,1978,2002]" italics="true" pageId="95" pageNumber="96">Tanyscelis</emphasis>
<bibRefCitation id="EF8A26830A70460D2D6DF837FE2DD095" author="Hardy &amp; Gullan" box="[262,486,1977,2002]" firstAuthor="Hardy" pageId="95" pageNumber="96" pagination="1 - 74" refId="ref202999" refString="Hardy, N. B. &amp; Gullan, P. J. (2010) Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species. ZooKeys, 58, 1 - 74. https: // doi. org / 10.3897 / zookeys. 58.507" type="journal article" year="2010">Hardy &amp; Gullan 2010</bibRefCitation>
</taxonomicName>
, 34–35.
<typeStatus id="54A0E5D00A70460D2E57F834FDA5D095" box="[572,622,1978,2002]" pageId="95" pageNumber="96">Type</typeStatus>
species:
<taxonomicName id="4C1B20F10A70460D2EA0F834FBE8D095" ID-CoL="6SR8W" authority="Froggatt." authorityName="Froggatt." box="[715,1059,1978,2002]" class="Insecta" family="Eriococcidae" genus="Opisthoscelis" kingdom="Animalia" order="Hemiptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="pisiformis">
<emphasis id="B96F87600A70460D2EA0F834FC74D095" box="[715,959,1978,2002]" italics="true" pageId="95" pageNumber="96">Opisthoscelis pisiformis</emphasis>
Froggatt.
</taxonomicName>
<typeStatus id="54A0E5D00A70460D2843F834FB91D095" box="[1064,1114,1978,2002]" pageId="95" pageNumber="96">Type</typeStatus>
designation unknown.
<emphasis id="B96F87600A4F46322CFCFF16FEF2D7F4" bold="true" box="[151,313,152,179]" pageId="96" pageNumber="97">Introduction.</emphasis>
This genus is restricted to
<collectingCountry id="F30C1BE20A4F46322E02FF17FD19D7F4" box="[617,722,153,179]" name="Australia" pageId="96" pageNumber="97">Australia</collectingCountry>
and currently contains 12 species (
<bibRefCitation id="EF8A26830A4F4632280BFF17FA95D7F4" author="Hardy &amp; Gullan" box="[1120,1374,153,179]" firstAuthor="Hardy" pageId="96" pageNumber="97" pagination="1 - 74" refId="ref202999" refString="Hardy, N. B. &amp; Gullan, P. J. (2010) Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species. ZooKeys, 58, 1 - 74. https: // doi. org / 10.3897 / zookeys. 58.507" type="journal article" year="2010">Hardy &amp; Gullan 2010</bibRefCitation>
). The females induce galls on leaves and stems of
<taxonomicName id="4C1B20F10A4F46322EE5FF32FCC7D791" ID-CoL="4F2P" box="[654,780,188,214]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="96" pageNumber="97" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F87600A4F46322EE5FF32FCC7D791" box="[654,780,188,214]" italics="true" pageId="96" pageNumber="97">Eucalyptus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C1B20F10A4F46322F28FF32FC09D791" ID-CoL="D48" box="[835,962,188,214]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="96" pageNumber="97" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
), each gall either globular, hemispherical, conical mammiform or thorn-like. Most species were originally placed in the genus
<taxonomicName id="4C1B20F10A4F4632283BFF51FA5CD7BE" ID-CoL="696K" authority="Schrader 1863" authorityName="Schrader" authorityYear="1863" box="[1104,1431,223,249]" class="Insecta" family="Eriococcidae" genus="Opisthoscelis" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F4632283BFF51FB21D7BE" box="[1104,1258,223,249]" italics="true" pageId="96" pageNumber="97">Opisthoscelis</emphasis>
<bibRefCitation id="EF8A26830A4F4632289BFF51FA5CD7BE" author="Schrader" box="[1264,1431,223,249]" firstAuthor="Schrader" pageId="96" pageNumber="97" pagination="1 - 6" refId="ref210938" refString="Schrader, H. L. (1863) Observations on certain gall-making Coccidae of Australia. Transactions of the Entomological Society of New South Wales, 1, 1 - 6." type="journal article" year="1863">Schrader 1863</bibRefCitation>
</taxonomicName>
, but adult male
<taxonomicName id="4C1B20F10A4F46322D28FE8CFE72D65B" authorityName="Hardy &amp; Gullan" authorityYear="2010" box="[323,441,258,284]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F46322D28FE8CFE72D65B" box="[323,441,258,284]" italics="true" pageId="96" pageNumber="97">Tanyscelis</emphasis>
</taxonomicName>
have 9-segmented antennae whereas those of
<taxonomicName id="4C1B20F10A4F46322FA5FE8CFBA3D65B" ID-CoL="696K" authorityName="Schrader" authorityYear="1863" box="[974,1128,258,284]" class="Insecta" family="Eriococcidae" genus="Opisthoscelis" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F46322FA5FE8CFBA3D65B" box="[974,1128,258,284]" italics="true" pageId="96" pageNumber="97">Opisthoscelis</emphasis>
</taxonomicName>
species have 10 segments. In addition, the abdomen of adult male
<taxonomicName id="4C1B20F10A4F46322E0CFEA8FD16D607" authorityName="Hardy &amp; Gullan" authorityYear="2010" box="[615,733,294,320]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F46322E0CFEA8FD16D607" box="[615,733,294,320]" italics="true" pageId="96" pageNumber="97">Tanyscelis</emphasis>
</taxonomicName>
species is attenuated, becoming very narrow near apex, and they lack glandular pouches, whereas the abdomen of adult male
<taxonomicName id="4C1B20F10A4F46322F14FEC7FBD2D624" ID-CoL="696K" authorityName="Schrader" authorityYear="1863" box="[895,1049,329,355]" class="Insecta" family="Eriococcidae" genus="Opisthoscelis" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F46322F14FEC7FBD2D624" box="[895,1049,329,355]" italics="true" pageId="96" pageNumber="97">Opisthoscelis</emphasis>
</taxonomicName>
species is not attenuated and they have a pair of glandular pouches (
<bibRefCitation id="EF8A26830A4F46322E7EFEE2FCD9D6C1" author="Hardy &amp; Gullan" box="[533,786,364,390]" firstAuthor="Hardy" pageId="96" pageNumber="97" pagination="1 - 74" refId="ref202999" refString="Hardy, N. B. &amp; Gullan, P. J. (2010) Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species. ZooKeys, 58, 1 - 74. https: // doi. org / 10.3897 / zookeys. 58.507" type="journal article" year="2010">Hardy &amp; Gullan 2010</bibRefCitation>
). Thus, the detailed description of adult male
<taxonomicName id="4C1B20F10A4F4632297FFEE2FF79D6EE" ID-CoL="696K" authorityName="Schrader" authorityYear="1863" class="Insecta" family="Eriococcidae" genus="Opisthoscelis" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F4632297FFEE2FF79D6EE" italics="true" pageId="96" pageNumber="97">Opisthoscelis</emphasis>
</taxonomicName>
spp. by
<bibRefCitation id="EF8A26830A4F46322D79FE01FE70D6ED" author="Theron" box="[274,443,399,426]" firstAuthor="Theron" pageId="96" pageNumber="97" pagination="87 - 99" refId="ref212291" refString="Theron, J. G. (1968) Studies on the morphology and relationships of male Apiomorpha and Opisthoscelis (Hemiptera: Coccoidea). Australian Journal of Zoology, Melbourne, 16, 87 - 99. https: // doi. org / 10.1071 / ZO 9680087" type="journal article" year="1968">Theron (1968)</bibRefCitation>
, based on specimens of two unidentified species collected from Keith,
<collectingRegion id="49DF95900A4F4632288EFE01FA53D6EE" box="[1253,1432,399,425]" country="Australia" name="South Australia" pageId="96" pageNumber="97">South Australia</collectingRegion>
, actually refers to species of
<taxonomicName id="4C1B20F10A4F46322DBCFE3CFD86D68B" authorityName="Hardy &amp; Gullan" authorityYear="2010" box="[471,589,434,460]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600A4F46322DBCFE3CFD86D68B" box="[471,589,434,460]" italics="true" pageId="96" pageNumber="97">Tanyscelis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30108F90A4F46322CACFE5BFC49D433" pageId="96" pageNumber="97" type="diagnosis">
<paragraph id="8BA45B720A4F46322CACFE5BFC49D433" blockId="96.[151,1437,152,884]" pageId="96" pageNumber="97">
<emphasis id="B96F87600A4F46322CACFE5BFC37D6B7" bold="true" box="[199,1020,469,496]" pageId="96" pageNumber="97">
Generic diagnosis based on adult male morphology of
<taxonomicName id="4C1B20F10A4F46322F00FE58FC37D6B7" box="[875,1020,469,496]" class="Insecta" family="Eriococcidae" genus="Tanyscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="verrucula">
<emphasis id="B96F87600A4F46322F00FE58FC37D6B7" bold="true" box="[875,1020,469,496]" italics="true" pageId="96" pageNumber="97">T. verrucula</emphasis>
</taxonomicName>
</emphasis>
(described below) (
<figureCitation id="132047F70A4F4632288CFE58FAF4D6B7" box="[1255,1343,470,496]" captionStart="FIGURE 39" captionStartId="97.[151,250,1946,1971]" captionTargetBox="[226,1366,191,1920]" captionTargetId="figure@97.[221,1366,181,1920]" captionTargetPageId="97" captionText="FIGURE 39. Tanyscelis verrucula (Froggatt). Macropterous male. (Eriococcidae, Gondwanan group). Where A=cranial apophysis and X=pores on head." figureDoi="http://doi.org/10.5281/zenodo.3774252" httpUri="https://zenodo.org/record/3774252/files/figure.png" pageId="96" pageNumber="97">Fig. 39</figureCitation>
).
<emphasis id="B96F87600A4F4632293EFE58FA5FD6B7" bold="true" box="[1365,1428,470,496]" pageId="96" pageNumber="97">Body</emphasis>
: abdomen greatly attenuated, narrowing to a very narrow style; body fairly setose, with mainly long, flagellate fs.
<emphasis id="B96F87600A4F46322CD7FD92FECDD571" bold="true" box="[188,262,540,566]" pageId="96" pageNumber="97">Head:</emphasis>
simple pores present; ocular sclerite without striations or reticulations; dorsal mid-cranial ridge well developed;
<emphasis id="B96F87600A4F46322D77FDB1FDE8D51E" box="[284,547,575,601]" italics="true" pageId="96" pageNumber="97">preocular ridge absent</emphasis>
;
<emphasis id="B96F87600A4F46322E58FDB1FCDAD51E" box="[563,785,575,601]" italics="true" pageId="96" pageNumber="97">genal setae present</emphasis>
;
<emphasis id="B96F87600A4F46322F4AFDCEFBEBD51E" box="[801,1056,575,601]" italics="true" pageId="96" pageNumber="97">antennae 9 segmented</emphasis>
; flagellar segments with long fs;
<emphasis id="B96F87600A4F46322CFCFDEDFD7BD53B" box="[151,688,610,637]" italics="true" pageId="96" pageNumber="97">capitate setae restricted to 3 on apical segment</emphasis>
; scape with many setae.
<emphasis id="B96F87600A4F46322FA7FDEDFBE4D53A" bold="true" box="[972,1071,611,637]" pageId="96" pageNumber="97">Thorax:</emphasis>
prosternal median ridge absent;
<emphasis id="B96F87600A4F46322CFCFD08FCBCD5E7" box="[151,887,646,672]" italics="true" pageId="96" pageNumber="97">prosternal setae and antemesospiracular setae present, mainly fs</emphasis>
; scutal setae present laterally;
<emphasis id="B96F87600A4F463228B8FD09FE2CD584" italics="true" pageId="96" pageNumber="97">postmesospiracular setae abundant, mainly fs</emphasis>
;
<emphasis id="B96F87600A4F46322D93FD27FC5DD584" box="[504,918,681,707]" italics="true" pageId="96" pageNumber="97">basisternal setae present, mainly fs;</emphasis>
metasternum with abundant fs; postmetaspiracular setae few; wings without alar setae, alar lobes and alar sensoria; hamulohalteres absent; legs extremely long and hirsute; tibia and tarsi with mainly fleshy setae;
<emphasis id="B96F87600A4F46322F44FD7EFC32D44E" box="[815,1017,751,777]" italics="true" pageId="96" pageNumber="97">tarsi 1 segmented</emphasis>
; claw digitules capitate; claws short and sharply pointed.
<emphasis id="B96F87600A4F46322DEFFC9CFDCFD46A" bold="true" box="[388,516,786,813]" pageId="96" pageNumber="97">Abdomen:</emphasis>
<emphasis id="B96F87600A4F46322E60FC9CFC84D46B" box="[523,847,786,812]" italics="true" pageId="96" pageNumber="97">drawn out and very elongate</emphasis>
;
<emphasis id="B96F87600A4F46322F35FC9CFA5FD46B" box="[862,1428,786,813]" italics="true" pageId="96" pageNumber="97">long fs numerous dorsally, ventrally and pleurally</emphasis>
; glandular pouches absent; caudal extensions absent; entire penial sheath long and narrow; all setae posteriorly on style long; aedeagus blade-like (i.e., very narrow dorsoventrally).
</paragraph>
</subSubSection>
</treatment>
</document>
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