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<mods:title id="C8E0EC066F65110A81261CA495A06F5E">A new species of the genus Quantula Baker, 1941 (Eupulmonata: Dyakiidae) from the southern part of north-eastern Thailand</mods:title>
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<paragraph id="8BBB36E0FFC5E473FC10AADB205BF976" blockId="8.[941,1368,1719,1776]" box="[993,1317,1719,1744]" pageId="8" pageNumber="527">
<heading id="D0F3818CFFC5E473FC10AADB205BF976" bold="true" box="[993,1317,1719,1744]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="527" reason="2">
<emphasis id="B970EAF2FFC5E473FC10AADB205BF976" bold="true" box="[993,1317,1719,1744]" pageId="8" pageNumber="527">
<taxonomicName id="4C044D63FFC5E473FC10AADB21EEF976" ID-CoL="ac51f98e-4da0-46ab-a4e9-ac9ae850018f" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[993,1168,1719,1743]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC5E473FC10AADB21EEF976" bold="true" box="[993,1168,1719,1743]" italics="true" pageId="8" pageNumber="527">Quantula doma</emphasis>
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, 
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<paragraph id="8BBB36E0FFC5E473FC5CAAB42026F949" blockId="8.[941,1368,1719,1776]" box="[941,1368,1752,1776]" pageId="8" pageNumber="527">
(
<figureCitation id="133F2A65FFC5E473FC44AAB4214AF949" box="[949,1076,1752,1776]" captionStart="Fig" captionStartId="8.[135,171,1667,1689]" captionTargetBox="[135,774,189,1651]" captionTargetId="figure-7@8.[135,774,189,1651]" captionTargetPageId="8" captionText="Fig. 3. Living snails of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana (NHMSU-00035) from Phanom Sawai, Surin; B–C, Q. doma, new species: B, NHMSU-00049 from Khao Sala, Surin; C, NHMSU-00051 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399989" httpUri="https://zenodo.org/record/7399989/files/figure.png" pageId="8" pageNumber="527">Figs. 3B–C</figureCitation>
, 
<figureCitation id="133F2A65FFC5E473FBB0AAB421FCF949" box="[1089,1154,1752,1776]" captionStart="Fig" captionStartId="8.[835,873,1528,1550]" captionTargetBox="[835,1475,189,1512]" captionTargetId="figure-93@8.[835,1475,189,1512]" captionTargetPageId="8" captionText="Fig. 4. Shell morphology of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana, specimen NHMSU-00035 from Phanom Sawai, Surin; B–C, Q. doma, new species: B, holotype NHMSU-00048 from Khao Sala, Surin; C, NHMSU-00050 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399987" httpUri="https://zenodo.org/record/7399987/files/figure.png" pageId="8" pageNumber="527">4B–C</figureCitation>
, 
<figureCitation id="133F2A65FFC5E473FB7EAAB421B5F949" box="[1167,1227,1752,1776]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="8" pageNumber="527">6A–J</figureCitation>
; 
<tableCitation id="C686035BFFC5E473FB26AAB4204FF949" box="[1239,1329,1752,1776]" captionStart="Table 1" captionStartId="1.[111,166,517,540]" captionTargetBox="[151,416,248,2084]" captionTargetPageId="2" captionText="Table 1. Species, localities, and GenBank accession numbers for sequences used in this study. Sequences marked * are from Jirapatrasilp et al. (2021)." httpUri="http://table.plazi.org/id/DF7B6668FFCCE47AFF9EAE692340FDA5" pageId="8" pageNumber="527" tableUuid="DF7B6668FFCCE47AFF9EAE692340FDA5">Tables 1</tableCitation>
–3)
</paragraph>
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<subSubSection id="C31E656BFFC5E473FCB2AB7A20F7F84C" pageId="8" pageNumber="527" type="reference_group">
<paragraph id="8BBB36E0FFC5E473FCB2AB7A20BCF892" blockId="8.[835,1474,1814,2038]" box="[835,1474,1814,1836]" pageId="8" pageNumber="527">
<treatmentCitationGroup id="AB1411CEFFC5E473FCB2AB7A20BCF892" box="[835,1474,1814,1836]" pageId="8" pageNumber="527">
<emphasis id="B970EAF2FFC5E473FCB2AB7A26E0F892" box="[835,926,1814,1835]" italics="true" pageId="8" pageNumber="527">Quantula</emphasis>
sp. – 
<treatmentCitation id="0AA510F1FFC5E473FC2BAB7A2193F892" author="Jumlong P &amp; Tumpeesuwan C &amp; Tumpeesuwan S" box="[986,1261,1814,1836]" page="71" pageId="8" pageNumber="527" year="2013">
<bibRefCitation id="EF954B11FFC5E473FC2BAB7A2193F892" author="Jumlong P &amp; Tumpeesuwan C &amp; Tumpeesuwan S" box="[986,1261,1814,1836]" pageId="8" pageNumber="527" pagination="67 - 81" refId="ref8585" refString="Jumlong P, Tumpeesuwan C &amp; Tumpeesuwan S (2013) Species diversity and abundance of land snails in sandstone and volcanic hills in Surin. Burapha Science Journal, 18 (1): 67 - 81. (In Thai with English abstract)" type="journal article" year="2013">Jumlong et al., 2013: 71–72</bibRefCitation>
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, 74–76, 78–79, fig. 3.
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</paragraph>
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<taxonomicName id="4C044D63FFC5E473FCB2AB5F2153F8F1" ID-CoL="7SZ3C" authority="Worachak, 2014" box="[835,1069,1843,1864]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="weinkauffiana">
<emphasis id="B970EAF2FFC5E473FCB2AB5F2153F8F1" box="[835,1069,1843,1864]" italics="true" pageId="8" pageNumber="527">Quantula weinkauffiana</emphasis>
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– Worachak, 2014: 40–42, figs. 4.1A–B, 4.2 A–C, 4.3 B–C.
</paragraph>
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<taxonomicName id="4C044D63FFC5E473FCB2AB0126A5F83B" box="[835,987,1901,1922]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="undefined-1">
<emphasis id="B970EAF2FFC5E473FCB2AB0126DEF83B" box="[835,928,1901,1922]" italics="true" pageId="8" pageNumber="527">Quantula</emphasis>
sp. 1
</taxonomicName>
– Worachak, 2014: 43–45, figs. 4.4A, 4.5A–E, 4.6A–B.
</paragraph>
<paragraph id="8BBB36E0FFC5E473FCB2ABCA20F7F84C" blockId="8.[835,1474,1814,2038]" pageId="8" pageNumber="527">
<taxonomicName id="4C044D63FFC5E473FCB2ABCA26ABF802" box="[835,981,1958,1980]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="undefined-1">
<emphasis id="B970EAF2FFC5E473FCB2ABCA26E0F802" box="[835,926,1958,1979]" italics="true" pageId="8" pageNumber="527">Quantula</emphasis>
sp. 1
</taxonomicName>
– 
<bibRefCitation id="EF954B11FFC5E473FC01ABCA2070F802" author="Jirapatrasilp P &amp; Tongkord P &amp; Jeratthitikul E &amp; Liew TS &amp; Pholyotha A &amp; Sutcharit C &amp; Panha S" box="[1008,1294,1958,1980]" pageId="8" pageNumber="527" pagination="250 - 280" refId="ref8474" refString="Jirapatrasilp P, Tongkord P, Jeratthitikul E, Liew TS, Pholyotha A, Sutcharit C &amp; Panha S (2021) Molecular phylogeny of the limacoid snail family Dyakiidae in Southeast Asia, with the description of a new genus and species. Zoological Journal of the Linnean Society, 193 (1): 250 - 280." type="journal article" year="2021">Jirapatrasilp et al., 2021: 193</bibRefCitation>
(1), 262–263. 
<taxonomicName id="4C044D63FFC5E473FCB2ABAF269BF860" ID-CoL="78Q88" baseAuthorityName="Grey" baseAuthorityYear="1834" box="[835,997,1987,2009]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="striata">
<emphasis id="B970EAF2FFC5E473FCB2ABAF269BF860" box="[835,997,1987,2009]" italics="true" pageId="8" pageNumber="527">Quantula striata</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EF954B11FFC5E473FBF1ABA8218BF860" author="Hoompuay A" box="[1024,1269,1987,2009]" pageId="8" pageNumber="527" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016: 22–25</bibRefCitation>
, figs. 4.1–4.3. 
<taxonomicName id="4C044D63FFC5E473FCB2AB8C26ABF84F" box="[835,981,2016,2038]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="527" phylum="Mollusca" rank="species" species="undefined-6">
<emphasis id="B970EAF2FFC5E473FCB2AB8C26E0F84C" box="[835,926,2016,2037]" italics="true" pageId="8" pageNumber="527">Quantula</emphasis>
sp. 6
</taxonomicName>
– 
<bibRefCitation id="EF954B11FFC5E473FC01AB8C2198F84F" author="Hoompuay A" box="[1008,1254,2016,2038]" pageId="8" pageNumber="527" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016: 43–45</bibRefCitation>
, figs. 4.21–4.24.
</paragraph>
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Fig. 5. 
<taxonomicName id="4C044D63FFC4E472FF4FAB9F24D0F7B1" box="[190,430,2035,2056]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="528" phylum="Mollusca" rank="species" species="weinkauffiana">
<emphasis id="B970EAF2FFC4E472FF4FAB9F24D0F7B1" box="[190,430,2035,2056]" italics="true" pageId="9" pageNumber="528">Quantula weinkauffiana</emphasis>
</taxonomicName>
genital system and radula morphology, specimen NHMSU-00035. A, genital system (adapted after 
<bibRefCitation id="EF954B11FFC4E472FF80A47C2460F79F" author="Hoompuay A" box="[113,286,2064,2086]" pageId="9" pageNumber="528" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F, right side of central teeth (C) and lateral teeth (L); G, right side of marginal teeth (F &amp; G, adapted from 
<bibRefCitation id="EF954B11FFC4E472FC0DA44121D6F7FA" author="Hoompuay A" box="[1020,1192,2093,2115]" pageId="9" pageNumber="528" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
).
</paragraph>
</caption>
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Fig. 6. 
<taxonomicName id="4C044D63FFC7E471FF3EAF9B241BFBB5" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[207,357,1015,1036]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="529" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC7E471FF3EAF9B241BFBB5" box="[207,357,1015,1036]" italics="true" pageId="10" pageNumber="529">Quantula doma</emphasis>
</taxonomicName>
, 
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; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after 
<bibRefCitation id="EF954B11FFC7E471FEE7A87824BCFB93" author="Hoompuay A" box="[278,450,1044,1066]" pageId="10" pageNumber="529" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after 
<bibRefCitation id="EF954B11FFC7E471FB51A85D2032FBFE" author="Hoompuay A" box="[1184,1356,1073,1095]" pageId="10" pageNumber="529" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v).
</paragraph>
</caption>
<subSubSection id="C31E656BFFC7E471FF79A8F62442F92B" pageId="10" pageNumber="529" type="materials_examined">
<paragraph id="8BBB36E0FFC7E471FF79A8F62442F92B" blockId="10.[134,775,1178,1682]" pageId="10" pageNumber="529">
<emphasis id="B970EAF2FFC7E471FF79A8F62400FB0B" bold="true" box="[136,382,1178,1202]" pageId="10" pageNumber="529">Material examined.</emphasis>
<materialsCitation id="3B6C3CBDFFC7E471FE7BA8F724A2FAAB" ID-GBIF-Occurrence="4010534302" collectingDate="2017-10-14" latitude="14.430695" location="Buachet District" longLatPrecision="1" longitude="103.93767" municipality="Khao Sala" pageId="10" pageNumber="529" specimenCount="1" stateProvince="Surin" typeStatus="holotype">
<typeStatus id="54BF8842FFC7E471FE7BA8F72485FB0A" box="[394,507,1179,1203]" pageId="10" pageNumber="529" type="holotype">Holotype</typeStatus>
: NHMSU-00048 (
<figureCitation id="133F2A65FFC7E471FD29A8F725D7FB6A" captionStart="Fig" captionStartId="8.[835,873,1528,1550]" captionTargetBox="[835,1475,189,1512]" captionTargetId="figure-93@8.[835,1475,189,1512]" captionTargetPageId="8" captionText="Fig. 4. Shell morphology of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana, specimen NHMSU-00035 from Phanom Sawai, Surin; B–C, Q. doma, new species: B, holotype NHMSU-00048 from Khao Sala, Surin; C, NHMSU-00050 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399987" httpUri="https://zenodo.org/record/7399987/files/figure.png" pageId="10" pageNumber="529">Fig. 4B</figureCitation>
) from 
<collectingMunicipality id="6BDFAC9AFFC7E471FF0FA8D72409FB6A" box="[254,375,1211,1235]" pageId="10" pageNumber="529">Khao Sala</collectingMunicipality>
, 
<location id="8EDB603BFFC7E471FE76A8D7273BFB6A" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FE76A8D7273BFB6A" box="[391,581,1211,1235]" latitude="14.430695" longLatPrecision="1" longitude="103.93767" municipality="Khao Sala" name="Buachet District" pageId="10" pageNumber="529" stateProvince="Surin">Buachet District</location>
, 
<collectingRegion id="49C0F802FFC7E471FDA5A8D7267DFB6A" box="[596,771,1211,1235]" country="Thailand" name="Surin" pageId="10" pageNumber="529">Surin Province</collectingRegion>
; 
<geoCoordinate id="EE305027FFC7E471FF79A8B5245BFB4B" box="[136,293,1241,1266]" degrees="14" direction="north" minutes="25" orientation="latitude" pageId="10" pageNumber="529" precision="1" seconds="50.5" value="14.430695">14°25′50.5″N</geoCoordinate>
<geoCoordinate id="EE305027FFC7E471FEC0A8B524A3FB4B" box="[305,477,1241,1266]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="10" pageNumber="529" precision="1" seconds="15.6" value="103.93767">103°56′15.6″E</geoCoordinate>
; 
<date id="FFBA1020FFC7E471FE1FA8B727C7FB4A" box="[494,697,1243,1267]" pageId="10" pageNumber="529" value="2017-10-14">
<collectingDate id="EFFEE9C8FFC7E471FE1FA8B727C7FB4A" box="[494,697,1243,1267]" pageId="10" pageNumber="529" value="2017-10-14">14 October 2017</collectingDate>
</date>
; SH = 
<quantity id="4CFC9B05FFC7E471FF78A897247CFAAB" box="[137,258,1275,1299]" metricMagnitude="-2" metricUnit="m" metricValue="1.869" pageId="10" pageNumber="529" unit="mm" value="18.69">18.69 mm</quantity>
, SW = 
<quantity id="4CFC9B05FFC7E471FE93A89724A2FAAB" box="[354,476,1275,1299]" metricMagnitude="-2" metricUnit="m" metricValue="2.8609999999999998" pageId="10" pageNumber="529" unit="mm" value="28.61">28.61 mm</quantity>
</materialsCitation>
. 
<materialsCitation id="3B6C3CBDFFC7E471FE1AA89724DAFAEA" ID-GBIF-Occurrence="4010534307" collectingDate="2017-10-11" country="Thailand" county="Nong Phok District" latitude="14.322863" location="One" longLatPrecision="1" longitude="103.32594" municipality="Pha Nam Yoi" pageId="10" pageNumber="529" specimenCount="47" specimenCount-adult="25" specimenCount-juvenile="22" stateProvince="Mukdahan" typeStatus="paratype">
<typeStatus id="54BF8842FFC7E471FE1AA8972721FAAA" box="[491,607,1275,1299]" pageId="10" pageNumber="529" type="paratype">Paratypes</typeStatus>
: 47 shells (
<specimenCount id="9D02FD69FFC7E471FD1BA89625B6FA8A" pageId="10" pageNumber="529" type="adult">25 adults</specimenCount>
&amp; 
<specimenCount id="9D02FD69FFC7E471FF01A9772407FA8A" box="[240,377,1307,1331]" pageId="10" pageNumber="529" type="juvenile">22 juveniles</specimenCount>
), 28 living specimens; same data as holotype; NHMSU-00049.
</materialsCitation>
<materialsCitation id="3B6C3CBDFFC7E471FE59A9572679FA2A" ID-GBIF-Occurrence="4010534301" collectingDate="2017-10-11" country="Thailand" county="Nong Phok District" latitude="16.322863" location="One" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" pageId="10" pageNumber="529" specimenCount="47" specimenCount-adult="25" specimenCount-juvenile="22" stateProvince="Mukdahan" typeStatus="paratype">
<location id="8EDB603BFFC7E471FE59A95724AAFAEA" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FE59A95724AAFAEA" box="[424,468,1339,1363]" country="Thailand" county="Nong Phok District" latitude="16.322863" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" name="One" pageId="10" pageNumber="529" stateProvince="Mukdahan">One</location>
shell (NHMSU-00050) from 
<location id="8EDB603BFFC7E471FF76A937246AFACB" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FF76A937246AFACB" box="[135,276,1371,1395]" country="Thailand" county="Nong Phok District" latitude="16.322863" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" name="Phu Jor Kor" pageId="10" pageNumber="529" stateProvince="Mukdahan">Phu Jor Kor</location>
, 
<location id="8EDB603BFFC7E471FED2A9372771FACA" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FED2A9372771FACA" box="[291,527,1371,1395]" country="Thailand" county="Nong Phok District" latitude="16.322863" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" name="Nong Soong District" pageId="10" pageNumber="529" stateProvince="Mukdahan">Nong Soong District</location>
, 
<collectingRegion id="49C0F802FFC7E471FDEFA937267FFACA" box="[542,769,1371,1395]" country="Thailand" name="Mukdahan" pageId="10" pageNumber="529">Mukdahan Province</collectingRegion>
; (
<geoCoordinate id="EE305027FFC7E471FF61A9152458FA2B" box="[144,294,1401,1426]" degrees="16" direction="north" minutes="22" orientation="latitude" pageId="10" pageNumber="529" precision="1" seconds="22.1" value="16.372807">16°22′22.1″N</geoCoordinate>
, 
<geoCoordinate id="EE305027FFC7E471FEC0A91524ACFA2B" box="[305,466,1401,1426]" degrees="104" direction="east" minutes="22" orientation="longitude" pageId="10" pageNumber="529" precision="1" seconds="32.2" value="104.37561">104°22′32.2″E</geoCoordinate>
), 
<date id="FFBA1020FFC7E471FE15A91727E9FA2A" box="[484,663,1403,1427]" pageId="10" pageNumber="529" value="2017-10-11">
<collectingDate id="EFFEE9C8FFC7E471FE15A91727E9FA2A" box="[484,663,1403,1427]" pageId="10" pageNumber="529" value="2017-10-11">11 October 2017</collectingDate>
</date>
(
<figureCitation id="133F2A65FFC7E471FD57A9172784FA2A" box="[678,762,1403,1427]" captionStart="Fig" captionStartId="8.[835,873,1528,1550]" captionTargetBox="[835,1475,189,1512]" captionTargetId="figure-93@8.[835,1475,189,1512]" captionTargetPageId="8" captionText="Fig. 4. Shell morphology of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana, specimen NHMSU-00035 from Phanom Sawai, Surin; B–C, Q. doma, new species: B, holotype NHMSU-00048 from Khao Sala, Surin; C, NHMSU-00050 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399987" httpUri="https://zenodo.org/record/7399987/files/figure.png" pageId="10" pageNumber="529">Fig. 4C</figureCitation>
);
</materialsCitation>
<materialsCitation id="3B6C3CBDFFC7E471FF79A9F727B3FA6B" ID-GBIF-Occurrence="4010534308" collectingDate="2017-10-11" country="Thailand" county="Nong Phok District" latitude="14.322863" location="One" longLatPrecision="1" longitude="103.32594" municipality="Pha Nam Yoi" pageId="10" pageNumber="529" specimenCount="47" specimenCount-adult="25" specimenCount-juvenile="22" stateProvince="Mukdahan" typeStatus="paratype">
<location id="8EDB603BFFC7E471FF79A9F725C6FA0B" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FF79A9F725C6FA0B" box="[136,184,1435,1458]" country="Thailand" county="Nong Phok District" latitude="16.322863" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" name="Two" pageId="10" pageNumber="529" stateProvince="Mukdahan">Two</location>
adult and 
<specimenCount id="9D02FD69FFC7E471FED0A9F724D1FA0A" box="[289,431,1435,1459]" pageId="10" pageNumber="529" type="juvenile">three juvenile</specimenCount>
shells, and nine living specimens (NHMSU-00051) same locality as NHMSU-00050;
</materialsCitation>
<materialsCitation id="3B6C3CBDFFC7E471FD2BA9D72456F9EA" ID-GBIF-Occurrence="4010534306" collectingDate="2017-10-11" country="Thailand" county="Nong Phok District" latitude="14.322863" location="One" longLatPrecision="1" longitude="103.32594" municipality="Pha Nam Yoi" pageId="10" pageNumber="529" specimenCount="47" specimenCount-adult="25" specimenCount-juvenile="22" stateProvince="Mukdahan" typeStatus="paratype">
<location id="8EDB603BFFC7E471FD2BA9D72679FA6A" LSID="urn:lsid:plazi:treatment:03AD87F6FFC5E470FC10AADB2129F8BA:8EDB603BFFC7E471FD2BA9D72679FA6A" box="[730,775,1467,1491]" country="Thailand" county="Nong Phok District" latitude="16.322863" longLatPrecision="1" longitude="104.32594" municipality="Pha Nam Yoi" name="One" pageId="10" pageNumber="529" stateProvince="Mukdahan">One</location>
adult and 
<specimenCount id="9D02FD69FFC7E471FF0FA9B724F4FA4A" box="[254,394,1499,1523]" pageId="10" pageNumber="529" type="juvenile">one juvenile</specimenCount>
shells and two living specimens (NHMSU-00052); 
<collectingMunicipality id="6BDFAC9AFFC7E471FEA7A9972491F9AA" box="[342,495,1531,1555]" pageId="10" pageNumber="529">Pha Nam Yoi</collectingMunicipality>
, 
<collectingCounty id="62DA4E6CFFC7E471FE0DA99727AAF9AA" box="[508,724,1531,1555]" pageId="10" pageNumber="529">Nong Phok District</collectingCounty>
, 
<collectingRegion id="49C0F802FFC7E471FD11A997246EF98A" country="Thailand" name="Roi Et" pageId="10" pageNumber="529">Roi Et Province</collectingRegion>
, 
<collectingCountry id="F3137670FFC7E471FEEEAA7724FBF98A" box="[287,389,1563,1587]" name="Thailand" pageId="10" pageNumber="529">Thailand</collectingCountry>
; 
<geoCoordinate id="EE305027FFC7E471FE65AA752755F98B" box="[404,555,1561,1586]" degrees="16" direction="north" minutes="19" orientation="latitude" pageId="10" pageNumber="529" precision="1" seconds="22.3" value="16.322863">16°19′22.3″N</geoCoordinate>
<geoCoordinate id="EE305027FFC7E471FDC6AA7527A3F98B" box="[567,733,1561,1586]" degrees="104" direction="east" minutes="19" orientation="longitude" pageId="10" pageNumber="529" precision="1" seconds="33.4" value="104.32594">104°19′33.4″E</geoCoordinate>
; 
<date id="FFBA1020FFC7E471FD1DAA77245BF9EA" pageId="10" pageNumber="529" value="2017-10-11">
<collectingDate id="EFFEE9C8FFC7E471FD1DAA77245BF9EA" pageId="10" pageNumber="529" value="2017-10-11">11 October 2017</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B6C3CBDFFC7E471FEC2AA572446F92B" ID-GBIF-Occurrence="4010534303" collectingDate="2017-10-11" country="Thailand" county="Nong Phok District" latitude="14.322863" location="One" longLatPrecision="1" longitude="103.32594" municipality="Pha Nam Yoi" pageId="10" pageNumber="529" specimenCount="47" specimenCount-adult="25" specimenCount-juvenile="22" stateProvince="Mukdahan" typeStatus="paratype">
<specimenCount id="9D02FD69FFC7E471FEC2AA5724CFF9EA" box="[307,433,1595,1619]" pageId="10" pageNumber="529" type="adult">seven adult</specimenCount>
and 
<specimenCount id="9D02FD69FFC7E471FE1FAA5727F9F9EA" box="[494,647,1595,1619]" pageId="10" pageNumber="529" type="juvenile">three juvenile</specimenCount>
shells, and five living specimens (NHMSU-00053), same data with NHMSU-00052
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E656BFFC7E471FF79AAD62444F8AA" pageId="10" pageNumber="529" type="diagnosis">
<paragraph id="8BBB36E0FFC7E471FF79AAD62444F8AA" blockId="10.[136,774,1722,1811]" pageId="10" pageNumber="529">
<emphasis id="B970EAF2FFC7E471FF79AAD6247FF96B" bold="true" box="[136,257,1722,1746]" pageId="10" pageNumber="529">Diagnosis.</emphasis>
Last whorl angular with peripheral keel, not descending in front. Amatorial organ duct longer than amatorial organ.
</paragraph>
</subSubSection>
<subSubSection id="C31E656BFFC7E471FF7BAB572704F82A" pageId="10" pageNumber="529" type="etymology">
<paragraph id="8BBB36E0FFC7E471FF7BAB572704F82A" blockId="10.[136,775,1851,1939]" pageId="10" pageNumber="529">
<emphasis id="B970EAF2FFC7E471FF7BAB572472F8EA" bold="true" box="[138,268,1851,1875]" pageId="10" pageNumber="529">Etymology.</emphasis>
The specific epithet is derived from the Latin word ‘domus’ meaning roof, referring to the fact that its shell shape is similar to a hemispherical roof.
</paragraph>
</subSubSection>
<subSubSection id="C31E656BFFC7E470FF79ABD6274BF87A" lastPageId="11" lastPageNumber="530" pageId="10" pageNumber="529" type="description">
<paragraph id="8BBB36E0FFC7E471FF79ABD626D2FACA" blockId="10.[136,775,1978,2099]" lastBlockId="10.[835,1476,1179,1395]" pageId="10" pageNumber="529">
<emphasis id="B970EAF2FFC7E471FF79ABD62469F86B" bold="true" box="[136,279,1978,2002]" pageId="10" pageNumber="529">Description.</emphasis>
Shell (n=48, 
<specimenCount id="9D02FD69FFC7E471FE49ABD7276AF86A" box="[440,532,1979,2003]" pageId="10" pageNumber="529" type="adult">26 adult</specimenCount>
&amp; 
<specimenCount id="9D02FD69FFC7E471FDCFABD727B9F86A" box="[574,711,1979,2003]" pageId="10" pageNumber="529" type="juvenile">22 juveniles</specimenCount>
from Khao Sala (
<figureCitation id="133F2A65FFC7E471FEF4ABB72429F84A" box="[261,343,2011,2035]" captionStart="Fig" captionStartId="8.[835,873,1528,1550]" captionTargetBox="[835,1475,189,1512]" captionTargetId="figure-93@8.[835,1475,189,1512]" captionTargetPageId="8" captionText="Fig. 4. Shell morphology of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana, specimen NHMSU-00035 from Phanom Sawai, Surin; B–C, Q. doma, new species: B, holotype NHMSU-00048 from Khao Sala, Surin; C, NHMSU-00050 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399987" httpUri="https://zenodo.org/record/7399987/files/figure.png" pageId="10" pageNumber="529">Fig. 4B</figureCitation>
) and n=20, 
<specimenCount id="9D02FD69FFC7E471FE27ABB72749F84A" box="[470,567,2011,2035]" pageId="10" pageNumber="529" type="adult">11 adults</specimenCount>
&amp; 
<specimenCount id="9D02FD69FFC7E471FDA9ABB727B2F84A" box="[600,716,2011,2035]" pageId="10" pageNumber="529" type="juvenile">9 juveniles</specimenCount>
from Phu Jor Kor and Pha Nam Yoi (
<figureCitation id="133F2A65FFC7E471FE1CAB97273DF7AA" box="[493,579,2043,2067]" captionStart="Fig" captionStartId="8.[835,873,1528,1550]" captionTargetBox="[835,1475,189,1512]" captionTargetId="figure-93@8.[835,1475,189,1512]" captionTargetPageId="8" captionText="Fig. 4. Shell morphology of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana, specimen NHMSU-00035 from Phanom Sawai, Surin; B–C, Q. doma, new species: B, holotype NHMSU-00048 from Khao Sala, Surin; C, NHMSU-00050 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399987" httpUri="https://zenodo.org/record/7399987/files/figure.png" pageId="10" pageNumber="529">Fig. 4C</figureCitation>
)): Dextral, domeshaped to depressed-conic lenticular, with 6–6¼ whorls, suture shallow, growth line prominent. Last whorl angular with peripheral keel, but less sharp than 
<taxonomicName id="4C044D63FFC7E471FAF2A8D720C2FB6A" baseAuthorityName="Crosse &amp; P.Fischer" baseAuthorityYear="1863" box="[1283,1468,1211,1235]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="529" phylum="Mollusca" rank="species" species="wienkauffiana">
<emphasis id="B970EAF2FFC7E471FAF2A8D720C2FB6A" box="[1283,1468,1211,1235]" italics="true" pageId="10" pageNumber="529">Q. wienkauffiana</emphasis>
</taxonomicName>
. Shell height 
<quantity id="4CFC9B05FFC7E471FC29A8B72100FB4B" box="[984,1150,1243,1267]" metricMagnitude="-2" metricUnit="m" metricValue="1.835" metricValueMax="2.19" metricValueMin="1.48" pageId="10" pageNumber="529" unit="mm" value="18.35" valueMax="21.9" valueMin="14.8">14.8–21.9 mm</quantity>
, shell width 
<quantity id="4CFC9B05FFC7E471FAE6A8B620C0FB4B" box="[1303,1470,1242,1266]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" metricValueMax="3.53" metricValueMin="2.51" pageId="10" pageNumber="529" unit="mm" value="30.2" valueMax="35.3" valueMin="25.1">25.1–35.3 mm</quantity>
. Apex blunt, shell light brown, ventral side paler than dorsal side. Apertural lip simple and thick, and slightly expanded at columellar side in adult specimens. Umbilicus narrow and deep.
</paragraph>
<paragraph id="8BBB36E0FFC7E471FCB2A9F720BAF9AB" blockId="10.[835,1476,1435,1555]" pageId="10" pageNumber="529">
Body (n=28 from Khao Sala (
<figureCitation id="133F2A65FFC7E471FB72A9F721ABFA0A" box="[1155,1237,1435,1459]" captionStart="Fig" captionStartId="8.[135,171,1667,1689]" captionTargetBox="[135,774,189,1651]" captionTargetId="figure-7@8.[135,774,189,1651]" captionTargetPageId="8" captionText="Fig. 3. Living snails of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana (NHMSU-00035) from Phanom Sawai, Surin; B–C, Q. doma, new species: B, NHMSU-00049 from Khao Sala, Surin; C, NHMSU-00051 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399989" httpUri="https://zenodo.org/record/7399989/files/figure.png" pageId="10" pageNumber="529">Fig. 3B</figureCitation>
) and n = 11 from Phu Jor Kor and Pha Nam Yoi (
<figureCitation id="133F2A65FFC7E471FB89A9D721B3FA6A" box="[1144,1229,1467,1491]" captionStart="Fig" captionStartId="8.[135,171,1667,1689]" captionTargetBox="[135,774,189,1651]" captionTargetId="figure-7@8.[135,774,189,1651]" captionTargetPageId="8" captionText="Fig. 3. Living snails of Quantula spp. in the southern part of north-eastern Thailand. A, Q. weinkauffiana (NHMSU-00035) from Phanom Sawai, Surin; B–C, Q. doma, new species: B, NHMSU-00049 from Khao Sala, Surin; C, NHMSU-00051 from Phu Jor Kor, Mukdahan." figureDoi="http://doi.org/10.5281/zenodo.7399989" httpUri="https://zenodo.org/record/7399989/files/figure.png" pageId="10" pageNumber="529">Fig. 3C</figureCitation>
)): Living snail orange brown, with two lateral light brown or dark stripes running from base of each upper tentacle backward to pneumostome.
</paragraph>
<paragraph id="8BBB36E0FFC7E470FCB3AA57273DF99D" blockId="10.[834,1476,1595,2099]" lastBlockId="11.[113,754,1227,1572]" lastPageId="11" lastPageNumber="530" pageId="10" pageNumber="529">
Genitalia (n=13, comprising 
<specimenCount id="9D02FD69FFC7E471FB9EAA572069F9EA" box="[1135,1303,1595,1619]" pageId="10" pageNumber="529" type="generic">three specimens</specimenCount>
from Khao Sala (
<specimenCount id="9D02FD69FFC7E471FCB8AA37269EF9CA" box="[841,992,1627,1651]" pageId="10" pageNumber="529" type="generic">one specimen</specimenCount>
from 
<bibRefCitation id="EF954B11FFC7E471FBD6AA372198F9CA" author="Hoompuay A" box="[1063,1254,1627,1651]" pageId="10" pageNumber="529" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
) (
<figureCitation id="133F2A65FFC7E471FB0EAA37200BF9CA" box="[1279,1397,1627,1651]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="10" pageNumber="529">Fig. 6A–E</figureCitation>
), 
<specimenCount id="9D02FD69FFC7E471FA7AAA3726CCF92A" pageId="10" pageNumber="529" type="generic">three specimens</specimenCount>
from Phu Jor Kor, and 
<specimenCount id="9D02FD69FFC7E471FB58AA172025F92A" box="[1193,1371,1659,1683]" pageId="10" pageNumber="529" type="generic">seven specimens</specimenCount>
from Pha Nam Yoi (
<specimenCount id="9D02FD69FFC7E471FC5FAAF72137F90A" box="[942,1097,1691,1715]" pageId="10" pageNumber="529" type="generic">five specimens</specimenCount>
from 
<bibRefCitation id="EF954B11FFC7E471FB74AAF72045F90A" author="Hoompuay A" box="[1157,1339,1691,1715]" pageId="10" pageNumber="529" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
) (
<figureCitation id="133F2A65FFC7E471FABEAAF720C8F90A" box="[1359,1462,1691,1715]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="10" pageNumber="529">Fig. 6F–J</figureCitation>
): Penis short, cylindrical, inner wall with robust longitudinal penial pilaster alternating with 1–2 thin longitudinal pilasters (
<figureCitation id="133F2A65FFC7E471FCBBAA9726DEF8AA" box="[842,928,1787,1811]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="10" pageNumber="529">Fig. 6D</figureCitation>
), whereas specimens from Phu Jor Kor sculptured with irregular undulate penial pilasters arranged in inclined and longitudinal rows (
<figureCitation id="133F2A65FFC7E471FBBDAB5721D9F8EA" box="[1100,1191,1851,1875]" captionStart="Fig" captionStartId="11.[113,148,1093,1115]" captionTargetBox="[113,1451,192,1077]" captionTargetId="figure-69@11.[113,1451,186,1080]" captionTargetPageId="11" captionText="Fig. 7. Radula morphology of Quantula doma, new species. A–B, paratype NHMSU-00049 from Khao Sala, Surin Province; A, left side of marginal teeth; B, left side of central teeth and lateral teeth. C–D, NHMSU-00051 from Phu Jor Kor, Mukdahan Province; C, right side of central teeth and lateral teeth. D, right side of marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7399997" httpUri="https://zenodo.org/record/7399997/files/figure.png" pageId="10" pageNumber="529">Fig. 7D</figureCitation>
). Epiphallus shorter than penis. Vas deferens longer than penis. Gametolytic duct connecting to basal part of amatorial organ, basally bulbous, distally with short and small duct. Tapering gametolytic sac long, robust (
<figureCitation id="133F2A65FFC7E471FBF1ABD72129F86A" box="[1024,1111,1979,2003]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="10" pageNumber="529">Fig. 6A</figureCitation>
). Amatorial organ elliptical, with inner wall smooth. Papilla of amatorial organ strong, long, reaching junction of gametolytic duct. Corneous spike present (
<figureCitation id="133F2A65FFC7E471FCBBA47726BDF78A" box="[842,963,2075,2099]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="10" pageNumber="529">Fig. 6B–C</figureCitation>
). Amatorial organ duct longer than amatorial organ. Vagina and free oviduct shorter than penis. Inner wall of vagina smooth, with 3–4 thin longitudinal pilasters on median part (
<figureCitation id="133F2A65FFC6E470FED4A9672407FA9A" box="[293,377,1291,1315]" captionStart="Fig" captionStartId="10.[136,171,1015,1037]" captionTargetBox="[136,1476,190,999]" captionTargetId="figure-210@10.[136,1488,182,999]" captionTargetPageId="10" captionText="Fig. 6. Quantula doma, new species; A–E, genital system of paratype NHMSU-00049 from Khao Sala, Surin Province. A, genital system (adapted after Hoompuay, 2016); B–E, inner structure of genital system; B and C, amatorial organ (am); D, penis (p); E, vagina (v); F–J, genital system of NHMSU-00051 from Phu Jor Kor, Mukdahan Province. F, genital system (adapted after Hoompuay, 2016); G–J, inner structure of genital system; G and H, amatorial organ (am); I, penis (p); J, vagina (v)." figureDoi="http://doi.org/10.5281/zenodo.7399993" httpUri="https://zenodo.org/record/7399993/files/figure.png" pageId="11" pageNumber="530">Fig. 6E</figureCitation>
). Specimens from Phu Jor Kor has gametolytic duct connecting to middle part of amatorial organ. Gametolytic sac cylindrical, long. Amatorial organ long, cylindrical, with inner wall sculptured with irregular undulated short pilasters. Papilla of amatorial organ robust, cylindrical, distally finger-shaped, without terminal corneous spike. Vagina longer than penis, internally with four to five longitudinal pilasters. Amatorial organ duct nearly as long as, or slightly longer than amatorial organ.
</paragraph>
<caption id="DF7B6668FFC6E470FF80A8292669FB2D" ID-DOI="http://doi.org/10.5281/zenodo.7399997" ID-Zenodo-Dep="7399997" httpUri="https://zenodo.org/record/7399997/files/figure.png" pageId="11" pageNumber="530" startId="11.[113,148,1093,1115]" targetBox="[113,1451,192,1077]" targetPageId="11">
<paragraph id="8BBB36E0FFC6E470FF80A8292669FB2D" blockId="11.[113,1453,1093,1172]" pageId="11" pageNumber="530">
Fig. 7. Radula morphology of 
<taxonomicName id="4C044D63FFC6E470FE6AA829274CFBE3" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[411,562,1093,1114]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC6E470FE6AA829274CFBE3" box="[411,562,1093,1114]" italics="true" pageId="11" pageNumber="530">Quantula doma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2435789FFC6E470FDB1A82927C9FBE2" box="[576,695,1093,1115]" pageId="11" pageNumber="530" rank="species">new species</taxonomicNameLabel>
. A–B, paratype NHMSU-00049 from Khao Sala, Surin Province; A, left side of marginal teeth; B, left side of central teeth and lateral teeth. C–D, NHMSU-00051 from Phu Jor Kor, Mukdahan Province; C, right side of central teeth and lateral teeth. D, right side of marginal teeth.
</paragraph>
</caption>
<paragraph id="8BBB36E0FFC6E470FF80AA27274BF87A" blockId="11.[112,754,1611,1987]" pageId="11" pageNumber="530">
Radula (n=13, comprising 
<specimenCount id="9D02FD69FFC6E470FE63AA272740F9DA" box="[402,574,1611,1635]" pageId="11" pageNumber="530" type="generic">three specimens</specimenCount>
from Khao Sala (
<specimenCount id="9D02FD69FFC6E470FF8BAA07246FF93A" box="[122,273,1643,1667]" pageId="11" pageNumber="530" type="generic">one specimen</specimenCount>
from 
<bibRefCitation id="EF954B11FFC6E470FEABAA002767F93A" author="Hoompuay A" box="[346,537,1643,1668]" pageId="11" pageNumber="530" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
) (
<figureCitation id="133F2A65FFC6E470FDC3AA0727D7F93D" box="[562,681,1643,1668]" captionStart="Fig" captionStartId="11.[113,148,1093,1115]" captionTargetBox="[113,1451,192,1077]" captionTargetId="figure-69@11.[113,1451,186,1080]" captionTargetPageId="11" captionText="Fig. 7. Radula morphology of Quantula doma, new species. A–B, paratype NHMSU-00049 from Khao Sala, Surin Province; A, left side of marginal teeth; B, left side of central teeth and lateral teeth. C–D, NHMSU-00051 from Phu Jor Kor, Mukdahan Province; C, right side of central teeth and lateral teeth. D, right side of marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7399997" httpUri="https://zenodo.org/record/7399997/files/figure.png" pageId="11" pageNumber="530">Fig. 7A–B</figureCitation>
) 
<specimenCount id="9D02FD69FFC6E470FD4AAA07259CF91A" pageId="11" pageNumber="530" type="generic">three specimens</specimenCount>
from Phu Jor Kor, and 
<specimenCount id="9D02FD69FFC6E470FE09AAE027CFF91A" box="[504,689,1675,1699]" pageId="11" pageNumber="530" type="generic">seven specimens</specimenCount>
from Pha Nam Yoi (
<specimenCount id="9D02FD69FFC6E470FEE0AAC724CFF97A" box="[273,433,1707,1731]" pageId="11" pageNumber="530" type="generic">five specimens</specimenCount>
from 
<bibRefCitation id="EF954B11FFC6E470FE02AAC027D1F97A" author="Hoompuay A" box="[499,687,1707,1732]" pageId="11" pageNumber="530" refId="ref8324" refString="Hoompuay A (2016) Taxonomy of land snail family Dyakiidae Gudde &amp; Woodward, 1921 in the northeastern of Thailand. Unpublished Master's Thesis. Department of Biology, Faculty of Science, Mahasarakham University, Thailand, 123 pp. (In Thai with English abstract)" type="book" year="2016">Hoompuay, 2016</bibRefCitation>
) (
<figureCitation id="133F2A65FFC6E470FD34AAC725C8F95A" captionStart="Fig" captionStartId="11.[113,148,1093,1115]" captionTargetBox="[113,1451,192,1077]" captionTargetId="figure-69@11.[113,1451,186,1080]" captionTargetPageId="11" captionText="Fig. 7. Radula morphology of Quantula doma, new species. A–B, paratype NHMSU-00049 from Khao Sala, Surin Province; A, left side of marginal teeth; B, left side of central teeth and lateral teeth. C–D, NHMSU-00051 from Phu Jor Kor, Mukdahan Province; C, right side of central teeth and lateral teeth. D, right side of marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7399997" httpUri="https://zenodo.org/record/7399997/files/figure.png" pageId="11" pageNumber="530">Fig. 7C–D</figureCitation>
): with 83–90 rows of teeth. Each row with 83–123 teeth, formula: (24–42) + (17–19) + 1 + (17–19) + (24–42) in specimens from Khao Sala, and approximately 70–82 rows of teeth, each of which comprises 91–123 teeth. Fomula: (31–43) + (14–18) + 1 + (14–18) + (31–43) in specimens from Phu Jor Kor and Pha Nam Yoi. Central and lateral teeth lanceolate, tricuspid. Marginal teeth bicuspid and gradually changing to unicuspid and sword shaped.
</paragraph>
</subSubSection>
<subSubSection id="C31E656BFFC6E470FF82AB87279EF7FA" pageId="11" pageNumber="530" type="distribution">
<paragraph id="8BBB36E0FFC6E470FF82AB87279EF7FA" blockId="11.[113,752,2027,2115]" pageId="11" pageNumber="530">
<emphasis id="B970EAF2FFC6E470FF82AB872472F7BA" bold="true" box="[115,268,2027,2051]" pageId="11" pageNumber="530">Distribution.</emphasis>
<taxonomicName id="4C044D63FFC6E470FEE4AB8724BBF7BA" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[277,453,2027,2051]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC6E470FEE4AB8724BBF7BA" box="[277,453,2027,2051]" italics="true" pageId="11" pageNumber="530">Quantula doma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2435789FFC6E470FE26AB80271DF7BA" box="[471,611,2027,2051]" pageId="11" pageNumber="530" rank="species">new species</taxonomicNameLabel>
, is found in sandstone mountains of the southern part of north-eastern 
<collectingCountry id="F3137670FFC6E470FF80A44725ADF7FA" box="[113,211,2091,2115]" name="Thailand" pageId="11" pageNumber="530">Thailand</collectingCountry>
, in the 
<collectingRegion id="49C0F802FFC6E470FED7A447241CF7FA" box="[294,354,2091,2115]" country="Thailand" name="Surin" pageId="11" pageNumber="530">Surin</collectingRegion>
, 
<collectingRegion id="49C0F802FFC6E470FE9EA44724C6F7FA" box="[367,440,2091,2115]" country="Thailand" name="Roi Et" pageId="11" pageNumber="530">Roi Et</collectingRegion>
, and 
<collectingRegion id="49C0F802FFC6E470FE04A4472719F7FA" box="[501,615,2091,2115]" country="Thailand" name="Mukdahan" pageId="11" pageNumber="530">Mukdahan</collectingRegion>
Provinces.
</paragraph>
</subSubSection>
<subSubSection id="C31E656BFFC6E470FCDCA8A72129F8BA" pageId="11" pageNumber="530" type="discussion">
<paragraph id="8BBB36E0FFC6E470FCDCA8A72129F8BA" blockId="11.[811,1452,1227,1795]" pageId="11" pageNumber="530">
<emphasis id="B970EAF2FFC6E470FCDCA8A726E7FB5A" bold="true" box="[813,921,1227,1251]" pageId="11" pageNumber="530">Remarks.</emphasis>
According to the genetic distance and phylogenetic analysis, 
<taxonomicName id="4C044D63FFC6E470FC7EA887268AFABA" authorityName="Baker" authorityYear="1941" box="[911,1012,1259,1283]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="genus">
<emphasis id="B970EAF2FFC6E470FC7EA887268AFABA" box="[911,1012,1259,1283]" italics="true" pageId="11" pageNumber="530">Quantula</emphasis>
</taxonomicName>
“sp. 1” from Khao Sala in 
<bibRefCitation id="EF954B11FFC6E470FAE1A88726EEFA9A" author="Jirapatrasilp P &amp; Tongkord P &amp; Jeratthitikul E &amp; Liew TS &amp; Pholyotha A &amp; Sutcharit C &amp; Panha S" pageId="11" pageNumber="530" pagination="250 - 280" refId="ref8474" refString="Jirapatrasilp P, Tongkord P, Jeratthitikul E, Liew TS, Pholyotha A, Sutcharit C &amp; Panha S (2021) Molecular phylogeny of the limacoid snail family Dyakiidae in Southeast Asia, with the description of a new genus and species. Zoological Journal of the Linnean Society, 193 (1): 250 - 280." type="journal article" year="2021">Jirapatrasilp et al. (2021)</bibRefCitation>
(specimen no. H60, COI and 28s accession numbers 
<accessionNumber id="9457AB03FFC6E470FCDAA94726DDFAFA" box="[811,931,1323,1347]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT803088" pageId="11" pageNumber="530">MT803088</accessionNumber>
and 
<accessionNumber id="9457AB03FFC6E470FC2BA947212AFAFA" box="[986,1108,1323,1347]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT741936" pageId="11" pageNumber="530">MT741936</accessionNumber>
) is clustered with the 
<taxonomicName id="4C044D63FFC6E470FAB8A94720DBFAFA" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[1353,1445,1323,1347]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC6E470FAB8A94720DBFAFA" box="[1353,1445,1323,1347]" italics="true" pageId="11" pageNumber="530">Q. doma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2435789FFC6E470FCDCA92726D4FADA" box="[813,938,1355,1379]" pageId="11" pageNumber="530" rank="species">new species</taxonomicNameLabel>
(specimen no. KSH and KSL). Genetic distances within the Khao Sala population are around 0.5% (
<tableCitation id="C686035BFFC6E470FAB9A90720E2FA3A" box="[1352,1436,1387,1411]" captionStart="Table 2" captionStartId="5.[111,166,523,546]" captionTargetPageId="5" captionText="Table 2. Pairwise genetic distance values for different species in this study." httpUri="http://table.plazi.org/id/DF7B6668FFC8E47EFF9EAE672628FD9B" pageId="11" pageNumber="530" tableUuid="DF7B6668FFC8E47EFF9EAE672628FD9B">Table 2</tableCitation>
), and all specimens have rather strong support (
<figureCitation id="133F2A65FFC6E470FAC3A9E7200BFA1A" box="[1330,1397,1419,1443]" captionStart="Fig" captionStartId="7.[114,149,1031,1053]" captionTargetBox="[115,1452,190,1017]" captionTargetId="figure-13@7.[101,1465,178,1024]" captionTargetPageId="7" captionText="Fig. 2. Bayesian inference (BI) tree of Quantula in this study (Blue colour) determined from 28s and COI genes. Numbers at nodes indicate branch support based on posterior probability (BI)/bootstrapping (ML)/bootstrapping (NJ). # Indicates branches that received &lt;0.50 of posterior probability and received &lt;50% of bootstrap support. - indicates non-supported branches by bootstrap. Scale bar = 0.1 substitutions/site." figureDoi="http://doi.org/10.5281/zenodo.7399985" httpUri="https://zenodo.org/record/7399985/files/figure.png" pageId="11" pageNumber="530">Fig. 2</figureCitation>
) (BI posterior probability of 1.00 and 100% for ML, and 99% for NJ bootstrap replicate), based on which we conclude that specimen H60, KSH and KSL represent the same species as 
<taxonomicName id="4C044D63FFC6E470FCDDAA6726F5F99A" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[812,907,1547,1571]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC6E470FCDDAA6726F5F99A" box="[812,907,1547,1571]" italics="true" pageId="11" pageNumber="530">Q. doma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2435789FFC6E470FC6AAA67215FF99A" box="[923,1057,1547,1571]" pageId="11" pageNumber="530" rank="species">new species</taxonomicNameLabel>
. Although conchologically this new species looks similar to 
<taxonomicName id="4C044D63FFC6E470FBCDAA4720D5F9FA" authority="Thach &amp; Huber 2018" authorityName="Thach &amp; Huber" authorityYear="2018" box="[1084,1451,1579,1603]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="simonei">
<emphasis id="B970EAF2FFC6E470FBCDAA472131F9FB" box="[1084,1103,1579,1602]" italics="true" pageId="11" pageNumber="530">Q</emphasis>
. 
<emphasis id="B970EAF2FFC6E470FB91AA4021CDF9FA" box="[1120,1203,1580,1603]" italics="true" pageId="11" pageNumber="530">simonei</emphasis>
Thach &amp; Huber 2018
</taxonomicName>
from 
<collectingCountry id="F3137670FFC6E470FC96AA2726E5F9DB" box="[871,923,1611,1634]" name="Laos" pageId="11" pageNumber="530">Laos</collectingCountry>
(see 
<bibRefCitation id="EF954B11FFC6E470FC23AA272125F9DA" author="Thach NN" box="[978,1115,1611,1635]" pageId="11" pageNumber="530" refId="ref9596" refString="Thach NN (2018) New shells of South Asia. Seashells - Freshwater &amp; Land Snails. 48 HR Books Company, Akron, 173 pp." type="book" year="2018">Thach, 2018</bibRefCitation>
: plate 49, figs. 580, 581 top), in 
<taxonomicName id="4C044D63FFC6E470FCDCAA0726F7F93A" authority="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan, 2022" authorityName="Deeprom &amp; Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2022" box="[813,905,1643,1667]" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="doma" status="sp. nov.">
<emphasis id="B970EAF2FFC6E470FCDCAA0726F7F93A" box="[813,905,1643,1667]" italics="true" pageId="11" pageNumber="530">Q. doma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2435789FFC6E470FC66AA072165F93A" box="[919,1051,1643,1667]" pageId="11" pageNumber="530" rank="species">new species</taxonomicNameLabel>
, the last whorl is angulate with weak peripheral keel and has an open umbilicus, whereas in 
<taxonomicName id="4C044D63FFC6E470FA60AAE726FEF97A" authorityName="Thach &amp; Huber" authorityYear="2018" class="Gastropoda" family="Dyakiidae" genus="Quantula" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="530" phylum="Mollusca" rank="species" species="simonei">
<emphasis id="B970EAF2FFC6E470FA60AAE720DAF91B" box="[1425,1444,1675,1698]" italics="true" pageId="11" pageNumber="530">Q</emphasis>
. 
<emphasis id="B970EAF2FFC6E470FCDCAAC026FEF97A" box="[813,896,1708,1731]" italics="true" pageId="11" pageNumber="530">simonei</emphasis>
</taxonomicName>
, the last whorl is well-inflated without peripheral keel, and the umbilicus is mostly covered by the columella portion of the apertural lip.
</paragraph>
</subSubSection>
</treatment>
</document>