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<document id="70068BDE264CE9403A038846855A6FCF" ID-DOI="10.5281/zenodo.214602" ID-GBIF-Dataset="f0096116-c79f-41cd-8376-35587cbe9fcd" ID-ISSN="1175-5326" ID-Zenodo-Dep="214602" ID-ZooBank="AF03126F-70F3-4696-A73B-0DF6B6C494CE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460520320751" checkinUser="plazi" docAuthor="M, Alexander Feijoo &amp; Celis, Liliana V." docDate="2012" docId="03AD87DC521DFFC511EFFF64FA1DFDC3" docLanguage="en" docName="zt03458p119.pdf" docOrigin="Zootaxa 3458" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Andiodrilus" docType="key" docVersion="10" lastPageNumber="117" masterDocId="FF94FFA45210FFCB1178FFF3FF81FFA7" masterDocTitle="New species of earthworms (Oligochaeta: Glossoscolecidae) in the Amazon region of Colombia" masterLastPageNumber="119" masterPageNumber="103" pageNumber="116" updateTime="1698314519981" updateUser="plazi">
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<mods:title id="19241152C689D6F01605B05E9EF8DA00">New species of earthworms (Oligochaeta: Glossoscolecidae) in the Amazon region of Colombia</mods:title>
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<mods:namePart id="F991D5306E26872C004E3D3F6E191B25">M, Alexander Feijoo</mods:namePart>
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<mods:namePart id="EBD4F6016F977BA9BF9AC133EFCD0716">Celis, Liliana V.</mods:namePart>
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<mods:date id="12BA70DB312BE71BC4021D9833A61936">2012</mods:date>
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<treatment id="03AD87DC521DFFC511EFFF64FA1DFDC3" ID-DOI="http://doi.org/10.5281/zenodo.5625210" ID-GBIF-Taxon="119525617" ID-Zenodo-Dep="5625210" LSID="urn:lsid:plazi:treatment:03AD87DC521DFFC511EFFF64FA1DFDC3" httpUri="http://treatment.plazi.org/id/03AD87DC521DFFC511EFFF64FA1DFDC3" lastPageId="14" lastPageNumber="117" pageId="13" pageNumber="116">
<subSubSection id="C31E6541521DFFC611EFFF64FE5CFF16" box="[151,477,151,177]" pageId="13" pageNumber="116" type="nomenclature">
<paragraph id="8BBB36CA521DFFC611EFFF64FE5CFF16" blockId="13.[151,477,151,177]" box="[151,477,151,177]" pageId="13" pageNumber="116">
<heading id="D0F381A6521DFFC611EFFF64FE5CFF16" bold="true" box="[151,477,151,177]" fontSize="11" level="1" pageId="13" pageNumber="116" reason="1">
<emphasis id="B970EAD8521DFFC611EFFF64FE5CFF16" bold="true" box="[151,477,151,177]" pageId="13" pageNumber="116">
Key to
<taxonomicName id="4C044D49521DFFC61188FF64FEFEFF16" box="[240,383,151,177]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="genus">
<emphasis id="B970EAD8521DFFC61188FF64FEFEFF16" bold="true" box="[240,383,151,177]" italics="true" pageId="13" pageNumber="116">Andiodrilus</emphasis>
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species
</emphasis>
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</paragraph>
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<subSubSection id="C31E6541521DFFC511EFFF2DFA1DFDC3" lastPageId="14" lastPageNumber="117" pageId="13" pageNumber="116" type="key">
<keyStep id="30F02E6F521DFFC611EFFF2DFA1DFEB7" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFF2DFA1DFF54" blockId="13.[151,1436,222,2033]" box="[151,1436,222,243]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFF2DFA1DFF54" box="[151,1436,222,243]" pageId="13" pageNumber="116">1 One pair of spermathecae in 8/9......................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFF08FA1DFEB7" blockId="13.[151,1436,222,2033]" box="[151,1436,251,272]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFF08FA1DFEB7" box="[151,1436,251,272]" pageId="13" pageNumber="116">- Three pairs of spermathecae in 6/7-8/9................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFEEAFA1DFEEC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFEEAFA1DFE89" blockId="13.[151,1436,222,2033]" box="[151,1436,281,302]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFEEAFA1DFE89" box="[151,1436,281,302]" pageId="13" pageNumber="116">
2 Spermathecal pores in line ab.............................................................
<taxonomicName id="4C044D49521DFFC6159EFEEAFA1DFE89" authority="Zicsi, 1989" authorityName="Zicsi" authorityYear="1989" box="[1254,1436,281,302]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="lacteus">
<emphasis id="B970EAD8521DFFC6159EFEEAFAABFE89" box="[1254,1322,281,302]" italics="true" pageId="13" pageNumber="116">lacteus</emphasis>
<bibRefCitation id="EF954B3B521DFFC61448FEEAFA1DFE89" author="Zicsi" box="[1328,1436,281,302]" pageId="13" pageNumber="116" refString="Zicsi, A. (1989) Uber drei neue Andiodrilus - Arten aus Ecuador (Oligochaeta: Glossoscolecidae) (Regenwurmer aus Sudamerika 12). Revue Suisse de Zoologie, 96, 4, 771 - 777." type="journal article" year="1989">Zicsi, 1989</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFEC5FA1DFEEC" blockId="13.[151,1436,222,2033]" box="[151,1436,310,331]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFEC5FA1DFEEC" box="[151,1436,310,331]" pageId="13" pageNumber="116">- Spermathecal pores in line c............................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFEA0FA1DFE21" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFEA0FA1DFECF" blockId="13.[151,1436,222,2033]" box="[151,1436,339,361]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFEA0FA1DFECF" box="[151,1436,339,361]" pageId="13" pageNumber="116">
3 Tubercula pubertatis in ½ 20- ½ 24...................................................
<taxonomicName id="4C044D49521DFFC615DBFEA7FA1DFECF" authority="Michaelsen, 1900" authorityName="Michaelsen" authorityYear="1900" box="[1187,1436,339,361]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="affinis">
<emphasis id="B970EAD8521DFFC615DBFEA7FB5EFECE" box="[1187,1247,340,361]" italics="true" pageId="13" pageNumber="116">affinis</emphasis>
<bibRefCitation id="EF954B3B521DFFC6159DFEA0FA1DFECF" author="Michaelsen" box="[1253,1436,339,360]" pageId="13" pageNumber="116" refString="Michaelsen, W. (1900 b) Die Terricolenfauna Columbiens. Archiv fur Naturgeschichte, 66, 231 - 266." type="journal article" year="1900" yearSuffix="b">Michaelsen, 1900b</bibRefCitation>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFE82FA1DFE21" blockId="13.[151,1436,222,2033]" box="[151,1436,369,390]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFE82FA1DFE21" box="[151,1436,369,390]" pageId="13" pageNumber="116">- Tubercula pubertatis in 21-22........................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFE7DFA1DFE67" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFE7DFA1DFE04" blockId="13.[151,1436,222,2033]" box="[151,1436,398,419]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFE7DFA1DFE04" box="[151,1436,398,419]" pageId="13" pageNumber="116">
4 Clitellum between 16-25................................................................
<taxonomicName id="4C044D49521DFFC615A7FE7DFA1DFE04" authority="Zicsi, 1989" authorityName="Zicsi" authorityYear="1989" box="[1247,1436,398,419]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="szekelyi">
<emphasis id="B970EAD8521DFFC615A7FE7DFAABFE04" box="[1247,1322,398,419]" italics="true" pageId="13" pageNumber="116">szekelyi</emphasis>
<bibRefCitation id="EF954B3B521DFFC61448FE7DFA1DFE04" author="Zicsi" box="[1328,1436,398,419]" pageId="13" pageNumber="116" refString="Zicsi, A. (1989) Uber drei neue Andiodrilus - Arten aus Ecuador (Oligochaeta: Glossoscolecidae) (Regenwurmer aus Sudamerika 12). Revue Suisse de Zoologie, 96, 4, 771 - 777." type="journal article" year="1989">Zicsi, 1989</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFE58FA1DFE67" blockId="13.[151,1436,222,2033]" box="[151,1436,427,448]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFE58FA1DFE67" box="[151,1436,427,448]" pageId="13" pageNumber="116">- Clitellum between 16-26.............................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFE3AFA1DFE5C" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFE3AFA1DFE79" blockId="13.[151,1436,222,2033]" box="[151,1436,457,478]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFE3AFA1DFE79" box="[151,1436,457,478]" pageId="13" pageNumber="116">
5 Clitellum between 16-26................................................................
<taxonomicName id="4C044D49521DFFC61599FE3AFA1DFE79" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1249,1436,457,478]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="boruka">
<emphasis id="B970EAD8521DFFC61599FE3AFAA4FE79" box="[1249,1317,457,478]" italics="true" pageId="13" pageNumber="116">boruka</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FE3AFA1DFE79" author="Righi" box="[1323,1436,457,478]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFE15FA1DFE5C" blockId="13.[151,1436,222,2033]" box="[151,1436,486,507]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFE15FA1DFE5C" box="[151,1436,486,507]" pageId="13" pageNumber="116">- Clitellum between 17-29.............................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFDF0FA1DFD91" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFDF0FA1DFDBF" blockId="13.[151,1436,222,2033]" box="[151,1436,515,537]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFDF0FA1DFDBF" box="[151,1436,515,537]" pageId="13" pageNumber="116">
<specimenCount id="9D02FD43521DFFC611EFFDF0FE93FDBF" box="[151,274,515,536]" pageId="13" pageNumber="116" type="male">6 Male</specimenCount>
pores in 20..................................................................
<taxonomicName id="4C044D49521DFFC615DEFDF7FA1DFDBF" authority="Michaelsen, 1900" authorityName="Michaelsen" authorityYear="1900" box="[1190,1436,515,537]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="major">
<emphasis id="B970EAD8521DFFC615DEFDF7FB5EFDBE" box="[1190,1247,516,537]" italics="true" pageId="13" pageNumber="116">major</emphasis>
<bibRefCitation id="EF954B3B521DFFC6159DFDF0FA1DFDBF" author="Michaelsen" box="[1253,1436,515,536]" pageId="13" pageNumber="116" refString="Michaelsen, W. (1900 b) Die Terricolenfauna Columbiens. Archiv fur Naturgeschichte, 66, 231 - 266." type="journal article" year="1900" yearSuffix="b">Michaelsen, 1900b</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFDD2FA1DFD91" blockId="13.[151,1436,222,2033]" box="[151,1436,545,566]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFDD2FA1DFD91" box="[151,1436,545,566]" pageId="13" pageNumber="116">- Male pores in 20/21.................................................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFDCDFA1AFDD6" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFDCDFA1DFDF4" blockId="13.[151,1436,222,2033]" box="[151,1436,574,595]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFDCDFA1DFDF4" box="[151,1436,574,595]" pageId="13" pageNumber="116">
7 Caecum absent..........................................................................
<taxonomicName id="4C044D49521DFFC61587FDCDFA1DFDF4" authority="Righi, 1984" authorityName="Righi" authorityYear="1984" box="[1279,1436,574,595]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="pipi">
<emphasis id="B970EAD8521DFFC61587FDCDFAA5FDF4" box="[1279,1316,574,595]" italics="true" pageId="13" pageNumber="116">pipi</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FDCDFA1DFDF4" author="Righi" box="[1323,1436,574,595]" pageId="13" pageNumber="116" refString="Righi, G. (1984) Pontoscolex (Oligochaeta, Glossoscolecidae) a new evaluation. Studies on Neotropical Fauna and Environment, 19, 159 - 177." type="journal article" year="1984">Righi, 1984</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFDAFFA1AFDD6" blockId="13.[151,1436,222,2033]" box="[151,1435,604,625]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFDAFFA1AFDD6" box="[151,1435,604,625]" pageId="13" pageNumber="116">- Caecum present in 22................................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFD8AFA1DFD0C" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFD8AFA1AFD29" blockId="13.[151,1436,222,2033]" box="[151,1435,633,654]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFD8AFA1AFD29" box="[151,1435,633,654]" pageId="13" pageNumber="116">8 Tubercula pubertatis between 20-24, ½ 25............................................................... 11</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFD65FA1DFD0C" blockId="13.[151,1436,222,2033]" box="[151,1436,662,683]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFD65FA1DFD0C" box="[151,1436,662,683]" pageId="13" pageNumber="116">
- Tubercula pubertatis between 2/
<date id="FFBA100A521DFFC6136EFD65FDD3FD0C" box="[534,594,662,683]" pageId="13" pageNumber="116" value="1920-03-02">3 20-2</date>
/3 23............................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFD47FA1DFD41" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFD47FA1DFD6E" blockId="13.[151,1436,222,2033]" box="[151,1436,692,713]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFD47FA1DFD6E" box="[151,1436,692,713]" pageId="13" pageNumber="116">
9 Seminal vesicles absent..................................................................
<taxonomicName id="4C044D49521DFFC61597FD47FA1DFD6E" authority="Righi, 1971" authorityName="Righi" authorityYear="1971" box="[1263,1436,692,713]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="icomi">
<emphasis id="B970EAD8521DFFC61597FD47FAA5FD6E" box="[1263,1316,692,713]" italics="true" pageId="13" pageNumber="116">icomi</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FD47FA1DFD6E" author="Righi" box="[1323,1436,692,713]" pageId="13" pageNumber="116" refString="Righi, G. (1971) Sobre la familia Glossoscolecidae (Oligochaeta) no Brasil. Arquivos de Zoologia Museo de Zoologia da Universidade de Sao Paulo, 20, 1 - 95." type="journal article" year="1971">Righi, 1971</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFD22FA1DFD41" blockId="13.[151,1436,222,2033]" box="[151,1436,721,742]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFD22FA1DFD41" box="[151,1436,721,742]" pageId="13" pageNumber="116">- Seminal vesicles present.............................................................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFD1DFA1DFC86" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFD1DFA1DFCA4" blockId="13.[151,1436,222,2033]" box="[151,1436,750,771]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFD1DFA1DFCA4" box="[151,1436,750,771]" pageId="13" pageNumber="116">
10 Spermathecae withou diverticulum.......................................................
<taxonomicName id="4C044D49521DFFC615AEFD1DFA1DFCA4" authority="Zicsi, 1988" authorityName="Zicsi" authorityYear="1988" box="[1238,1436,750,771]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="vaucheri">
<emphasis id="B970EAD8521DFFC615AEFD1DFAABFCA4" box="[1238,1322,750,771]" italics="true" pageId="13" pageNumber="116">vaucheri</emphasis>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFCFFFA1DFC86" blockId="13.[151,1436,222,2033]" box="[151,1436,780,801]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFCFFFA1DFC86" box="[151,1436,780,801]" pageId="13" pageNumber="116">- Spermathecae with diverticulum....................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFCDAFA1DFCFC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFCDAFA1DFC99" blockId="13.[151,1436,222,2033]" box="[151,1436,809,830]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFCDAFA1DFC99" box="[151,1436,809,830]" pageId="13" pageNumber="116">
<specimenCount id="9D02FD43521DFFC611EFFCDAFE93FC99" box="[151,274,809,830]" pageId="13" pageNumber="116" type="male">11 Male</specimenCount>
pores between 19/20................................................................
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<emphasis id="B970EAD8521DFFC6158CFCDAFAA5FC99" box="[1268,1316,809,830]" italics="true" pageId="13" pageNumber="116">katio</emphasis>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFCB5FA1DFCFC" blockId="13.[151,1436,222,2033]" box="[151,1436,838,859]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFCB5FA1DFCFC" box="[151,1436,838,859]" pageId="13" pageNumber="116">- Male pores between in ½ 21........................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFC97FA1DFC31" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFC97FA1DFCDE" blockId="13.[151,1436,222,2033]" box="[151,1436,868,889]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC97FA1DFCDE" box="[151,1436,868,889]" pageId="13" pageNumber="116">
12 Clitellum between 15, 16-26, 27........................................................
<taxonomicName id="4C044D49521DFFC615B2FC97FA1DFCDE" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1226,1436,868,889]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="achagua">
<emphasis id="B970EAD8521DFFC615B2FC97FA9CFCDE" box="[1226,1309,868,889]" italics="true" pageId="13" pageNumber="116">achagua</emphasis>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFC72FA1DFC31" blockId="13.[151,1436,222,2033]" box="[151,1436,897,918]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC72FA1DFC31" box="[151,1436,897,918]" pageId="13" pageNumber="116">- Clitellum between 16-27............................................................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFC6DFA1DFC76" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFC6DFA1DFC14" blockId="13.[151,1436,222,2033]" box="[151,1436,926,947]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC6DFA1DFC14" box="[151,1436,926,947]" pageId="13" pageNumber="116">
13 Setal ratio: 5.0:1.0:6.3:0.7:15.8...........................................................
<taxonomicName id="4C044D49521DFFC6159BFC6DFA1DFC14" authority="Righi, 1984" authorityName="Righi" authorityYear="1984" box="[1251,1436,926,947]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="argous">
<emphasis id="B970EAD8521DFFC6159BFC6DFAA4FC14" box="[1251,1317,926,947]" italics="true" pageId="13" pageNumber="116">argous</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FC6DFA1DFC14" author="Righi" box="[1323,1436,926,947]" pageId="13" pageNumber="116" refString="Righi, G. (1984) Pontoscolex (Oligochaeta, Glossoscolecidae) a new evaluation. Studies on Neotropical Fauna and Environment, 19, 159 - 177." type="journal article" year="1984">Righi, 1984</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFC4FFA1DFC76" blockId="13.[151,1436,222,2033]" box="[151,1436,956,977]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC4FFA1DFC76" box="[151,1436,956,977]" pageId="13" pageNumber="116">- Setal ratio 3.1:1.0:3.3:0.3:15.6......................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFC2AFA1DFBAC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFC2AFA1DFC49" blockId="13.[151,1436,222,2033]" box="[151,1436,985,1006]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC2AFA1DFC49" box="[151,1436,985,1006]" pageId="13" pageNumber="116">
14 Tubercula pubertatis between 20,
<date id="FFBA100A521DFFC6135DFC2AFD03FC49" box="[549,642,985,1006]" pageId="13" pageNumber="116" value="1925-02-21" valueMax="1925-02-01" valueMin="1925-02-21">21-1/2 25</date>
...............................................
<emphasis id="B970EAD8521DFFC615B1FC2AFAA4FC49" box="[1225,1317,985,1006]" italics="true" pageId="13" pageNumber="116">borbakoa</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FC2AFA1DFC49" author="Righi" box="[1323,1436,985,1006]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFC05FA1DFBAC" blockId="13.[151,1436,222,2033]" box="[151,1436,1014,1035]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFC05FA1DFBAC" box="[151,1436,1014,1035]" pageId="13" pageNumber="116">- Tubercula pubertatis between 19-22.................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFBE7FA1DFBE1" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFBE7FA1DFB8E" blockId="13.[151,1436,222,2033]" box="[151,1436,1044,1065]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFBE7FA1DFB8E" box="[151,1436,1044,1065]" pageId="13" pageNumber="116">
15 Setal ratio: 2.0:1.0:3.1:0.25:11.0........................................................
<taxonomicName id="4C044D49521DFFC615B7FBE7FA1DFB8E" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1231,1436,1044,1065]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="piapoco">
<emphasis id="B970EAD8521DFFC615B7FBE7FA9DFB8E" box="[1231,1308,1044,1065]" italics="true" pageId="13" pageNumber="116">piapoco</emphasis>
<bibRefCitation id="EF954B3B521DFFC6145BFBE7FA1DFB8E" author="Feijoo" box="[1315,1436,1044,1065]" pageId="13" pageNumber="116" refString="Feijoo, A. (2008) Lombrices de tierra (Annelida, Oligochaeta) del Parque Nacional Sumapaz, Colombia. In: Van der Hammen (Ed.), Studies on Tropical Andean Ecosystems, Vol. 7. J. Cramer, Berlin, pp. 615 - 675." type="book" year="2008">Feijoo, 2008</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFBC2FA1DFBE1" blockId="13.[151,1436,222,2033]" box="[151,1436,1073,1094]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFBC2FA1DFBE1" box="[151,1436,1073,1094]" pageId="13" pageNumber="116">- Setal ratio: 1.3:0.8:2.0:0.8:8.2......................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFBBDFA1AFB26" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFBBDFA1DFBC4" blockId="13.[151,1436,222,2033]" box="[151,1436,1102,1123]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFBBDFA1DFBC4" box="[151,1436,1102,1123]" pageId="13" pageNumber="116">
16 Spermathecal pores in 6/7-8/9........................................................
<taxonomicName id="4C044D49521DFFC615C0FBBDFA1DFBC4" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1208,1436,1102,1123]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="masiguare">
<emphasis id="B970EAD8521DFFC615C0FBBDFA9DFBC4" box="[1208,1308,1102,1123]" italics="true" pageId="13" pageNumber="116">masiguare</emphasis>
<bibRefCitation id="EF954B3B521DFFC6145BFBBDFA1DFBC4" author="Feijoo" box="[1315,1436,1102,1123]" pageId="13" pageNumber="116" refString="Feijoo, A. (2008) Lombrices de tierra (Annelida, Oligochaeta) del Parque Nacional Sumapaz, Colombia. In: Van der Hammen (Ed.), Studies on Tropical Andean Ecosystems, Vol. 7. J. Cramer, Berlin, pp. 615 - 675." type="book" year="2008">Feijoo, 2008</bibRefCitation>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFB9FFA1AFB26" blockId="13.[151,1436,222,2033]" box="[151,1435,1132,1153]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB9FFA1AFB26" box="[151,1435,1132,1153]" pageId="13" pageNumber="116">- Spermathecal pores in 7/8-8/9.......................................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFB7AFA1DFB1C" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFB7AFA1DFB39" blockId="13.[151,1436,222,2033]" box="[151,1436,1161,1182]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB7AFA1DFB39" box="[151,1436,1161,1182]" pageId="13" pageNumber="116">
17 Clitellum saddle-shaped in 17, 18-26.......................................................
<taxonomicName id="4C044D49521DFFC6159CFB7AFA1DFB39" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1252,1436,1161,1182]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="mokoa">
<emphasis id="B970EAD8521DFFC6159CFB7AFAA5FB39" box="[1252,1316,1161,1182]" italics="true" pageId="13" pageNumber="116">mokoa</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FB7AFA1DFB39" author="Righi" box="[1323,1436,1161,1182]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFB55FA1DFB1C" blockId="13.[151,1436,222,2033]" box="[151,1436,1190,1211]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB55FA1DFB1C" box="[151,1436,1190,1211]" pageId="13" pageNumber="116">- Clitellum annular in 16-25............................................................................ 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFB37FA1DFB51" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFB37FA1DFB7E" blockId="13.[151,1436,222,2033]" box="[151,1436,1220,1241]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB37FA1DFB7E" box="[151,1436,1220,1241]" pageId="13" pageNumber="116">
18 Tubercula pubertatis between ½ 20-23.....................................................
<taxonomicName id="4C044D49521DFFC6159EFB37FA1DFB7E" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1254,1436,1220,1241]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="sambu">
<emphasis id="B970EAD8521DFFC6159EFB37FAA4FB7E" box="[1254,1317,1220,1241]" italics="true" pageId="13" pageNumber="116">sambu</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453FB37FA1DFB7E" author="Righi" box="[1323,1436,1220,1241]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFB12FA1DFB51" blockId="13.[151,1436,222,2033]" box="[151,1436,1249,1270]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB12FA1DFB51" box="[151,1436,1249,1270]" pageId="13" pageNumber="116">
- Tubercula pubertatis between ½ 19- ½ 23.............................................
<taxonomicName id="4C044D49521DFFC615F5FB12FA1DFB51" authority="Michaelsen, 1895" authorityName="Michaelsen" authorityYear="1895" box="[1165,1436,1249,1270]" class="Clitellata" family="Rhinodrilidae" genus="Anteus" higherTaxonomySource="CoL" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="schuetti">
<emphasis id="B970EAD8521DFFC615F5FB12FB5DFB51" box="[1165,1244,1249,1270]" italics="true" pageId="13" pageNumber="116">schuetti</emphasis>
(
<bibRefCitation id="EF954B3B521DFFC61590FB12FA14FB51" author="Michaelsen" box="[1256,1429,1249,1270]" pageId="13" pageNumber="116" refString="Michaelsen, W. (1895) Zur Kenntnis der Oligochaeten. Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein Hamburg, 13, 1 - 35." type="journal article" year="1895">Michaelsen, 1895</bibRefCitation>
)
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</paragraph>
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<keyStep id="30F02E6F521DFFC611EFFB0DFA1DFA96" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFB0DFA1DFAB4" blockId="13.[151,1436,222,2033]" box="[151,1436,1278,1299]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFB0DFA1DFAB4" box="[151,1436,1278,1299]" pageId="13" pageNumber="116">
19 Two pairs of spermathecae in 7/8, 8/9......................................................
<taxonomicName id="4C044D49521DFFC6159FFB0DFA1DFAB4" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1255,1436,1278,1299]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="cuiba">
<emphasis id="B970EAD8521DFFC6159FFB0DFA9DFAB4" box="[1255,1308,1278,1299]" italics="true" pageId="13" pageNumber="116">cuiba</emphasis>
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</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFAEFFA1DFA96" blockId="13.[151,1436,222,2033]" box="[151,1436,1308,1329]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFAEFFA1DFA96" box="[151,1436,1308,1329]" pageId="13" pageNumber="116">- Three pairs of spermathecae in 6/7- 8/9.................................................................. 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFACAFA1AFACC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFACAFA1DFAE9" blockId="13.[151,1436,222,2033]" box="[151,1436,1337,1358]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFACAFA1DFAE9" box="[151,1436,1337,1358]" pageId="13" pageNumber="116">
20 Tubercula pubertatis between 20-22................................................
<taxonomicName id="4C044D49521DFFC615F3FACAFA1DFAE9" authority="Zicsi, 1989" authorityName="Zicsi" authorityYear="1989" box="[1163,1436,1337,1358]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="reventadoriensis">
<emphasis id="B970EAD8521DFFC615F3FACAFAABFAE9" box="[1163,1322,1337,1358]" italics="true" pageId="13" pageNumber="116">reventadoriensis</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFAA5FA1AFACC" blockId="13.[151,1436,222,2033]" box="[151,1435,1366,1387]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFAA5FA1AFACC" box="[151,1435,1366,1387]" pageId="13" pageNumber="116">- Tubercula pubertatis between 19-21..................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFA87FA1AFA01" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFA87FA1DFA2E" blockId="13.[151,1436,222,2033]" box="[151,1436,1396,1417]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFA87FA1DFA2E" box="[151,1436,1396,1417]" pageId="13" pageNumber="116">
21 Spermathecae in the clitellum.............................................................
<taxonomicName id="4C044D49521DFFC6158CFA87FA1DFA2E" authority="Zicsi, 1993" authorityName="Zicsi" authorityYear="1993" box="[1268,1436,1396,1417]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="graffi">
<emphasis id="B970EAD8521DFFC6158CFA87FAA8FA2E" box="[1268,1321,1396,1417]" italics="true" pageId="13" pageNumber="116">graffi</emphasis>
<bibRefCitation id="EF954B3B521DFFC61448FA87FA1DFA2E" author="Zicsi" box="[1328,1436,1396,1417]" pageId="13" pageNumber="116" refString="Zicsi, A. (1993) Revision der Gattung Andiodrilus Michaelsen, 1900 (Oligochaeta, Glossoscolecidae) Regenwurmer aus Sudamerika. Acta Zoologica Hungarica, 39, 1 - 4, 311 - 342." type="journal article" year="1993">Zicsi, 1993</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFA62FA1AFA01" blockId="13.[151,1436,222,2033]" box="[151,1435,1425,1446]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFA62FA1AFA01" box="[151,1435,1425,1446]" pageId="13" pageNumber="116">- Spermathecae pre-clitellar............................................................................. 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFA5DFA1DFA46" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFA5DFA1DFA64" blockId="13.[151,1436,222,2033]" box="[151,1436,1454,1475]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFA5DFA1DFA64" box="[151,1436,1454,1475]" pageId="13" pageNumber="116">
22 Spermathecal pores in line with c d between 6/7-8/9.................................
<taxonomicName id="4C044D49521DFFC61515FA5DFA1DFA64" authority="Michaelsen, 1900" authorityName="Michaelsen" authorityYear="1900" box="[1133,1436,1454,1475]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="bogotaensis">
<emphasis id="B970EAD8521DFFC61515FA5DFB61FA64" box="[1133,1248,1454,1475]" italics="true" pageId="13" pageNumber="116">bogotaensis</emphasis>
<bibRefCitation id="EF954B3B521DFFC6159DFA5DFA1DFA64" author="Michaelsen" box="[1253,1436,1454,1475]" pageId="13" pageNumber="116" refString="Michaelsen, W. (1900 b) Die Terricolenfauna Columbiens. Archiv fur Naturgeschichte, 66, 231 - 266." type="journal article" year="1900" yearSuffix="b">Michaelsen, 1900b</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFFA3FFA1DFA46" blockId="13.[151,1436,222,2033]" box="[151,1436,1484,1505]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFA3FFA1DFA46" box="[151,1436,1484,1505]" pageId="13" pageNumber="116">- Spermathecal pores in the dorsal midline................................................................ 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFFA1AFA1AF9BC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFFA1AFA1DFA59" blockId="13.[151,1436,222,2033]" box="[151,1436,1513,1534]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFFA1AFA1DFA59" box="[151,1436,1513,1534]" pageId="13" pageNumber="116">
23 Clitellum between segments 15-24, ½25.................................................
<taxonomicName id="4C044D49521DFFC615CFFA1AFA1DFA59" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1207,1436,1513,1534]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="guayabero">
<emphasis id="B970EAD8521DFFC615CFFA1AFA9CFA59" box="[1207,1309,1513,1534]" italics="true" pageId="13" pageNumber="116">guayabero</emphasis>
<bibRefCitation id="EF954B3B521DFFC6145BFA1AFA1DFA59" author="Feijoo" box="[1315,1436,1513,1534]" pageId="13" pageNumber="116" refString="Feijoo, A. (2008) Lombrices de tierra (Annelida, Oligochaeta) del Parque Nacional Sumapaz, Colombia. In: Van der Hammen (Ed.), Studies on Tropical Andean Ecosystems, Vol. 7. J. Cramer, Berlin, pp. 615 - 675." type="book" year="2008">Feijoo, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF9F5FA1AF9BC" blockId="13.[151,1436,222,2033]" box="[151,1435,1542,1563]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF9F5FA1AF9BC" box="[151,1435,1542,1563]" pageId="13" pageNumber="116">- Clitellum between segments 18-26...................................................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF9D7FA1DF9F1" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF9D7FA1DF99E" blockId="13.[151,1436,222,2033]" box="[151,1436,1572,1593]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF9D7FA1DF99E" box="[151,1436,1572,1593]" pageId="13" pageNumber="116">
24 Tubercula pubertatis between ½19- ½ 24......................................................
<taxonomicName id="4C044D49521DFFC6147BF9D7FA1DF99E" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1283,1436,1572,1593]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="kiri">
<emphasis id="B970EAD8521DFFC6147BF9D7FAA5F99E" box="[1283,1316,1572,1593]" italics="true" pageId="13" pageNumber="116">kiri</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453F9D7FA1DF99E" author="Righi" box="[1323,1436,1572,1593]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF9B2FA1DF9F1" blockId="13.[151,1436,222,2033]" box="[151,1436,1601,1622]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF9B2FA1DF9F1" box="[151,1436,1601,1622]" pageId="13" pageNumber="116">
- Tubercula pubertatis between 2/
<date id="FFBA100A521DFFC6136EF9B2FDDEF9F1" box="[534,607,1601,1622]" pageId="13" pageNumber="116" value="1918-03-01">3 18 – 1</date>
/3 24............................................................. 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF9ADFA1AF936" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF9ADFA1DF9D4" blockId="13.[151,1436,222,2033]" box="[151,1436,1630,1651]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF9ADFA1DF9D4" box="[151,1436,1630,1651]" pageId="13" pageNumber="116">
<specimenCount id="9D02FD43521DFFC611EFF9ADFE93F9D4" box="[151,274,1630,1651]" pageId="13" pageNumber="116" type="male">25 Male</specimenCount>
pores in 22/23....................................................................
<taxonomicName id="4C044D49521DFFC6159BF9ADFA1DF9D4" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1251,1436,1630,1651]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="patoko">
<emphasis id="B970EAD8521DFFC6159BF9ADFAA5F9D4" box="[1251,1316,1630,1651]" italics="true" pageId="13" pageNumber="116">patoko</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453F9ADFA1DF9D4" author="Righi" box="[1323,1436,1630,1651]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF98FFA1AF936" blockId="13.[151,1436,222,2033]" box="[151,1435,1660,1681]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF98FFA1AF936" box="[151,1435,1660,1681]" pageId="13" pageNumber="116">- Male pores in 19..................................................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF96AFA1DF96C" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF96AFA1DF909" blockId="13.[151,1436,222,2033]" box="[151,1436,1689,1710]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF96AFA1DF909" box="[151,1436,1689,1710]" pageId="13" pageNumber="116">
26 Last pair of intestinal hearts in 13.......................................................
<taxonomicName id="4C044D49521DFFC615B6F96AFA1DF909" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1230,1436,1689,1710]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="guahibo">
<emphasis id="B970EAD8521DFFC615B6F96AFA9DF909" box="[1230,1308,1689,1710]" italics="true" pageId="13" pageNumber="116">guahibo</emphasis>
<bibRefCitation id="EF954B3B521DFFC6145BF96AFA1DF909" author="Feijoo" box="[1315,1436,1689,1710]" pageId="13" pageNumber="116" refString="Feijoo, A. (2008) Lombrices de tierra (Annelida, Oligochaeta) del Parque Nacional Sumapaz, Colombia. In: Van der Hammen (Ed.), Studies on Tropical Andean Ecosystems, Vol. 7. J. Cramer, Berlin, pp. 615 - 675." type="book" year="2008">Feijoo, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF945FA1DF96C" blockId="13.[151,1436,222,2033]" box="[151,1436,1718,1739]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF945FA1DF96C" box="[151,1436,1718,1739]" pageId="13" pageNumber="116">- Last pair of intestinal hearts in 12...................................................................... 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF927FA1AF8A1" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF927FA1DF94E" blockId="13.[151,1436,222,2033]" box="[151,1436,1748,1769]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF927FA1DF94E" box="[151,1436,1748,1769]" pageId="13" pageNumber="116">
27 Intestinal hearts in 10-12.............................................................
<taxonomicName id="4C044D49521DFFC615C3F927FA1DF94E" authority="Zicsi, 1988" authorityName="Zicsi" authorityYear="1988" box="[1211,1436,1748,1769]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="paramensis">
<emphasis id="B970EAD8521DFFC615C3F927FAABF94E" box="[1211,1322,1748,1769]" italics="true" pageId="13" pageNumber="116">paramensis</emphasis>
<bibRefCitation id="EF954B3B521DFFC61448F927FA1DF94E" author="Zicsi" box="[1328,1436,1748,1769]" pageId="13" pageNumber="116" refString="Zicsi, A. (1988) Neue Andiodrilus - Arten aus Kolumbien (Oligochaeta: Glossoscolecidae) Regenwurmer aus Sudamerika 5. Revue Suisse de Zoologie, 95, 3, 715 - 722." type="journal article" year="1988">Zicsi, 1988</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF902FA1AF8A1" blockId="13.[151,1436,222,2033]" box="[151,1435,1777,1798]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF902FA1AF8A1" box="[151,1435,1777,1798]" pageId="13" pageNumber="116">- Intestinal hearts in 10-13.............................................................................. 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF8FDFA1DF8E6" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF8FDFA1DF884" blockId="13.[151,1436,222,2033]" box="[151,1436,1806,1827]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF8FDFA1DF884" box="[151,1436,1806,1827]" pageId="13" pageNumber="116">
28 Clitellum between 16-26.............................................................
<taxonomicName id="4C044D49521DFFC615C7F8FDFA1DF884" authority="Cognetti, 1904" authorityName="Cognetti" authorityYear="1904" box="[1215,1436,1806,1827]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="biolleyi">
<emphasis id="B970EAD8521DFFC615C7F8FDFA86F884" box="[1215,1287,1806,1827]" italics="true" pageId="13" pageNumber="116">biolleyi</emphasis>
<bibRefCitation id="EF954B3B521DFFC61475F8FDFA1DF884" author="Cognetti" box="[1293,1436,1806,1827]" pageId="13" pageNumber="116" refString="Cognetti De Martiis, L. (1904) Descrizione di un nuovo lombrico cavernicola. Bollettinno dei Musei di Zoologia et Anatomia Comparata della Reale Universita di Torino, 20, 466, 1 - 6." type="journal article" year="1904">Cognetti, 1904</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF8DFFA1DF8E6" blockId="13.[151,1436,222,2033]" box="[151,1436,1836,1857]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF8DFFA1DF8E6" box="[151,1436,1836,1857]" pageId="13" pageNumber="116">- Clitellum between 1/n 16, 17-25....................................................................... 36</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF8BAFA1AF8DC" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF8BAFA1DF8F9" blockId="13.[151,1436,222,2033]" box="[151,1436,1865,1886]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF8BAFA1DF8F9" box="[151,1436,1865,1886]" pageId="13" pageNumber="116">
29 Spermathecae with seminal chambers.......................................................
<taxonomicName id="4C044D49521DFFC61597F8BAFA1DF8F9" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1263,1436,1865,1886]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="pixao">
<emphasis id="B970EAD8521DFFC61597F8BAFAA5F8F9" box="[1263,1316,1865,1886]" italics="true" pageId="13" pageNumber="116">pixao</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453F8BAFA1DF8F9" author="Righi" box="[1323,1436,1865,1886]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF895FA1AF8DC" blockId="13.[151,1436,222,2033]" box="[151,1435,1894,1915]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF895FA1AF8DC" box="[151,1435,1894,1915]" pageId="13" pageNumber="116">- Spermathecae without seminal chambers................................................................. 30</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF877FA1AF811" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF877FA1DF83E" blockId="13.[151,1436,222,2033]" box="[151,1436,1924,1945]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF877FA1DF83E" box="[151,1436,1924,1945]" pageId="13" pageNumber="116">
30 Tubercula pubertatis in ½ 20- ¾ 24..................................................
<taxonomicName id="4C044D49521DFFC615E1F877FA1DF83E" authority="Michaelsen, 1913" authorityName="Michaelsen" authorityYear="1913" box="[1177,1436,1924,1945]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="ruizanus">
<emphasis id="B970EAD8521DFFC615E1F877FB6DF83E" box="[1177,1260,1924,1945]" italics="true" pageId="13" pageNumber="116">ruizanus</emphasis>
<bibRefCitation id="EF954B3B521DFFC61589F877FA1DF83E" author="Michaelsen" box="[1265,1436,1924,1945]" pageId="13" pageNumber="116" refString="Michaelsen, W. (1913) Die Oligochaeten Columbias. In: Fuhrmann O. &amp; Mayor, E. (Eds.), Voyage d'Exploration Scientifique en Columbie: Memoires de la Societe des Sciences Naturelles de Neuchatel, 5, 202 - 252." type="journal article" year="1913">Michaelsen, 1913</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF852FA1AF811" blockId="13.[151,1436,222,2033]" box="[151,1435,1953,1974]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF852FA1AF811" box="[151,1435,1953,1974]" pageId="13" pageNumber="116">- Tubercula pubertatis in ½ 19-23......................................................................... 31</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521DFFC611EFF84DFA1DF856" pageId="13" pageNumber="116">
<paragraph id="8BBB36CA521DFFC611EFF84DFA1DF874" blockId="13.[151,1436,222,2033]" box="[151,1436,1982,2003]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF84DFA1DF874" box="[151,1436,1982,2003]" pageId="13" pageNumber="116">
31 Clitellum between 15, 16- ½ 26........................................................
<taxonomicName id="4C044D49521DFFC615C7F84DFA1DF874" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1215,1436,1982,2003]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="13" pageNumber="116" phylum="Annelida" rank="species" species="cabalensis">
<emphasis id="B970EAD8521DFFC615C7F84DFAA5F874" box="[1215,1316,1982,2003]" italics="true" pageId="13" pageNumber="116">cabalensis</emphasis>
<bibRefCitation id="EF954B3B521DFFC61453F84DFA1DF874" author="Righi" box="[1323,1436,1982,2003]" pageId="13" pageNumber="116" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521DFFC611EFF82FFA1DF856" blockId="13.[151,1436,222,2033]" box="[151,1436,2012,2033]" pageId="13" pageNumber="116">
<keyLead id="30F595FF521DFFC611EFF82FFA1DF856" box="[151,1436,2012,2033]" pageId="13" pageNumber="116">- Clitellum between 16 – 25............................................................................ 32</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFF64FA1AFF6D" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFF64FA1DFF0B" blockId="14.[151,1436,151,613]" box="[151,1436,151,173]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFF64FA1DFF0B" box="[151,1436,151,173]" pageId="14" pageNumber="117">
32 Caecum present in 26 ó 27...............................................................
<taxonomicName id="4C044D49521EFFC5159FFF6BFA1DFF0B" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1255,1436,151,173]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="xaropi">
<emphasis id="B970EAD8521EFFC5159FFF6BFAA4FF0A" box="[1255,1317,152,173]" italics="true" pageId="14" pageNumber="117">xaropi</emphasis>
<bibRefCitation id="EF954B3B521EFFC51453FF64FA1DFF0B" author="Righi" box="[1323,1436,151,172]" pageId="14" pageNumber="117" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFF46FA1AFF6D" blockId="14.[151,1436,151,613]" box="[151,1435,181,202]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFF46FA1AFF6D" box="[151,1435,181,202]" pageId="14" pageNumber="117">- Caecum absent...................................................................................... 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFF21FA1DFEA3" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFF21FA1DFF40" blockId="14.[151,1436,151,613]" box="[151,1436,210,231]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFF21FA1DFF40" box="[151,1436,210,231]" pageId="14" pageNumber="117">
33 Spermathecae in
<date id="FFBA100A521EFFC510FEFF21FE56FF40" box="[390,471,210,231]" pageId="14" pageNumber="117" value="2010-07-06" valueMax="2010-09-06" valueMin="2010-07-06">6/7-9/10</date>
.............................................................
<taxonomicName id="4C044D49521EFFC515CFFF21FA1DFF40" authority="Zicsi, 1993" authorityName="Zicsi" authorityYear="1993" box="[1207,1436,210,231]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="michaelseni">
<emphasis id="B970EAD8521EFFC515CFFF21FAABFF40" box="[1207,1322,210,231]" italics="true" pageId="14" pageNumber="117">michaelseni</emphasis>
<bibRefCitation id="EF954B3B521EFFC51448FF21FA1DFF40" author="Zicsi" box="[1328,1436,210,231]" pageId="14" pageNumber="117" refString="Zicsi, A. (1993) Revision der Gattung Andiodrilus Michaelsen, 1900 (Oligochaeta, Glossoscolecidae) Regenwurmer aus Sudamerika. Acta Zoologica Hungarica, 39, 1 - 4, 311 - 342." type="journal article" year="1993">Zicsi, 1993</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFF1CFA1DFEA3" blockId="14.[151,1436,151,613]" box="[151,1436,239,260]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFF1CFA1DFEA3" box="[151,1436,239,260]" pageId="14" pageNumber="117">- Spermathecae in 6/7-8/9.............................................................................. 34</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFEFEFA1AFE98" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFEFEFA1DFE85" blockId="14.[151,1436,151,613]" box="[151,1436,269,290]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFEFEFA1DFE85" box="[151,1436,269,290]" pageId="14" pageNumber="117">
34 Setal ratio: 10:2:10:2:50..................................................................
<emphasis id="B970EAD8521EFFC51595FEFEFAABFE85" box="[1261,1322,269,290]" italics="true" pageId="14" pageNumber="117">sturmi</emphasis>
<bibRefCitation id="EF954B3B521EFFC51448FEFEFA1DFE85" author="Zicsi" box="[1328,1436,269,290]" pageId="14" pageNumber="117" refString="Zicsi, A. (1988) Neue Andiodrilus - Arten aus Kolumbien (Oligochaeta: Glossoscolecidae) Regenwurmer aus Sudamerika 5. Revue Suisse de Zoologie, 95, 3, 715 - 722." type="journal article" year="1988">Zicsi, 1988</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFED9FA1AFE98" blockId="14.[151,1436,151,613]" box="[151,1435,298,319]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFED9FA1AFE98" box="[151,1435,298,319]" pageId="14" pageNumber="117">- Setal ratio: 4.2:0.35:4.0:0.39:10......................................................................... 35</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFEB4FA1DFEDD" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFEB4FA1DFEFB" blockId="14.[151,1436,151,613]" box="[151,1436,327,349]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFEB4FA1DFEFB" box="[151,1436,327,349]" pageId="14" pageNumber="117">
35 Clitellum between 15-24, 25.........................................................
<taxonomicName id="4C044D49521EFFC515CBFEBBFA1DFEFB" authority="Feijoo, 2008" authorityName="Feijoo" authorityYear="2008" box="[1203,1436,327,349]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="makaguaje">
<emphasis id="B970EAD8521EFFC515CBFEBBFA9DFEFA" box="[1203,1308,328,349]" italics="true" pageId="14" pageNumber="117">makaguaje</emphasis>
<bibRefCitation id="EF954B3B521EFFC5145BFEB4FA1DFEFB" author="Feijoo" box="[1315,1436,327,348]" pageId="14" pageNumber="117" refString="Feijoo, A. (2008) Lombrices de tierra (Annelida, Oligochaeta) del Parque Nacional Sumapaz, Colombia. In: Van der Hammen (Ed.), Studies on Tropical Andean Ecosystems, Vol. 7. J. Cramer, Berlin, pp. 615 - 675." type="book" year="2008">Feijoo, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFE96FA1DFEDD" blockId="14.[151,1436,151,613]" box="[151,1436,357,378]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE96FA1DFEDD" box="[151,1436,357,378]" pageId="14" pageNumber="117">
- Clitellum between 16-26...........................................................
<taxonomicName id="4C044D49521EFFC515DCFE96FA1DFEDD" authority="Righi, 1989" authorityName="Righi" authorityYear="1989" box="[1188,1436,357,378]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="venezuelanus">
<emphasis id="B970EAD8521EFFC515DCFE96FAA5FEDD" box="[1188,1316,357,378]" italics="true" pageId="14" pageNumber="117">venezuelanus</emphasis>
<bibRefCitation id="EF954B3B521EFFC51453FE96FA1DFEDD" author="Righi" box="[1323,1436,357,378]" pageId="14" pageNumber="117" refString="Righi, G. (1989) Adicao ao conhecimento dos Oligochaeta da Venezuela. Revista Brasileira de Biologia, 49, 4, 1065 - 1084." type="journal article" year="1989">Righi, 1989</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFE71FA1DFE13" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFE71FA1DFE30" blockId="14.[151,1436,151,613]" box="[151,1436,386,407]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE71FA1DFE30" box="[151,1436,386,407]" pageId="14" pageNumber="117">
36 Spermathecal pores in line
<emphasis id="B970EAD8521EFFC5109EFE71FE7DFE30" box="[486,508,386,407]" italics="true" pageId="14" pageNumber="117">cd</emphasis>
......................................................
<taxonomicName id="4C044D49521EFFC515F3FE71FA1DFE30" authority="Michaelsen, 1912" authorityName="Michaelsen" authorityYear="1912" box="[1163,1436,386,407]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="orosiensis">
<emphasis id="B970EAD8521EFFC515F3FE71FB6AFE30" box="[1163,1259,386,407]" italics="true" pageId="14" pageNumber="117">orosiensis</emphasis>
<bibRefCitation id="EF954B3B521EFFC51589FE71FA1DFE30" author="Michaelsen" box="[1265,1436,386,407]" pageId="14" pageNumber="117" refString="Michaelsen, W. (1912) Uber einige zentralamenrikanische Oligochaeten. Archiv fur Naturgeschichte, 78, 112 - 129." type="journal article" year="1912">Michaelsen, 1912</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFE6CFA1DFE13" blockId="14.[151,1436,151,613]" box="[151,1436,415,436]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE6CFA1DFE13" box="[151,1436,415,436]" pageId="14" pageNumber="117">
- Spermathecal pores between 6/7-8/
<quantity id="4CFC9B2F521EFFC5134AFE6CFDD6FE13" box="[562,599,415,436]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="14" pageNumber="117" unit="in" value="9.0">9 in</quantity>
line ab............................................................ 37
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFE4EFA1AFE48" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFE4EFA1DFE75" blockId="14.[151,1436,151,613]" box="[151,1436,445,466]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE4EFA1DFE75" box="[151,1436,445,466]" pageId="14" pageNumber="117">
37 Tubercula pubertatis ¼ 19, 20-25.................................................
<taxonomicName id="4C044D49521EFFC5150DFE4EFA1DFE75" authority="Michaelsen, 1900" authorityName="Michaelsen" authorityYear="1900" box="[1141,1436,445,466]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="pachoensis">
<emphasis id="B970EAD8521EFFC5150DFE4EFB61FE75" box="[1141,1248,445,466]" italics="true" pageId="14" pageNumber="117">pachoensis</emphasis>
<bibRefCitation id="EF954B3B521EFFC5159DFE4EFA1DFE75" author="Michaelsen" box="[1253,1436,445,466]" pageId="14" pageNumber="117" refString="Michaelsen, W. (1900 b) Die Terricolenfauna Columbiens. Archiv fur Naturgeschichte, 66, 231 - 266." type="journal article" year="1900" yearSuffix="b">Michaelsen, 1900b</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFE29FA1AFE48" blockId="14.[151,1436,151,613]" box="[151,1435,474,495]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE29FA1AFE48" box="[151,1435,474,495]" pageId="14" pageNumber="117">- Tubercula pubertatis 21-24, ½25........................................................................ 38</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFE04FA1AFD8D" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFE04FA1DFDAB" blockId="14.[151,1436,151,613]" box="[151,1436,503,525]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFE04FA1DFDAB" box="[151,1436,503,525]" pageId="14" pageNumber="117">
38 Clitellum between 17-24...............................................................
<taxonomicName id="4C044D49521EFFC515A4FE0BFA1DFDAB" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1244,1436,503,525]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="terraba">
<emphasis id="B970EAD8521EFFC515A4FE0BFAA5FDAA" box="[1244,1316,504,525]" italics="true" pageId="14" pageNumber="117">terraba</emphasis>
<bibRefCitation id="EF954B3B521EFFC51453FE04FA1DFDAB" author="Righi" box="[1323,1436,503,524]" pageId="14" pageNumber="117" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
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</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFDE6FA1AFD8D" blockId="14.[151,1436,151,613]" box="[151,1435,533,554]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFDE6FA1AFD8D" box="[151,1435,533,554]" pageId="14" pageNumber="117">- Clitellum between 15–25.............................................................................. 39</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F02E6F521EFFC511EFFDC1FA1DFDC3" pageId="14" pageNumber="117">
<paragraph id="8BBB36CA521EFFC511EFFDC1FA1AFDE0" blockId="14.[151,1436,151,613]" box="[151,1435,562,583]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFDC1FA1AFDE0" box="[151,1435,562,583]" pageId="14" pageNumber="117">
39 Tubercula pubertatis between ¼ 19 – ¼ 24.....................................................
<emphasis id="B970EAD8521EFFC51586FDC1FA1AFDE0" bold="true" box="[1278,1435,562,583]" pageId="14" pageNumber="117">
<taxonomicName id="4C044D49521EFFC51586FDC1FAC8FDE0" box="[1278,1353,562,583]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="nonuya" status="sp. nov.">
<emphasis id="B970EAD8521EFFC51586FDC1FAC8FDE0" bold="true" box="[1278,1353,562,583]" italics="true" pageId="14" pageNumber="117">nonuya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357A3521EFFC51437FDC1FA1AFDE0" box="[1359,1435,562,583]" pageId="14" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BBB36CA521EFFC511EFFDBCFA1DFDC3" blockId="14.[151,1436,151,613]" box="[151,1436,591,613]" pageId="14" pageNumber="117">
<keyLead id="30F595FF521EFFC511EFFDBCFA1DFDC3" box="[151,1436,591,613]" pageId="14" pageNumber="117">
- Tubercula pubertatis between ½ 20-½ 25..................................................
<taxonomicName id="4C044D49521EFFC515B3FDA3FA1DFDC3" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[1227,1436,591,613]" class="Clitellata" family="Glossoscolecidae" genus="Andiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="14" pageNumber="117" phylum="Annelida" rank="species" species="talamank">
<emphasis id="B970EAD8521EFFC515B3FDA3FAA5FDC2" box="[1227,1316,592,613]" italics="true" pageId="14" pageNumber="117">talamank</emphasis>
<bibRefCitation id="EF954B3B521EFFC51453FDBCFA1DFDC3" author="Righi" box="[1323,1436,591,612]" pageId="14" pageNumber="117" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Dos Santos, A. G. (Eds), Studies on Tropical Andean Ecosystems / Estudios de Ecosistemas Tropandinos 4. Cramer, Berlin, pp. 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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