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<document id="ECC987D39A1F201FA85BB0DCE54A50FA" ID-CLB-Dataset="88683" ID-DOI="10.5281/zenodo.6448815" ID-GBIF-Dataset="bbbf94d9-a910-4cda-97df-7eca124163ed" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6448815" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" checkinTime="1635825784914" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03AD87FAFFA7F649896B3279F689FAC0" docLanguage="en" docName="hbmw_9_Pteropodidae_16.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Pteropus lylei K. Andersen 1908" docType="treatment" docVersion="14" lastPageNumber="157" masterDocId="FF94FF82FFC4F62A891E341CFFA5FF9B" masterDocTitle="Pteropodidae" masterLastPageNumber="162" masterPageNumber="16" pageNumber="157" updateTime="1719592658732" updateUser="admin">
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<mods:title id="6D01D72F8FD5F2797C15DCD82ADBB0A7">Pteropodidae</mods:title>
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<mods:namePart id="4B66CFF45BA79923CA6BD597C9984AC0">Don E. Wilson</mods:namePart>
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<mods:namePart id="ABF99DF90E02F434030003FC19D6CA85">Russell A. Mittermeier</mods:namePart>
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<paragraph id="8BBB36ECFFA7F649896B3279FF61F90C" blockId="99.[112,966,1637,1731]" box="[117,196,1637,1687]" pageId="99" pageNumber="157">
<heading id="D0F38180FFA7F649896B3279FF61F90C" box="[117,196,1637,1687]" pageId="99" pageNumber="157">
<figureCitation id="133F2A69FFA7F649896B3279FF61F90C" box="[117,196,1637,1687]" captionStart="Plate 11: Pteropodidae" captionStartId="97.[117,147,3321,3346]" captionTargetBox="[11,2741,16,3661]" captionTargetPageId="96" captionText="174. Livingstones Flying Fox (Pteropus livingstonii), 175. Pemba Flying Fox (Pleropus voeltzkowi), 176. Ryukyu Flying Fox (Pleropus dasymallus), 177. Bonin Flying Fox (Pteropus pselaphon), 178. Little Golden-mantled Flying Fox (Pleropus pumilus), 179. Lyles Flying Fox (Pteropus lylei), 180. Indian Flying Fox (Pleropus medius), 181. Rodrigues Flying Fox (Pteropus rodricensus), 182. Large Flying Fox (Pteropus vampyrus), 183. Aldabra Flying Fox (Pleropus aldabrensis), 184. Malagasy Flying Fox (Pteropus rufus), 185. Seychelles Flying Fox (Pleropus seychellensis), 186. Greater Mascarene Flying Fox (Pteropus niger), 187. Aru Flying Fox (Pteropus aruensis), 188. Kai Flying Fox (Pleropus keyensis), 189. Black-bearded Flying Fox (Pleropus melanopogon), 190. South Moluccan Flying Fox (Pteropus chrysoproctus), 191. North Moluccan Flying Fox (Pteropus caniceps)" figureDoi="http://doi.org/10.5281/zenodo.6448911" httpUri="https://zenodo.org/record/6448911/files/figure.png" pageId="99" pageNumber="157">179.</figureCitation>
</heading>
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<subSubSection id="C31E6567FFA7F64989CB3279FD95F90C" box="[213,560,1637,1687]" pageId="99" pageNumber="157" type="vernacular_names">
<paragraph id="8BBB36ECFFA7F64989CB3279FD95F90C" blockId="99.[112,966,1637,1731]" box="[213,560,1637,1687]" pageId="99" pageNumber="157">
<heading id="D0F38180FFA7F64989CB3279FD95F90C" box="[213,560,1637,1687]" pageId="99" pageNumber="157">
<vernacularName id="050746C2FFA7F64989CB3279FD95F90C" ID-CoL="4Q32K" authorityName="K. Andersen" authorityYear="1908" box="[213,560,1637,1687]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" language="eng" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Lyles Flying Fox</vernacularName>
</heading>
</paragraph>
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<paragraph id="8BBB36ECFFA7F6498B6D3279FCF2F90C" blockId="99.[112,966,1637,1731]" box="[627,855,1637,1687]" pageId="99" pageNumber="157">
<heading id="D0F38180FFA7F6498B6D3279FCF2F90C" box="[627,855,1637,1687]" pageId="99" pageNumber="157">
<taxonomicName id="4C044D6FFFA7F6498B6D3279FCF2F90C" ID-CoL="4Q32K" authorityName="K. Andersen" authorityYear="1908" box="[627,855,1637,1687]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">
<emphasis id="B970EAFEFFA7F6498B6D3279FCF2F90C" box="[627,855,1637,1687]" italics="true" pageId="99" pageNumber="157">Pteropus lylei</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31E6567FFA7F649896F32B5FC63F925" box="[113,966,1705,1726]" pageId="99" pageNumber="157" type="vernacular_names">
<paragraph id="8BBB36ECFFA7F649896F32B5FC63F925" blockId="99.[112,966,1637,1731]" box="[113,966,1705,1726]" pageId="99" pageNumber="157">
<heading id="D0F38180FFA7F649896F32B5FC63F925" box="[113,966,1705,1726]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F649896F32B5FF18F925" bold="true" box="[113,189,1705,1726]" pageId="99" pageNumber="157">French:</emphasis>
<vernacularName id="050746C2FFA7F64989D932B5FEDCF925" ID-CoL="4Q32K" authorityName="K. Andersen" authorityYear="1908" box="[199,377,1705,1726]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" language="fra" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Roussette de Lyle</vernacularName>
/
<emphasis id="B970EAFEFFA7F649889332B5FE4DF925" bold="true" box="[397,488,1705,1726]" pageId="99" pageNumber="157">German:</emphasis>
<vernacularName id="050746C2FFA7F64988EC32B5FDDEF925" ID-CoL="4Q32K" authorityName="K. Andersen" authorityYear="1908" box="[498,635,1705,1726]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" language="deu" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Lyle-Flughund</vernacularName>
/
<emphasis id="B970EAFEFFA7F6498B8E32B5FD49F925" bold="true" box="[656,748,1705,1726]" pageId="99" pageNumber="157">Spanish:</emphasis>
<vernacularName id="050746C2FFA7F6498BEB32B5FC63F925" ID-CoL="4Q32K" authorityName="K. Andersen" authorityYear="1908" box="[757,966,1705,1726]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" language="esp" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Zorro volador de Lyle</vernacularName>
</heading>
</paragraph>
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<subSubSection id="C31E6567FFA7F6498BC832F0FC85F8AE" pageId="99" pageNumber="157" type="reference_group">
<paragraph id="8BBB36ECFFA7F6498BC832F0FC85F8AE" blockId="99.[726,1319,1772,2200]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498BC832F0FCD4F896" bold="true" box="[726,881,1772,1805]" pageId="99" pageNumber="157">Taxonomy.</emphasis>
<taxonomicName id="4C044D6FFFA7F6498A9032F0FCBFF8AE" ID-CoL="4Q32K" authority="K. Andersen, 1908" authorityName="K. Andersen" authorityYear="1908" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Pteropus lylei K. Andersen, 1908</taxonomicName>
,
</paragraph>
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<paragraph id="8BBB36ECFFA7F6498A323304FB78F8AE" blockId="99.[726,1319,1772,2200]" box="[812,1245,1816,1845]" pageId="99" pageNumber="157">
<materialsCitation id="3B6C3CB1FFA7F6498A323304FB78F8AE" ID-GBIF-Occurrence="3828391396" box="[812,1245,1816,1845]" pageId="99" pageNumber="157">
<collectingRegion id="49C0F80EFFA7F6498A283304FC10F8AE" box="[822,949,1816,1845]" country="Thailand" name="Krung Thep Mahanakhon" pageId="99" pageNumber="157">Bangkok</collectingRegion>
,
<collectingCountry id="F313767CFFA7F6498AD83304FBAFF8AE" box="[966,1034,1816,1845]" name="Thailand" pageId="99" pageNumber="157">Siam</collectingCountry>
[=
<collectingCountry id="F313767CFFA7F6498D253304FB60F8AE" box="[1083,1221,1816,1845]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
].”
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</paragraph>
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<subSubSection id="C31E6567FFA7F6498BC83327FC7FF818" pageId="99" pageNumber="157" type="discussion">
<paragraph id="8BBB36ECFFA7F6498BC83327FC7FF818" blockId="99.[726,1319,1772,2200]" pageId="99" pageNumber="157">
<taxonomicName id="4C044D6FFFA7F6498BC83327FC22F8C7" authorityName="K. Andersen" authorityYear="1908" box="[726,903,1851,1884]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" order="Chiroptera" pageId="99" pageNumber="157" phylum="Chordata" rank="species" species="lylei">Pteropus lylei</taxonomicName>
1s in the
<taxonomicName id="4C044D6FFFA7F6498D293327FB17F8C7" authorityName="Linnaeus" authorityYear="1758" box="[1079,1202,1851,1884]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="99" pageNumber="157" phylum="Ascomycota" rank="species" species="vampyrus">vampyrus</taxonomicName>
species group. Monotypic.
</paragraph>
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<caption id="DF7B6664FFA7F6498BC83392FC85F7D2" ID-DOI="http://doi.org/10.5281/zenodo.6803131" ID-Zenodo-Dep="6803131" httpUri="https://zenodo.org/record/6803131/files/figure.png" inLine="true" pageId="99" pageNumber="157" startId="99.[726,898,1934,1963]" targetBox="[110,702,1786,2200]" targetPageId="99">
<paragraph id="8BBB36ECFFA7F6498BC83392FC85F7D2" blockId="99.[726,1319,1772,2200]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498BC83392FC23F830" bold="true" box="[726,902,1934,1963]" pageId="99" pageNumber="157">Distribution.</emphasis>
S
<collectingCountry id="F313767CFFA7F6498AB63392FB8CF830" box="[936,1065,1934,1963]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
, S
<collectingCountry id="F313767CFFA7F6498D4E3392FB47F830" box="[1104,1250,1934,1963]" name="Cambodia" pageId="99" pageNumber="157">Cambodia</collectingCountry>
, and S
<collectingCountry id="F313767CFFA7F6498BF233A9FCC1F849" box="[748,868,1973,2002]" name="Vietnam" pageId="99" pageNumber="157">Vietnam</collectingCountry>
around Gulf of
<collectingCountry id="F313767CFFA7F6498D5B33A9FB62F849" box="[1093,1223,1973,2002]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
, entering inland C
<collectingCountry id="F313767CFFA7F6498A8633C5FB8FF861" box="[920,1066,2009,2042]" name="Cambodia" pageId="99" pageNumber="157">Cambodia</collectingCountry>
. Single specimen recorded in
<collectingRegion id="49C0F80EFFA7F6498A933C18FC5EF7BA" box="[909,1019,2052,2081]" country="China" name="Yunnan" pageId="99" pageNumber="157">Yunnan</collectingRegion>
(
<collectingCountry id="F313767CFFA7F6498D093C18FBCBF7BA" box="[1047,1134,2052,2081]" name="China" pageId="99" pageNumber="157">China</collectingCountry>
) considered alien.
</paragraph>
</caption>
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<paragraph id="8BBB36ECFFA7F6498BC83C53FC39F436" blockId="99.[726,1319,1772,2200]" lastBlockId="99.[111,1320,2206,3462]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498BC83C53FC7FF7EB" bold="true" box="[726,986,2127,2160]" pageId="99" pageNumber="157">Descriptive notes.</emphasis>
Head-body
<quantity id="4CFC9B09FFA7F6498DB73C53FA80F7EB" box="[1193,1317,2127,2160]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="99" pageNumber="157" unit="mm" value="200.0">200 mm</quantity>
(tailless), ear
<quantity id="4CFC9B09FFA7F6498AB53C67FBEFF703" box="[939,1098,2171,2200]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.6" metricValueMin="3.4" pageId="99" pageNumber="157" unit="mm" value="35.0" valueMax="36.0" valueMin="34.0">34-36 mm</quantity>
, hindfoot
<quantity id="4CFC9B09FFA7F6498DEB3C67FF72F724" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="99" pageNumber="157" unit="mm" value="52.0" valueMax="54.0" valueMin="50.0">50-54 mm</quantity>
, forearm
<quantity id="4CFC9B09FFA7F64988723C82FD81F724" box="[364,548,2206,2239]" metricMagnitude="-1" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="99" pageNumber="157" unit="mm" value="152.5" valueMax="160.0" valueMin="145.0">145-160 mm</quantity>
; weight
<quantity id="4CFC9B09FFA7F6498BBC3C82FC90F724" box="[674,821,2206,2239]" metricMagnitude="-1" metricUnit="kg" metricValue="4.35" metricValueMax="4.8" metricValueMin="3.9" pageId="99" pageNumber="157" unit="g" value="435.0" valueMax="480.0" valueMin="390.0">390-480 g</quantity>
. Lyles Flying Fox is similar to the Indian Flying Fox (
<taxonomicName id="4C044D6FFFA7F64988983CD9FDA1F77D" authorityName="Temminck" authorityYear="1825" box="[390,516,2245,2278]" class="Dothideomycetes" family="Pteropodidae" genus="Pteropus" kingdom="Fungi" order="Chiroptera" pageId="99" pageNumber="157" phylum="Ascomycota" rank="species" species="medius">P. medius</taxonomicName>
) but smaller. Muzzle is long, tapering, dark brown, and sparsely haired; nostrils are shortly tubular. Eyes are large, with brown irises. Ears are long and pointed. Head pelage is dark brown around eyes; cap and sides of head are reddish to yellowish brown. Nape and mantle are distinctly yellowish to buff or ocherous buff, orange in some individuals, with longer and hirsute hairs, extending to sides of neck; mantle ends abruptly on back, which is dark brown to rump, with closely adpressed, distinctly grizzled hairs. Throat is dark; collar is reddish brown to russet, separate from seal-brown, grizzled belly, often washed in reddish tinge; and in some individuals, belly is russet or orange-brown down to black genitals. Wing membranes are dark brown from slightly above medial plane, index claw is present, and all claws are black. Forearm and tibia are naked dorsally. Uropatagium is narrow at center; calcaris short. Skull is typical pteropine, with strong basicranial deflection; rostrum is long and tapering; orbit is large; zygoma is thin, weak, and gently arched; braincase is domed; postorbital foramina are large; postorbital constriction is poorly marked; temporal lines do notjoin in raised sagittal crest; nuchal crest is obvious but low; palate is flat and long; tooth rows are slightly divergent posteriorly; and post-dental palate is relatively short. Mandible is rather delicate; symphysis is long and sloping. Dentition is typical pteropine but generally weak, with rather rounded cusps; M,is small and peglike. Chromosomal complement has 2n = 40 and FN = 76.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F649896E3FAFFBF8F3D1" pageId="99" pageNumber="157" type="biology_ecology">
<paragraph id="8BBB36ECFFA7F649896E3FAFFBF8F3D1" blockId="99.[111,1320,2206,3462]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F649896E3FAFFF7AF44F" bold="true" box="[112,223,2995,3028]" pageId="99" pageNumber="157">Habitat.</emphasis>
Mangrove forests, freshwater and peat swamp forests, lowland deciduous dry forests, mountain rainforests, and rainforests. Roosting habitat of Lyles Flying Fox can be in close proximity to human settlements, with marked preference (53% of roosts) for trees growing in Buddhist temples, and is usually near watercourses.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F649896E384DFD08F2AC" pageId="99" pageNumber="157" type="food_feeding">
<paragraph id="8BBB36ECFFA7F649896E384DFD08F2AC" blockId="99.[111,1320,2206,3462]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F649896E384DFED0F3E9" bold="true" box="[112,373,3153,3186]" pageId="99" pageNumber="157">Food and Feeding.</emphasis>
Liyles Flying Fox is primarily frugivorous. In
<collectingCountry id="F313767CFFA7F6498AED384DFBC8F3E9" box="[1011,1133,3153,3186]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
,it eats fruits of 34 plant genera, predominantly figs (
<taxonomicName id="4C044D6FFFA7F6498BD63864FCA8F302" box="[712,781,3192,3225]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="99" pageNumber="157" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
,
<taxonomicName id="4C044D6FFFA7F6498A3D3864FC08F302" box="[803,941,3192,3225]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="99" pageNumber="157" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
) and cultivated plants in orchards. Plants visited for flowers and leaves include
<taxonomicName id="4C044D6FFFA7F6498A9838B8FB22F35A" authority="" box="[902,1159,3236,3265]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="99" pageNumber="98" phylum="Tracheophyta" rank="subGenus" subGenus="Fabaceae">Acacia (Fabaceae)</taxonomicName>
,
<taxonomicName id="4C044D6FFFA7F6498D8538B8FEB8F373" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="99" pageNumber="157" phylum="Tracheophyta" rank="genus">Eucalyptus (Myrtaceae)</taxonomicName>
, and
<taxonomicName id="4C044D6FFFA7F649886A38DBFD50F373" box="[372,757,3271,3304]" class="Magnoliopsida" family="Bignoniaceae" genus="Markhamia" kingdom="Plantae" order="Lamiales" pageId="99" pageNumber="157" phylum="Tracheophyta" rank="genus">Markhamia (Bignoniaceae)</taxonomicName>
. Lyles Flying Foxes ingest ¢.50% their body weight/day. It is an important seed disperser and a central-place forager that returns to the same roosts after foraging.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F64989713922F6E1FEC2" pageId="99" pageNumber="157" type="breeding">
<paragraph id="8BBB36ECFFA7F64989713922F6E1FEC2" blockId="99.[111,1320,2206,3462]" lastBlockId="99.[1389,2603,272,1376]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F64989713922FF53F2C4" bold="true" box="[111,246,3390,3423]" pageId="99" pageNumber="157">Breeding.</emphasis>
Lyles Flying Fox is seasonally monoestrous. Mating and courtship occur in morning and late afternoon at roosts. Litter size is one, occasionally two. Young are more common in colonies in April-May (hot inter-monsoonal period). Young remains attached to, and flies with, its mother for most of the lactation period.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F6498C733543F886FDF6" pageId="99" pageNumber="157" type="activity">
<paragraph id="8BBB36ECFFA7F6498C733543F886FDF6" blockId="99.[1389,2603,272,1376]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498C733543F9F9FE1B" bold="true" box="[1389,1628,351,384]" pageId="99" pageNumber="157">Activity patterns.</emphasis>
Lyles Flying Foxes are crepuscular and nocturnal. They leave day roosts in the evening to forage and return in early morning (03:00-05:00 h). Lunar phase had no influence on timing of colony-wide emergence; during breeding season, females were the last to emerge. During daytime, sleeping, grooming, and wing flapping are the most frequent behaviors of roosting bats. Wing spreading, movement, and courtship are more frequent in males than females; the opposite is true for grooming and antagonistic behaviors.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F6498C70366FF862FC33" pageId="99" pageNumber="157" type="biology_ecology">
<paragraph id="8BBB36ECFFA7F6498C70366FF862FC33" blockId="99.[1389,2603,272,1376]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498C70366FF797FD0F" bold="true" box="[1390,2098,627,660]" pageId="99" pageNumber="157">Movements, Home range and Social organization.</emphasis>
Lyles Flying Fox is highly colonial and roosts in tall trees; in
<collectingCountry id="F313767CFFA7F6498FC93683F8FCFD27" box="[1751,1881,671,700]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
, roosts contain 100-1000 individuals of both sexes, with one colony of more than 11,000 individuals. It is highly mobile and can travel up to
<quantity id="4CFC9B09FFA7F6498C8936F5FA50FC91" box="[1431,1525,745,778]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="99" pageNumber="157" unit="km" value="24.0">24 km</quantity>
/night from day roosts to foraging areas; cumulative distances traveled per night are up to
<quantity id="4CFC9B09FFA7F6498F53370CF900FCAA" box="[1613,1701,784,817]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="99" pageNumber="157" unit="km" value="50.0">50 km</quantity>
. Fidelity for roosting and feeding sites is high. Lyles Flying Fox spent 5—-8% of time commuting to foraging sites, 58-78% foraging, and 12-37% roosting depending on season (more foraging in November, more roosting in April-May); 62-65% of roosting time was spent resting.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F6498C7137B2F723FB78" pageId="99" pageNumber="157" type="conservation">
<paragraph id="8BBB36ECFFA7F6498C7137B2F723FB78" blockId="99.[1389,2603,272,1376]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498C7137B2F970FC54" bold="true" box="[1391,1749,942,975]" pageId="99" pageNumber="157">Status and Conservation.</emphasis>
CITES Appendix II. Classified as Vulnerable on The IUCN Red List. Lyles Flying Fox is perceived to have decline of more than 30% over the past 15 years due to increase hunting and trade, decline in habitat quality and extent due to deforestation, and loss of tall roosting trees. It occurs in protected areas. It is tolerant of human presence; roosting sites are in close proximity to settlements, particularly Buddhist temples where it is protected by monks (58% of 30 known roosts in
<collectingCountry id="F313767CFFA7F6498C713087FA52FB27" box="[1391,1527,1179,1212]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
). The Wildlife Preservation and Protection Act protects it in
<collectingCountry id="F313767CFFA7F64980423087F67AFB27" box="[2396,2527,1179,1212]" name="Thailand" pageId="99" pageNumber="157">Thailand</collectingCountry>
, and overall population is estimated to be 75,000 individuals.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFA7F6498C7130EFF689FAC0" pageId="99" pageNumber="157" type="bibRefCitation_list">
<paragraph id="8BBB36ECFFA7F6498C7130EFF689FAC0" blockId="99.[1389,2603,272,1376]" pageId="99" pageNumber="157">
<emphasis id="B970EAFEFFA7F6498C7130EFF9ACFA97" bold="true" box="[1391,1545,1267,1292]" pageId="99" pageNumber="157">Bibliography.</emphasis>
Almeida et al. (2014), Andersen (1912b), Boonneung (1977), Bumrungsri, Suyanto &amp; Francis (2008), Chaiyes et al. (2017), Hengjan (2011), Hondo et al. (2010), Hood et al. (1988), Smith, A.T. &amp; Xie Yan (2008), Smith, J.D.L. &amp; Gregory (2009), Weber et al. (2015), Zhang Jinshuo et al. (2010).
</paragraph>
</subSubSection>
</treatment>
</document>