treatments-xml/data/03/B1/29/03B12920A57BFF84C2EA8C2196A2FDF5.xml

<document id="819EDC07D0ED63D8F4D1E3D2D1FC9A68" ID="10.11646/zootaxa.4250.1.2SLASH10700" ID-CLB-Dataset="33319" ID-DOI="10.11646/zootaxa.4250.1.2" ID-GBIF-Dataset="1c5bce12-4b99-4480-a230-9fe3cd5e0fee" ID-ISSN="1175-5326" ID-Zenodo-Dep="439780" ID-ZooBank="CDE1CCE1-71E1-4947-A7BE-9C9F98B26C81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491204228329" checkinUser="plazi" docAuthor="Vargovitsh, Robert S." docDate="2017" docId="03B12920A57BFF84C2EA8C2196A2FDF5" docLanguage="en" docName="zootaxa.4250.1.2SLASH10700.pdf" docOrigin="Zootaxa 4250 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygmarrhopalites dbari Vargovitsh, 2017, sp. nov." docType="treatment" docUuid="5F1FC036-FF43-4334-8638-5908AF219115" docUuidSource="ZooBank" docVersion="10" lastPageNumber="40" masterDocId="FF885158A57AFF95C27D881D935CFF80" masterDocTitle="Two new troglobiont Pygmarrhopalites species of the principalis group (Collembola: Arrhopalitidae) from the West Caucasus" masterLastPageNumber="42" masterPageNumber="23" pageNumber="24" updateTime="1698438633354" updateUser="ExternalLinkService">
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<mods:title id="24AAC487C9723FFFA0EB30C92D3CA253">Two new troglobiont Pygmarrhopalites species of the principalis group (Collembola: Arrhopalitidae) from the West Caucasus</mods:title>
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<mods:namePart id="01A3B72A74B0525B614251B0CB25315A">Vargovitsh, Robert S.</mods:namePart>
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<mods:date id="CF19C91B7677775E9167F673182015EE">2017</mods:date>
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<treatment id="03B12920A57BFF84C2EA8C2196A2FDF5" ID-DOI="http://doi.org/10.5281/zenodo.5679213" ID-GBIF-Taxon="128090467" ID-Zenodo-Dep="5679213" LSID="urn:lsid:zoobank.org:act:5F1FC036-FF43-4334-8638-5908AF219115" httpUri="http://treatment.plazi.org/id/03B12920A57BFF84C2EA8C2196A2FDF5" lastPageId="17" lastPageNumber="40" pageId="1" pageNumber="24">
<subSubSection id="C302CBBDA57BFF94C2EA8C2192D2FBF7" pageId="1" pageNumber="24" type="nomenclature">
<paragraph id="8BA79836A57BFF94C2EA8C219145FBD6" blockId="1.[151,537,1084,1143]" box="[151,537,1084,1110]" pageId="1" pageNumber="24">
<heading id="D0EF2F5AA57BFF94C2EA8C219145FBD6" bold="true" box="[151,537,1084,1110]" fontSize="11" level="1" pageId="1" pageNumber="24" reason="1">
<emphasis id="B96C4424A57BFF94C2EA8C219145FBD6" bold="true" box="[151,537,1084,1110]" pageId="1" pageNumber="24">
<taxonomicName id="4C18E3B5A57BFF94C2EA8C2192E8FBD6" box="[151,436,1084,1110]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A57BFF94C2EA8C2192E8FBD6" bold="true" box="[151,436,1084,1110]" italics="true" pageId="1" pageNumber="24">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25FF95FA57BFF94C3C18C219145FBD6" box="[444,537,1084,1110]" pageId="1" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA79836A57BFF94C2EA8C4392D2FBF7" blockId="1.[151,537,1084,1143]" box="[151,398,1118,1143]" pageId="1" pageNumber="24">
<figureCitation id="132384B3A57BFF94C2EA8C439254FBF7" box="[151,264,1118,1143]" captionStart-0="FIGURES 1 – 2. 1" captionStart-1="FIGURES 3 – 7" captionStart-2="FIGURES 8 – 19" captionStart-3="FIGURES 20 – 24" captionStart-4="FIGURES 25 – 30" captionStart-5="FIGURES 31 – 33" captionStartId-0="4.[151,264,1975,1998]" captionStartId-1="5.[151,264,1952,1975]" captionStartId-2="6.[151,264,1952,1975]" captionStartId-3="7.[151,264,1938,1961]" captionStartId-4="8.[151,264,1953,1976]" captionStartId-5="9.[151,264,1460,1483]" captionTargetBox-0="[317,1260,226,1922]" captionTargetBox-1="[213,1395,193,1895]" captionTargetBox-2="[224,1360,222,1893]" captionTargetBox-3="[197,1397,193,1879]" captionTargetBox-4="[184,1402,213,1899]" captionTargetBox-5="[260,1350,222,1407]" captionTargetId-0="figure@4.[284,1297,195,1940]" captionTargetId-1="figure@5.[188,1399,193,1932]" captionTargetId-2="figure@6.[187,1399,193,1932]" captionTargetId-3="figure@7.[192,1397,193,1921]" captionTargetId-4="figure@8.[165,1422,193,1932]" captionTargetId-5="figure@9.[213,1373,175,1457]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURES 1 – 2. 1, type localities of Pygmarrhopalites dbari sp. nov. (1 — Psyrtskha Cave), and P. kovali sp. nov. (2 — Tsebeldinskaya Cave, 3 — Tshalintshur Cave, 4 — Verkhnyaya Shakuranskaya Cave); 2, habitus of mounted male." captionText-1="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." captionText-2="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." captionText-3="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." captionText-4="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." captionText-5="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri-0="https://zenodo.org/record/439781/files/figure.png" httpUri-1="https://zenodo.org/record/439782/files/figure.png" httpUri-2="https://zenodo.org/record/439783/files/figure.png" httpUri-3="https://zenodo.org/record/439784/files/figure.png" httpUri-4="https://zenodo.org/record/439785/files/figure.png" httpUri-5="https://zenodo.org/record/439786/files/figure.png" pageId="1" pageNumber="24">Figs 2–33</figureCitation>
, 
<tableCitation id="C69AAD8DA57BFF94C3698C439232FBF7" box="[276,366,1118,1143]" captionStart="TABLE 1" captionStartId="3.[151,239,512,535]" captionText="TABLE 1. Measurements (in µm) for body parts of types of Pygmarrhopalites dbari sp. nov. and P. kovali sp. nov." pageId="1" pageNumber="24">Tables 1</tableCitation>
–3
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA57BFF94C2EA8CBB91ACFA1A" pageId="1" pageNumber="24" type="diagnosis">
<paragraph id="8BA79836A57BFF94C2EA8CBB91ACFA1A" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
<emphasis id="B96C4424A57BFF94C2EA8CBB924DFB3F" bold="true" box="[151,273,1190,1215]" pageId="1" pageNumber="24">Diagnosis.</emphasis>
Body up to 
<quantity id="4CE035D3A57BFF94C3E18CBA92AEFB3E" box="[412,498,1191,1215]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="24" unit="mm" value="1.1">1.1 mm</quantity>
length with elongated appendages (
<figureCitation id="132384B3A57BFF94C1F98CBB9096FB3F" box="[900,970,1190,1215]" captionStart="FIGURES 1 – 2. 1" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[317,1260,226,1922]" captionTargetId="figure@4.[284,1297,195,1940]" captionTargetPageId="4" captionText="FIGURES 1 – 2. 1, type localities of Pygmarrhopalites dbari sp. nov. (1 — Psyrtskha Cave), and P. kovali sp. nov. (2 — Tsebeldinskaya Cave, 3 — Tshalintshur Cave, 4 — Verkhnyaya Shakuranskaya Cave); 2, habitus of mounted male." httpUri="https://zenodo.org/record/439781/files/figure.png" pageId="1" pageNumber="24">Fig. 2</figureCitation>
). Eyes 1 + 1. Trichobothria ABC form an angle 150–160o and AB&lt;BC. Antenna about 3× of the head, Ant IV with 7 distinctly separated subsegments and 13 whorls of setae. Head dorsum with 5 + 1 + 5 weakly spine-like setae. Femur II with 13 and III with 12 setae. All claws slender, without tunica and with inner tooth. Tips of empodia not reaching tips of corresponding claws, empodium I–II with and III without corner tooth. Manubrium with 7 + 7 posterior setae; dens with 2 outer and 1 inner spine-like setae. Tip of mucro rounded, slightly broadened. Abd VI with broadened circumanal setae and palmate appendices anales with secondary branching.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA57BFF94C2BA8DB89173F9CE" pageId="1" pageNumber="24" type="materials_examined">
<paragraph id="8BA79836A57BFF94C2BA8DB890DAF986" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
<materialsCitation id="3B70926BA57BFF94C2BA8DB890DAF986" ID-GBIF-Occurrence="1455395223" collectingDate="2011-08-25" collectorName="Abd VI &amp; R. S. Vargovitsh" county="West Caucasus" location="Psyrtskha River" municipality="Novy Afon" pageId="1" pageNumber="24" specimenCount="2" stateProvince="Abkhazia" typeStatus="holotype">
<emphasis id="B96C4424A57BFF94C2BA8DB89232FA3E" bold="true" box="[199,366,1445,1470]" pageId="1" pageNumber="24">
<typeStatus id="54A32694A57BFF94C2BA8DB8925EFA3E" box="[199,258,1445,1470]" pageId="1" pageNumber="24">Type</typeStatus>
material
</emphasis>
. 
<emphasis id="B96C4424A57BFF94C3008DB892AFFA3E" bold="true" box="[381,499,1445,1470]" pageId="1" pageNumber="24">
<typeStatus id="54A32694A57BFF94C3008DB892B2FA3E" box="[381,494,1445,1470]" pageId="1" pageNumber="24" type="holotype">Holotype</typeStatus>
:
</emphasis>
female, dissected and mounted on four slides (C-801a-2a: head, C-801a-2b: furca, C-801a-2c: 
<collectorName id="26EDFDE0A57BFF94C3638DD49224FA61" box="[286,376,1481,1506]" pageId="1" pageNumber="24">Abd VI</collectorName>
, C-801a-2d: great abdomen with legs), 
<collectingCounty id="62C6E0BAA57BFF94C13E8DD790ACFA62" box="[835,1008,1482,1506]" pageId="1" pageNumber="24">West Caucasus</collectingCounty>
, 
<collectingRegion id="49DC56D4A57BFF94C1838DD49730FA62" box="[1022,1132,1481,1506]" country="Georgia" name="Abkhazia" pageId="1" pageNumber="24">Abkhazia</collectingRegion>
, 
<collectingMunicipality id="6BC3024CA57BFF94C6078DD797A2FA62" box="[1146,1278,1481,1506]" pageId="1" pageNumber="24">Novy Afon</collectingMunicipality>
, valley of the 
<location id="8EC7CEEDA57BFF94C2EA8DF09217F986" LSID="urn:lsid:plazi:treatment:03B12920A57BFF84C2EA8C2196A2FDF5:8EC7CEEDA57BFF94C2EA8DF09217F986" box="[151,331,1517,1542]" county="West Caucasus" municipality="Novy Afon" name="Psyrtskha River" pageId="1" pageNumber="24" stateProvince="Abkhazia">Psyrtskha River</location>
, 
<location id="8EC7CEEDA57BFF94C32B8DF0915AF986" LSID="urn:lsid:plazi:treatment:03B12920A57BFF84C2EA8C2196A2FDF5:8EC7CEEDA57BFF94C32B8DF0915AF986" box="[342,518,1517,1542]" county="West Caucasus" municipality="Novy Afon" name="Psyrtskha Cave" pageId="1" pageNumber="24" stateProvince="Abkhazia">Psyrtskha Cave</location>
, 
<date id="FFA6BEF6A57BFF94C06C8DF091C6F986" box="[529,666,1517,1542]" pageId="1" pageNumber="24" value="2011-08-25">
<collectingDate id="EFE2471EA57BFF94C06C8DF091C6F986" box="[529,666,1517,1542]" pageId="1" pageNumber="24" value="2011-08-25">25.viii.2011</collectingDate>
</date>
, 
<collectorName id="26EDFDE0A57BFF94C0DB8DF3900AF986" box="[678,854,1517,1542]" pageId="1" pageNumber="24">R.S. Vargovitsh</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BA79836A57BFF94C2BA8E0C9173F9CE" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
<emphasis id="B96C4424A57BFF94C2BA8E0C9263F9AA" bold="true" box="[199,319,1553,1578]" pageId="1" pageNumber="24">Paratypes</emphasis>
on slides: 4 males, 2 females and 3 juv. collected together with holotype; same cave and collector, 
<date id="FFA6BEF6A57BFF94C2EA8E28924DF9CE" box="[151,273,1589,1614]" pageId="1" pageNumber="24" value="2006-06-17">17.vi.2006</date>
, 2 males, 1 female, 1 juv.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA57BFF9FC2BA8E44918BFEC0" lastPageId="10" lastPageNumber="33" pageId="1" pageNumber="24" type="description">
<paragraph id="8BA79836A57BFF94C2BA8E449785F916" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
<emphasis id="B96C4424A57BFF94C2BA8E4492E1F9F2" bold="true" box="[199,445,1625,1650]" pageId="1" pageNumber="24">Description. Female.</emphasis>
Total length without appendages up to 
<quantity id="4CE035D3A57BFF94C10F8E479097F9F1" box="[882,971,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="24" unit="mm" value="1.1">1.1 mm</quantity>
. Pigmentation absent in young specimens and present in adults as reddish spots, more intensive on the dorsum of head and thorax (
<figureCitation id="132384B3A57BFF94C6048E609791F916" box="[1145,1229,1661,1686]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="1" pageNumber="24">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BA79836A57BFF94C2BA8EBC971FF93A" blockId="1.[151,1437,1190,2014]" box="[199,1091,1697,1722]" pageId="1" pageNumber="24">
Head (
<figureCitation id="132384B3A57BFF94C36C8EBC9205F93A" box="[273,345,1697,1722]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="1" pageNumber="24">Figs 3</figureCitation>
, 
<figureCitation id="132384B3A57BFF94C3188EBF92D9F93A" box="[357,389,1698,1722]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="1" pageNumber="24">28</figureCitation>
). Eyes 1 + 1, about 8 µm in diameter, reddish or unpigmented.
</paragraph>
<paragraph id="8BA79836A57BFF94C2BA8ED89227F8EE" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
Clypeal area: row 
<emphasis id="B96C4424A57BFF94C3E58ED892FAF95E" bold="true" box="[408,422,1733,1758]" pageId="1" pageNumber="24">a</emphasis>
: 4 + axial + 4 setae, rows 
<emphasis id="B96C4424A57BFF94C0AB8ED891B9F95E" bold="true" box="[726,741,1733,1758]" pageId="1" pageNumber="24">b</emphasis>
, 
<emphasis id="B96C4424A57BFF94C0898ED8905CF95E" bold="true" box="[756,768,1733,1758]" pageId="1" pageNumber="24">c</emphasis>
and 
<emphasis id="B96C4424A57BFF94C1478ED8901AF95E" bold="true" box="[826,838,1733,1758]" pageId="1" pageNumber="24">e</emphasis>
with 5 + 5 setae each, row 
<emphasis id="B96C4424A57BFF94C6FF8ED897CEF95E" bold="true" box="[1154,1170,1733,1758]" pageId="1" pageNumber="24">d</emphasis>
with 7 + 7 setae, row 
<emphasis id="B96C4424A57BFF94C7EE8ED896C0F95E" bold="true" box="[1427,1436,1733,1758]" pageId="1" pageNumber="24">f</emphasis>
with 6 + 6 setae, median region M with 2 asymmetrically arranged setae (marked with oval in 
<figureCitation id="132384B3A57BFF94C6CC8EF497AFF882" box="[1201,1267,1769,1794]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="1" pageNumber="24">Fig. 3</figureCitation>
). Interantennal area: row 
<emphasis id="B96C4424A57BFF94C3758F109244F8A6" bold="true" box="[264,280,1805,1830]" pageId="1" pageNumber="24">α</emphasis>
: 2 + 2, row 
<emphasis id="B96C4424A57BFF94C3DE8F1092EEF8A6" bold="true" box="[419,434,1805,1830]" pageId="1" pageNumber="24">β</emphasis>
: 1 + axial + 1. Dorsal area (rows 
<emphasis id="B96C4424A57BFF94C14D8F109018F8A6" bold="true" box="[816,836,1805,1830]" pageId="1" pageNumber="24">A</emphasis>
to 
<emphasis id="B96C4424A57BFF94C1148F109021F8A6" bold="true" box="[873,893,1805,1830]" pageId="1" pageNumber="24">D</emphasis>
): axial seta present in rows 
<emphasis id="B96C4424A57BFF94C6C68F109793F8A6" bold="true" box="[1211,1231,1805,1830]" pageId="1" pageNumber="24">A</emphasis>
, 
<emphasis id="B96C4424A57BFF94C6A08F1097ACF8A6" bold="true" box="[1245,1264,1805,1830]" pageId="1" pageNumber="24">B</emphasis>
and 
<emphasis id="B96C4424A57BFF94C75A8F109667F8A6" bold="true" box="[1319,1339,1805,1830]" pageId="1" pageNumber="24">C</emphasis>
; weakly spine-like setae (
<figureCitation id="132384B3A57BFF94C3298F2C92FFF8CA" box="[340,419,1841,1866]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="1" pageNumber="24">Fig. 11</figureCitation>
) in rows 
<emphasis id="B96C4424A57BFF94C0708F2C917CF8CA" bold="true" box="[525,544,1841,1866]" pageId="1" pageNumber="24">B</emphasis>
(1 + 1), 
<emphasis id="B96C4424A57BFF94C0FC8F2C91C9F8CA" bold="true" box="[641,661,1841,1866]" pageId="1" pageNumber="24">C</emphasis>
(2 + 1 + 2) and 
<emphasis id="B96C4424A57BFF94C1378F2C9002F8CA" bold="true" box="[842,862,1841,1866]" pageId="1" pageNumber="24">D</emphasis>
(2 + 2). Chaetotaxy of the head posterior side as in 
<figureCitation id="132384B3A57BFF94C2EA8F489384F8EE" box="[151,216,1877,1902]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="1" pageNumber="24">Fig. 3</figureCitation>
(dotted lines).
</paragraph>
<paragraph id="8BA79836A57BFF94C2BA8F6496C0F85A" blockId="1.[151,1437,1190,2014]" pageId="1" pageNumber="24">
Chaetotaxy of mouth region. Labrum (
<figureCitation id="132384B3A57BFF94C0088F6491E5F812" box="[629,697,1913,1938]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="1" pageNumber="24">Fig. 3</figureCitation>
): labral / prelabral chaetotaxy: 4 5 5 / 6. Labium: submentum with 4 setae, mentum with 5 setae; labial palp (
<figureCitation id="132384B3A57BFF94C0FD8F809199F836" box="[640,709,1949,1974]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="1" pageNumber="24">Fig. 4</figureCitation>
) with 5 proximal setae, 5 papillae (
<emphasis id="B96C4424A57BFF94C61C8F809729F836" bold="true" box="[1121,1141,1949,1974]" pageId="1" pageNumber="24">A</emphasis>
, 
<emphasis id="B96C4424A57BFF94C6F88F8097C4F836" bold="true" box="[1157,1176,1949,1974]" pageId="1" pageNumber="24">B</emphasis>
, 
<emphasis id="B96C4424A57BFF94C6D58F8097E0F836" bold="true" box="[1192,1212,1949,1974]" pageId="1" pageNumber="24">C</emphasis>
, 
<emphasis id="B96C4424A57BFF94C6B08F8097BDF836" bold="true" box="[1229,1249,1949,1974]" pageId="1" pageNumber="24">D</emphasis>
, 
<emphasis id="B96C4424A57BFF94C68C8F809658F836" bold="true" box="[1265,1284,1949,1974]" pageId="1" pageNumber="24">E</emphasis>
) with deeply inserted setae and with 11 associated guard setae (hooked 
<emphasis id="B96C4424A57BFF94C1408FDC9008F85E" bold="true" box="[829,852,1985,2014]" pageId="1" pageNumber="24">a1</emphasis>
, 
<emphasis id="B96C4424A57BFF94C1198FDC90D4F85E" bold="true" box="[868,904,1985,2014]" pageId="1" pageNumber="24">b1-4</emphasis>
, 
<emphasis id="B96C4424A57BFF94C1E48FDC90E1F85E" bold="true" box="[921,957,1985,2014]" pageId="1" pageNumber="24">d1-3</emphasis>
, 
<emphasis id="B96C4424A57BFF94C1B38FDC90ACF85E" bold="true" box="[974,1008,1985,2014]" pageId="1" pageNumber="24">e1-3</emphasis>
) and 3 hypostomal setae (
<emphasis id="B96C4424A57BFF94C7588FDC9667F85A" bold="true" box="[1317,1339,1985,2010]" pageId="1" pageNumber="24">H</emphasis>
, 
<emphasis id="B96C4424A57BFF94C7318FDC9638F85E" bold="true" box="[1356,1380,1985,2014]" pageId="1" pageNumber="24">h1</emphasis>
, 
<emphasis id="B96C4424A57BFF94C7098FDC96D0F85E" bold="true" box="[1396,1420,1985,2014]" pageId="1" pageNumber="24">h2</emphasis>
).
</paragraph>
<paragraph id="8BA79836A578FF97C2EA888A91ADFF54" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Maxilla: apical seta of the maxillary outer lobe (
<figureCitation id="132384B3A578FF97C0C8888A91ABFF30" box="[693,759,151,176]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="2" pageNumber="25">Fig. 5</figureCitation>
) with short and thin subparallel branch at the base, sublobal plate with three sublobal hairs. Oral fold with 2 setae.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA88FD938CFE74" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Antenna (
<figureCitation id="132384B3A578FF97C34988C292C4FF78" box="[308,408,223,248]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="2" pageNumber="25">Figs 6–7</figureCitation>
) as long as body (~ 
<quantity id="4CE035D3A578FF97C00988FD91EFFF78" box="[628,691,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="25" unit="mm" value="1.0">1 mm</quantity>
) and about 3× of head. Length ratio of antennal segments I: II: III: IV = 1: 3.1: 5.2: 14.1; basal subsegment of Ant IV subequal with Ant III. Ant I with 7 setae, 
<emphasis id="B96C4424A578FF97C6BE8919978EFE9D" bold="true" box="[1219,1234,260,285]" pageId="2" pageNumber="25">p</emphasis>
as microseta. Ant II with 15 setae, two interior ones about twice longer than others. Ant III with 18 setae and 2 scabrous (not smooth) and relatively long (9–10 µm) sense rods in separate pits (
<figureCitation id="132384B3A578FF97C151895190DEFEE5" box="[812,898,332,357]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 26</figureCitation>
); antennal III organ as in figs 6 and 26. Ant IV (
<figureCitation id="132384B3A578FF97C2E2897293BFFE08" box="[159,227,367,392]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="2" pageNumber="25">Fig. 7</figureCitation>
) bears 13 whorls of setae located in 7 distinct subsegments; subsegmental formula: 1 + 5 + 1 = (
<emphasis id="B96C4424A578FF97C74F897293E3FE2D" bold="true" pageId="2" pageNumber="25">A1–A3 + M1</emphasis>
) + (
<emphasis id="B96C4424A578FF97C2858989920BFE2D" bold="true" box="[248,343,404,429]" pageId="2" pageNumber="25">M2–M6</emphasis>
) + (
<emphasis id="B96C4424A578FF97C3EC898991F3FE2D" bold="true" box="[401,687,404,429]" pageId="2" pageNumber="25">BA + BM1–BM2 + BB</emphasis>
). 1–2 small intercalar pseudosubsegments without setae present between normal subsegments. Apical subsegment with knobbed subapical organite and short blunt apical sensillum 
<emphasis id="B96C4424A578FF97C2EA89C19394FE75" bold="true" box="[151,200,476,501]" pageId="2" pageNumber="25">
A 
<specimenCount id="9D1E53BFA578FF97C2D189C19394FE75" box="[172,200,476,501]" pageId="2" pageNumber="25" type="generic">1p</specimenCount>
</emphasis>
.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA89E29616FD70" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Foreleg (
<figureCitation id="132384B3A578FF97C35089E2922AFD98" box="[301,374,511,536]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 8</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3F88A1D92FFFD98" box="[389,419,512,536]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">20</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3CD8A1D9293FD98" box="[432,463,512,536]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">27</figureCitation>
a): precoxae 1, 2 and coxa with 1, 0, 1 setae respectively (
<figureCitation id="132384B3A578FF97C61A89E297E1FD98" box="[1127,1213,511,536]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
). Trochanter with 3 anterior and 1 posterior setae. Femur with 14 setae, 
<emphasis id="B96C4424A578FF97C1788A39907EFDBD" bold="true" box="[773,802,548,573]" pageId="2" pageNumber="25">a4</emphasis>
turned perpendicularly to the longitudinal axis of the segment, 
<emphasis id="B96C4424A578FF97C3798A5A927FFDE0" bold="true" box="[260,291,583,608]" pageId="2" pageNumber="25">p1</emphasis>
and 
<emphasis id="B96C4424A578FF97C3208A5A9227FDE0" bold="true" box="[349,379,583,608]" pageId="2" pageNumber="25">p3</emphasis>
thin and short. Tibiotarsus with 45 setae: whorl I with 9 setae among which 
<emphasis id="B96C4424A578FF97C6858A5A9648FDE0" bold="true" box="[1272,1300,583,608]" pageId="2" pageNumber="25">Ja</emphasis>
curved and somewhat thickened, each of whorls II–V with 8 setae and region 
<emphasis id="B96C4424A578FF97C1D48A7190E6FD05" bold="true" box="[937,954,620,645]" pageId="2" pageNumber="25">F</emphasis>
with 3 primary 
<emphasis id="B96C4424A578FF97C6038A7197FCFD05" bold="true" box="[1150,1184,620,645]" pageId="2" pageNumber="25">FP</emphasis>
setae (
<emphasis id="B96C4424A578FF97C6858A719658FD05" bold="true" box="[1272,1284,620,645]" pageId="2" pageNumber="25">e</emphasis>
, 
<emphasis id="B96C4424A578FF97C76A8A71966DFD05" bold="true" box="[1303,1329,620,645]" pageId="2" pageNumber="25">ae</emphasis>
, 
<emphasis id="B96C4424A578FF97C73E8A719603FD05" bold="true" box="[1347,1375,620,645]" pageId="2" pageNumber="25">pe</emphasis>
) and secondary seta 
<emphasis id="B96C4424A578FF97C3398A92922FFD28" bold="true" box="[324,371,655,680]" pageId="2" pageNumber="25">FSa</emphasis>
. Pretarsus with 1 anterior and 1 posterior setulae. Foot complex (
<figureCitation id="132384B3A578FF97C6258A9297F1FD28" box="[1112,1197,655,680]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 27</figureCitation>
a): claw thin, without tunica, with inner tooth and two pairs of indistinct lateral teeth, about 5.8× shorter than tibiotarsus; empodium about 1.7× shorter than claw, thin, with corner tooth, with short apical filament not reaching the tip of claw.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8AE19201FC6D" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Mid leg (
<figureCitation id="132384B3A578FF97C3538AE1922BFC94" box="[302,375,764,789]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 9</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3F98AE192FEFC94" box="[388,418,764,788]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">20</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3D38AE19291FC95" box="[430,461,764,789]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">27</figureCitation>
b): precoxae 1 and 2 with 1, 1 setae respectively, precoxal process present, coxa with 3 setae and microsensillum (
<figureCitation id="132384B3A578FF97C3B08B029179FCB8" box="[461,549,799,824]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
). Trochanter with 5 setae: 3 simple anterior, trochanteral organ and 1simple posterior. Femur with 13 setae (
<emphasis id="B96C4424A578FF97C3868B59917AFCDD" bold="true" box="[507,550,836,861]" pageId="2" pageNumber="25">pe2</emphasis>
and 
<emphasis id="B96C4424A578FF97C0218B5991D9FCDD" bold="true" box="[604,645,836,861]" pageId="2" pageNumber="25">ae3</emphasis>
absent), 
<emphasis id="B96C4424A578FF97C0978B599054FCDD" bold="true" box="[746,776,836,861]" pageId="2" pageNumber="25">p1</emphasis>
and 
<emphasis id="B96C4424A578FF97C1438B599000FCDD" bold="true" box="[830,860,836,861]" pageId="2" pageNumber="25">p3</emphasis>
very thin. Tibiotarsus with 44 setae: whorl I with 9 setae whorls II–IV with 8 setae, whorl V with 7 setae (
<emphasis id="B96C4424A578FF97C16D8B7A9068FC00" bold="true" box="[784,820,871,896]" pageId="2" pageNumber="25">Vp</emphasis>
absent), region 
<emphasis id="B96C4424A578FF97C1908B7A90A2FC00" bold="true" box="[1005,1022,871,896]" pageId="2" pageNumber="25">F</emphasis>
with 3 
<emphasis id="B96C4424A578FF97C6248B7A9727FC00" bold="true" box="[1113,1147,871,896]" pageId="2" pageNumber="25">FP</emphasis>
setae and 
<emphasis id="B96C4424A578FF97C68B8B7A9679FC00" bold="true" box="[1270,1317,871,896]" pageId="2" pageNumber="25">FSa</emphasis>
seta. Foot complex (
<figureCitation id="132384B3A578FF97C37A8B919206FC25" box="[263,346,908,933]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 27</figureCitation>
b): claw thin, without tunica, with inner tooth, with 2 pairs of small lateral teeth, about 6× shorter than tibiotarsus; empodium somewhat broader and distinctly shorter than in foreleg, with corner tooth, about 2.2× shorter than claw.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8BEA92CBFB45" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Hind leg (
<figureCitation id="132384B3A578FF97C3448BEA92CCFB90" box="[313,400,1015,1040]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 10</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3E08BE592E7FB90" box="[413,443,1016,1040]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">20</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3BA8BE592BBFB90" box="[455,487,1016,1040]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">27</figureCitation>
c): precoxae 1 and 2 with 1, 1 setae respectively, process on precoxa 1 present, coxa with 3 setae and microsensillum (
<figureCitation id="132384B3A578FF97C0698C019135FBB4" box="[532,617,1052,1077]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
). Trochanter chaetotaxy as in midleg. Femur with 12 setae (
<emphasis id="B96C4424A578FF97C76C8C019660FBB5" bold="true" box="[1297,1340,1052,1077]" pageId="2" pageNumber="25">pe2</emphasis>
and 
<emphasis id="B96C4424A578FF97C7098C0196C7FBB5" bold="true" box="[1396,1435,1052,1077]" pageId="2" pageNumber="25">ae3</emphasis>
absent), 
<emphasis id="B96C4424A578FF97C28B8C229248FBD8" bold="true" box="[246,276,1087,1112]" pageId="2" pageNumber="25">p1</emphasis>
and 
<emphasis id="B96C4424A578FF97C3368C229234FBD8" bold="true" box="[331,360,1087,1112]" pageId="2" pageNumber="25">p3</emphasis>
as microsetae. Tibiotarsus with 44 setae as in mid leg. Foot complex (
<figureCitation id="132384B3A578FF97C6FC8C229788FBD8" box="[1153,1236,1087,1112]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 27</figureCitation>
c): claw relatively slender, without tunica, with inner tooth, with 2 pairs of small lateral teeth, about 8× shorter than tibiotarsus; empodium about 1.6× shorter than claw, broad, without corner tooth, with short apical filament not reaching the tip of corresponding claw.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8CD2979DFB68" blockId="2.[151,1437,151,2013]" box="[199,1217,1231,1256]" pageId="2" pageNumber="25">Lengths ratio of tibiotarsi I: II: III = 1: 1: 1.2. Tibiotarsus I about 1.2× shorter than head.</paragraph>
<paragraph id="8BA79836A578FF97C2BA8CE991A3FA8D" blockId="2.[151,1437,151,2013]" box="[199,767,1268,1293]" pageId="2" pageNumber="25">
Ventral tube (
<figureCitation id="132384B3A578FF97C3238CE992ECFA8C" box="[350,432,1268,1293]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
) with 1 + 1 subapical setulae.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8D0A9636FAB0" blockId="2.[151,1437,151,2013]" box="[199,1386,1303,1328]" pageId="2" pageNumber="25">
Tenaculum (
<figureCitation id="132384B3A578FF97C32F8D0A92FAFAB0" box="[338,422,1303,1328]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 22</figureCitation>
): with 2 apical setulae on the anterior lobe, 3 teeth and a basal process on each ramus.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8D2090F5FA3F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Furca (
<figureCitation id="132384B3A578FF97C3648D21922CFAD4" box="[281,368,1340,1365]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 21</figureCitation>
): manubrium with 7 + 7 posterior setae. Dens (23 setae): anterior side with 3, 2, 1, 1 setae; posterior side with 
<emphasis id="B96C4424A578FF97C3108D4292C8FAF8" bold="true" box="[365,404,1375,1400]" pageId="2" pageNumber="25">Ipe</emphasis>
and 
<emphasis id="B96C4424A578FF97C3B48D4292A7FAF8" bold="true" box="[457,507,1375,1400]" pageId="2" pageNumber="25">IIpe</emphasis>
as massive spines (
<figureCitation id="132384B3A578FF97C0A98D429008FAF8" box="[724,852,1375,1400]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 14, 15</figureCitation>
), 
<emphasis id="B96C4424A578FF97C1148D429020FAF8" bold="true" box="[873,892,1375,1400]" pageId="2" pageNumber="25">Ii</emphasis>
moderately spine-like, 
<emphasis id="B96C4424A578FF97C6FE8D4297E7FAF8" bold="true" box="[1155,1211,1375,1400]" pageId="2" pageNumber="25">IIIpi</emphasis>
and 
<emphasis id="B96C4424A578FF97C68D8D42967AFAF8" bold="true" box="[1264,1318,1375,1400]" pageId="2" pageNumber="25">IVpi</emphasis>
not spinelike. Mucro (
<figureCitation id="132384B3A578FF97C3498D9992D1FA1C" box="[308,397,1412,1437]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 30</figureCitation>
): posterior lamellae serrated (about 25–30 teeth), anterior lamella poorly developed; tip rounded and somewhat broadened. Dens about 1.8× as long as mucro.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8DD1938AF884" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Great abdomen (
<figureCitation id="132384B3A578FF97C3FF8DD1928BFA64" box="[386,471,1484,1509]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
): Th II bears single sensillum in row 
<emphasis id="B96C4424A578FF97C10B8DD190D8FA65" bold="true" box="[886,900,1484,1509]" pageId="2" pageNumber="25">a</emphasis>
and 3 setae in row 
<emphasis id="B96C4424A578FF97C6218DD1972FFA65" bold="true" box="[1116,1139,1484,1509]" pageId="2" pageNumber="25">m</emphasis>
with 
<emphasis id="B96C4424A578FF97C6CE8DD19784FA65" bold="true" box="[1203,1240,1484,1509]" pageId="2" pageNumber="25">m1</emphasis>
somewhat spinelike and longer than other setae of Th II (
<figureCitation id="132384B3A578FF97C01F8DF291E9F988" box="[610,693,1519,1544]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Fig. 12</figureCitation>
). Th III with sensillum in row 
<emphasis id="B96C4424A578FF97C6708DF29747F988" bold="true" box="[1037,1051,1519,1544]" pageId="2" pageNumber="25">a</emphasis>
and 3 setae in row 
<emphasis id="B96C4424A578FF97C6898DF29657F988" bold="true" box="[1268,1291,1519,1544]" pageId="2" pageNumber="25">m</emphasis>
. Abd I bears single row with 5 short setae (
<figureCitation id="132384B3A578FF97C3998E09916AF9AC" box="[484,566,1556,1581]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Fig. 13</figureCitation>
). Abd II with row of 4 setae anterior to trichobothrial complex. Trichobothrial complex (
<figureCitation id="132384B3A578FF97C37B8E2A9207F9D0" box="[262,347,1591,1616]" captionStart="FIGURES 25 – 30" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[184,1402,213,1899]" captionTargetId="figure@8.[165,1422,193,1932]" captionTargetPageId="8" captionText="FIGURES 25 – 30. Pygmarrhopalites dbari sp. nov.: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens." httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="2" pageNumber="25">Fig. 29</figureCitation>
): 
<emphasis id="B96C4424A578FF97C3128E2A92F6F9D0" bold="true" box="[367,426,1591,1616]" pageId="2" pageNumber="25">ABC</emphasis>
form an angle about 160° (157–161°) and 
<emphasis id="B96C4424A578FF97C1F68E2A90EEF9D0" bold="true" box="[907,946,1591,1616]" pageId="2" pageNumber="25">AB</emphasis>
is ~0.8× of 
<emphasis id="B96C4424A578FF97C6478E2A973DF9D0" bold="true" box="[1082,1121,1591,1616]" pageId="2" pageNumber="25">BC</emphasis>
. Seta 
<emphasis id="B96C4424A578FF97C6DB8E2A97E9F9D0" bold="true" box="[1190,1205,1591,1616]" pageId="2" pageNumber="25">p</emphasis>
is located below the level of trichobothrium 
<emphasis id="B96C4424A578FF97C3DC8E4192E8F9F5" bold="true" box="[417,436,1628,1653]" pageId="2" pageNumber="25">B</emphasis>
; seta 
<emphasis id="B96C4424A578FF97C3888E41914FF9F5" bold="true" box="[501,531,1628,1653]" pageId="2" pageNumber="25">b1</emphasis>
lies almost in the middle of line 
<emphasis id="B96C4424A578FF97C1FE8E4190F6F9F5" bold="true" box="[899,938,1628,1653]" pageId="2" pageNumber="25">BC</emphasis>
; seta 
<emphasis id="B96C4424A578FF97C1968E41975AF9F5" bold="true" box="[1003,1030,1628,1653]" pageId="2" pageNumber="25">c1</emphasis>
(~27 µm) somewhat shorter than 
<emphasis id="B96C4424A578FF97C7FF8E4196C0F9F5" bold="true" box="[1410,1436,1628,1653]" pageId="2" pageNumber="25">c2</emphasis>
but not reduced. Posterior lateral complex with 6 and furca base complex with 9 setae. Central dorsal complex with three setae of which seta 
<emphasis id="B96C4424A578FF97C3C68EB99295F93D" bold="true" box="[443,457,1700,1725]" pageId="2" pageNumber="25">3</emphasis>
is shorter and thinner than 
<emphasis id="B96C4424A578FF97C17B8EB99048F93D" bold="true" box="[774,788,1700,1725]" pageId="2" pageNumber="25">1</emphasis>
and 
<emphasis id="B96C4424A578FF97C1328EB99001F93D" bold="true" box="[847,861,1700,1725]" pageId="2" pageNumber="25">2</emphasis>
(
<figureCitation id="132384B3A578FF97C1108EB99098F93C" box="[877,964,1700,1725]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
). Posterior dorsal complex with about 18 setae the longest of which (
<emphasis id="B96C4424A578FF97C3B68EDA92A1F960" bold="true" box="[459,509,1735,1760]" pageId="2" pageNumber="25">dI-1</emphasis>
~100 µm) is 2–2.4× longer than hind claw (
<figureCitation id="132384B3A578FF97C1908EDA971DF960" box="[1005,1089,1735,1760]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Fig. 16</figureCitation>
). Ventral complex with up to 3 setae.
</paragraph>
<paragraph id="8BA79836A578FF97C2BA8F1290A3F8CC" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="25">
Fifth abdominal segment (
<figureCitation id="132384B3A578FF97C38C8F12911BF8A8" box="[497,583,1807,1832]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Fig. 20</figureCitation>
): with 2 setae and trichobothrium 
<emphasis id="B96C4424A578FF97C1B68F129083F8A8" bold="true" box="[971,991,1807,1832]" pageId="2" pageNumber="25">D</emphasis>
in row 
<emphasis id="B96C4424A578FF97C6448F12971BF8A8" bold="true" box="[1081,1095,1807,1832]" pageId="2" pageNumber="25">a</emphasis>
, and 2 setae in row 
<emphasis id="B96C4424A578FF97C7508F129660F8A8" bold="true" box="[1325,1340,1807,1832]" pageId="2" pageNumber="25">p</emphasis>
; seta 
<emphasis id="B96C4424A578FF97C7FD8F1296C0F8A8" bold="true" box="[1408,1436,1807,1832]" pageId="2" pageNumber="25">a3</emphasis>
thinner and somewhat shorter than others. Genital field with 2 + 2 short setae.
</paragraph>
<paragraph id="8BA79836A578FF96C2BA8F4A9281FF54" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1437,151,464]" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
Sixth abdominal segment (
<figureCitation id="132384B3A578FF97C38A8F4A9113F8F0" box="[503,591,1879,1904]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="2" pageNumber="25">Figs 23</figureCitation>
, 
<figureCitation id="132384B3A578FF97C0208F459121F8F0" box="[605,637,1880,1904]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="2" pageNumber="25">33</figureCitation>
): dorsal valve with 11 + 2 axial + 11 setae, of which circumanal setae 
<emphasis id="B96C4424A578FF97C2EA8F61939BF815" bold="true" box="[151,199,1916,1941]" pageId="2" pageNumber="25">ms1</emphasis>
(
<figureCitation id="132384B3A578FF97C2AB8F619275F814" box="[214,297,1916,1941]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Fig. 17</figureCitation>
), 
<emphasis id="B96C4424A578FF97C3438F619221F815" bold="true" box="[318,381,1916,1941]" pageId="2" pageNumber="25">mps1</emphasis>
, 
<emphasis id="B96C4424A578FF97C3F18F619291F815" bold="true" box="[396,461,1916,1941]" pageId="2" pageNumber="25">mps2</emphasis>
and 
<emphasis id="B96C4424A578FF97C07F8F61911EF815" bold="true" box="[514,578,1916,1941]" pageId="2" pageNumber="25">mps3</emphasis>
(
<figureCitation id="132384B3A578FF97C02C8F6191F4F814" box="[593,680,1916,1941]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 18</figureCitation>
, 
<figureCitation id="132384B3A578FF97C0C98F61918DF814" box="[692,721,1916,1940]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="2" pageNumber="25">33</figureCitation>
) are moderately broadened. Each of lateral valve bears 20 setae with broadened 
<emphasis id="B96C4424A578FF97C3378FBD92DAF839" bold="true" box="[330,390,1952,1977]" pageId="2" pageNumber="25">mpi1</emphasis>
and 
<emphasis id="B96C4424A578FF97C3C08FBD92A5F839" bold="true" box="[445,505,1952,1977]" pageId="2" pageNumber="25">mpi2</emphasis>
(
<figureCitation id="132384B3A578FF97C0758FBD9107F838" box="[520,603,1952,1977]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="2" pageNumber="25">Fig. 33</figureCitation>
). Some of circumanal setae (e.g. 
<emphasis id="B96C4424A578FF97C1B08FBD9751F839" bold="true" box="[973,1037,1952,1977]" pageId="2" pageNumber="25">mps3</emphasis>
, 
<emphasis id="B96C4424A578FF97C6668FBD970BF839" bold="true" box="[1051,1111,1952,1977]" pageId="2" pageNumber="25">mpi2</emphasis>
) slightly serrated near bases. Appendices anales (
<figureCitation id="132384B3A578FF97C3018FD99289F85C" box="[380,469,1988,2013]" captionStart="FIGURES 8 – 19" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[224,1360,222,1893]" captionTargetId="figure@6.[187,1399,193,1932]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Pygmarrhopalites dbari sp. nov.: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11 – 19, shape of setae: 11, spine-like seta of head vertex; 12, m 1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI- 1 of great abdomen; 17, ms 1 of Abd VI; 18, mps 3 of abd VI; 19, appendices anales." httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="2" pageNumber="25">Figs 19</figureCitation>
, 
<figureCitation id="132384B3A578FF97C3998FD9915DF85C" box="[484,513,1988,2012]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="2" pageNumber="25">33</figureCitation>
) palmate and broadened in distal third, with secondary branches present (length about 27 µm and width of the palmated part about 15 µm in dorsoventral position). In total, normally 64 setae on Abd VI are present (
<figureCitation id="132384B3A579FF96C30188A19293FF54" box="[380,463,188,213]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="3" pageNumber="26">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BA79836A579FF96C2BA88C292C8FE50" blockId="3.[151,1437,151,464]" pageId="3" pageNumber="26">
<emphasis id="B96C4424A579FF96C2BA88C29257FF78" bold="true" box="[199,267,223,248]" pageId="3" pageNumber="26">Male.</emphasis>
Habitus as in 
<figureCitation id="132384B3A579FF96C3BD88C29156FF78" box="[448,522,223,248]" captionStart="FIGURES 1 – 2. 1" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[317,1260,226,1922]" captionTargetId="figure@4.[284,1297,195,1940]" captionTargetPageId="4" captionText="FIGURES 1 – 2. 1, type localities of Pygmarrhopalites dbari sp. nov. (1 — Psyrtskha Cave), and P. kovali sp. nov. (2 — Tsebeldinskaya Cave, 3 — Tshalintshur Cave, 4 — Verkhnyaya Shakuranskaya Cave); 2, habitus of mounted male." httpUri="https://zenodo.org/record/439781/files/figure.png" pageId="3" pageNumber="26">Fig. 2</figureCitation>
. Body length about 
<quantity id="4CE035D3A579FF96C17C88FD9002FF77" box="[769,862,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="26" unit="mm" value="0.8">0.8 mm</quantity>
, measurements are given in 
<tableCitation id="C69AAD8DA579FF96C6CA88C29653FF78" box="[1207,1295,223,248]" captionStart="TABLE 1" captionStartId="3.[151,239,512,535]" captionText="TABLE 1. Measurements (in µm) for body parts of types of Pygmarrhopalites dbari sp. nov. and P. kovali sp. nov." pageId="3" pageNumber="26">Table 1</tableCitation>
, chaetotaxy resembles the one in female, except for small abdomen. Ant IV has a stronger tendency than females to separate terminal part of subsegments and thus bears up to 5 short naked intercalar pseudosubsegments between normal subsegments (
<figureCitation id="132384B3A579FF96C33E895192F8FEE4" box="[323,420,332,357]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="3" pageNumber="26">Fig. 31</figureCitation>
). Besides, the basal subsegment is often secondarily subdivided into up to 5 pseudosubsegments by several annuli between them (
<figureCitation id="132384B3A579FF96C08D8972901FFE08" box="[752,835,367,392]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="3" pageNumber="26">Fig. 31</figureCitation>
). Abd VI (
<figureCitation id="132384B3A579FF96C1C189729748FE08" box="[956,1044,367,392]" captionStart="FIGURES 20 – 24" captionStartId="7.[151,264,1938,1961]" captionTargetBox="[197,1397,193,1879]" captionTargetId="figure@7.[192,1397,193,1921]" captionTargetPageId="7" captionText="FIGURES 20 – 24. Pygmarrhopalites dbari sp. nov.: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI." httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="3" pageNumber="26">Figs 24</figureCitation>
, 
<figureCitation id="132384B3A579FF96C662896D9760FE08" box="[1055,1084,368,392]" captionStart="FIGURES 31 – 33" captionStartId="9.[151,264,1460,1483]" captionTargetBox="[260,1350,222,1407]" captionTargetId="figure@9.[213,1373,175,1457]" captionTargetPageId="9" captionText="FIGURES 31 – 33. Pygmarrhopalites dbari sp. nov.: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view." httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="3" pageNumber="26">32</figureCitation>
) without broadened setae, with 7 + 2 + 7 setae on dorsal valve and 14 + 14 setae on lateral valves: 
<quantity id="4CE035D3A579FF96C1F38989909BFE2D" box="[910,967,404,429]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" pageId="3" pageNumber="26" unit="in" value="44.0">44 in</quantity>
total. Genital opening is surrounded with about 14 short setulae.
</paragraph>
<caption id="DF67C8BEA579FF96C2EA8A1D963BFD97" box="[151,1383,511,535]" pageId="3" pageNumber="26">
<paragraph id="8BA79836A579FF96C2EA8A1D963BFD97" blockId="3.[151,1383,511,535]" box="[151,1383,511,535]" pageId="3" pageNumber="26">
<emphasis id="B96C4424A579FF96C2EA8A1D925FFD97" bold="true" box="[151,259,512,535]" pageId="3" pageNumber="26">TABLE 1</emphasis>
. Measurements (in µm) for body parts of types of 
<taxonomicName id="4C18E3B5A579FF96C16289E2977CFD97" box="[799,1056,511,535]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A579FF96C16289E2977CFD97" box="[799,1056,511,535]" italics="true" pageId="3" pageNumber="26">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A579FF96C65B8A1D9720FD97" bold="true" box="[1062,1148,512,535]" pageId="3" pageNumber="26">
<taxonomicNameLabel id="A25FF95FA579FF96C65B8A1D9720FD97" box="[1062,1148,512,535]" pageId="3" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C18E3B5A579FF96C6D28A1D9657FD97" box="[1199,1291,511,535]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A579FF96C6D28A1D9657FD97" box="[1199,1291,511,535]" italics="true" pageId="3" pageNumber="26">P. kovali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25FF95FA579FF96C76C8A1D963BFD97" box="[1297,1383,512,535]" pageId="3" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA79836A579FF96C2E28A2F90BDFDC8" blockId="3.[159,1385,562,708]" box="[159,993,562,584]" pageId="3" pageNumber="26">
Body part 
<emphasis id="B96C4424A579FF96C3DE8A2F90BDFDC8" box="[419,993,562,584]" italics="true" pageId="3" pageNumber="26">
<taxonomicName id="4C18E3B5A579FF96C3DE8A2F9286FDC8" box="[419,474,562,584]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="dbari">dbari</taxonomicName>
<taxonomicName id="4C18E3B5A579FF96C1DE8A2F90BDFDC8" box="[931,993,562,584]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="kovali">kovali</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA79836A579FF96C3DE8A419649FDF2" blockId="3.[159,1385,562,708]" box="[419,1301,604,626]" pageId="3" pageNumber="26">
Females, n= 
<specimenCount id="9D1E53BFA579FF96C0628A4191BFFDF2" box="[543,739,604,626]" pageId="3" pageNumber="26" type="male">4 Males</specimenCount>
, n= 
<specimenCount id="9D1E53BFA579FF96C1748A4190A6FDF2" box="[777,1018,604,626]" pageId="3" pageNumber="26" type="female">6 Females</specimenCount>
, n= 
<specimenCount id="9D1E53BFA579FF96C65D8A4197BFFDF2" box="[1056,1251,604,626]" pageId="3" pageNumber="26" type="male">3 Males</specimenCount>
, n=2
</paragraph>
<paragraph id="8BA79836A579FF96C3DE8A989767F865" blockId="3.[159,1385,562,708]" lastBlockId="3.[151,1437,1889,2022]" pageId="3" pageNumber="26">
min–max mean min–max mean min–max mean min–max mean Total 
<date id="FFA6BEF6A579FF96C3DE8AB3914AFD44" box="[419,534,686,708]" pageId="3" pageNumber="26" value="1000" valueMax="1088">1000–1088</date>
1065 772–880 831 920–960 937 830–? 830 
<emphasis id="B96C4424A579FF96C2BA8F7C9211F8FA" bold="true" box="[199,333,1889,1914]" pageId="3" pageNumber="26">Variability.</emphasis>
A number of short naked intercalar pseudosubsegments on Ant IV varies: in females from 1 (usually) to 2, in males from 2 to 5 (most often 4). Secondary pseudosubsegments of the Ant IV basal subsegment usual in males and vary from 2 to 5. Among 14 examined specimens, in one female, a small corner tooth was observed on one empodim III. Ventral complex of great abdomen with 3 or 2 setae.
</paragraph>
<paragraph id="8BA79836A579FF96C2E28AC59604F8A0" pageId="3" pageNumber="26">
<table id="F9186A96A579006AC2E28AC59635F8A0" box="[159,1385,728,1824]" gridcols="9" gridrows="27" pageId="3" pageNumber="26">
<tr id="35289A74A579006AC2E28AC59635FD6E" box="[159,1385,728,750]" gridrow="0" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28AC59234FD6E" box="[159,360,728,750]" gridcol="0" gridrow="0" pageId="3" pageNumber="26">Body</th>
<th id="76F9F308A579006AC3DE8AC59154FD6E" box="[419,520,728,750]" gridcol="1" gridrow="0" pageId="3" pageNumber="26">670–740</th>
<th id="76F9F308A579006AC0468AC59132FD6E" box="[571,622,728,750]" gridcol="2" gridrow="0" pageId="3" pageNumber="26">717</th>
<th id="76F9F308A579006AC0DE8AC591A0FD6E" box="[675,764,728,750]" gridcol="3" gridrow="0" pageId="3" pageNumber="26">512–640</th>
<th id="76F9F308A579006AC14F8AC59004FD6E" box="[818,856,728,750]" gridcol="4" gridrow="0" pageId="3" pageNumber="26">584</th>
<th id="76F9F308A579006AC1DE8AC590A0FD6E" box="[931,1020,728,750]" gridcol="5" gridrow="0" pageId="3" pageNumber="26">610–677</th>
<th id="76F9F308A579006AC6468AC5973DFD6E" box="[1083,1121,728,750]" gridcol="6" gridrow="0" pageId="3" pageNumber="26">636</th>
<th id="76F9F308A579006AC6DE8AC597A0FD6E" box="[1187,1276,728,750]" gridcol="7" gridrow="0" pageId="3" pageNumber="26">580–?</th>
<th id="76F9F308A579006AC74F8AC59635FD6E" box="[1330,1385,728,750]" gridcol="8" gridrow="0" pageId="3" pageNumber="26">580</th>
</tr>
<tr id="35289A74A579006AC2E28B1C9635FC97" box="[159,1385,769,791]" gridrow="1" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28B1C9234FC97" box="[159,360,769,791]" gridcol="0" gridrow="1" pageId="3" pageNumber="26">Head</th>
<td id="76F9F308A579006AC3DE8B1C9154FC97" box="[419,520,769,791]" gridcol="1" gridrow="1" pageId="3" pageNumber="26">345–368</td>
<td id="76F9F308A579006AC0468B1C9132FC97" box="[571,622,769,791]" gridcol="2" gridrow="1" pageId="3" pageNumber="26">357</td>
<td id="76F9F308A579006AC0DE8B1C91A0FC97" box="[675,764,769,791]" gridcol="3" gridrow="1" pageId="3" pageNumber="26">265–300</td>
<td id="76F9F308A579006AC14F8B1C9004FC97" box="[818,856,769,791]" gridcol="4" gridrow="1" pageId="3" pageNumber="26">280</td>
<td id="76F9F308A579006AC1DE8B1C90A0FC97" box="[931,1020,769,791]" gridcol="5" gridrow="1" pageId="3" pageNumber="26">310–360</td>
<td id="76F9F308A579006AC6468B1C973DFC97" box="[1083,1121,769,791]" gridcol="6" gridrow="1" pageId="3" pageNumber="26">336</td>
<td id="76F9F308A579006AC6DE8B1C97A0FC97" box="[1187,1276,769,791]" gridcol="7" gridrow="1" pageId="3" pageNumber="26">296–307</td>
<td id="76F9F308A579006AC74F8B1C9635FC97" box="[1330,1385,769,791]" gridcol="8" gridrow="1" pageId="3" pageNumber="26">302</td>
</tr>
<tr id="35289A74A579006AC2E28B379635FCC0" box="[159,1385,810,832]" gridrow="2" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28B379234FCC0" box="[159,360,810,832]" gridcol="0" gridrow="2" pageId="3" pageNumber="26">Head longest spine</th>
<td id="76F9F308A579006AC3DE8B379154FCC0" box="[419,520,810,832]" gridcol="1" gridrow="2" pageId="3" pageNumber="26">23–35</td>
<td id="76F9F308A579006AC0468B379132FCC0" box="[571,622,810,832]" gridcol="2" gridrow="2" pageId="3" pageNumber="26">29</td>
<td id="76F9F308A579006AC0DE8B3791A0FCC0" box="[675,764,810,832]" gridcol="3" gridrow="2" pageId="3" pageNumber="26">22–27</td>
<td id="76F9F308A579006AC14F8B379004FCC0" box="[818,856,810,832]" gridcol="4" gridrow="2" pageId="3" pageNumber="26">23</td>
<td id="76F9F308A579006AC1DE8B3790A0FCC0" box="[931,1020,810,832]" gridcol="5" gridrow="2" pageId="3" pageNumber="26">24–26</td>
<td id="76F9F308A579006AC6468B37973DFCC0" box="[1083,1121,810,832]" gridcol="6" gridrow="2" pageId="3" pageNumber="26">25</td>
<td id="76F9F308A579006AC6DE8B3797A0FCC0" box="[1187,1276,810,832]" gridcol="7" gridrow="2" pageId="3" pageNumber="26">22–23</td>
<td id="76F9F308A579006AC74F8B379635FCC0" box="[1330,1385,810,832]" gridcol="8" gridrow="2" pageId="3" pageNumber="26">22</td>
</tr>
<tr id="35289A74A579006AC2E28B499635FCEA" box="[159,1385,852,874]" gridrow="3" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28B499234FCEA" box="[159,360,852,874]" gridcol="0" gridrow="3" pageId="3" pageNumber="26">Eye diameter</th>
<td id="76F9F308A579006AC3DE8B499154FCEA" box="[419,520,852,874]" gridcol="1" gridrow="3" pageId="3" pageNumber="26">7.0–9.0</td>
<td id="76F9F308A579006AC0468B499132FCEA" box="[571,622,852,874]" gridcol="2" gridrow="3" pageId="3" pageNumber="26">8.0</td>
<td id="76F9F308A579006AC0DE8B4991A0FCEA" box="[675,764,852,874]" gridcol="3" gridrow="3" pageId="3" pageNumber="26">6.0–6.7</td>
<td id="76F9F308A579006AC14F8B499004FCEA" box="[818,856,852,874]" gridcol="4" gridrow="3" pageId="3" pageNumber="26">6.4</td>
<td id="76F9F308A579006AC1DE8B4990A0FCEA" box="[931,1020,852,874]" gridcol="5" gridrow="3" pageId="3" pageNumber="26">9.3–10.4</td>
<td id="76F9F308A579006AC6468B49973DFCEA" box="[1083,1121,852,874]" gridcol="6" gridrow="3" pageId="3" pageNumber="26">9.8</td>
<td id="76F9F308A579006AC6DE8B4997A0FCEA" box="[1187,1276,852,874]" gridcol="7" gridrow="3" pageId="3" pageNumber="26">9.2–9.3</td>
<td id="76F9F308A579006AC74F8B499635FCEA" box="[1330,1385,852,874]" gridcol="8" gridrow="3" pageId="3" pageNumber="26">9.3</td>
</tr>
<tr id="35289A74A579006AC2E28B609635FC13" box="[159,1385,893,915]" gridrow="4" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28B609234FC13" box="[159,360,893,915]" gridcol="0" gridrow="4" pageId="3" pageNumber="26">Antenna</th>
<td id="76F9F308A579006AC3DE8B609154FC13" box="[419,520,893,915]" gridcol="1" gridrow="4" pageId="3" pageNumber="26">98 9–1033</td>
<td id="76F9F308A579006AC0468B609132FC13" box="[571,622,893,915]" gridcol="2" gridrow="4" pageId="3" pageNumber="26">1017</td>
<td id="76F9F308A579006AC0DE8B6091A0FC13" box="[675,764,893,915]" gridcol="3" gridrow="4" pageId="3" pageNumber="26">837–856</td>
<td id="76F9F308A579006AC14F8B609004FC13" box="[818,856,893,915]" gridcol="4" gridrow="4" pageId="3" pageNumber="26">850</td>
<td id="76F9F308A579006AC1DE8B6090A0FC13" box="[931,1020,893,915]" gridcol="5" gridrow="4" pageId="3" pageNumber="26">740–777</td>
<td id="76F9F308A579006AC6468B60973DFC13" box="[1083,1121,893,915]" gridcol="6" gridrow="4" pageId="3" pageNumber="26">756</td>
<td id="76F9F308A579006AC6DE8B6097A0FC13" box="[1187,1276,893,915]" gridcol="7" gridrow="4" pageId="3" pageNumber="26">693–745</td>
<td id="76F9F308A579006AC74F8B609635FC13" box="[1330,1385,893,915]" gridcol="8" gridrow="4" pageId="3" pageNumber="26">719</td>
</tr>
<tr id="35289A74A579006AC2E28BBB9635FC3C" box="[159,1385,934,956]" gridrow="5" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28BBB9234FC3C" box="[159,360,934,956]" gridcol="0" gridrow="5" pageId="3" pageNumber="26">Ant I</th>
<td id="76F9F308A579006AC3DE8BBB9154FC3C" box="[419,520,934,956]" gridcol="1" gridrow="5" pageId="3" pageNumber="26">42–47</td>
<td id="76F9F308A579006AC0468BBB9132FC3C" box="[571,622,934,956]" gridcol="2" gridrow="5" pageId="3" pageNumber="26">45</td>
<td id="76F9F308A579006AC0DE8BBB91A0FC3C" box="[675,764,934,956]" gridcol="3" gridrow="5" pageId="3" pageNumber="26">36–38</td>
<td id="76F9F308A579006AC14F8BBB9004FC3C" box="[818,856,934,956]" gridcol="4" gridrow="5" pageId="3" pageNumber="26">37</td>
<td id="76F9F308A579006AC1DE8BBB90A0FC3C" box="[931,1020,934,956]" gridcol="5" gridrow="5" pageId="3" pageNumber="26">42–45</td>
<td id="76F9F308A579006AC6468BBB973DFC3C" box="[1083,1121,934,956]" gridcol="6" gridrow="5" pageId="3" pageNumber="26">44</td>
<td id="76F9F308A579006AC6DE8BBB97A0FC3C" box="[1187,1276,934,956]" gridcol="7" gridrow="5" pageId="3" pageNumber="26">36–37</td>
<td id="76F9F308A579006AC74F8BBB9635FC3C" box="[1330,1385,934,956]" gridcol="8" gridrow="5" pageId="3" pageNumber="26">37</td>
</tr>
<tr id="35289A74A579006AC2E28BCD9635FC66" box="[159,1385,976,998]" gridrow="6" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28BCD9234FC66" box="[159,360,976,998]" gridcol="0" gridrow="6" pageId="3" pageNumber="26">Ant II</th>
<td id="76F9F308A579006AC3DE8BCD9154FC66" box="[419,520,976,998]" gridcol="1" gridrow="6" pageId="3" pageNumber="26">13 6–140</td>
<td id="76F9F308A579006AC0468BCD9132FC66" box="[571,622,976,998]" gridcol="2" gridrow="6" pageId="3" pageNumber="26">137</td>
<td id="76F9F308A579006AC0DE8BCD91A0FC66" box="[675,764,976,998]" gridcol="3" gridrow="6" pageId="3" pageNumber="26">103–124</td>
<td id="76F9F308A579006AC14F8BCD9004FC66" box="[818,856,976,998]" gridcol="4" gridrow="6" pageId="3" pageNumber="26">109</td>
<td id="76F9F308A579006AC1DE8BCD90A0FC66" box="[931,1020,976,998]" gridcol="5" gridrow="6" pageId="3" pageNumber="26">101–105</td>
<td id="76F9F308A579006AC6468BCD973DFC66" box="[1083,1121,976,998]" gridcol="6" gridrow="6" pageId="3" pageNumber="26">102</td>
<td id="76F9F308A579006AC6DE8BCD97A0FC66" box="[1187,1276,976,998]" gridcol="7" gridrow="6" pageId="3" pageNumber="26">95–100</td>
<td id="76F9F308A579006AC74F8BCD9635FC66" box="[1330,1385,976,998]" gridcol="8" gridrow="6" pageId="3" pageNumber="26">98</td>
</tr>
<tr id="35289A74A579006AC2E28BE49635FB8F" box="[159,1385,1017,1039]" gridrow="7" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28BE49234FB8F" box="[159,360,1017,1039]" gridcol="0" gridrow="7" pageId="3" pageNumber="26">Ant III</th>
<td id="76F9F308A579006AC3DE8BE49154FB8F" box="[419,520,1017,1039]" gridcol="1" gridrow="7" pageId="3" pageNumber="26">21 2–243</td>
<td id="76F9F308A579006AC0468BE49132FB8F" box="[571,622,1017,1039]" gridcol="2" gridrow="7" pageId="3" pageNumber="26">234</td>
<td id="76F9F308A579006AC0DE8BE491A0FB8F" box="[675,764,1017,1039]" gridcol="3" gridrow="7" pageId="3" pageNumber="26">174–200</td>
<td id="76F9F308A579006AC14F8BE49004FB8F" box="[818,856,1017,1039]" gridcol="4" gridrow="7" pageId="3" pageNumber="26">188</td>
<td id="76F9F308A579006AC1DE8BE490A0FB8F" box="[931,1020,1017,1039]" gridcol="5" gridrow="7" pageId="3" pageNumber="26">160–176</td>
<td id="76F9F308A579006AC6468BE4973DFB8F" box="[1083,1121,1017,1039]" gridcol="6" gridrow="7" pageId="3" pageNumber="26">167</td>
<td id="76F9F308A579006AC6DE8BE497A0FB8F" box="[1187,1276,1017,1039]" gridcol="7" gridrow="7" pageId="3" pageNumber="26">136–162</td>
<td id="76F9F308A579006AC74F8BE49635FB8F" box="[1330,1385,1017,1039]" gridcol="8" gridrow="7" pageId="3" pageNumber="26">149</td>
</tr>
<tr id="35289A74A579006AC2E28C3F9635FBB8" box="[159,1385,1058,1080]" gridrow="8" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28C3F9234FBB8" box="[159,360,1058,1080]" gridcol="0" gridrow="8" pageId="3" pageNumber="26">Ant IV</th>
<td id="76F9F308A579006AC3DE8C3F9154FBB8" box="[419,520,1058,1080]" gridcol="1" gridrow="8" pageId="3" pageNumber="26">61 6–650</td>
<td id="76F9F308A579006AC0468C3F9132FBB8" box="[571,622,1058,1080]" gridcol="2" gridrow="8" pageId="3" pageNumber="26">631</td>
<td id="76F9F308A579006AC0DE8C3F91A0FBB8" box="[675,764,1058,1080]" gridcol="3" gridrow="8" pageId="3" pageNumber="26">522–545</td>
<td id="76F9F308A579006AC14F8C3F9004FBB8" box="[818,856,1058,1080]" gridcol="4" gridrow="8" pageId="3" pageNumber="26">530</td>
<td id="76F9F308A579006AC1DE8C3F90A0FBB8" box="[931,1020,1058,1080]" gridcol="5" gridrow="8" pageId="3" pageNumber="26">444–463</td>
<td id="76F9F308A579006AC6468C3F973DFBB8" box="[1083,1121,1058,1080]" gridcol="6" gridrow="8" pageId="3" pageNumber="26">451</td>
<td id="76F9F308A579006AC6DE8C3F97A0FBB8" box="[1187,1276,1058,1080]" gridcol="7" gridrow="8" pageId="3" pageNumber="26">434–445</td>
<td id="76F9F308A579006AC74F8C3F9635FBB8" box="[1330,1385,1058,1080]" gridcol="8" gridrow="8" pageId="3" pageNumber="26">440</td>
</tr>
<tr id="35289A74A579006AC2E28C519635FBE2" box="[159,1385,1100,1122]" gridrow="9" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28C519234FBE2" box="[159,360,1100,1122]" gridcol="0" gridrow="9" pageId="3" pageNumber="26">Ant IV basal subs.</th>
<td id="76F9F308A579006AC3DE8C519154FBE2" box="[419,520,1100,1122]" gridcol="1" gridrow="9" pageId="3" pageNumber="26">220–240</td>
<td id="76F9F308A579006AC0468C519132FBE2" box="[571,622,1100,1122]" gridcol="2" gridrow="9" pageId="3" pageNumber="26">235</td>
<td id="76F9F308A579006AC0DE8C5191A0FBE2" box="[675,764,1100,1122]" gridcol="3" gridrow="9" pageId="3" pageNumber="26">180–208</td>
<td id="76F9F308A579006AC14F8C519004FBE2" box="[818,856,1100,1122]" gridcol="4" gridrow="9" pageId="3" pageNumber="26">196</td>
<td id="76F9F308A579006AC1DE8C5190A0FBE2" box="[931,1020,1100,1122]" gridcol="5" gridrow="9" pageId="3" pageNumber="26">199–210</td>
<td id="76F9F308A579006AC6468C51973DFBE2" box="[1083,1121,1100,1122]" gridcol="6" gridrow="9" pageId="3" pageNumber="26">203</td>
<td id="76F9F308A579006AC6DE8C5197A0FBE2" box="[1187,1276,1100,1122]" gridcol="7" gridrow="9" pageId="3" pageNumber="26">194–203</td>
<td id="76F9F308A579006AC74F8C519635FBE2" box="[1330,1385,1100,1122]" gridcol="8" gridrow="9" pageId="3" pageNumber="26">199</td>
</tr>
<tr id="35289A74A579006AC2E28C689635FB0B" box="[159,1385,1141,1163]" gridrow="10" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28C689234FB0B" box="[159,360,1141,1163]" gridcol="0" gridrow="10" pageId="3" pageNumber="26">Ant III organ rods</th>
<td id="76F9F308A579006AC3DE8C689154FB0B" box="[419,520,1141,1163]" gridcol="1" gridrow="10" pageId="3" pageNumber="26">8.0–10.0</td>
<td id="76F9F308A579006AC0468C689132FB0B" box="[571,622,1141,1163]" gridcol="2" gridrow="10" pageId="3" pageNumber="26">9.3</td>
<td id="76F9F308A579006AC0DE8C6891A0FB0B" box="[675,764,1141,1163]" gridcol="3" gridrow="10" pageId="3" pageNumber="26">9.0–9.5</td>
<td id="76F9F308A579006AC14F8C689004FB0B" box="[818,856,1141,1163]" gridcol="4" gridrow="10" pageId="3" pageNumber="26">9.3</td>
<td id="76F9F308A579006AC1DE8C6890A0FB0B" box="[931,1020,1141,1163]" gridcol="5" gridrow="10" pageId="3" pageNumber="26">4.4–4.5</td>
<td id="76F9F308A579006AC6468C68973DFB0B" box="[1083,1121,1141,1163]" gridcol="6" gridrow="10" pageId="3" pageNumber="26">4.5</td>
<td id="76F9F308A579006AC6DE8C6897A0FB0B" box="[1187,1276,1141,1163]" gridcol="7" gridrow="10" pageId="3" pageNumber="26">3.9–4.3</td>
<td id="76F9F308A579006AC74F8C689635FB0B" box="[1330,1385,1141,1163]" gridcol="8" gridrow="10" pageId="3" pageNumber="26">4.1</td>
</tr>
<tr id="35289A74A579006AC2E28C839635FB34" box="[159,1385,1182,1204]" gridrow="11" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28C839234FB34" box="[159,360,1182,1204]" gridcol="0" gridrow="11" pageId="3" pageNumber="26">Tita I</th>
<td id="76F9F308A579006AC3DE8C839154FB34" box="[419,520,1182,1204]" gridcol="1" gridrow="11" pageId="3" pageNumber="26">280–306</td>
<td id="76F9F308A579006AC0468C839132FB34" box="[571,622,1182,1204]" gridcol="2" gridrow="11" pageId="3" pageNumber="26">292</td>
<td id="76F9F308A579006AC0DE8C8391A0FB34" box="[675,764,1182,1204]" gridcol="3" gridrow="11" pageId="3" pageNumber="26">225–241</td>
<td id="76F9F308A579006AC14F8C839004FB34" box="[818,856,1182,1204]" gridcol="4" gridrow="11" pageId="3" pageNumber="26">232</td>
<td id="76F9F308A579006AC1DE8C8390A0FB34" box="[931,1020,1182,1204]" gridcol="5" gridrow="11" pageId="3" pageNumber="26">244–259</td>
<td id="76F9F308A579006AC6468C83973DFB34" box="[1083,1121,1182,1204]" gridcol="6" gridrow="11" pageId="3" pageNumber="26">250</td>
<td id="76F9F308A579006AC6DE8C8397A0FB34" box="[1187,1276,1182,1204]" gridcol="7" gridrow="11" pageId="3" pageNumber="26">238–238</td>
<td id="76F9F308A579006AC74F8C839635FB34" box="[1330,1385,1182,1204]" gridcol="8" gridrow="11" pageId="3" pageNumber="26">238</td>
</tr>
<tr id="35289A74A579006AC2E28CD59635FB5E" box="[159,1385,1224,1246]" gridrow="12" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28CD59234FB5E" box="[159,360,1224,1246]" gridcol="0" gridrow="12" pageId="3" pageNumber="26">Tita II</th>
<td id="76F9F308A579006AC3DE8CD59154FB5E" box="[419,520,1224,1246]" gridcol="1" gridrow="12" pageId="3" pageNumber="26">280–305</td>
<td id="76F9F308A579006AC0468CD59132FB5E" box="[571,622,1224,1246]" gridcol="2" gridrow="12" pageId="3" pageNumber="26">293</td>
<td id="76F9F308A579006AC0DE8CD591A0FB5E" box="[675,764,1224,1246]" gridcol="3" gridrow="12" pageId="3" pageNumber="26">225–237</td>
<td id="76F9F308A579006AC14F8CD59004FB5E" box="[818,856,1224,1246]" gridcol="4" gridrow="12" pageId="3" pageNumber="26">232</td>
<td id="76F9F308A579006AC1DE8CD590A0FB5E" box="[931,1020,1224,1246]" gridcol="5" gridrow="12" pageId="3" pageNumber="26">240–262</td>
<td id="76F9F308A579006AC6468CD5973DFB5E" box="[1083,1121,1224,1246]" gridcol="6" gridrow="12" pageId="3" pageNumber="26">249</td>
<td id="76F9F308A579006AC6DE8CD597A0FB5E" box="[1187,1276,1224,1246]" gridcol="7" gridrow="12" pageId="3" pageNumber="26">245–245</td>
<td id="76F9F308A579006AC74F8CD59635FB5E" box="[1330,1385,1224,1246]" gridcol="8" gridrow="12" pageId="3" pageNumber="26">245</td>
</tr>
<tr id="35289A74A579006AC2E28CEC9635FA87" box="[159,1385,1265,1287]" gridrow="13" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28CEC9234FA87" box="[159,360,1265,1287]" gridcol="0" gridrow="13" pageId="3" pageNumber="26">Tita III</th>
<td id="76F9F308A579006AC3DE8CEC9154FA87" box="[419,520,1265,1287]" gridcol="1" gridrow="13" pageId="3" pageNumber="26">346–368</td>
<td id="76F9F308A579006AC0468CEC9132FA87" box="[571,622,1265,1287]" gridcol="2" gridrow="13" pageId="3" pageNumber="26">355</td>
<td id="76F9F308A579006AC0DE8CEC91A0FA87" box="[675,764,1265,1287]" gridcol="3" gridrow="13" pageId="3" pageNumber="26">275–295</td>
<td id="76F9F308A579006AC14F8CEC9004FA87" box="[818,856,1265,1287]" gridcol="4" gridrow="13" pageId="3" pageNumber="26">285</td>
<td id="76F9F308A579006AC1DE8CEC90A0FA87" box="[931,1020,1265,1287]" gridcol="5" gridrow="13" pageId="3" pageNumber="26">278–313</td>
<td id="76F9F308A579006AC6468CEC973DFA87" box="[1083,1121,1265,1287]" gridcol="6" gridrow="13" pageId="3" pageNumber="26">296</td>
<td id="76F9F308A579006AC6DE8CEC97A0FA87" box="[1187,1276,1265,1287]" gridcol="7" gridrow="13" pageId="3" pageNumber="26">288–290</td>
<td id="76F9F308A579006AC74F8CEC9635FA87" box="[1330,1385,1265,1287]" gridcol="8" gridrow="13" pageId="3" pageNumber="26">289</td>
</tr>
<tr id="35289A74A579006AC2E28D079635FAB0" box="[159,1385,1306,1328]" gridrow="14" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28D079234FAB0" box="[159,360,1306,1328]" gridcol="0" gridrow="14" pageId="3" pageNumber="26">Claw I</th>
<td id="76F9F308A579006AC3DE8D079154FAB0" box="[419,520,1306,1328]" gridcol="1" gridrow="14" pageId="3" pageNumber="26">49–51</td>
<td id="76F9F308A579006AC0468D079132FAB0" box="[571,622,1306,1328]" gridcol="2" gridrow="14" pageId="3" pageNumber="26">50</td>
<td id="76F9F308A579006AC0DE8D0791A0FAB0" box="[675,764,1306,1328]" gridcol="3" gridrow="14" pageId="3" pageNumber="26">37–42</td>
<td id="76F9F308A579006AC14F8D079004FAB0" box="[818,856,1306,1328]" gridcol="4" gridrow="14" pageId="3" pageNumber="26">40</td>
<td id="76F9F308A579006AC1DE8D0790A0FAB0" box="[931,1020,1306,1328]" gridcol="5" gridrow="14" pageId="3" pageNumber="26">52–55</td>
<td id="76F9F308A579006AC6468D07973DFAB0" box="[1083,1121,1306,1328]" gridcol="6" gridrow="14" pageId="3" pageNumber="26">54</td>
<td id="76F9F308A579006AC6DE8D0797A0FAB0" box="[1187,1276,1306,1328]" gridcol="7" gridrow="14" pageId="3" pageNumber="26">48–50</td>
<td id="76F9F308A579006AC74F8D079635FAB0" box="[1330,1385,1306,1328]" gridcol="8" gridrow="14" pageId="3" pageNumber="26">49</td>
</tr>
<tr id="35289A74A579006AC2E28D599635FADA" box="[159,1385,1348,1370]" gridrow="15" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28D599234FADA" box="[159,360,1348,1370]" gridcol="0" gridrow="15" pageId="3" pageNumber="26">Claw II</th>
<td id="76F9F308A579006AC3DE8D599154FADA" box="[419,520,1348,1370]" gridcol="1" gridrow="15" pageId="3" pageNumber="26">48–49</td>
<td id="76F9F308A579006AC0468D599132FADA" box="[571,622,1348,1370]" gridcol="2" gridrow="15" pageId="3" pageNumber="26">49</td>
<td id="76F9F308A579006AC0DE8D5991A0FADA" box="[675,764,1348,1370]" gridcol="3" gridrow="15" pageId="3" pageNumber="26">36–41</td>
<td id="76F9F308A579006AC14F8D599004FADA" box="[818,856,1348,1370]" gridcol="4" gridrow="15" pageId="3" pageNumber="26">38</td>
<td id="76F9F308A579006AC1DE8D5990A0FADA" box="[931,1020,1348,1370]" gridcol="5" gridrow="15" pageId="3" pageNumber="26">50–55</td>
<td id="76F9F308A579006AC6468D59973DFADA" box="[1083,1121,1348,1370]" gridcol="6" gridrow="15" pageId="3" pageNumber="26">52</td>
<td id="76F9F308A579006AC6DE8D5997A0FADA" box="[1187,1276,1348,1370]" gridcol="7" gridrow="15" pageId="3" pageNumber="26">48–49</td>
<td id="76F9F308A579006AC74F8D599635FADA" box="[1330,1385,1348,1370]" gridcol="8" gridrow="15" pageId="3" pageNumber="26">48</td>
</tr>
<tr id="35289A74A579006AC2E28D709635FA03" box="[159,1385,1389,1411]" gridrow="16" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28D709234FA03" box="[159,360,1389,1411]" gridcol="0" gridrow="16" pageId="3" pageNumber="26">Claw III</th>
<td id="76F9F308A579006AC3DE8D709154FA03" box="[419,520,1389,1411]" gridcol="1" gridrow="16" pageId="3" pageNumber="26">44–46</td>
<td id="76F9F308A579006AC0468D709132FA03" box="[571,622,1389,1411]" gridcol="2" gridrow="16" pageId="3" pageNumber="26">45</td>
<td id="76F9F308A579006AC0DE8D7091A0FA03" box="[675,764,1389,1411]" gridcol="3" gridrow="16" pageId="3" pageNumber="26">36–39</td>
<td id="76F9F308A579006AC14F8D709004FA03" box="[818,856,1389,1411]" gridcol="4" gridrow="16" pageId="3" pageNumber="26">37</td>
<td id="76F9F308A579006AC1DE8D7090A0FA03" box="[931,1020,1389,1411]" gridcol="5" gridrow="16" pageId="3" pageNumber="26">49–54</td>
<td id="76F9F308A579006AC6468D70973DFA03" box="[1083,1121,1389,1411]" gridcol="6" gridrow="16" pageId="3" pageNumber="26">52</td>
<td id="76F9F308A579006AC6DE8D7097A0FA03" box="[1187,1276,1389,1411]" gridcol="7" gridrow="16" pageId="3" pageNumber="26">46–46</td>
<td id="76F9F308A579006AC74F8D709635FA03" box="[1330,1385,1389,1411]" gridcol="8" gridrow="16" pageId="3" pageNumber="26">46</td>
</tr>
<tr id="35289A74A579006AC2E28D8B9635FA2C" box="[159,1385,1430,1452]" gridrow="17" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28D8B9234FA2C" box="[159,360,1430,1452]" gridcol="0" gridrow="17" pageId="3" pageNumber="26">Emp I</th>
<td id="76F9F308A579006AC3DE8D8B9154FA2C" box="[419,520,1430,1452]" gridcol="1" gridrow="17" pageId="3" pageNumber="26">28–31</td>
<td id="76F9F308A579006AC0468D8B9132FA2C" box="[571,622,1430,1452]" gridcol="2" gridrow="17" pageId="3" pageNumber="26">30</td>
<td id="76F9F308A579006AC0DE8D8B91A0FA2C" box="[675,764,1430,1452]" gridcol="3" gridrow="17" pageId="3" pageNumber="26">18–24</td>
<td id="76F9F308A579006AC14F8D8B9004FA2C" box="[818,856,1430,1452]" gridcol="4" gridrow="17" pageId="3" pageNumber="26">22</td>
<td id="76F9F308A579006AC1DE8D8B90A0FA2C" box="[931,1020,1430,1452]" gridcol="5" gridrow="17" pageId="3" pageNumber="26">29–35</td>
<td id="76F9F308A579006AC6468D8B973DFA2C" box="[1083,1121,1430,1452]" gridcol="6" gridrow="17" pageId="3" pageNumber="26">31</td>
<td id="76F9F308A579006AC6DE8D8B97A0FA2C" box="[1187,1276,1430,1452]" gridcol="7" gridrow="17" pageId="3" pageNumber="26">25–31</td>
<td id="76F9F308A579006AC74F8D8B9635FA2C" box="[1330,1385,1430,1452]" gridcol="8" gridrow="17" pageId="3" pageNumber="26">28</td>
</tr>
<tr id="35289A74A579006AC2E28DDD9635FA56" box="[159,1385,1472,1494]" gridrow="18" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28DDD9234FA56" box="[159,360,1472,1494]" gridcol="0" gridrow="18" pageId="3" pageNumber="26">Emp II</th>
<td id="76F9F308A579006AC3DE8DDD9154FA56" box="[419,520,1472,1494]" gridcol="1" gridrow="18" pageId="3" pageNumber="26">20–24</td>
<td id="76F9F308A579006AC0468DDD9132FA56" box="[571,622,1472,1494]" gridcol="2" gridrow="18" pageId="3" pageNumber="26">23</td>
<td id="76F9F308A579006AC0DE8DDD91A0FA56" box="[675,764,1472,1494]" gridcol="3" gridrow="18" pageId="3" pageNumber="26">15–18</td>
<td id="76F9F308A579006AC14F8DDD9004FA56" box="[818,856,1472,1494]" gridcol="4" gridrow="18" pageId="3" pageNumber="26">16</td>
<td id="76F9F308A579006AC1DE8DDD90A0FA56" box="[931,1020,1472,1494]" gridcol="5" gridrow="18" pageId="3" pageNumber="26">27–28</td>
<td id="76F9F308A579006AC6468DDD973DFA56" box="[1083,1121,1472,1494]" gridcol="6" gridrow="18" pageId="3" pageNumber="26">28</td>
<td id="76F9F308A579006AC6DE8DDD97A0FA56" box="[1187,1276,1472,1494]" gridcol="7" gridrow="18" pageId="3" pageNumber="26">25–25</td>
<td id="76F9F308A579006AC74F8DDD9635FA56" box="[1330,1385,1472,1494]" gridcol="8" gridrow="18" pageId="3" pageNumber="26">25</td>
</tr>
<tr id="35289A74A579006AC2E28DF49635FA7F" box="[159,1385,1513,1535]" gridrow="19" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28DF49234FA7F" box="[159,360,1513,1535]" gridcol="0" gridrow="19" pageId="3" pageNumber="26">Emp III</th>
<td id="76F9F308A579006AC3DE8DF49154FA7F" box="[419,520,1513,1535]" gridcol="1" gridrow="19" pageId="3" pageNumber="26">27–29</td>
<td id="76F9F308A579006AC0468DF49132FA7F" box="[571,622,1513,1535]" gridcol="2" gridrow="19" pageId="3" pageNumber="26">28</td>
<td id="76F9F308A579006AC0DE8DF491A0FA7F" box="[675,764,1513,1535]" gridcol="3" gridrow="19" pageId="3" pageNumber="26">19–24</td>
<td id="76F9F308A579006AC14F8DF49004FA7F" box="[818,856,1513,1535]" gridcol="4" gridrow="19" pageId="3" pageNumber="26">22</td>
<td id="76F9F308A579006AC1DE8DF490A0FA7F" box="[931,1020,1513,1535]" gridcol="5" gridrow="19" pageId="3" pageNumber="26">29–30</td>
<td id="76F9F308A579006AC6468DF4973DFA7F" box="[1083,1121,1513,1535]" gridcol="6" gridrow="19" pageId="3" pageNumber="26">29</td>
<td id="76F9F308A579006AC6DE8DF497A0FA7F" box="[1187,1276,1513,1535]" gridcol="7" gridrow="19" pageId="3" pageNumber="26">26–28</td>
<td id="76F9F308A579006AC74F8DF49635FA7F" box="[1330,1385,1513,1535]" gridcol="8" gridrow="19" pageId="3" pageNumber="26">27</td>
</tr>
<tr id="35289A74A579006AC2E28E0F9635F9A8" box="[159,1385,1554,1576]" gridrow="20" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28E0F9234F9A8" box="[159,360,1554,1576]" gridcol="0" gridrow="20" pageId="3" pageNumber="26">Abd dI-1</th>
<td id="76F9F308A579006AC3DE8E0F9154F9A8" box="[419,520,1554,1576]" gridcol="1" gridrow="20" pageId="3" pageNumber="26">90–104</td>
<td id="76F9F308A579006AC0468E0F9132F9A8" box="[571,622,1554,1576]" gridcol="2" gridrow="20" pageId="3" pageNumber="26">97</td>
<td id="76F9F308A579006AC0DE8E0F91A0F9A8" box="[675,764,1554,1576]" gridcol="3" gridrow="20" pageId="3" pageNumber="26">80–84</td>
<td id="76F9F308A579006AC14F8E0F9004F9A8" box="[818,856,1554,1576]" gridcol="4" gridrow="20" pageId="3" pageNumber="26">82</td>
<td id="76F9F308A579006AC1DE8E0F90A0F9A8" box="[931,1020,1554,1576]" gridcol="5" gridrow="20" pageId="3" pageNumber="26">70–85</td>
<td id="76F9F308A579006AC6468E0F973DF9A8" box="[1083,1121,1554,1576]" gridcol="6" gridrow="20" pageId="3" pageNumber="26">76</td>
<td id="76F9F308A579006AC6DE8E0F97A0F9A8" box="[1187,1276,1554,1576]" gridcol="7" gridrow="20" pageId="3" pageNumber="26">81–?</td>
<td id="76F9F308A579006AC74F8E0F9635F9A8" box="[1330,1385,1554,1576]" gridcol="8" gridrow="20" pageId="3" pageNumber="26">81?</td>
</tr>
<tr id="35289A74A579006AC2E28E219635F9D2" box="[159,1385,1596,1618]" gridrow="21" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28E219234F9D2" box="[159,360,1596,1618]" gridcol="0" gridrow="21" pageId="3" pageNumber="26">Abd VI longest seta</th>
<td id="76F9F308A579006AC3DE8E219154F9D2" box="[419,520,1596,1618]" gridcol="1" gridrow="21" pageId="3" pageNumber="26">63–65</td>
<td id="76F9F308A579006AC0468E219132F9D2" box="[571,622,1596,1618]" gridcol="2" gridrow="21" pageId="3" pageNumber="26">64</td>
<td id="76F9F308A579006AC0DE8E2191A0F9D2" box="[675,764,1596,1618]" gridcol="3" gridrow="21" pageId="3" pageNumber="26">41–47</td>
<td id="76F9F308A579006AC14F8E219004F9D2" box="[818,856,1596,1618]" gridcol="4" gridrow="21" pageId="3" pageNumber="26">43</td>
<td id="76F9F308A579006AC1DE8E2190A0F9D2" box="[931,1020,1596,1618]" gridcol="5" gridrow="21" pageId="3" pageNumber="26">62–69</td>
<td id="76F9F308A579006AC6468E21973DF9D2" box="[1083,1121,1596,1618]" gridcol="6" gridrow="21" pageId="3" pageNumber="26">66</td>
<td id="76F9F308A579006AC6DE8E2197A0F9D2" box="[1187,1276,1596,1618]" gridcol="7" gridrow="21" pageId="3" pageNumber="26">40–40</td>
<td id="76F9F308A579006AC74F8E219635F9D2" box="[1330,1385,1596,1618]" gridcol="8" gridrow="21" pageId="3" pageNumber="26">40</td>
</tr>
<tr id="35289A74A579006AC2E28E789635F9FB" box="[159,1385,1637,1659]" gridrow="22" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28E789234F9FB" box="[159,360,1637,1659]" gridcol="0" gridrow="22" pageId="3" pageNumber="26">App an, length</th>
<td id="76F9F308A579006AC3DE8E789154F9FB" box="[419,520,1637,1659]" gridcol="1" gridrow="22" pageId="3" pageNumber="26">26–27</td>
<td id="76F9F308A579006AC0468E789132F9FB" box="[571,622,1637,1659]" gridcol="2" gridrow="22" pageId="3" pageNumber="26">27</td>
<td id="76F9F308A579006AC0DE8E7891A0F9FB" box="[675,764,1637,1659]" gridcol="3" gridrow="22" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC14F8E789004F9FB" box="[818,856,1637,1659]" gridcol="4" gridrow="22" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC1DE8E7890A0F9FB" box="[931,1020,1637,1659]" gridcol="5" gridrow="22" pageId="3" pageNumber="26">28–35</td>
<td id="76F9F308A579006AC6468E78973DF9FB" box="[1083,1121,1637,1659]" gridcol="6" gridrow="22" pageId="3" pageNumber="26">31</td>
<td id="76F9F308A579006AC6DE8E7897A0F9FB" box="[1187,1276,1637,1659]" gridcol="7" gridrow="22" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC74F8E789635F9FB" box="[1330,1385,1637,1659]" gridcol="8" gridrow="22" pageId="3" pageNumber="26">-</td>
</tr>
<tr id="35289A74A579006AC2E28E939635F924" box="[159,1385,1678,1700]" gridrow="23" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28E939234F924" box="[159,360,1678,1700]" gridcol="0" gridrow="23" pageId="3" pageNumber="26">App an, max. width</th>
<td id="76F9F308A579006AC3DE8E939154F924" box="[419,520,1678,1700]" gridcol="1" gridrow="23" pageId="3" pageNumber="26">14–15</td>
<td id="76F9F308A579006AC0468E939132F924" box="[571,622,1678,1700]" gridcol="2" gridrow="23" pageId="3" pageNumber="26">15</td>
<td id="76F9F308A579006AC0DE8E9391A0F924" box="[675,764,1678,1700]" gridcol="3" gridrow="23" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC14F8E939004F924" box="[818,856,1678,1700]" gridcol="4" gridrow="23" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC1DE8E9390A0F924" box="[931,1020,1678,1700]" gridcol="5" gridrow="23" pageId="3" pageNumber="26">15–17</td>
<td id="76F9F308A579006AC6468E93973DF924" box="[1083,1121,1678,1700]" gridcol="6" gridrow="23" pageId="3" pageNumber="26">16</td>
<td id="76F9F308A579006AC6DE8E9397A0F924" box="[1187,1276,1678,1700]" gridcol="7" gridrow="23" pageId="3" pageNumber="26">-</td>
<td id="76F9F308A579006AC74F8E939635F924" box="[1330,1385,1678,1700]" gridcol="8" gridrow="23" pageId="3" pageNumber="26">-</td>
</tr>
<tr id="35289A74A579006AC2E28EA59635F94E" box="[159,1385,1720,1742]" gridrow="24" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28EA59234F94E" box="[159,360,1720,1742]" gridcol="0" gridrow="24" pageId="3" pageNumber="26">Manubrium</th>
<td id="76F9F308A579006AC3DE8EA59154F94E" box="[419,520,1720,1742]" gridcol="1" gridrow="24" pageId="3" pageNumber="26">209–222</td>
<td id="76F9F308A579006AC0468EA59132F94E" box="[571,622,1720,1742]" gridcol="2" gridrow="24" pageId="3" pageNumber="26">214</td>
<td id="76F9F308A579006AC0DE8EA591A0F94E" box="[675,764,1720,1742]" gridcol="3" gridrow="24" pageId="3" pageNumber="26">155–171</td>
<td id="76F9F308A579006AC14F8EA59004F94E" box="[818,856,1720,1742]" gridcol="4" gridrow="24" pageId="3" pageNumber="26">162</td>
<td id="76F9F308A579006AC1DE8EA590A0F94E" box="[931,1020,1720,1742]" gridcol="5" gridrow="24" pageId="3" pageNumber="26">207–226</td>
<td id="76F9F308A579006AC6468EA5973DF94E" box="[1083,1121,1720,1742]" gridcol="6" gridrow="24" pageId="3" pageNumber="26">217</td>
<td id="76F9F308A579006AC6DE8EA597A0F94E" box="[1187,1276,1720,1742]" gridcol="7" gridrow="24" pageId="3" pageNumber="26">194–197</td>
<td id="76F9F308A579006AC74F8EA59635F94E" box="[1330,1385,1720,1742]" gridcol="8" gridrow="24" pageId="3" pageNumber="26">196</td>
</tr>
<tr id="35289A74A579006AC2E28EFC9635F977" box="[159,1385,1761,1783]" gridrow="25" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28EFC9234F977" box="[159,360,1761,1783]" gridcol="0" gridrow="25" pageId="3" pageNumber="26">Dens</th>
<td id="76F9F308A579006AC3DE8EFC9154F977" box="[419,520,1761,1783]" gridcol="1" gridrow="25" pageId="3" pageNumber="26">231–250</td>
<td id="76F9F308A579006AC0468EFC9132F977" box="[571,622,1761,1783]" gridcol="2" gridrow="25" pageId="3" pageNumber="26">242</td>
<td id="76F9F308A579006AC0DE8EFC91A0F977" box="[675,764,1761,1783]" gridcol="3" gridrow="25" pageId="3" pageNumber="26">181–204</td>
<td id="76F9F308A579006AC14F8EFC9004F977" box="[818,856,1761,1783]" gridcol="4" gridrow="25" pageId="3" pageNumber="26">193</td>
<td id="76F9F308A579006AC1DE8EFC90A0F977" box="[931,1020,1761,1783]" gridcol="5" gridrow="25" pageId="3" pageNumber="26">212–236</td>
<td id="76F9F308A579006AC6468EFC973DF977" box="[1083,1121,1761,1783]" gridcol="6" gridrow="25" pageId="3" pageNumber="26">221</td>
<td id="76F9F308A579006AC6DE8EFC97A0F977" box="[1187,1276,1761,1783]" gridcol="7" gridrow="25" pageId="3" pageNumber="26">205–218</td>
<td id="76F9F308A579006AC74F8EFC9635F977" box="[1330,1385,1761,1783]" gridcol="8" gridrow="25" pageId="3" pageNumber="26">212</td>
</tr>
<tr id="35289A74A579006AC2E28F179635F8A0" box="[159,1385,1802,1824]" gridrow="26" pageId="3" pageNumber="26">
<th id="76F9F308A579006AC2E28F179234F8A0" box="[159,360,1802,1824]" gridcol="0" gridrow="26" pageId="3" pageNumber="26">Mucro</th>
<td id="76F9F308A579006AC3DE8F179154F8A0" box="[419,520,1802,1824]" gridcol="1" gridrow="26" pageId="3" pageNumber="26">133–138</td>
<td id="76F9F308A579006AC0468F179132F8A0" box="[571,622,1802,1824]" gridcol="2" gridrow="26" pageId="3" pageNumber="26">136</td>
<td id="76F9F308A579006AC0DE8F1791A0F8A0" box="[675,764,1802,1824]" gridcol="3" gridrow="26" pageId="3" pageNumber="26">110–122</td>
<td id="76F9F308A579006AC14F8F179004F8A0" box="[818,856,1802,1824]" gridcol="4" gridrow="26" pageId="3" pageNumber="26">116</td>
<td id="76F9F308A579006AC1DE8F1790A0F8A0" box="[931,1020,1802,1824]" gridcol="5" gridrow="26" pageId="3" pageNumber="26">130–149</td>
<td id="76F9F308A579006AC6468F17973DF8A0" box="[1083,1121,1802,1824]" gridcol="6" gridrow="26" pageId="3" pageNumber="26">137</td>
<td id="76F9F308A579006AC6DE8F1797A0F8A0" box="[1187,1276,1802,1824]" gridcol="7" gridrow="26" pageId="3" pageNumber="26">128–130</td>
<td id="76F9F308A579006AC74F8F179635F8A0" box="[1330,1385,1802,1824]" gridcol="8" gridrow="26" pageId="3" pageNumber="26">129</td>
</tr>
</table>
</paragraph>
<caption id="DF67C8BEA57EFF91C2EA8FAA9651F86D" httpUri="https://zenodo.org/record/439781/files/figure.png" pageId="4" pageNumber="27" targetBox="[317,1260,226,1922]" targetPageId="4">
<paragraph id="8BA79836A57EFF91C2EA8FAA9651F86D" blockId="4.[151,1436,1975,2029]" pageId="4" pageNumber="27">
<emphasis id="B96C4424A57EFF91C2EA8FAA9263F84D" bold="true" box="[151,319,1975,1998]" pageId="4" pageNumber="27">FIGURES 1–2.</emphasis>
1, type localities of 
<taxonomicName id="4C18E3B5A57EFF91C05B8FA59043F84E" box="[550,799,1976,1998]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A57EFF91C05B8FA59043F84E" box="[550,799,1976,1998]" italics="true" pageId="4" pageNumber="27">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A57EFF91C1578FA590DCF84D" bold="true" box="[810,896,1976,1997]" pageId="4" pageNumber="27">
<taxonomicNameLabel id="A25FF95FA57EFF91C1578FA590DCF84D" box="[810,896,1976,1997]" pageId="4" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B96C4424A57EFF91C1E98FA590FDF84E" box="[916,929,1976,1998]" italics="true" pageId="4" pageNumber="27">1</emphasis>
—Psyrtskha Cave), and 
<taxonomicName id="4C18E3B5A57EFF91C6D88FA4965EF84E" box="[1189,1282,1976,1998]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A57EFF91C6D88FA4965EF84E" box="[1189,1282,1976,1998]" italics="true" pageId="4" pageNumber="27">P. kovali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25FF95FA57EFF91C7718FA5963EF84E" box="[1292,1378,1976,1998]" pageId="4" pageNumber="27" rank="species">
<emphasis id="B96C4424A57EFF91C7718FA59600F84D" bold="true" box="[1292,1372,1976,1997]" pageId="4" pageNumber="27">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<emphasis id="B96C4424A57EFF91C70B8FA596DFF84E" box="[1398,1411,1976,1998]" italics="true" pageId="4" pageNumber="27">2</emphasis>
— Tsebeldinskaya Cave, 
<emphasis id="B96C4424A57EFF91C3048FCA92DAF86D" box="[377,390,2007,2029]" italics="true" pageId="4" pageNumber="27">3</emphasis>
—Tshalintshur Cave, 
<emphasis id="B96C4424A57EFF91C01E8FCA9133F86D" box="[611,623,2007,2029]" italics="true" pageId="4" pageNumber="27">4</emphasis>
—Verkhnyaya Shakuranskaya Cave); 2, habitus of mounted male.
</paragraph>
</caption>
<caption id="DF67C8BEA57FFF90C2EA8FBD90FDF856" httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="5" pageNumber="28" targetBox="[213,1395,193,1895]" targetPageId="5">
<paragraph id="8BA79836A57FFF90C2EA8FBD90FDF856" blockId="5.[151,1436,1952,2006]" pageId="5" pageNumber="28">
<emphasis id="B96C4424A57FFF90C2EA8FBD9266F836" bold="true" box="[151,314,1952,1975]" pageId="5" pageNumber="28">FIGURES 3–7.</emphasis>
<taxonomicName id="4C18E3B5A57FFF90C3428FBC916EF837" box="[319,562,1953,1975]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A57FFF90C3428FBC916EF837" box="[319,562,1953,1975]" italics="true" pageId="5" pageNumber="28">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A57FFF90C0458FBC91D4F836" bold="true" box="[568,648,1953,1974]" pageId="5" pageNumber="28">
<taxonomicNameLabel id="A25FF95FA57FFF90C0458FBC91D4F836" box="[568,648,1953,1974]" pageId="5" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I–III; 7, chaetotaxy of antennal segment IV.
</paragraph>
</caption>
<caption id="DF67C8BEA57CFF93C2EA8FBD97EEF874" httpUri="https://zenodo.org/record/439783/files/figure.png" pageId="6" pageNumber="29" targetBox="[224,1360,222,1893]" targetPageId="6">
<paragraph id="8BA79836A57CFF93C2EA8FBD97EEF874" blockId="6.[151,1436,1952,2036]" pageId="6" pageNumber="29">
<emphasis id="B96C4424A57CFF93C2EA8FBD9214F836" bold="true" box="[151,328,1952,1975]" pageId="6" pageNumber="29">FIGURES 8–19.</emphasis>
<taxonomicName id="4C18E3B5A57CFF93C3328FBC9118F837" box="[335,580,1953,1975]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A57CFF93C3328FBC9118F837" box="[335,580,1953,1975]" italics="true" pageId="6" pageNumber="29">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A57CFF93C0368FBC91C1F836" bold="true" box="[587,669,1953,1974]" pageId="6" pageNumber="29">
<taxonomicNameLabel id="A25FF95FA57CFF93C0368FBC91C1F836" box="[587,669,1953,1974]" pageId="6" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 8, chaetotaxy of foreleg; 9, chaetotaxy of mid leg; 10, chaetotaxy of hind leg; 11–19, shape of setae: 11, spine-like seta of head vertex; 12, m1 of mesothorax; 13, dorsal sea of Abd I; 14, Ie of dens; 15, IIpe of dens; 16, dI-1 of great abdomen; 17, ms1 of Abd VI; 18, mps3 of abd VI; 19, appendices anales.
</paragraph>
</caption>
<caption id="DF67C8BEA57DFF92C2EA8F8F92A3F866" httpUri="https://zenodo.org/record/439784/files/figure.png" pageId="7" pageNumber="30" targetBox="[197,1397,193,1879]" targetPageId="7">
<paragraph id="8BA79836A57DFF92C2EA8F8F92A3F866" blockId="7.[151,1436,1938,2022]" pageId="7" pageNumber="30">
<emphasis id="B96C4424A57DFF92C2EA8F8F9208F828" bold="true" box="[151,340,1938,1961]" pageId="7" pageNumber="30">FIGURES 20–24.</emphasis>
<taxonomicName id="4C18E3B5A57DFF92C3248F8E9110F829" box="[345,588,1939,1961]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="30" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A57DFF92C3248F8E9110F829" box="[345,588,1939,1961]" italics="true" pageId="7" pageNumber="30">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A57DFF92C02E8F8E91FFF828" bold="true" box="[595,675,1939,1960]" pageId="7" pageNumber="30">
<taxonomicNameLabel id="A25FF95FA57DFF92C02E8F8E91FFF828" box="[595,675,1939,1960]" pageId="7" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 20, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 21, chaetotaxy of furca; 22, tenaculum; 23, chaetotaxy of female Abd VI; 24, chaetotaxy of male Abd VI.
</paragraph>
</caption>
<caption id="DF67C8BEA572FF9DC2EA8FBC9222F874" httpUri="https://zenodo.org/record/439785/files/figure.png" pageId="8" pageNumber="31" targetBox="[184,1402,213,1899]" targetPageId="8">
<paragraph id="8BA79836A572FF9DC2EA8FBC9222F874" blockId="8.[151,1436,1953,2036]" pageId="8" pageNumber="31">
<emphasis id="B96C4424A572FF9DC2EA8FBC9208F836" bold="true" box="[151,340,1953,1976]" pageId="8" pageNumber="31">FIGURES 25–30.</emphasis>
<taxonomicName id="4C18E3B5A572FF9DC3278FBC9112F837" box="[346,590,1953,1975]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="31" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A572FF9DC3278FBC9112F837" box="[346,590,1953,1975]" italics="true" pageId="8" pageNumber="31">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A572FF9DC0298FBF91F9F837" bold="true" box="[596,677,1954,1975]" pageId="8" pageNumber="31">
<taxonomicNameLabel id="A25FF95FA572FF9DC0298FBF91F9F837" box="[596,677,1954,1975]" pageId="8" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 25, habitus of alcohol specimen; 26, chaetotaxy of antennal III organ; 27, foot complex (a—fore, b—mid, c—hind); 28, fragment of head dorsum; 29, chaetotaxy of trichobothrial complex; 30, mucro and distal part of dens.
</paragraph>
</caption>
<caption id="DF67C8BEA573FF9CC2EA8DA9918DFA6A" httpUri="https://zenodo.org/record/439786/files/figure.png" pageId="9" pageNumber="32" targetBox="[260,1350,222,1407]" targetPageId="9">
<paragraph id="8BA79836A573FF9CC2EA8DA9918DFA6A" blockId="9.[151,1435,1460,1514]" pageId="9" pageNumber="32">
<emphasis id="B96C4424A573FF9CC2EA8DA9920FFA4A" bold="true" box="[151,339,1460,1483]" pageId="9" pageNumber="32">FIGURES 31–33.</emphasis>
<taxonomicName id="4C18E3B5A573FF9CC3248DA89110FA4B" box="[345,588,1461,1483]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A573FF9CC3248DA89110FA4B" box="[345,588,1461,1483]" italics="true" pageId="9" pageNumber="32">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A573FF9CC02F8DA891FEFA4A" bold="true" box="[594,674,1461,1482]" pageId="9" pageNumber="32">
<taxonomicNameLabel id="A25FF95FA573FF9CC02F8DA891FEFA4A" box="[594,674,1461,1482]" pageId="9" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 31, Ant IV of male; 32, small abdomen of male, lateral view; 33, fragment of female Abd VI with circumanal setae, posterior view.
</paragraph>
</caption>
<paragraph id="8BA79836A573FF9CC2BA8E0B91EFF8AB" blockId="9.[151,1437,1558,2015]" pageId="9" pageNumber="32">
<emphasis id="B96C4424A573FF9CC2BA8E0B915FF9AF" bold="true" box="[199,515,1558,1583]" pageId="9" pageNumber="32">Bionomy and distribution.</emphasis>
<taxonomicName id="4C18E3B5A573FF9CC0778E0A913DF9AF" box="[522,609,1558,1583]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A573FF9CC0778E0A913DF9AF" box="[522,609,1558,1583]" italics="true" pageId="9" pageNumber="32">P. dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A573FF9CC0178E0B9198F9AF" bold="true" box="[618,708,1558,1583]" pageId="9" pageNumber="32">
<taxonomicNameLabel id="A25FF95FA573FF9CC0178E0B9198F9AF" box="[618,708,1558,1583]" pageId="9" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lives in the distant aphotic zone of the cave with air temperature 13.0°C and water temperature 12.6°C. Not numerous specimens were collected on the water surface of the siphon and nearby in 
<date id="FFA6BEF6A573FF9CC3488E439285F9F7" box="[309,473,1630,1655]" pageId="9" pageNumber="32" value="2006" valueMax="2011">2006 and 2011</date>
but were missing in 
<date id="FFA6BEF6A573FF9CC0BD8E439025F9F7" box="[704,889,1630,1655]" pageId="9" pageNumber="32" value="2015-09">September 2015</date>
. They occur together with two 
<taxonomicName id="4C18E3B5A573FF9CC6AF8E4396C0F9F7" box="[1234,1436,1630,1655]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B96C4424A573FF9CC6AF8E4396C0F9F7" box="[1234,1436,1630,1655]" italics="true" pageId="9" pageNumber="32">Pygmarrhopalites</emphasis>
</taxonomicName>
species from 
<taxonomicName id="4C18E3B5A573FF9CC3538E9992C2F91B" box="[302,414,1668,1691]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B96C4424A573FF9CC3538E9992C2F91B" box="[302,414,1668,1691]" italics="true" pageId="9" pageNumber="32">pygmaeus</emphasis>
</taxonomicName>
-group and 
<taxonomicName id="4C18E3B5A573FF9CC05C8E9F906EF91B" box="[545,818,1666,1691]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B96C4424A573FF9CC05C8E9F906EF91B" box="[545,818,1666,1691]" italics="true" pageId="9" pageNumber="32">Arrhopalites abchasicus</emphasis>
</taxonomicName>
. Thus, relatively small Psyrtskha Cave is inhabited by four sympatric species of the family. Unlike 
<taxonomicName id="4C18E3B5A573FF9CC0F58EBA907FF93F" box="[648,803,1702,1727]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B96C4424A573FF9CC0F58EBA907FF93F" box="[648,803,1702,1727]" italics="true" pageId="9" pageNumber="32">A. abchasicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C18E3B5A573FF9CC14C8EBA90D5F93F" box="[817,905,1702,1727]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="dbari">
<emphasis id="B96C4424A573FF9CC14C8EBA90D5F93F" box="[817,905,1702,1727]" italics="true" pageId="9" pageNumber="32">P. dbari</emphasis>
</taxonomicName>
was not found in neighbour and hydrologically connected huge Novoafonskaya Cave. Due to its morphology (much elongated antennae, well-developed antennal organ, thin claws with short empodia, poor or absent pigmentation) and topical preferences, this species can be classified as a highly troglomorphic troglobiont.
</paragraph>
<paragraph id="8BA79836A573FF9FC2BA8F2B918BFEC0" blockId="9.[151,1437,1558,2015]" lastBlockId="10.[151,1437,151,680]" lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32">
<emphasis id="B96C4424A573FF9CC2BA8F2B92E1F8CF" bold="true" box="[199,445,1846,1871]" pageId="9" pageNumber="32">Conservation status.</emphasis>
The new species demonstrates extremely restricted distribution and is confined to the only cave: no findings in other nearby caves of Novy Afon as well as in numerous W Caucasian caves of other karst massifs and also in epigean environment were registered. Troglomorphic appearance confirms that the species is ecologically specialized cave/subterranean inhabitant. Number of individuals of this species in Psyrtskha Cave, located in active touristic zone, is visually much lower than of cohabiting 
<taxonomicName id="4C18E3B5A573FF9CC6558FDB9665F85F" box="[1064,1337,1990,2015]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="abchasicus">
<emphasis id="B96C4424A573FF9CC6558FDB9665F85F" box="[1064,1337,1990,2015]" italics="true" pageId="9" pageNumber="32">Arrhopalites abchasicus</emphasis>
</taxonomicName>
and two other representatives of 
<taxonomicName id="4C18E3B5A570FF9FC3D6888A910EFF30" box="[427,594,151,176]" class="Entognatha" family="Arrhopalitidae" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="family">Arrhopalitidae</taxonomicName>
. According to 
<bibRefCitation id="EF89E5C7A570FF9FC081888590C9FF30" author="IUCN" box="[764,917,152,176]" pageId="10" pageNumber="33" refString="IUCN. (2012) IUCN Red List Categories and Criteria: Version 3.1. 2 nd Edition. IUCN, Gland &amp; Cambridge, iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
classification 
<taxonomicName id="4C18E3B5A570FF9FC647888497C9FF30" box="[1082,1173,151,176]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC647888497C9FF30" box="[1082,1173,151,176]" italics="true" pageId="10" pageNumber="33">P. dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC6E0888A97A5FF30" bold="true" box="[1181,1273,151,176]" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC6E0888A97A5FF30" box="[1181,1273,151,176]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most credible corresponds to the category 
<emphasis id="B96C4424A570FF9FC3A188A19103FF55" bold="true" box="[476,607,188,213]" pageId="10" pageNumber="33">Vulnerable</emphasis>
and criterion 
<emphasis id="B96C4424A570FF9FC17F88A19079FF55" bold="true" box="[770,805,188,213]" pageId="10" pageNumber="33">D2</emphasis>
: “Restricted area of occupancy or number of locations with a plausible future threat that could drive the taxon to Critically Endangered or even Extinct in a very short time”. Therefore, 
<taxonomicName id="4C18E3B5A570FF9FC3178918929BFE9D" box="[362,455,260,285]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC3178918929BFE9D" box="[362,455,260,285]" italics="true" pageId="10" pageNumber="33">P. dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC3AE8919916EFE9D" bold="true" box="[467,562,260,285]" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC3AE8919916EFE9D" box="[467,562,260,285]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
should be included into IUCN Red List of Threatened Species and meet adequate protection of its habitat—Psyrtskha Cave.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA570FF9FC2BA8951908CFE08" pageId="10" pageNumber="33" type="etymology">
<paragraph id="8BA79836A570FF9FC2BA8951908CFE08" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="33">
<emphasis id="B96C4424A570FF9FC2BA89519212FEE5" bold="true" box="[199,334,332,357]" pageId="10" pageNumber="33">Etymology.</emphasis>
The new species is dedicated to Dr. Roman Dbar (Sukhum) for his great contribution to study and protection of Abkhazian nature, including amazing Novoafonskaya Cave.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA570FF9FC2BA89899772FD28" pageId="10" pageNumber="33" type="discussion">
<paragraph id="8BA79836A570FF9FC2BA89899772FD28" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="33">
<emphasis id="B96C4424A570FF9FC2BA89899267FE2D" bold="true" box="[199,315,404,429]" pageId="10" pageNumber="33">Remarks.</emphasis>
Among about 30 described species of the 
<taxonomicName id="4C18E3B5A570FF9FC13F898990E7FE2D" box="[834,955,404,429]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A570FF9FC13F898990E7FE2D" box="[834,955,404,429]" italics="true" pageId="10" pageNumber="33">principalis</emphasis>
</taxonomicName>
group with palmated appendices anales, Nearctic 
<taxonomicName id="4C18E3B5A570FF9FC37D89A49187FE50" authority="Zeppelini &amp; Christiansen, 2003" authorityName="Zeppelini &amp; Christiansen" authorityYear="2003" box="[256,731,439,464]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="arca">
<emphasis id="B96C4424A570FF9FC37D89A49211FE50" box="[256,333,441,464]" italics="true" pageId="10" pageNumber="33">P. arca</emphasis>
(
<bibRefCitation id="EF89E5C7A570FF9FC31F89AA918FFE50" author="Zeppelini" box="[354,723,439,464]" pageId="10" pageNumber="33" refString="Zeppelini, D. &amp; Christiansen, K. (2003) Arrhopalites (Collembola: Arrhopalitidae) in U. S. caves with the description of seven new species. Journal of cave and karst studies, 65 (1), 36 - 42." type="journal article" year="2003">Zeppelini &amp; Christiansen, 2003</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C18E3B5A570FF9FC09789A49735FE50" authority="Christiansen, 1966" authorityName="Christiansen" authorityYear="1966" box="[746,1129,439,464]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="bellingeri">
<emphasis id="B96C4424A570FF9FC09789A49029FE50" box="[746,885,439,464]" italics="true" pageId="10" pageNumber="33">P. bellingeri</emphasis>
(
<bibRefCitation id="EF89E5C7A570FF9FC1FA89AA973EFE50" author="Christiansen" box="[903,1122,439,464]" pageId="10" pageNumber="33" refString="Christiansen, K. (1966) The genus Arrhopalites (Collembola, Sminthuridae) in the United States and Canada. International Journal of Speleology, 2, 43 - 73." type="journal article" year="1966">Christiansen, 1966</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C18E3B5A570FF9FC60489A49227FE74" authority="Zeppelini &amp; Christiansen, 2003" authorityName="Zeppelini &amp; Christiansen" authorityYear="2003" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="hubbardi">
<emphasis id="B96C4424A570FF9FC60489A497A0FE50" box="[1145,1276,439,464]" italics="true" pageId="10" pageNumber="33">P. hubbardi</emphasis>
(
<bibRefCitation id="EF89E5C7A570FF9FC77389AA9228FE74" author="Zeppelini" pageId="10" pageNumber="33" refString="Zeppelini, D. &amp; Christiansen, K. (2003) Arrhopalites (Collembola: Arrhopalitidae) in U. S. caves with the description of seven new species. Journal of cave and karst studies, 65 (1), 36 - 42." type="journal article" year="2003">Zeppelini &amp; Christiansen, 2003</bibRefCitation>
)
</taxonomicName>
and Japanese 
<taxonomicName id="4C18E3B5A570FF9FC04B89C090D3FE74" authority="Yosii, 1970" authorityName="Yosii" authorityYear="1970" box="[566,911,476,501]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="octacanthus">
<emphasis id="B96C4424A570FF9FC04B89C091BEFE75" box="[566,738,476,501]" italics="true" pageId="10" pageNumber="33">P. octacanthus</emphasis>
(
<bibRefCitation id="EF89E5C7A570FF9FC08489C190D4FE74" author="Yosii" box="[761,904,476,501]" pageId="10" pageNumber="33" refString="Yosii, R. (1970) On some Collembola of Japan and adjacent countries II. Contributions from the Biological Laboratory, Kyoto University, 23, 1 - 32." type="journal article" year="1970">Yosii, 1970</bibRefCitation>
)
</taxonomicName>
, as well as new species, normally bear 7 subsegments on Ant IV. Among them and within 
<taxonomicName id="4C18E3B5A570FF9FC0BC89E29066FD98" box="[705,826,511,536]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A570FF9FC0BC89E29066FD98" box="[705,826,511,536]" italics="true" pageId="10" pageNumber="33">principalis</emphasis>
</taxonomicName>
group only 
<taxonomicName id="4C18E3B5A570FF9FC1BE8A1C9751FD98" box="[963,1037,513,536]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="arca">
<emphasis id="B96C4424A570FF9FC1BE8A1C9751FD98" box="[963,1037,513,536]" italics="true" pageId="10" pageNumber="33">P. arca</emphasis>
</taxonomicName>
shares with the new species muchelongated antennae, 3× of head. Differential characters for species with 7 subsegments on Ant IV are given in 
<tableCitation id="C69AAD8DA570FF9FC2EA8A5A93B1FDE0" box="[151,237,583,608]" captionStart="TABLE 2" captionStartId="10.[151,239,725,748]" captionTargetBox="[151,1376,808,1407]" captionTargetPageId="10" captionText="TABLE 2. Differences between Pygmarrhopalites dbari sp. nov. and congeners of principalis group with 7 subsegments on Ant IV." httpUri="http://table.plazi.org/id/DF67C8BEA570FF9FC2EA8AC8925BFC8C" pageId="10" pageNumber="33" tableUuid="DF67C8BEA570FF9FC2EA8AC8925BFC8C">Table 2</tableCitation>
. Additionally, it is notable that the number of setae on femur II (13 setae) and III (12 setae) in 
<taxonomicName id="4C18E3B5A570FF9FC7678A54962FFDE0" box="[1306,1395,583,608]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC7678A54962FFDE0" box="[1306,1395,583,608]" italics="true" pageId="10" pageNumber="33">P. dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC7078A5A9394FD05" bold="true" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC7078A5A9394FD05" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is less than in other representatives of the 
<taxonomicName id="4C18E3B5A570FF9FC0B98A719061FD05" box="[708,829,620,645]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A570FF9FC0B98A719061FD05" box="[708,829,620,645]" italics="true" pageId="10" pageNumber="33">principalis</emphasis>
</taxonomicName>
group for which this character is known to date (
<taxonomicName id="4C18E3B5A570FF9FC7FA8A70923CFD28" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="pseudoprincipalis">
<emphasis id="B96C4424A570FF9FC7FA8A70923CFD28" italics="true" pageId="10" pageNumber="33">P. pseudoprincipalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C18E3B5A570FF9FC3108A8C913EFD28" box="[365,610,655,680]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="principalis" subSpecies="skelicus">
<emphasis id="B96C4424A570FF9FC3108A8C913EFD28" box="[365,610,655,680]" italics="true" pageId="10" pageNumber="33">P. principalis skelicus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C18E3B5A570FF9FC0E28A8C91A3FD28" box="[671,767,655,680]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC0E28A8C91A3FD28" box="[671,767,655,680]" italics="true" pageId="10" pageNumber="33">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC17B8A92903CFD28" bold="true" box="[774,864,655,680]" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC17B8A92903CFD28" box="[774,864,655,680]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
described below).
</paragraph>
</subSubSection>
<caption id="DF67C8BEA570FF9FC2EA8AC8925BFC8C" ID-Table-UUID="DF67C8BEA570FF9FC2EA8AC8925BFC8C" httpUri="http://table.plazi.org/id/DF67C8BEA570FF9FC2EA8AC8925BFC8C" pageId="10" pageNumber="33" targetBox="[151,1376,808,1407]" targetIsTable="true" targetPageId="10">
<paragraph id="8BA79836A570FF9FC2EA8AC8925BFC8C" blockId="10.[151,1435,724,780]" pageId="10" pageNumber="33">
<emphasis id="B96C4424A570FF9FC2EA8AC89255FD6C" bold="true" box="[151,265,725,748]" pageId="10" pageNumber="33">TABLE 2.</emphasis>
Differences between 
<taxonomicName id="4C18E3B5A570FF9FC38D8AC991ADFD6C" box="[496,753,724,748]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC38D8AC991ADFD6C" box="[496,753,724,748]" italics="true" pageId="10" pageNumber="33">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC08B8AC89010FD6C" bold="true" box="[758,844,725,748]" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC08B8AC89010FD6C" box="[758,844,725,748]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and congeners of 
<taxonomicName id="4C18E3B5A570FF9FC6718AC99723FD6C" box="[1036,1151,724,748]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A570FF9FC6718AC99723FD6C" box="[1036,1151,724,748]" italics="true" pageId="10" pageNumber="33">principalis</emphasis>
</taxonomicName>
group with 7 subsegments on Ant IV.
</paragraph>
</caption>
<paragraph id="8BA79836A570FF9FC2E28B35917FFADE" pageId="10" pageNumber="33">
<table id="F9186A96A570006AC2EA8B35963CFAFF" box="[151,1376,808,1407]" gridcols="6" gridrows="12" pageId="10" pageNumber="33">
<tr id="35289A74A570006AC2EA8B35963CFCBE" box="[151,1376,808,830]" gridrow="0" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8B3592E2FCBE" box="[151,446,808,830]" gridcol="0" gridrow="0" pageId="10" pageNumber="33">Character / Species</th>
<th id="76F9F308A570006AC3B88B359130FCBE" box="[453,620,808,830]" gridcol="1" gridrow="0" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC3A38B359149FCBE" box="[478,533,808,830]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A570FF9FC3A38B359149FCBE" box="[478,533,808,830]" italics="true" pageId="10" pageNumber="33">dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A570FF9FC0668B359130FCBD" bold="true" box="[539,620,808,829]" pageId="10" pageNumber="33">
<taxonomicNameLabel id="A25FF95FA570FF9FC0668B359130FCBD" box="[539,620,808,829]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76F9F308A570006AC0E68B359181FCBE" box="[667,733,808,830]" gridcol="2" gridrow="0" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC0E68B349194FCBE" box="[667,712,809,830]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="arca">
<emphasis id="B96C4424A570FF9FC0E68B349194FCBE" box="[667,712,809,830]" italics="true" pageId="10" pageNumber="33">arca</emphasis>
</taxonomicName>
</th>
<th id="76F9F308A570006AC1288B3590E2FCBE" box="[853,958,808,830]" gridcol="3" gridrow="0" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC1288B3590EBFCBE" box="[853,951,808,830]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="bellingeri">
<emphasis id="B96C4424A570FF9FC1288B3590EBFCBE" box="[853,951,808,830]" italics="true" pageId="10" pageNumber="33">bellingeri</emphasis>
</taxonomicName>
</th>
<th id="76F9F308A570006AC6728B3597CFFCBE" box="[1039,1171,808,830]" gridcol="4" gridrow="0" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC6728B359737FCBE" box="[1039,1131,808,830]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="hubbardi">
<emphasis id="B96C4424A570FF9FC6728B359737FCBE" box="[1039,1131,808,830]" italics="true" pageId="10" pageNumber="33">hubbardi</emphasis>
</taxonomicName>
</th>
<th id="76F9F308A570006AC6A78B35963CFCBE" box="[1242,1376,808,830]" gridcol="5" gridrow="0" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC6A78B359608FCBE" box="[1242,1364,808,830]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="octacanthus">
<emphasis id="B96C4424A570FF9FC6A78B359608FCBE" box="[1242,1364,808,830]" italics="true" pageId="10" pageNumber="33">octacanthus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35289A74A570006AC2EA8B4C963CFCE7" box="[151,1376,849,871]" gridrow="1" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8B4C92E2FCE7" box="[151,446,849,871]" gridcol="0" gridrow="1" pageId="10" pageNumber="33">Ant/head</th>
<td id="76F9F308A570006AC3B88B4C9130FCE7" box="[453,620,849,871]" gridcol="1" gridrow="1" pageId="10" pageNumber="33">3</td>
<td id="76F9F308A570006AC0E68B4C9181FCE7" box="[667,733,849,871]" gridcol="2" gridrow="1" pageId="10" pageNumber="33">2.7-3</td>
<td id="76F9F308A570006AC1288B4C90E2FCE7" box="[853,958,849,871]" gridcol="3" gridrow="1" pageId="10" pageNumber="33">?</td>
<td id="76F9F308A570006AC6728B4C97CFFCE7" box="[1039,1171,849,871]" gridcol="4" gridrow="1" pageId="10" pageNumber="33">2</td>
<td id="76F9F308A570006AC6A78B4C963CFCE7" box="[1242,1376,849,871]" gridcol="5" gridrow="1" pageId="10" pageNumber="33">1.7</td>
</tr>
<tr id="35289A74A570006AC2EA8B67963CFC10" box="[151,1376,890,912]" gridrow="2" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8B6792E2FC10" box="[151,446,890,912]" gridcol="0" gridrow="2" pageId="10" pageNumber="33">Axial setae of head dorsum</th>
<td id="76F9F308A570006AC3B88B679130FC10" box="[453,620,890,912]" gridcol="1" gridrow="2" pageId="10" pageNumber="33">4</td>
<td id="76F9F308A570006AC0E68B679181FC10" box="[667,733,890,912]" gridcol="2" gridrow="2" pageId="10" pageNumber="33">4</td>
<td id="76F9F308A570006AC1288B6790E2FC10" box="[853,958,890,912]" gridcol="3" gridrow="2" pageId="10" pageNumber="33">4</td>
<td id="76F9F308A570006AC6728B6797CFFC10" box="[1039,1171,890,912]" gridcol="4" gridrow="2" pageId="10" pageNumber="33">5</td>
<td id="76F9F308A570006AC6A78B67963CFC10" box="[1242,1376,890,912]" gridcol="5" gridrow="2" pageId="10" pageNumber="33">5</td>
</tr>
<tr id="35289A74A570006AC2EA8BB9963CFC3A" box="[151,1376,932,954]" gridrow="3" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8BB992E2FC3A" box="[151,446,932,954]" gridcol="0" gridrow="3" pageId="10" pageNumber="33">Seta ms1 of Abd VI</th>
<td id="76F9F308A570006AC3B88BB99130FC3A" box="[453,620,932,954]" gridcol="1" gridrow="3" pageId="10" pageNumber="33">not forked</td>
<td id="76F9F308A570006AC0E68BB99181FC3A" box="[667,733,932,954]" gridcol="2" gridrow="3" pageId="10" pageNumber="33">forked</td>
<td id="76F9F308A570006AC1288BB990E2FC3A" box="[853,958,932,954]" gridcol="3" gridrow="3" pageId="10" pageNumber="33">not forked</td>
<td id="76F9F308A570006AC6728BB997CFFC3A" box="[1039,1171,932,954]" gridcol="4" gridrow="3" pageId="10" pageNumber="33">forked</td>
<td id="76F9F308A570006AC6A78BB9963CFC3A" box="[1242,1376,932,954]" gridcol="5" gridrow="3" pageId="10" pageNumber="33">not forked</td>
</tr>
<tr id="35289A74A570006AC2EA8BD0963CFC63" box="[151,1376,973,995]" gridrow="4" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8BD092E2FC63" box="[151,446,973,995]" gridcol="0" gridrow="4" pageId="10" pageNumber="33">Claw tooth I, II, III</th>
<td id="76F9F308A570006AC3B88BD09130FC63" box="[453,620,973,995]" gridcol="1" gridrow="4" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC0E68BD09181FC63" box="[667,733,973,995]" gridcol="2" gridrow="4" pageId="10" pageNumber="33">— + +</td>
<td id="76F9F308A570006AC1288BD090E2FC63" box="[853,958,973,995]" gridcol="3" gridrow="4" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC6728BD097CFFC63" box="[1039,1171,973,995]" gridcol="4" gridrow="4" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC6A78BD0963CFC63" box="[1242,1376,973,995]" gridcol="5" gridrow="4" pageId="10" pageNumber="33">— + +</td>
</tr>
<tr id="35289A74A570006AC2EA8BEB963CFB8C" box="[151,1376,1014,1036]" gridrow="5" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8BEB92E2FB8C" box="[151,446,1014,1036]" gridcol="0" gridrow="5" pageId="10" pageNumber="33">Emp tooth I, II, III</th>
<td id="76F9F308A570006AC3B88BEB9130FB8C" box="[453,620,1014,1036]" gridcol="1" gridrow="5" pageId="10" pageNumber="33">+ + —</td>
<td id="76F9F308A570006AC0E68BEB9181FB8C" box="[667,733,1014,1036]" gridcol="2" gridrow="5" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC1288BEB90E2FB8C" box="[853,958,1014,1036]" gridcol="3" gridrow="5" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC6728BEB97CFFB8C" box="[1039,1171,1014,1036]" gridcol="4" gridrow="5" pageId="10" pageNumber="33">+ + +</td>
<td id="76F9F308A570006AC6A78BEB963CFB8C" box="[1242,1376,1014,1036]" gridcol="5" gridrow="5" pageId="10" pageNumber="33">— — —</td>
</tr>
<tr id="35289A74A570006AC2EA8C3D963CFBB6" box="[151,1376,1056,1078]" gridrow="6" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8C3D92E2FBB6" box="[151,446,1056,1078]" gridcol="0" gridrow="6" pageId="10" pageNumber="33">Emp tip: claw tip I, II, III</th>
<td id="76F9F308A570006AC3B88C3D9130FBB6" box="[453,620,1056,1078]" gridcol="1" gridrow="6" pageId="10" pageNumber="33">&lt;&lt;&lt;</td>
<td id="76F9F308A570006AC0E68C3D9181FBB6" box="[667,733,1056,1078]" gridcol="2" gridrow="6" pageId="10" pageNumber="33">&gt;&gt; =</td>
<td id="76F9F308A570006AC1288C3D90E2FBB6" box="[853,958,1056,1078]" gridcol="3" gridrow="6" pageId="10" pageNumber="33">&gt; = =</td>
<td id="76F9F308A570006AC6728C3D97CFFBB6" box="[1039,1171,1056,1078]" gridcol="4" gridrow="6" pageId="10" pageNumber="33">&gt; = =</td>
<td id="76F9F308A570006AC6A78C3D963CFBB6" box="[1242,1376,1056,1078]" gridcol="5" gridrow="6" pageId="10" pageNumber="33">&gt;&gt;&gt; (&gt; = =)</td>
</tr>
<tr id="35289A74A570006AC2EA8C54963CFBDF" box="[151,1376,1097,1119]" gridrow="7" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8C5492E2FBDF" box="[151,446,1097,1119]" gridcol="0" gridrow="7" pageId="10" pageNumber="33">Basal papilla on Ant III</th>
<td id="76F9F308A570006AC3B88C549130FBDF" box="[453,620,1097,1119]" gridcol="1" gridrow="7" pageId="10" pageNumber="33">—</td>
<td id="76F9F308A570006AC0E68C549181FBDF" box="[667,733,1097,1119]" gridcol="2" gridrow="7" pageId="10" pageNumber="33">—</td>
<td id="76F9F308A570006AC1288C5490E2FBDF" box="[853,958,1097,1119]" gridcol="3" gridrow="7" pageId="10" pageNumber="33">—</td>
<td id="76F9F308A570006AC6728C5497CFFBDF" box="[1039,1171,1097,1119]" gridcol="4" gridrow="7" pageId="10" pageNumber="33">+</td>
<td id="76F9F308A570006AC6A78C54963CFBDF" box="[1242,1376,1097,1119]" gridcol="5" gridrow="7" pageId="10" pageNumber="33">+</td>
</tr>
<tr id="35289A74A570006AC2EA8C6F963CFB08" box="[151,1376,1138,1160]" gridrow="8" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8C6F92E2FB08" box="[151,446,1138,1160]" gridcol="0" gridrow="8" pageId="10" pageNumber="33">eyes per side</th>
<td id="76F9F308A570006AC3B88C6F9130FB08" box="[453,620,1138,1160]" gridcol="1" gridrow="8" pageId="10" pageNumber="33">1</td>
<td id="76F9F308A570006AC0E68C6F9181FB08" box="[667,733,1138,1160]" gridcol="2" gridrow="8" pageId="10" pageNumber="33">1</td>
<td id="76F9F308A570006AC1288C6F90E2FB08" box="[853,958,1138,1160]" gridcol="3" gridrow="8" pageId="10" pageNumber="33">2</td>
<td id="76F9F308A570006AC6728C6F97CFFB08" box="[1039,1171,1138,1160]" gridcol="4" gridrow="8" pageId="10" pageNumber="33">1+1 vestigial</td>
<td id="76F9F308A570006AC6A78C6F963CFB08" box="[1242,1376,1138,1160]" gridcol="5" gridrow="8" pageId="10" pageNumber="33">1</td>
</tr>
<tr id="35289A74A570006AC2EA8C81963CFB32" box="[151,1376,1180,1202]" gridrow="9" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8C8192E2FB32" box="[151,446,1180,1202]" gridcol="0" gridrow="9" pageId="10" pageNumber="33">spiny setae on dens</th>
<td id="76F9F308A570006AC3B88C819130FB32" box="[453,620,1180,1202]" gridcol="1" gridrow="9" pageId="10" pageNumber="33">3</td>
<td id="76F9F308A570006AC0E68C819181FB32" box="[667,733,1180,1202]" gridcol="2" gridrow="9" pageId="10" pageNumber="33">5</td>
<td id="76F9F308A570006AC1288C8190E2FB32" box="[853,958,1180,1202]" gridcol="3" gridrow="9" pageId="10" pageNumber="33">5</td>
<td id="76F9F308A570006AC6728C8197CFFB32" box="[1039,1171,1180,1202]" gridcol="4" gridrow="9" pageId="10" pageNumber="33">5</td>
<td id="76F9F308A570006AC6A78C81963CFB32" box="[1242,1376,1180,1202]" gridcol="5" gridrow="9" pageId="10" pageNumber="33">6</td>
</tr>
<tr id="35289A74A570006AC2EA8CD8963CFB5B" box="[151,1376,1221,1243]" gridrow="10" pageId="10" pageNumber="33">
<th id="76F9F308A570006AC2EA8CD892E2FB5B" box="[151,446,1221,1243]" gridcol="0" gridrow="10" pageId="10" pageNumber="33">spiny setae on head</th>
<td id="76F9F308A570006AC3B88CD89130FB5B" box="[453,620,1221,1243]" gridcol="1" gridrow="10" pageId="10" pageNumber="33">5+1+5 weak</td>
<td id="76F9F308A570006AC0E68CD89181FB5B" box="[667,733,1221,1243]" gridcol="2" gridrow="10" pageId="10" pageNumber="33">—</td>
<td id="76F9F308A570006AC1288CD890E2FB5B" box="[853,958,1221,1243]" gridcol="3" gridrow="10" pageId="10" pageNumber="33">+</td>
<td id="76F9F308A570006AC6728CD897CFFB5B" box="[1039,1171,1221,1243]" gridcol="4" gridrow="10" pageId="10" pageNumber="33">—</td>
<td id="76F9F308A570006AC6A78CD8963CFB5B" box="[1242,1376,1221,1243]" gridcol="5" gridrow="10" pageId="10" pageNumber="33">4+4 heavy</td>
</tr>
<tr id="35289A74A570006AC2EA8D59963CFAFF" box="[151,1376,1348,1407]" gridrow="11" pageId="10" pageNumber="33" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76F9F308A570006AC2EA8D5992E2FAFF" box="[151,446,1348,1407]" gridcol="0" gridrow="11" pageId="10" pageNumber="33">
<taxonomicName id="4C18E3B5A570FF9FC2EA8D5992E2FADE" box="[151,446,1348,1374]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="kovali">
<emphasis id="B96C4424A570FF9FC2EA8D5992E2FADE" bold="true" box="[151,446,1348,1374]" italics="true" pageId="10" pageNumber="33">Pygmarrhopalites kovali</emphasis>
</taxonomicName>
Figs 34–61, Tables 1, 3
</th>
<td id="76F9F308A570006AC3B88D599130FAFF" box="[453,620,1348,1407]" gridcol="1" gridrow="11" pageId="10" pageNumber="33">sp. nov.</td>
</tr>
</table>
</paragraph>
<subSubSection id="C302CBBDA570FF9FC2EA8DB3929CF91F" pageId="10" pageNumber="33" type="diagnosis">
<paragraph id="8BA79836A570FF9FC2EA8DB3929CF91F" blockId="10.[151,1437,1454,2019]" pageId="10" pageNumber="33">
<emphasis id="B96C4424A570FF9FC2EA8DB3924DFA47" bold="true" box="[151,273,1454,1479]" pageId="10" pageNumber="33">Diagnosis.</emphasis>
Body up to 
<quantity id="4CE035D3A570FF9FC3D18DB292ADFA46" box="[428,497,1455,1479]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="33" unit="mm" value="1.0">1 mm</quantity>
length. Eyes 1 + 1. Trichobothria ABC form a very obtuse angle and AB&lt;BC. Antenna about 2.3× of head, Ant IV with 5 distinctly separated subsegments and 13 whorls of setae. Head dorsum with 4 + 4 moderately spine-like setae. Femur II with 15 and III with 14 setae. All claws slender, without tunica and with inner tooth. Tips of empodia not reaching tips of corresponding claws, empodium I–II with and III without corner tooth. Manubrium with 7 + 7 posterior setae; dens with 2 strongly spine-like outer and 3 moderately spinelike inner setae. Tip of mucro rounded. Abd VI with broadened circumanal setae and palmate appendices anales with secondary branching.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA570FF9FC2BA8EB79755F83F" pageId="10" pageNumber="33" type="materials_examined">
<paragraph id="8BA79836A570FF9FC2BA8EB79793F88B" blockId="10.[151,1437,1454,2019]" pageId="10" pageNumber="33">
<materialsCitation id="3B70926BA570FF9FC2BA8EB79711F967" ID-GBIF-Occurrence="1455395226" collectingDate="2009-08-19" collectorName="R. S. Vargovitsh" elevation="510" location="Gulripsh District" municipality="West Caucasus" pageId="10" pageNumber="33" specimenCount="2" stateProvince="Abkhazia" typeStatus="holotype">
<emphasis id="B96C4424A570FF9FC2BA8EB79229F943" bold="true" box="[199,373,1706,1731]" pageId="10" pageNumber="33">
<typeStatus id="54A32694A570FF9FC2BA8EB7925EF943" box="[199,258,1706,1731]" pageId="10" pageNumber="33">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54A32694A570FF9FC3018EB792B8F943" box="[380,484,1706,1731]" pageId="10" pageNumber="33" type="holotype">Holotype</typeStatus>
on slide C-689-1 (
<figureCitation id="132384B3A570FF9FC0C48EB79052F943" box="[697,782,1706,1731]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="10" pageNumber="33">Fig. 57</figureCitation>
): female, the 
<collectingMunicipality id="6BC3024CA570FF9FC1DA8EB6970FF943" box="[935,1107,1707,1731]" pageId="10" pageNumber="33">West Caucasus</collectingMunicipality>
, 
<collectingRegion id="49DC56D4A570FF9FC6238EB79790F943" box="[1118,1228,1706,1731]" country="Georgia" name="Abkhazia" pageId="10" pageNumber="33">Abkhazia</collectingRegion>
, 
<location id="8EC7CEEDA570FF9FC6A58EB796C4F943" LSID="urn:lsid:plazi:treatment:03B12920A57BFF84C2EA8C2196A2FDF5:8EC7CEEDA570FF9FC6A58EB796C4F943" box="[1240,1432,1706,1731]" municipality="West Caucasus" name="Gulripsh District" pageId="10" pageNumber="33" stateProvince="Abkhazia">Gulripsh District</location>
, Tsebelda Vill., 
<elevation id="00357F05A570FF9FC33E8ED39296F967" box="[323,458,1742,1767]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="33" unit="m" value="510.0">
<quantity id="4CE035D3A570FF9FC33E8ED392D0F966" box="[323,396,1742,1767]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="33" unit="m" value="510.0">510 m</quantity>
a.s.l.
</elevation>
, Tsebeldinskaya Cave, 
<date id="FFA6BEF6A570FF9FC0AB8ED3903CF967" box="[726,864,1742,1767]" pageId="10" pageNumber="33" value="2009-08-19">
<collectingDate id="EFE2471EA570FF9FC0AB8ED3903CF967" box="[726,864,1742,1767]" pageId="10" pageNumber="33" value="2009-08-19">19.viii.2009</collectingDate>
</date>
, 
<collectorName id="26EDFDE0A570FF9FC1108ED29743F967" box="[877,1055,1742,1767]" pageId="10" pageNumber="33">R.S. Vargovitsh</collectorName>
leg
</materialsCitation>
. Paratypes on slides: 2 males (one of them dissected and mounted on 4 slides) and 2 females collected together with holotype.
</paragraph>
<paragraph id="8BA79836A570FF9FC2BA8F0B9755F83F" blockId="10.[151,1437,1454,2019]" pageId="10" pageNumber="33">
<materialsCitation id="3B70926BA570FF9FC2BA8F0B9241F8F7" ID-GBIF-Occurrence="1455395224" collectingDate="2010-08-18" collectorName="W Caucasus" county="Other" elevation="280" location="Gulripsh District" municipality="On" pageId="10" pageNumber="33" specimenCount="5" specimenCount-female="5" stateProvince="Abkhazia">
<emphasis id="B96C4424A570FF9FC2BA8F0B92DAF8AF" bold="true" box="[199,390,1814,1839]" pageId="10" pageNumber="33">
<collectingCounty id="62C6E0BAA570FF9FC2BA8F0B9253F8AF" box="[199,271,1814,1839]" pageId="10" pageNumber="33">Other</collectingCounty>
material.
</emphasis>
<collectingMunicipality id="6BC3024CA570FF9FC3ED8F0A92EEF8AF" box="[400,434,1815,1839]" pageId="10" pageNumber="33">On</collectingMunicipality>
slides: male and 
<specimenCount id="9D1E53BFA570FF9FC0F88F0B91ABF8AF" box="[645,759,1814,1839]" pageId="10" pageNumber="33" type="female">5 females</specimenCount>
, the 
<collectorName id="26EDFDE0A570FF9FC14E8F0A909AF8AF" box="[819,966,1815,1839]" pageId="10" pageNumber="33">W Caucasus</collectorName>
, 
<collectingRegion id="49DC56D4A570FF9FC1A98F0B971EF8AF" box="[980,1090,1814,1839]" country="Georgia" name="Abkhazia" pageId="10" pageNumber="33">Abkhazia</collectingRegion>
, 
<location id="8EC7CEEDA570FF9FC62F8F0B9649F8AF" LSID="urn:lsid:plazi:treatment:03B12920A57BFF84C2EA8C2196A2FDF5:8EC7CEEDA570FF9FC62F8F0B9649F8AF" box="[1106,1301,1814,1839]" county="Other" municipality="On" name="Gulripsh District" pageId="10" pageNumber="33" stateProvince="Abkhazia">Gulripsh District</location>
, vicinity of Amtkel Vill., 
<elevation id="00357F05A570FF9FC33A8F2692BDF8D3" box="[327,481,1850,1875]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="33" unit="m" value="280.0">
<quantity id="4CE035D3A570FF9FC33A8F2692C5F8D2" box="[327,409,1851,1875]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="33" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
, Verkhnyaya Shakuranskaya Cave, surface of Barber’s traps exposed during 
<date id="FFA6BEF6A570FF9FC2EA8F439241F8F7" box="[151,285,1886,1911]" pageId="10" pageNumber="33" value="2010-08-18">
<collectingDate id="EFE2471EA570FF9FC2EA8F439241F8F7" box="[151,285,1886,1911]" pageId="10" pageNumber="33" value="2010-08-18">18.viii.2010</collectingDate>
</date>
</materialsCitation>
– 
<materialsCitation id="3B70926BA570FF9FC3508F43928FF83F" ID-GBIF-Occurrence="1455395225" collectingDate="2010-08-20" collectingDateMax="2011-08-20" collectingDateMin="2010-08-20" collectorName="R. S. Vargovitsh &amp; A. G. Koval &amp; W Caucasus" elevation="300" location="Gulripsh District" pageId="10" pageNumber="33" specimenCount="1" stateProvince="Abkhazia">
<date id="FFA6BEF6A570FF9FC3508F4392E9F8F7" box="[301,437,1886,1911]" pageId="10" pageNumber="33" value="2011-08-20">
<collectingDate id="EFE2471EA570FF9FC3508F4392E9F8F7" box="[301,437,1886,1911]" pageId="10" pageNumber="33" value="2011-08-20">20.viii.2011</collectingDate>
</date>
, 70 m from entrance, 
<collectorName id="26EDFDE0A570FF9FC0C28F42902FF8F7" box="[703,883,1886,1911]" pageId="10" pageNumber="33">R.S. Vargovitsh</collectorName>
and 
<collectorName id="26EDFDE0A570FF9FC1CC8F42976FF8F7" box="[945,1075,1886,1911]" pageId="10" pageNumber="33">A.G. Koval</collectorName>
leg.; female, the 
<collectorName id="26EDFDE0A570FF9FC77B8F4296C5F8F7" box="[1286,1433,1887,1911]" pageId="10" pageNumber="33">W Caucasus</collectorName>
, 
<collectingRegion id="49DC56D4A570FF9FC2EA8F9F9259F81B" box="[151,261,1922,1947]" country="Georgia" name="Abkhazia" pageId="10" pageNumber="33">Abkhazia</collectingRegion>
, 
<location id="8EC7CEEDA570FF9FC36E8F9F928AF81B" LSID="urn:lsid:plazi:treatment:03B12920A57BFF84C2EA8C2196A2FDF5:8EC7CEEDA570FF9FC36E8F9F928AF81B" box="[275,470,1922,1947]" name="Gulripsh District" pageId="10" pageNumber="33" stateProvince="Abkhazia">Gulripsh District</location>
, vicinity of Amtkel Vill., 
<elevation id="00357F05A570FF9FC17A8F9E90CCF81B" box="[775,912,1922,1947]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="33" unit="m" value="300.0">
<quantity id="4CE035D3A570FF9FC17A8F9E900DF81A" box="[775,849,1923,1947]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="33" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
, Tshalintshur Cave, surface of Barber’s traps exposed during 
<date id="FFA6BEF6A570FF9FC3378FBB928FF83F" box="[330,467,1958,1983]" pageId="10" pageNumber="33" value="2010-08-20">
<collectingDate id="EFE2471EA570FF9FC3378FBB928FF83F" box="[330,467,1958,1983]" pageId="10" pageNumber="33" value="2010-08-20">20.viii.2010</collectingDate>
</date>
</materialsCitation>
– 
<date id="FFA6BEF6A570FF9FC39E8FBB9134F83F" box="[483,616,1958,1983]" pageId="10" pageNumber="33" value="2011-08-21">21.viii.2011</date>
, R.S. Vargovitsh and A.G. Koval leg.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA570FF99C2BA8FD79759FB68" lastPageId="12" lastPageNumber="35" pageId="10" pageNumber="33" type="description">
<paragraph id="8BA79836A570FF9FC2BA8FD796C0F863" blockId="10.[151,1437,1454,2019]" box="[199,1436,1994,2019]" pageId="10" pageNumber="33">
<emphasis id="B96C4424A570FF9FC2BA8FD7929AF863" bold="true" box="[199,454,1994,2019]" pageId="10" pageNumber="33">Description. Female.</emphasis>
Body length without appendages up to 
<quantity id="4CE035D3A570FF9FC1C28FD69757F862" box="[959,1035,1995,2019]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="33" unit="mm" value="1.0">1 mm</quantity>
, habitus of 
<typeStatus id="54A32694A570FF9FC6DB8FD79654F863" box="[1190,1288,1994,2019]" pageId="10" pageNumber="33" type="holotype">holotype</typeStatus>
in 
<figureCitation id="132384B3A570FF9FC7408FD796C4F863" box="[1341,1432,1994,2019]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="10" pageNumber="33">Fig. 57</figureCitation>
.
</paragraph>
<paragraph id="8BA79836A571FF9EC2EA888A9291FF55" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">Pigmentation of alcohol specimens varies from absence to presence of reddish spots of pigment on head dorsum and interantennal area only.</paragraph>
<paragraph id="8BA79836A571FF9EC2BA88C29389FE50" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Head (
<figureCitation id="132384B3A571FF9EC36F88C29237FF78" box="[274,363,223,248]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Figs 34</figureCitation>
, 
<figureCitation id="132384B3A571FF9EC30588C292C4FF78" box="[376,408,223,248]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="11" pageNumber="34">58</figureCitation>
). Eyes 1 + 1, in alcohol as very small dark pigmented spots or unpigmented. Clypeal area (rows 
<emphasis id="B96C4424A571FF9EC2A3891993B0FE9D" bold="true" box="[222,236,260,285]" pageId="11" pageNumber="34">a</emphasis>
to 
<emphasis id="B96C4424A571FF9EC36D89199245FE9D" bold="true" box="[272,281,260,285]" pageId="11" pageNumber="34">f</emphasis>
) with axial seta in row 
<emphasis id="B96C4424A571FF9EC0598919916EFE9D" bold="true" box="[548,562,260,285]" pageId="11" pageNumber="34">a</emphasis>
and median region M (marked with oval in 
<figureCitation id="132384B3A571FF9EC65889199729FE9C" box="[1061,1141,260,285]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 34</figureCitation>
) with two asymmetrically arranged setae. Interantennal area (rows 
<emphasis id="B96C4424A571FF9EC019893A9128FEC0" bold="true" box="[612,628,295,320]" pageId="11" pageNumber="34">α</emphasis>
and 
<emphasis id="B96C4424A571FF9EC0D1893A91E7FEC0" bold="true" box="[684,699,295,320]" pageId="11" pageNumber="34">β</emphasis>
) with axial seta in row 
<emphasis id="B96C4424A571FF9EC1B1893A9087FEC0" bold="true" box="[972,987,295,320]" pageId="11" pageNumber="34">β</emphasis>
. Dorsal area (rows 
<emphasis id="B96C4424A571FF9EC6C1893A978CFEC0" bold="true" box="[1212,1232,295,320]" pageId="11" pageNumber="34">A</emphasis>
to 
<emphasis id="B96C4424A571FF9EC68A893A9657FEC0" bold="true" box="[1271,1291,295,320]" pageId="11" pageNumber="34">D</emphasis>
): axial setae present in rows 
<emphasis id="B96C4424A571FF9EC33689519203FEE5" bold="true" box="[331,351,332,357]" pageId="11" pageNumber="34">A</emphasis>
, 
<emphasis id="B96C4424A571FF9EC313895192DDFEE5" bold="true" box="[366,385,332,357]" pageId="11" pageNumber="34">B</emphasis>
and 
<emphasis id="B96C4424A571FF9EC3C489519291FEE5" bold="true" box="[441,461,332,357]" pageId="11" pageNumber="34">C</emphasis>
; 4 + 4 moderately spine-like setae in rows 
<emphasis id="B96C4424A571FF9EC1C589519090FEE5" bold="true" box="[952,972,332,357]" pageId="11" pageNumber="34">C</emphasis>
(2 + 2) and 
<emphasis id="B96C4424A571FF9EC62789519732FEE5" bold="true" box="[1114,1134,332,357]" pageId="11" pageNumber="34">D</emphasis>
(2 + 2). Chaetotaxy of the head posterior side as in 
<figureCitation id="132384B3A571FF9EC3D1897292A7FE08" box="[428,507,367,392]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 34</figureCitation>
(dotted lines). Labral / prelabral chaetotaxy: 4 5 5 / 6. Maxillary outer lobe as in 
<taxonomicName id="4C18E3B5A571FF9EC7FA896C938FFE2D" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="dbari">
<emphasis id="B96C4424A571FF9EC7FA896C938FFE2D" italics="true" pageId="11" pageNumber="34">P. dbari</emphasis>
</taxonomicName>
(
<figureCitation id="132384B3A571FF9EC29B89899271FE2D" box="[230,301,404,429]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[213,1395,193,1895]" captionTargetId="figure@5.[188,1399,193,1932]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Pygmarrhopalites dbari sp. nov.: 3, chaetotaxy of head; 4, chaetotaxy of labial palp; 5, maxillary outer lobe; 6, chaetotaxy of antennal segments I – III; 7, chaetotaxy of antennal segment IV." httpUri="https://zenodo.org/record/439782/files/figure.png" pageId="11" pageNumber="34">Fig. 5</figureCitation>
): apical seta with short and thin subparallel branch at the base, sublobal plate with three sublobal hairs.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA89C191C7FD04" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Antenna (
<figureCitation id="132384B3A571FF9EC34B89C192D0FE75" box="[310,396,476,501]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 35</figureCitation>
): length 2.2–2.4× of head. Length ratio of antennal segments I: II: III: IV = 1: 2.2: 3.6: 9.9; basal subsegment of Ant IV 1.2 times longer than Ant III. Ant I with 7 setae of which subapical one is microseta. Ant II with 15 setae. Ant III with 18 setae and 2 relatively short (4.5 µm) sense rods in separate pits. Ant IV bears 13 whorls of setae located in 5 distinct subsegments; subsegmental formula: 1 + 3 + 1 = (
<emphasis id="B96C4424A571FF9EC6898A5A93E3FD05" bold="true" pageId="11" pageNumber="34">A1–A3 + M1– M2</emphasis>
) + (
<emphasis id="B96C4424A571FF9EC28D8A719212FD05" bold="true" box="[240,334,620,645]" pageId="11" pageNumber="34">M3–M5</emphasis>
) + (
<emphasis id="B96C4424A571FF9EC3028A7191D7FD05" bold="true" box="[383,651,620,645]" pageId="11" pageNumber="34">BA + BM1–BM3 + BB</emphasis>
).
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8A929168FC00" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Foreleg (
<figureCitation id="132384B3A571FF9EC3518A9292DFFD28" box="[300,387,655,680]" captionStart="FIGURES 49 – 51" captionStartId="14.[151,264,1967,1990]" captionTargetBox="[163,1430,247,1890]" captionTargetId="figure@14.[153,1433,193,1947]" captionTargetPageId="14" captionText="FIGURES 49 – 51. Pygmarrhopalites kovali sp. nov., chaetotaxy of legs: 49, foreleg; 50, mid leg; 51, hind leg." httpUri="https://zenodo.org/record/439788/files/figure.png" pageId="11" pageNumber="34">Figs 49</figureCitation>
, 
<figureCitation id="132384B3A571FF9EC3ED8A9292ECFD28" box="[400,432,655,680]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">52</figureCitation>
): precoxae 1, 2 and coxa with 1, 0, 1 setae respectively. Trochanter with 3 anterior and 1 posterior setae; femur with 14 setae, 
<emphasis id="B96C4424A571FF9EC04B8AA9910FFD4D" bold="true" box="[566,595,692,717]" pageId="11" pageNumber="34">a4</emphasis>
turned perpendicularly to the longitudinal axis of the segment. Tibiotarsus with 45 setae: whorl I with 9 setae among which 
<emphasis id="B96C4424A571FF9EC0B48ACA91B9FD70" bold="true" box="[713,741,727,752]" pageId="11" pageNumber="34">Ja</emphasis>
somewhat curved and thickened, each of whorls II–V with 8 setae and region 
<emphasis id="B96C4424A571FF9EC3268AE19230FC95" bold="true" box="[347,364,764,789]" pageId="11" pageNumber="34">F</emphasis>
with 3 primary 
<emphasis id="B96C4424A571FF9EC0568AE19111FC95" bold="true" box="[555,589,764,789]" pageId="11" pageNumber="34">FP</emphasis>
setae (
<emphasis id="B96C4424A571FF9EC0DC8AE191F1FC95" bold="true" box="[673,685,764,789]" pageId="11" pageNumber="34">e</emphasis>
, 
<emphasis id="B96C4424A571FF9EC0C38AE19185FC95" bold="true" box="[702,729,764,789]" pageId="11" pageNumber="34">ae</emphasis>
, 
<emphasis id="B96C4424A571FF9EC0948AE19059FC95" bold="true" box="[745,773,764,789]" pageId="11" pageNumber="34">pe</emphasis>
) and secondary seta 
<emphasis id="B96C4424A571FF9EC1848AE19774FC95" bold="true" box="[1017,1064,764,789]" pageId="11" pageNumber="34">FSa</emphasis>
. Pretarsus with 1 anterior and 1 posterior setulae. Foot complex: claw thin, without tunica, with inner tooth and two pairs of indistinct lateral teeth, 4.5–4.8× shorter than tibiotarsus; empodium thin, with corner tooth, about 1.8× shorter than claw, tip of empodial filament not reaching the tip of claw.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8B9192CBFBFD" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Mid leg (
<figureCitation id="132384B3A571FF9EC3518B9192DFFC25" box="[300,387,908,933]" captionStart="FIGURES 49 – 51" captionStartId="14.[151,264,1967,1990]" captionTargetBox="[163,1430,247,1890]" captionTargetId="figure@14.[153,1433,193,1947]" captionTargetPageId="14" captionText="FIGURES 49 – 51. Pygmarrhopalites kovali sp. nov., chaetotaxy of legs: 49, foreleg; 50, mid leg; 51, hind leg." httpUri="https://zenodo.org/record/439788/files/figure.png" pageId="11" pageNumber="34">Figs 50</figureCitation>
, 
<figureCitation id="132384B3A571FF9EC3F38B9192F2FC25" box="[398,430,908,933]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">52</figureCitation>
): precoxae 1 and 2 with 1, 1 setae respectively, precoxal process present, coxa with 3 setae and microsensillum. Trochanter with 5 setae: 3 simple anterior, trochanteral organ and 1 simple posterior. Femur with 15 setae. Tibiotarsus with 44 setae: whorl I with 9 setae whorls II–IV with 8 setae, whorl V with 7 setae (
<emphasis id="B96C4424A571FF9EC7058BC996C0FC6D" bold="true" box="[1400,1436,980,1005]" pageId="11" pageNumber="34">Vp</emphasis>
absent), region 
<emphasis id="B96C4424A571FF9EC33B8BEA920BFB90" bold="true" box="[326,343,1015,1040]" pageId="11" pageNumber="34">F</emphasis>
with 3 
<emphasis id="B96C4424A571FF9EC3D28BEA928EFB90" bold="true" box="[431,466,1015,1040]" pageId="11" pageNumber="34">FP</emphasis>
setae and 
<emphasis id="B96C4424A571FF9EC0378BEA9125FB90" bold="true" box="[586,633,1015,1040]" pageId="11" pageNumber="34">FSa</emphasis>
seta. Foot complex: claw thin, without tunica, with inner tooth, with 2 pairs of small and indistinct lateral teeth, 4.7–4.8× shorter than tibiotarsus; empodium somewhat broader than in foreleg, with corner tooth, with very short apical filament not reaching the tip of corresponding claw, and about 1.9× shorter than claw.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8C9A90AEFAD5" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Hind leg (
<figureCitation id="132384B3A571FF9EC34B8C9A92D0FB20" box="[310,396,1159,1184]" captionStart="FIGURES 49 – 51" captionStartId="14.[151,264,1967,1990]" captionTargetBox="[163,1430,247,1890]" captionTargetId="figure@14.[153,1433,193,1947]" captionTargetPageId="14" captionText="FIGURES 49 – 51. Pygmarrhopalites kovali sp. nov., chaetotaxy of legs: 49, foreleg; 50, mid leg; 51, hind leg." httpUri="https://zenodo.org/record/439788/files/figure.png" pageId="11" pageNumber="34">Figs 51</figureCitation>
, 
<figureCitation id="132384B3A571FF9EC3E58C9A92E4FB20" box="[408,440,1159,1184]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">52</figureCitation>
): precoxae 1 and 2 with 1, 1 setae respectively, process on precoxa 1 present, coxa with 3 setae and microsensillum. Trochanter with anterior trochanteral organ, 3 anterior and 1 posterior simple setae. Femur with 14 setae, among them 2 posterior microsetae. Tibiotarsus with 44 setae as in mid leg. Foot complex: claw relatively slender, without tunica, with inner tooth, with 2 pairs of small and indistinct subbasal and subapical lateral teeth, 5.5–5.7× shorter than tibiotarsus; empodium broad, without corner tooth, with short apical filament not reaching the tip of corresponding claw, and about 1.8× shorter than claw.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8D429787FAF8" blockId="11.[151,1437,151,2013]" box="[199,1243,1375,1400]" pageId="11" pageNumber="34">Lengths ratio of tibiotarsi I: II: III = 1: 1: 1.2. Tibiotarsus I is about 1.3× shorter than head.</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8D9991A3FA1D" blockId="11.[151,1437,151,2013]" box="[199,767,1412,1437]" pageId="11" pageNumber="34">
Ventral tube (
<figureCitation id="132384B3A571FF9EC3238D9992ECFA1D" box="[350,432,1412,1437]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Fig. 52</figureCitation>
) with 1 + 1 subapical setulae.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8DBA9636FA3F" blockId="11.[151,1437,151,2013]" box="[199,1386,1447,1472]" pageId="11" pageNumber="34">
Tenaculum (
<figureCitation id="132384B3A571FF9EC32F8DBA92FAFA40" box="[338,422,1447,1472]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Fig. 54</figureCitation>
): with 2 apical setulae on the anterior lobe, 3 teeth and a basal process on each ramus.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8DD0972DF9AC" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Furca (
<figureCitation id="132384B3A571FF9EC3648DD1922CFA65" box="[281,368,1484,1509]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Fig. 53</figureCitation>
): manubrium with 7 + 7 posterior setae. Dens (23 setae): anterior side with 3, 2, 1, 1 setae; posterior side with 
<emphasis id="B96C4424A571FF9EC3138DF292D9F988" bold="true" box="[366,389,1519,1544]" pageId="11" pageNumber="34">Ie</emphasis>
(
<figureCitation id="132384B3A571FF9EC3E98DF292BAF988" box="[404,486,1519,1544]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 39</figureCitation>
) and 
<emphasis id="B96C4424A571FF9EC0598DF2910AF988" bold="true" box="[548,598,1519,1544]" pageId="11" pageNumber="34">IIpe</emphasis>
(
<figureCitation id="132384B3A571FF9EC0188DF291EBF988" box="[613,695,1519,1544]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 40</figureCitation>
) as massive spines, 
<emphasis id="B96C4424A571FF9EC1E58DF290F7F988" bold="true" box="[920,939,1519,1544]" pageId="11" pageNumber="34">Ii</emphasis>
, 
<emphasis id="B96C4424A571FF9EC1C48DF290ADF988" bold="true" box="[953,1009,1519,1544]" pageId="11" pageNumber="34">IIIpi</emphasis>
(
<figureCitation id="132384B3A571FF9EC67D8DF297DCF988" box="[1024,1152,1519,1544]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Figs 41, 42</figureCitation>
) and 
<emphasis id="B96C4424A571FF9EC6C38DF297A8F988" bold="true" box="[1214,1268,1519,1544]" pageId="11" pageNumber="34">IVpi</emphasis>
— moderately spine-like. Mucro: posterior lamellae serrated, tip rounded. Dens 1.6× as long as mucro.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8E25975AF8EF" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Great abdomen (
<figureCitation id="132384B3A571FF9EC3FF8E2A928AF9D0" box="[386,470,1591,1616]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Fig. 52</figureCitation>
): Th II bears single sensillum in row 
<emphasis id="B96C4424A571FF9EC1058E2A90DAF9D0" bold="true" box="[888,902,1591,1616]" pageId="11" pageNumber="34">a</emphasis>
and 3 setae in row 
<emphasis id="B96C4424A571FF9EC61D8E2A972BF9D0" bold="true" box="[1120,1143,1591,1616]" pageId="11" pageNumber="34">m</emphasis>
with 
<emphasis id="B96C4424A571FF9EC6CA8E2A9780F9D0" bold="true" box="[1207,1244,1591,1616]" pageId="11" pageNumber="34">m1</emphasis>
shaped similarly to posterior cephalic spine-like setae and longer than other setae of Th II (
<figureCitation id="132384B3A571FF9EC1A18E41976DF9F4" box="[988,1073,1628,1653]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 37</figureCitation>
). Th III with sensillum in row 
<emphasis id="B96C4424A571FF9EC7F38E4196C0F9F5" bold="true" box="[1422,1436,1628,1653]" pageId="11" pageNumber="34">a</emphasis>
and 3 simple short setae in row 
<emphasis id="B96C4424A571FF9EC3828E62914AF918" bold="true" box="[511,534,1663,1688]" pageId="11" pageNumber="34">m</emphasis>
(
<figureCitation id="132384B3A571FF9EC05A8E629127F918" box="[551,635,1663,1688]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 38</figureCitation>
). Abd I bears single row with 5 short setae. Abd II with row of 4 setae anterior to trichobothrial complex. Trichobothrial complex usual for the genus (
<figureCitation id="132384B3A571FF9EC6558EB99722F93D" box="[1064,1150,1700,1725]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="11" pageNumber="34">Fig. 59</figureCitation>
): 
<emphasis id="B96C4424A571FF9EC6E98EB99793F93D" bold="true" box="[1172,1231,1700,1725]" pageId="11" pageNumber="34">ABC</emphasis>
form very obtuse angle (150°–170°) and 
<emphasis id="B96C4424A571FF9EC3E08EDA9298F960" bold="true" box="[413,452,1735,1760]" pageId="11" pageNumber="34">AB</emphasis>
is about 0.7× of 
<emphasis id="B96C4424A571FF9EC0F88EDA91F0F960" bold="true" box="[645,684,1735,1760]" pageId="11" pageNumber="34">BC</emphasis>
; 
<emphasis id="B96C4424A571FF9EC0C18EDA9197F960" bold="true" box="[700,715,1735,1760]" pageId="11" pageNumber="34">p</emphasis>
seta is located below the level of trichobothrium 
<emphasis id="B96C4424A571FF9EC6868EDA9652F960" bold="true" box="[1275,1294,1735,1760]" pageId="11" pageNumber="34">B</emphasis>
; seta 
<emphasis id="B96C4424A571FF9EC72D8EDA9631F960" bold="true" box="[1360,1389,1735,1760]" pageId="11" pageNumber="34">b1</emphasis>
lies almost on line 
<emphasis id="B96C4424A571FF9EC33C8EF19234F885" bold="true" box="[321,360,1772,1797]" pageId="11" pageNumber="34">BC</emphasis>
, somewhat closer to 
<emphasis id="B96C4424A571FF9EC0218EF1912CF885" bold="true" box="[604,624,1772,1797]" pageId="11" pageNumber="34">C</emphasis>
; setae 
<emphasis id="B96C4424A571FF9EC0BC8EF19187F885" bold="true" box="[705,731,1772,1797]" pageId="11" pageNumber="34">c1</emphasis>
and 
<emphasis id="B96C4424A571FF9EC1698EF19073F885" bold="true" box="[788,815,1772,1797]" pageId="11" pageNumber="34">c2</emphasis>
located below the level of trichobothrium 
<emphasis id="B96C4424A571FF9EC7658EF19670F885" bold="true" box="[1304,1324,1772,1797]" pageId="11" pageNumber="34">C</emphasis>
and 
<emphasis id="B96C4424A571FF9EC71B8EF196DDF885" bold="true" box="[1382,1409,1772,1797]" pageId="11" pageNumber="34">c1</emphasis>
is higher than 
<emphasis id="B96C4424A571FF9EC35C8F129267F8A8" bold="true" box="[289,315,1807,1832]" pageId="11" pageNumber="34">c2</emphasis>
. Posterior lateral complex with 6 setae. Furca base complex with 9 setae. Central dorsal complex with 3 subequal setae. Posterior dorsal complex with 22–23 setae the longest of which (
<emphasis id="B96C4424A571FF9EC60E8F2997F9F8CD" bold="true" box="[1139,1189,1844,1869]" pageId="11" pageNumber="34">dI-1</emphasis>
= 72 µm in 
<typeStatus id="54A32694A571FF9EC74D8F2996C8F8CD" box="[1328,1428,1844,1869]" pageId="11" pageNumber="34" type="holotype">holotype</typeStatus>
) is 1.3–1.7× longer than hind claw (
<figureCitation id="132384B3A571FF9EC0638F4A912EF8F0" box="[542,626,1879,1904]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="11" pageNumber="34">Fig. 44</figureCitation>
). Ventral complex with 1 or 2 setae.
</paragraph>
<paragraph id="8BA79836A571FF9EC2BA8F61915DF838" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="34">
Fifth abdominal segment (
<figureCitation id="132384B3A571FF9EC3928F61911DF815" box="[495,577,1916,1941]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Fig. 52</figureCitation>
) with 2 thin setae and trichobothrium 
<emphasis id="B96C4424A571FF9EC18C8F619759F815" bold="true" box="[1009,1029,1916,1941]" pageId="11" pageNumber="34">D</emphasis>
in row 
<emphasis id="B96C4424A571FF9EC6208F619737F815" bold="true" box="[1117,1131,1916,1941]" pageId="11" pageNumber="34">a</emphasis>
, and 2 thicker setae in row 
<emphasis id="B96C4424A571FF9EC2EA8FBD93FAF839" bold="true" box="[151,166,1952,1977]" pageId="11" pageNumber="34">p</emphasis>
. Genital field with 2 + 2 setae.
</paragraph>
<paragraph id="8BA79836A571FF99C2BA8FD992CBFEBF" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,1904]" lastPageId="12" lastPageNumber="35" pageId="11" pageNumber="34">
Sixth abdominal segment (
<figureCitation id="132384B3A571FF9EC38A8FD99113F85D" box="[503,591,1988,2013]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="11" pageNumber="34">Figs 56</figureCitation>
, 
<figureCitation id="132384B3A571FF9EC0208FD99121F85D" box="[605,637,1988,2013]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="11" pageNumber="34">60</figureCitation>
): dorsal valve with 11 + 2 axial + 11 setae, of which circumanal setae 
<emphasis id="B96C4424A576FF99C2EA888A939BFF30" bold="true" box="[151,199,151,176]" pageId="12" pageNumber="35">ms1</emphasis>
(
<figureCitation id="132384B3A576FF99C2AA888A9277FF30" box="[215,299,151,176]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="12" pageNumber="35">Fig. 45</figureCitation>
), 
<emphasis id="B96C4424A576FF99C33D888A9223FF30" bold="true" box="[320,383,151,176]" pageId="12" pageNumber="35">mps1</emphasis>
, 
<emphasis id="B96C4424A576FF99C3F3888A9293FF30" bold="true" box="[398,463,151,176]" pageId="12" pageNumber="35">mps2</emphasis>
and 
<emphasis id="B96C4424A576FF99C078888A9119FF30" bold="true" box="[517,581,151,176]" pageId="12" pageNumber="35">mps3</emphasis>
(
<figureCitation id="132384B3A576FF99C028888A91F4FF30" box="[597,680,151,176]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="12" pageNumber="35">Fig. 46</figureCitation>
) are distinctly broadened. Each of lateral valve bears 18 setae with broadened 
<emphasis id="B96C4424A576FF99C36888A1920FFF55" bold="true" box="[277,339,188,213]" pageId="12" pageNumber="35">mpi1</emphasis>
and 
<emphasis id="B96C4424A576FF99C3F288A19297FF55" bold="true" box="[399,459,188,213]" pageId="12" pageNumber="35">mpi2</emphasis>
(
<figureCitation id="132384B3A576FF99C3A388A19169FF54" box="[478,565,188,213]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="12" pageNumber="35">Fig. 47</figureCitation>
). Some of circumanal setae (e.g. 
<emphasis id="B96C4424A576FF99C1C188A190A7FF55" bold="true" box="[956,1019,188,213]" pageId="12" pageNumber="35">mps3</emphasis>
, 
<emphasis id="B96C4424A576FF99C67088A19715FF55" bold="true" box="[1037,1097,188,213]" pageId="12" pageNumber="35">mpi1</emphasis>
, 
<emphasis id="B96C4424A576FF99C62688A197CBFF55" bold="true" box="[1115,1175,188,213]" pageId="12" pageNumber="35">mpi2</emphasis>
) slightly serrated near bases (
<figureCitation id="132384B3A576FF99C29888C29267FF78" box="[229,315,223,248]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="12" pageNumber="35">Fig. 47</figureCitation>
). Appendices anales (
<figureCitation id="132384B3A576FF99C04988C291D1FF78" box="[564,653,223,248]" captionStart="FIGURES 34 – 48" captionStartId="13.[151,264,1913,1936]" captionTargetBox="[234,1372,206,1874]" captionTargetId="figure@13.[193,1372,195,1883]" captionTargetPageId="13" captionText="FIGURES 34 – 48. Pygmarrhopalites kovali sp. nov.: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36 – 48, shape of setae: 36, spine-like seta of head vertex; 37, m 1 of mesothorax; 38, m 1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI- 1 of great abdomen; 45, ms 1 of Abd VI; 46, mps 3 of abd VI; 47, mpi 2 of Abd VI; 48, appendices anales." httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="12" pageNumber="35">Figs 48</figureCitation>
, 
<figureCitation id="132384B3A576FF99C0E688C291E4FF78" box="[667,696,223,248]" captionStart="FIGURES 57 – 61" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[270,1296,203,1910]" captionTargetId="figure@16.[239,1347,193,1932]" captionTargetPageId="16" captionText="FIGURES 57 – 61. Pygmarrhopalites kovali sp. nov.: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales." httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="12" pageNumber="35">61</figureCitation>
) palmate and broadened in distal third, with secondary branches present (length about 30 µm and width of the palmate part about 17 µm in dorsoventral position). In total, 60 setae are present on Abd VI.
</paragraph>
<paragraph id="8BA79836A576FF99C2BA89519157FD4D" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA89519257FEE5" bold="true" box="[199,267,332,357]" pageId="12" pageNumber="35">Male.</emphasis>
The length of body parts is slightly lesser than in female (
<tableCitation id="C69AAD8DA576FF99C1C989519757FEE4" box="[948,1035,332,357]" captionStart="TABLE 1" captionStartId="3.[151,239,512,535]" captionText="TABLE 1. Measurements (in µm) for body parts of types of Pygmarrhopalites dbari sp. nov. and P. kovali sp. nov." pageId="12" pageNumber="35">Table 1</tableCitation>
). Chaetotaxy resembles the one in female, except for small abdomen (
<figureCitation id="132384B3A576FF99C056897291DCFE08" box="[555,640,367,392]" captionStart="FIGURES 52 – 56" captionStartId="15.[151,264,1952,1975]" captionTargetBox="[178,1408,193,1932]" captionTargetId="figure@15.[178,1408,193,1932]" captionTargetPageId="15" captionText="FIGURES 52 – 56. Pygmarrhopalites kovali sp. nov.: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI." httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="12" pageNumber="35">Fig. 55</figureCitation>
). Chaetotaxy of Abd VI differs from females by shape (no broadened setae), and number (44 setae = 7 + 2 axial + 7 on dorsal valve and 14 + 14 on lateral valves). The genital opening is surrounded by about 16 setulae. Unlike in examined females, ventral complex of great abdomen bears 3 setae, but this character is rather variable and cannot be recognized as representing sexual dimorphism. Length ratio of antennal segments I: II: III: IV = 1: 2.6: 3.7: 11.7. Ant IV bears 5 well-separated subsegments (as in female) but along with this, the basal subsegment is subdivided into 5 (pseudo)subsegments separated from each other by up to 4 more or less developed annuli. Thus, every setal whorl of basal subsegment is allocated in the separate pseudosubsegment. Tip of mucro is somewhat spoon-like, whereas in females it is rounded but rather not broadened. Chaetotaxy of small abdomen and arrangement of Ant IV subsegmentation should be considered as characters of sexual dimorphism.
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8ACA9028FC95" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA8ACA9211FD70" bold="true" box="[199,333,727,752]" pageId="12" pageNumber="35">Variability.</emphasis>
In one female appendices anales are deeply split into two main branches and each of them is palmate. Chaetotaxy of ventral complex of great abdomen varies.
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8B0292BCFBB5" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA8B029157FCB8" bold="true" box="[199,523,799,824]" pageId="12" pageNumber="35">Bionomy and distribution.</emphasis>
<taxonomicName id="4C18E3B5A576FF99C06A8B3C9127FCB8" box="[535,635,799,824]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A576FF99C06A8B3C9127FCB8" box="[535,635,799,824]" italics="true" pageId="12" pageNumber="35">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C0FA8B0291B9FCB8" bold="true" box="[647,741,799,824]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C0FA8B0291B9FCB8" box="[647,741,799,824]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
inhabits the aphotic zone of Tsebeldinskaya Cave with air temperature 11.2–13.2°C (min-max annual) and water temperature 12.0–12.3°C. Specimens were taken from the wooden debris and from the water surface. In this cave, the new species cohabits with three other 
<taxonomicName id="4C18E3B5A576FF99C6848B7A96C0FC00" box="[1273,1436,871,896]" class="Entognatha" family="Arrhopalitidae" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="family">Arrhopalitidae</taxonomicName>
species from the genera 
<taxonomicName id="4C18E3B5A576FF99C3CA8B9191DDFC25" box="[439,641,908,933]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B96C4424A576FF99C3CA8B9191DDFC25" box="[439,641,908,933]" italics="true" pageId="12" pageNumber="35">Pygmarrhopalites</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C18E3B5A576FF99C0C28B91900FFC25" box="[703,851,908,933]" class="Entognatha" family="Arrhopalitidae" genus="Troglopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B96C4424A576FF99C0C28B91900FFC25" box="[703,851,908,933]" italics="true" pageId="12" pageNumber="35">Troglopalites</emphasis>
</taxonomicName>
. Due to its morphology (elongated antennae, thin claws with short empodia, poor or absent pigmentation) and topical preferences, this species can be classified as a moderately troglomorphic troglobiont. Besides 
<typeStatus id="54A32694A576FF99C0D48BC89185FC6D" box="[681,729,981,1005]" pageId="12" pageNumber="35">type</typeStatus>
locality, the species was found in some other caves of Tsebelda Karst Massif: Tshalintshur Cave and Verkhnyaya Shakuranskaya Cave (
<figureCitation id="132384B3A576FF99C1C08BEA90A2FB90" box="[957,1022,1015,1040]" captionStart="FIGURES 1 – 2. 1" captionStartId="4.[151,264,1975,1998]" captionTargetBox="[317,1260,226,1922]" captionTargetId="figure@4.[284,1297,195,1940]" captionTargetPageId="4" captionText="FIGURES 1 – 2. 1, type localities of Pygmarrhopalites dbari sp. nov. (1 — Psyrtskha Cave), and P. kovali sp. nov. (2 — Tsebeldinskaya Cave, 3 — Tshalintshur Cave, 4 — Verkhnyaya Shakuranskaya Cave); 2, habitus of mounted male." httpUri="https://zenodo.org/record/439781/files/figure.png" pageId="12" pageNumber="35">Fig. 1</figureCitation>
). Most probably 
<taxonomicName id="4C18E3B5A576FF99C6BF8BE4967DFB90" box="[1218,1313,1015,1040]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A576FF99C6BF8BE4967DFB90" box="[1218,1313,1015,1040]" italics="true" pageId="12" pageNumber="35">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C7548BEA96DEFB90" bold="true" box="[1321,1410,1015,1040]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C7548BEA96DEFB90" box="[1321,1410,1015,1040]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the West Caucasian endemic.
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8C229759FB68" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA8C2292E3FBD8" bold="true" box="[199,447,1087,1112]" pageId="12" pageNumber="35">Conservation status.</emphasis>
The new species shows restricted distribution possibly limited by caves of Tsebelda Karst Massif. According to 
<bibRefCitation id="EF89E5C7A576FF99C3948C7991DAFBFC" author="IUCN" box="[489,646,1124,1148]" pageId="12" pageNumber="35" refString="IUCN. (2012) IUCN Red List Categories and Criteria: Version 3.1. 2 nd Edition. IUCN, Gland &amp; Cambridge, iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
classification the species most likely corresponds to the category 
<emphasis id="B96C4424A576FF99C2EA8C9A9246FB20" bold="true" box="[151,282,1159,1184]" pageId="12" pageNumber="35">Vulnerable</emphasis>
and criterion 
<emphasis id="B96C4424A576FF99C3C48C9A9287FB20" bold="true" box="[441,475,1159,1184]" pageId="12" pageNumber="35">D2</emphasis>
. Therefore, 
<taxonomicName id="4C18E3B5A576FF99C01A8C949194FB20" box="[615,712,1159,1184]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A576FF99C01A8C949194FB20" box="[615,712,1159,1184]" italics="true" pageId="12" pageNumber="35">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C0AD8C9A9077FB20" bold="true" box="[720,811,1159,1184]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C0AD8C9A9077FB20" box="[720,811,1159,1184]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is recommended to be included into IUCN Red List of Threatened Species along with protection of its habitats—Tsebeldinskaya Cave and complex of Shakuran caves which are 
<typeStatus id="54A32694A576FF99C3718CCC9260FB69" box="[268,316,1233,1257]" pageId="12" pageNumber="35">type</typeStatus>
and the only localities for several invertebrate troglobiont taxa.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA576FF99C2BA8CE9909DFAD4" pageId="12" pageNumber="35" type="etymology">
<paragraph id="8BA79836A576FF99C2BA8CE9909DFAD4" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA8CE99212FA8D" bold="true" box="[199,334,1268,1293]" pageId="12" pageNumber="35">Etymology.</emphasis>
The new species is dedicated to my colleague entomologist and biospeleologist Dr. Alexander Koval (St. Petersburg, All-Russia Institute of Plant Protection) who has made many efforts on the exploration of the West Caucasian cave fauna including fauna of Tsebeldinskaya Cave.
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA576FF99C2BA8D4290F2F9F4" pageId="12" pageNumber="35" type="discussion">
<paragraph id="8BA79836A576FF99C2BA8D42966FF988" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<emphasis id="B96C4424A576FF99C2BA8D429267FAF8" bold="true" box="[199,315,1375,1400]" pageId="12" pageNumber="35">Remarks.</emphasis>
Short empodia, not exceeding tip of corresponding claws, is a character that does not often occur in the species of 
<taxonomicName id="4C18E3B5A576FF99C3388D9992E2FA1D" box="[325,446,1412,1437]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A576FF99C3388D9992E2FA1D" box="[325,446,1412,1437]" italics="true" pageId="12" pageNumber="35">principalis</emphasis>
</taxonomicName>
group, so the range of comparable congeners is limited by five of them (Table 3): European 
<taxonomicName id="4C18E3B5A576FF99C3778DB4913DFA40" authority="Loksa, 1964" authorityName="Loksa" authorityYear="1964" box="[266,609,1447,1472]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="thermophilus">
<emphasis id="B96C4424A576FF99C3778DB492E6FA40" box="[266,442,1447,1472]" italics="true" pageId="12" pageNumber="35">P. thermophilus</emphasis>
(
<bibRefCitation id="EF89E5C7A576FF99C3B48DBA9106FA40" author="Loksa" box="[457,602,1447,1472]" pageId="12" pageNumber="35" refString="Loksa, I. (1964) Einige neue und weniger bekannte Collembolen-Arten aus ungarischen Flaumeichen-Buschwaldern. Opuscula Zoologica Budapest, 5 (1), 83 - 98." type="journal article" year="1964">Loksa, 1964</bibRefCitation>
)
</taxonomicName>
, Japanese 
<taxonomicName id="4C18E3B5A576FF99C0A48DB490A3FA40" authority="Yosii, 1956" authorityName="Yosii" authorityYear="1956" box="[729,1023,1447,1472]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B96C4424A576FF99C0A48DB4903EFA40" box="[729,866,1449,1472]" italics="true" pageId="12" pageNumber="35">P. japonicus</emphasis>
(
<bibRefCitation id="EF89E5C7A576FF99C10C8DBA90A4FA40" author="Yosii" box="[881,1016,1447,1472]" pageId="12" pageNumber="35" refString="Yosii, R. (1956) Monographie zur Hohlencollembolen Japans. Contributions from the Biological Laboratory Kyoto University, 3, 1 - 109." type="journal article" year="1956">Yosii, 1956</bibRefCitation>
)
</taxonomicName>
, Nearctic 
<taxonomicName id="4C18E3B5A576FF99C60F8DB492D8FA64" authority="Zeppelini, Taylor &amp; Slay, 2009" authorityName="Zeppelini, Taylor &amp; Slay" authorityYear="2009" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="ashcraftensis">
<emphasis id="B96C4424A576FF99C60F8DB4967EFA40" box="[1138,1314,1447,1472]" italics="true" pageId="12" pageNumber="35">P. ashcraftensis</emphasis>
<bibRefCitation id="EF89E5C7A576FF99C7578DBA92D8FA64" author="Zeppelini" pageId="12" pageNumber="35" refString="Zeppelini, D., Taylor, S. J. &amp; Slay, M. E. (2009) Cave Pygmarrhopalites Vargovitsh, 2009 (Collembola, Symphypleona, Arrhopalitidae) in United States. Zootaxa, 2204, 1 - 18." type="journal article" year="2009">Zeppelini, Taylor &amp; Slay, 2009</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C18E3B5A576FF99C3EF8DD090FCFA65" authority="Christiansen &amp; Bellinger, 1996" authorityName="Christiansen &amp; Bellinger" authorityYear="1996" box="[402,928,1484,1509]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="carolynae">
<emphasis id="B96C4424A576FF99C3EF8DD09143FA65" box="[402,543,1484,1509]" italics="true" pageId="12" pageNumber="35">P. carolynae</emphasis>
(
<bibRefCitation id="EF89E5C7A576FF99C04D8DD190C4FA65" author="Christiansen" box="[560,920,1484,1509]" pageId="12" pageNumber="35" refString="Christiansen, K. &amp; Bellinger, P. (1996) Cave Arrhopalites new to science. Journal of Cave and Karst Studies, 58 (3), 168 - 180." type="journal article" year="1996">Christiansen &amp; Bellinger, 1996</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C18E3B5A576FF99C1A18DD09387F988" authority="Christiansen &amp; Bellinger, 1996" authorityName="Christiansen &amp; Bellinger" authorityYear="1996" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B96C4424A576FF99C1A18DD09739FA65" box="[988,1125,1484,1509]" italics="true" pageId="12" pageNumber="35">P. marshalli</emphasis>
(
<bibRefCitation id="EF89E5C7A576FF99C60B8DD1938FF988" author="Christiansen" pageId="12" pageNumber="35" refString="Christiansen, K. &amp; Bellinger, P. (1996) Cave Arrhopalites new to science. Journal of Cave and Karst Studies, 58 (3), 168 - 180." type="journal article" year="1996">Christiansen &amp; Bellinger, 1996</bibRefCitation>
)
</taxonomicName>
. Only two of these species have claws of all legs without tunica (
<taxonomicName id="4C18E3B5A576FF99C1CB8DEC973AF988" box="[950,1126,1519,1544]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="ashcraftensis">
<emphasis id="B96C4424A576FF99C1CB8DEC973AF988" box="[950,1126,1519,1544]" italics="true" pageId="12" pageNumber="35">P. ashcraftensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C18E3B5A576FF99C6E18DEC967EF988" box="[1180,1314,1519,1544]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B96C4424A576FF99C6E18DEC967EF988" box="[1180,1314,1519,1544]" italics="true" pageId="12" pageNumber="35">P. marshalli</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8E0890F2F9F4" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<taxonomicName id="4C18E3B5A576FF99C2BA8E089275F9AD" box="[199,297,1556,1581]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A576FF99C2BA8E089275F9AD" box="[199,297,1556,1581]" italics="true" pageId="12" pageNumber="35">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C34C8E0992D0F9AD" bold="true" box="[305,396,1556,1581]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C34C8E0992D0F9AD" box="[305,396,1556,1581]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C18E3B5A576FF99C0588E08918BF9AD" box="[549,727,1556,1581]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="ashcraftensis">
<emphasis id="B96C4424A576FF99C0588E08918BF9AD" box="[549,727,1556,1581]" italics="true" pageId="12" pageNumber="35">P. ashcraftensis</emphasis>
</taxonomicName>
by: higher antenna/head ratio (2.3 vs. 1.8), number of Ant IV subsegments (5 vs. 6), presence of spiny setae on head (vs. usual setae), setae number on dorsal valve of Abd VI (13 vs. 12 per side), absence of corner tooth on claw III (vs. presence).
</paragraph>
</subSubSection>
<subSubSection id="C302CBBDA576FF84C2BA8E9D96A2FDF5" lastPageId="17" lastPageNumber="40" pageId="12" pageNumber="35" type="description">
<paragraph id="8BA79836A576FF99C2BA8E9D97B5F93C" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
From 
<taxonomicName id="4C18E3B5A576FF99C3768E9C92CEF918" box="[267,402,1663,1688]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="B96C4424A576FF99C3768E9C92CEF918" box="[267,402,1663,1688]" italics="true" pageId="12" pageNumber="35">P. marshalli</emphasis>
</taxonomicName>
the new species differs by: smaller body size (
<quantity id="4CE035D3A576FF99C1DC8E9D90A4F917" box="[929,1016,1664,1688]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="35" unit="mm" value="0.9">0.9 mm</quantity>
vs. 
<quantity id="4CE035D3A576FF99C65A8E9D97DDF918" box="[1063,1153,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="35" unit="mm" value="1.7">1.7 mm</quantity>
), absence of corner tooth on claw III (vs. presence), 7 + 7 setae on manubrium (vs. 6 + 6), presence of 1 + 1 eyes (vs. 2 + 2).
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8ED496DEF8CD" blockId="12.[151,1437,151,1904]" pageId="12" pageNumber="35">
<taxonomicName id="4C18E3B5A576FF99C2BA8ED49276F960" box="[199,298,1735,1760]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A576FF99C2BA8ED49276F960" box="[199,298,1735,1760]" italics="true" pageId="12" pageNumber="35">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C3488EDA92CFF960" bold="true" box="[309,403,1735,1760]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C3488EDA92CFF960" box="[309,403,1735,1760]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="4C18E3B5A576FF99C0498ED491CCF960" box="[564,656,1735,1760]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A576FF99C0498ED491CCF960" box="[564,656,1735,1760]" italics="true" pageId="12" pageNumber="35">P. dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A576FF99C0E78EDA91ABF960" bold="true" box="[666,759,1735,1760]" pageId="12" pageNumber="35">
<taxonomicNameLabel id="A25FF95FA576FF99C0E78EDA91ABF960" box="[666,759,1735,1760]" pageId="12" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, described above, in many respects but differs from it by antenna / head ratio (2.3 vs. 3), subsegments of Ant IV (5 vs. 7), length of sense rods of antennal III organ (4.5 vs. 10 µm), spiny setae on head dorsum (4 + 4 vs. 5 + 1 + 5), number of setae on femora II (15 vs. 13) and III (14 vs. 12), posterior dorsal seta 
<emphasis id="B96C4424A576FF99C3CE8F2992B9F8CD" bold="true" box="[435,485,1844,1869]" pageId="12" pageNumber="35">dI-1</emphasis>
/ claw III ratio (1.5 vs. 2.2), shape of dens setae 
<emphasis id="B96C4424A576FF99C67B8F299762F8CD" bold="true" box="[1030,1086,1844,1869]" pageId="12" pageNumber="35">IIIpi</emphasis>
and 
<emphasis id="B96C4424A576FF99C6098F2997F6F8CD" bold="true" box="[1140,1194,1844,1869]" pageId="12" pageNumber="35">IVpi</emphasis>
(spiny vs. simple).
</paragraph>
<paragraph id="8BA79836A576FF99C2BA8F4A973FF8F0" blockId="12.[151,1437,151,1904]" box="[199,1123,1879,1904]" pageId="12" pageNumber="35">Comparison of two species described here with others is also presented in Table 3.</paragraph>
<caption id="DF67C8BEA577FF98C2EA8F6491E2F86B" httpUri="https://zenodo.org/record/439787/files/figure.png" pageId="13" pageNumber="36" targetBox="[234,1372,206,1874]" targetPageId="13">
<paragraph id="8BA79836A577FF98C2EA8F6491E2F86B" blockId="13.[151,1437,1913,2027]" pageId="13" pageNumber="36">
<emphasis id="B96C4424A577FF98C2EA8F649208F80F" bold="true" box="[151,340,1913,1936]" pageId="13" pageNumber="36">FIGURES 34–48.</emphasis>
<taxonomicName id="4C18E3B5A577FF98C3248F679108F810" box="[345,596,1914,1936]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A577FF98C3248F679108F810" box="[345,596,1914,1936]" italics="true" pageId="13" pageNumber="36">Pygmarrhopalites kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A577FF98C0278F6791F6F80F" bold="true" box="[602,682,1914,1935]" pageId="13" pageNumber="36">
<taxonomicNameLabel id="A25FF95FA577FF98C0278F6791F6F80F" box="[602,682,1914,1935]" pageId="13" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 34, chaetotaxy of head; 35, chaetotaxy of antenna; 36–48, shape of setae: 36, spine-like seta of head vertex; 37, m1 of mesothorax; 38, m1 of metathorax; 39, Ie of dens; 40, IIpe of dens; 41, IIIpi of dens, lateral and 42, dorsal view; 43, trichobothrium B, proximal part; 44, dI-1 of great abdomen; 45, ms1 of Abd VI; 46, mps3 of abd VI; 47, mpi2 of Abd VI; 48, appendices anales.
</paragraph>
</caption>
<caption id="DF67C8BEA574FF9BC2EA8FB297A4F846" box="[151,1272,1967,1990]" httpUri="https://zenodo.org/record/439788/files/figure.png" pageId="14" pageNumber="37" targetBox="[163,1430,247,1890]" targetPageId="14">
<paragraph id="8BA79836A574FF9BC2EA8FB297A4F846" blockId="14.[151,1272,1967,1990]" box="[151,1272,1967,1990]" pageId="14" pageNumber="37">
<emphasis id="B96C4424A574FF9BC2EA8FB2920FF845" bold="true" box="[151,339,1967,1990]" pageId="14" pageNumber="37">FIGURES 49–51.</emphasis>
<taxonomicName id="4C18E3B5A574FF9BC3278FAD9108F846" box="[346,596,1968,1990]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A574FF9BC3278FAD9108F846" box="[346,596,1968,1990]" italics="true" pageId="14" pageNumber="37">Pygmarrhopalites kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A574FF9BC0278FAD91F7F845" bold="true" box="[602,683,1968,1989]" pageId="14" pageNumber="37">
<taxonomicNameLabel id="A25FF95FA574FF9BC0278FAD91F7F845" box="[602,683,1968,1989]" pageId="14" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, chaetotaxy of legs: 49, foreleg; 50, mid leg; 51, hind leg.
</paragraph>
</caption>
<caption id="DF67C8BEA575FF9AC2EA8FBD92B7F874" httpUri="https://zenodo.org/record/439789/files/figure.png" pageId="15" pageNumber="38" targetBox="[178,1408,193,1932]" targetPageId="15">
<paragraph id="8BA79836A575FF9AC2EA8FBD92B7F874" blockId="15.[151,1436,1952,2036]" pageId="15" pageNumber="38">
<emphasis id="B96C4424A575FF9AC2EA8FBD920FF836" bold="true" box="[151,339,1952,1975]" pageId="15" pageNumber="38">FIGURES 52–56.</emphasis>
<taxonomicName id="4C18E3B5A575FF9AC3258FBC910FF837" box="[344,595,1953,1975]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A575FF9AC3258FBC910FF837" box="[344,595,1953,1975]" italics="true" pageId="15" pageNumber="38">Pygmarrhopalites kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A575FF9AC0258FBC91FBF836" bold="true" box="[600,679,1953,1974]" pageId="15" pageNumber="38">
<taxonomicNameLabel id="A25FF95FA575FF9AC0258FBC91FBF836" box="[600,679,1953,1974]" pageId="15" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 52, chaetotaxy of great abdomen, red arrow directed downward points out a position of seta p below the level of trichobothrium B; 53, chaetotaxy of furca; 54, tenaculum; 55, chaetotaxy of male Abd VI; 56, chaetotaxy of female Abd VI.
</paragraph>
</caption>
<caption id="DF67C8BEA56AFF85C2EA8FBC9634F856" httpUri="https://zenodo.org/record/439790/files/figure.png" pageId="16" pageNumber="39" targetBox="[270,1296,203,1910]" targetPageId="16">
<paragraph id="8BA79836A56AFF85C2EA8FBC9634F856" blockId="16.[151,1436,1953,2006]" pageId="16" pageNumber="39">
<emphasis id="B96C4424A56AFF85C2EA8FBC920AF836" bold="true" box="[151,342,1953,1976]" pageId="16" pageNumber="39">FIGURES 57–61.</emphasis>
<taxonomicName id="4C18E3B5A56AFF85C3228FBC9100F837" box="[351,604,1953,1975]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A56AFF85C3228FBC9100F837" box="[351,604,1953,1975]" italics="true" pageId="16" pageNumber="39">Pygmarrhopalites kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A56AFF85C0198FBF91EBF837" bold="true" box="[612,695,1954,1975]" pageId="16" pageNumber="39">
<taxonomicNameLabel id="A25FF95FA56AFF85C0198FBF91EBF837" box="[612,695,1954,1975]" pageId="16" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 57, habitus of mounted holotype; 58, head dorsum; 59, chaetotaxy of trichobothrial complex; 60, fragment of female Abd VI with circumanal setae, lateral view; 61, female appendices anales.
</paragraph>
</caption>
<paragraph id="8BA79836A56BFF84C2008A4396A2FDF5" blockId="17.[125,1534,606,630]" box="[125,1534,606,630]" pageId="17" pageNumber="40">
<emphasis id="B96C4424A56BFF84C2008A439384FDF5" bold="true" box="[125,216,606,629]" pageId="17" pageNumber="40">TABLE ³</emphasis>
. Differences between 
<taxonomicName id="4C18E3B5A56BFF84C3E28A439128FDF6" box="[415,628,606,630]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="dbari" status="sp. nov.">
<emphasis id="B96C4424A56BFF84C3E28A439128FDF6" box="[415,628,606,630]" italics="true" pageId="17" pageNumber="40">Pygmarrhopalites dbari</emphasis>
</taxonomicName>
<emphasis id="B96C4424A56BFF84C0048A43919DFDF5" bold="true" box="[633,705,606,629]" pageId="17" pageNumber="40">
<taxonomicNameLabel id="A25FF95FA56BFF84C0048A43919DFDF5" box="[633,705,606,629]" pageId="17" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C18E3B5A56BFF84C0B18A439046FDF6" box="[716,794,606,630]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="kovali" status="sp. nov.">
<emphasis id="B96C4424A56BFF84C0B18A439046FDF6" box="[716,794,606,630]" italics="true" pageId="17" pageNumber="40">P. kovali</emphasis>
</taxonomicName>
<emphasis id="B96C4424A56BFF84C15D8A43903BFDF5" bold="true" box="[800,871,606,629]" pageId="17" pageNumber="40">
<taxonomicNameLabel id="A25FF95FA56BFF84C15D8A43903BFDF5" box="[800,871,606,629]" pageId="17" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
anđ congeners of 
<taxonomicName id="4C18E3B5A56BFF84C6748A439735FDF6" box="[1033,1129,606,630]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="B96C4424A56BFF84C6748A439735FDF6" box="[1033,1129,606,630]" italics="true" pageId="17" pageNumber="40">principalis</emphasis>
</taxonomicName>
group with empođia not exceeđing claws tips
</paragraph>
</subSubSection>
</treatment>
</document>