302 lines
39 KiB
XML
302 lines
39 KiB
XML
<document id="F94F582343793DE824DF98C1541CE6BE" ID-DOI="10.5281/zenodo.7300614" ID-ISSN="1942-1354" ID-Zenodo-Dep="7300614" ID-ZooBank="2425983D-0398-45D4-A728-3BF5991D07BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667844991072" checkinUser="felipe" docAuthor="Smith, Trevor Randall" docDate="2022" docId="03AD87DAFFCDAE4BFF0237B5FB7A4F0E" docLanguage="en" docName="InsectaMundi.2022.0950.1-35.pdf" docOrigin="Insecta Mundi 2022 (950)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Cybocephalus skelleyi Smith, 2022, new species" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="FF94FFA2FFD7AE56FFC23325FFDA4E5A" masterDocTitle="Review of the Cybocephalidae (Coleoptera) of North America and the West Indies with descriptions of two new species of Cybocephalus Erichson" masterLastPageNumber="35" masterPageNumber="1" pageNumber="25" updateTime="1699393177864" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="48B57863693D927947CA221FF2EB4EB7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8CF590E847B2924B1491FC2A3E916984">
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<mods:title id="8C7897C3C22663228B8DA60869FDCD9D">Review of the Cybocephalidae (Coleoptera) of North America and the West Indies with descriptions of two new species of Cybocephalus Erichson</mods:title>
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<mods:name id="AE2F2465614AF4B02538F047A7105C68" type="personal">
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<mods:roleTerm id="4B6CED2E31A769B07147D9316CADDDEE">Author</mods:roleTerm>
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<mods:namePart id="0AF5343289184C2BD96D0EE79EE3B21C">Smith, Trevor Randall</mods:namePart>
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<mods:affiliation id="C1A625B5861A32193A6C016AC13F8342">Division of Plant Industry, Florida Department of Agriculture and Consumer Services 1911 SW 34 th St. / PO Box 147100, Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:typeOfResource id="AD9E892A31747AF86BFF6472352F8EDC">text</mods:typeOfResource>
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<mods:title id="69AEBC3F06FE629C11C6A3072FC5D52F">Insecta Mundi</mods:title>
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<mods:part id="22717A9BEA71FED670C0C34059B21A6E">
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<mods:date id="AE9D55F4C3BA744FA831374B46BE14B1">2022</mods:date>
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<mods:number id="FF6E2D6DBDB505280AE67BB8633CF069">2022-08-26</mods:number>
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<mods:detail id="70B92662BB0E091A47B6E1A5198551C7" type="volume">
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<mods:number id="9A3201D1B39D5194BEBC4C0117F747D3">2022</mods:number>
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<mods:detail id="5F5379D5F1EDDC83B2E532C5FD8947A1" type="issue">
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<mods:number id="D975DE0521BF83D2341E838F07F210CE">950</mods:number>
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<mods:start id="B90B53A2DBF787508EC33F1A5F50035F">1</mods:start>
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<mods:classification id="6F02A142526EA97A1FDBEACF5F1EF626">journal article</mods:classification>
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<mods:identifier id="1899BFD17D4586C856D369CC64BDE513" type="DOI">10.5281/zenodo.7300614</mods:identifier>
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<mods:identifier id="69C24119BA184CFCF3926A0BC5338082" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="E0AA7CC134A7F162EFFCDA3D07CFBDBE" type="Zenodo-Dep">7300614</mods:identifier>
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<mods:identifier id="44409CEF91F7347A1301E5AB190AB772" type="ZooBank">2425983D-0398-45D4-A728-3BF5991D07BE</mods:identifier>
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<treatment id="03AD87DAFFCDAE4BFF0237B5FB7A4F0E" ID-DOI="http://doi.org/10.5281/zenodo.7300758" ID-Zenodo-Dep="7300758" LSID="urn:lsid:plazi:treatment:03AD87DAFFCDAE4BFF0237B5FB7A4F0E" httpUri="http://treatment.plazi.org/id/03AD87DAFFCDAE4BFF0237B5FB7A4F0E" lastPageId="29" lastPageNumber="28" pageId="26" pageNumber="25">
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<subSubSection id="C31E6547FFCDAE4CFF0237B5FCD34AF7" box="[192,777,1168,1197]" pageId="26" pageNumber="25" type="nomenclature">
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<paragraph id="8BBB36CCFFCDAE4CFF0237B5FCD34AF7" blockId="26.[192,777,1168,1231]" box="[192,777,1168,1197]" pageId="26" pageNumber="25">
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<heading id="D0F381A0FFCDAE4CFF0237B5FCD34AF7" box="[192,777,1168,1197]" fontSize="12" level="2" pageId="26" pageNumber="25" reason="2">
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<emphasis id="B970EADEFFCDAE4CFF0237B5FCD34AF7" bold="true" box="[192,777,1168,1197]" pageId="26" pageNumber="25">
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<taxonomicName id="4C044D4FFFCDAE4CFF0237B5FE154AF7" authorityName="Smith" authorityYear="2022" box="[192,463,1168,1197]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="skelleyi">
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<emphasis id="B970EADEFFCDAE4CFF0237B5FE154AF7" bold="true" box="[192,463,1168,1197]" italics="true" pageId="26" pageNumber="25">Cybocephalus skelleyi</emphasis>
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</taxonomicName>
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T. R.
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<collectingRegion id="49C0F82EFFCDAE4CFDDB37B5FDB24AF7" box="[537,616,1168,1197]" country="Bermuda" name="Smith's" pageId="26" pageNumber="25">Smith</collectingRegion>
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,
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<taxonomicNameLabel id="A24357A5FFCDAE4CFDB637B4FCD34AF7" box="[628,777,1168,1197]" pageId="26" pageNumber="25" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E6547FFCDAE4CFF023791FEB94A95" box="[192,355,1204,1231]" pageId="26" pageNumber="25" type="description">
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<paragraph id="8BBB36CCFFCDAE4CFF023791FEB94A95" blockId="26.[192,777,1168,1231]" box="[192,355,1204,1231]" pageId="26" pageNumber="25">
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(
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<figureCitation id="133F2A49FFCDAE4CFF0A3791FE814A95" box="[200,347,1204,1231]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="26" pageNumber="25">Fig. 135–143</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E6547FFCDAE4CFF0237FBFAF74B61" pageId="26" pageNumber="25" type="etymology">
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<paragraph id="8BBB36CCFFCDAE4CFF0237FBFAF74B61" blockId="26.[192,1440,1246,1339]" pageId="26" pageNumber="25">
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<emphasis id="B970EADEFFCDAE4CFF0237FBFE9E4AA2" bold="true" box="[192,324,1246,1272]" pageId="26" pageNumber="25">Etymology.</emphasis>
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This species is named in honor of Dr. Paul Skelley, whose passion for taxonomy, systematics and natural history is second to none. He is the most inspiring scientist and entomologist I have ever met, and it has been my honor to work alongside him in the
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<collectingRegion id="49C0F82EFFCDAE4CFD7A3604FC9C4B61" box="[696,838,1313,1339]" country="Uruguay" name="Florida" pageId="26" pageNumber="25">Florida State</collectingRegion>
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Collection of Arthropods for over 20 years.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E6547FFCDAE4CFF02366FFC1A4874" pageId="26" pageNumber="25" type="diagnosis">
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<paragraph id="8BBB36CCFFCDAE4CFF02366FFC1A4874" blockId="26.[191,1442,1354,1583]" pageId="26" pageNumber="25">
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<emphasis id="B970EADEFFCDAE4CFF02366FFEE64B3E" bold="true" box="[192,316,1354,1380]" pageId="26" pageNumber="25">Diagnosis.</emphasis>
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Body convex and ovate. Antennae with 11 antennomeres, with an asymmetrical, 3-antennomere club with serrated inner margin and a truncate terminal antennomere (
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<figureCitation id="133F2A49FFCDAE4CFC263649FB994BDD" box="[996,1091,1388,1415]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="26" pageNumber="25">Fig. 135</figureCitation>
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). Head large and wide. Elytron pointed at apex rather than truncate with a very small notch at the tip in both males and females. Hindwings present. Legs, both femora and tibiae not dilated (
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<figureCitation id="133F2A49FFCDAE4CFD02368AFC8F4B93" box="[704,853,1455,1481]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="26" pageNumber="25">
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Fig. 136
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<emphasis id="B970EADEFFCDAE4CFCD9368AFCF34B93" bold="true" box="[795,809,1455,1481]" pageId="26" pageNumber="25">–</emphasis>
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138
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</figureCitation>
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). Metaventrite almost the same length in the middle as the lateral margins, only slightly longer at the lateral edge. Metaventrite only slightly longer than mesoventrite. Abdominal ventrite I longer than abdominal ventrites II-IV, combined with a truncate apical projection between the metacoxae. Tarsi with distinctly lobed tarsomeres (
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<figureCitation id="133F2A49FFCDAE4CFCDD3531FC694874" box="[799,947,1556,1582]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="26" pageNumber="25">
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Fig. 136
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<emphasis id="B970EADEFFCDAE4CFCBB3531FC5D4874" bold="true" box="[889,903,1556,1582]" pageId="26" pageNumber="25">–</emphasis>
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138
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</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E6547FFCDAE4AFF023518FD8B4B36" lastPageId="28" lastPageNumber="27" pageId="26" pageNumber="25" type="description">
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<paragraph id="8BBB36CCFFCDAE4CFF023518FC4B4802" blockId="26.[192,1441,1597,1927]" box="[192,913,1597,1624]" pageId="26" pageNumber="25">
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<emphasis id="B970EADEFFCDAE4CFF023518FE88480D" bold="true" box="[192,338,1597,1623]" pageId="26" pageNumber="25">Description.</emphasis>
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Length (excluding head)
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<quantity id="4CFC9B29FFCDAE4CFDB2351BFD0F4802" box="[624,725,1598,1624]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="25" unit="mm" value="1.1">1.10 mm</quantity>
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, width
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<quantity id="4CFC9B29FFCDAE4CFCE5351BFC574802" box="[807,909,1598,1624]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="25" unit="mm" value="0.8">0.80 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BBB36CCFFCDAE4CFEC2357AFEC2495A" blockId="26.[192,1441,1597,1927]" pageId="26" pageNumber="25">
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<emphasis id="B970EADEFFCDAE4CFEC2357AFE9B4823" bold="true" box="[256,321,1631,1657]" pageId="26" pageNumber="25">Male.</emphasis>
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Body small and ovate. Coloration: head light brown at base and becoming darker brown towards the clypeus. Pronotum, scutellar shield and elytra brown. Antennae light brown. Underside brown and clothed in thick setae, palpi and legs brown. Translucent yellowish border along the lateral lobe of the pronotum. Translucent yellowish border beginning just after the epipleural fold and expanding in width towards the apices of the elytron.
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</paragraph>
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<paragraph id="8BBB36CCFFCDAE4AFEC23422FBCE4F86" blockId="26.[192,1441,1597,1927]" lastBlockId="28.[191,1443,214,1388]" lastPageId="28" lastPageNumber="27" pageId="26" pageNumber="25">
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Head wide and produced (width =
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<quantity id="4CFC9B29FFCDAE4CFD4B3422FD344978" box="[649,750,1799,1826]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="26" pageNumber="25" unit="mm" value="0.58">0.58 mm</quantity>
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; length =
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<quantity id="4CFC9B29FFCDAE4CFC9F3422FC19497B" box="[861,963,1799,1825]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="26" pageNumber="25" unit="mm" value="0.43">0.43 mm</quantity>
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), width 1.3× the length, deflexed; antennal fossae bordered and deeply emarginated, inwardly sloping for reception of scape. Surface alutaceous and impunctate in the posterior half, becoming minutely and sparsely punctate on frons and towards the apical portion of head, interspaces alutaceous. Margin of genae narrowly visible. Antennae with 11 antennomeres, asymmetrical, 3-antennomere club with serrated inner margin and a truncate terminal antennomere. Antennomere III longer than pedicel but shorter than antennomeres IV and V combined. Scape expanded medially and pedicel globular (
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<figureCitation id="133F2A49FFCBAE4AFF2C323CFE944F6E" box="[238,334,281,308]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 135</figureCitation>
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). Antennal length about 0.87× the width of the head. Clypeus long and produced, extending well beyond apex of eyes, with slightly convex apical margin. Labrum visible from above with apical margin bilobed. Eyes ovate, length
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<quantity id="4CFC9B29FFCBAE4AFE503278FE234F2D" box="[402,505,349,375]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="27" unit="mm" value="0.19">0.19 mm</quantity>
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, visible ventrally, with distinct ommatidia. Mandibles heavily built and moderately produced. Maxillary palpi dilated with ultimate palpomere conical and gradually narrowing towards apex and about the same length as the preceding palpomere. Labial palpi dilated with ultimate palpomere cylindrical with a truncate apical margin and about twice the length of preceding palpomere.
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</paragraph>
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<caption id="DF7B6644FFCCAE4DFF023422FEF749DC" ID-DOI="http://doi.org/10.5281/zenodo.7300650" ID-Zenodo-Dep="7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="27" pageNumber="26" startId="27.[192,279,1799,1825]" targetBox="[230,1402,316,1741]" targetPageId="27">
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<paragraph id="8BBB36CCFFCCAE4DFF023422FEF749DC" blockId="27.[192,1441,1799,1926]" pageId="27" pageNumber="26">
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<emphasis id="B970EADEFFCCAE4DFF023422FE53497B" bold="true" box="[192,393,1799,1825]" pageId="27" pageNumber="26">Figures 135–143.</emphasis>
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<taxonomicName id="4C044D4FFFCCAE4DFE503422FDA4497B" authorityName="Smith" authorityYear="2022" box="[402,638,1799,1825]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="skelleyi">
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<emphasis id="B970EADEFFCCAE4DFE503422FDA4497B" box="[402,638,1799,1825]" italics="true" pageId="27" pageNumber="26">Cybocephalus skelleyi</emphasis>
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</taxonomicName>
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T. R. Smith.
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<emphasis id="B970EADEFFCCAE4DFCDB3422FC97497B" bold="true" box="[793,845,1799,1825]" pageId="27" pageNumber="26">135)</emphasis>
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Antenna.
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<emphasis id="B970EADEFFCCAE4DFC053422FC20497B" bold="true" box="[967,1018,1799,1825]" pageId="27" pageNumber="26">136)</emphasis>
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Proleg.
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<emphasis id="B970EADEFFCCAE4DFB993422FB55497B" bold="true" box="[1115,1167,1799,1825]" pageId="27" pageNumber="26">137)</emphasis>
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Mesoleg.
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<emphasis id="B970EADEFFCCAE4DFAC63422FAED497B" bold="true" box="[1284,1335,1799,1825]" pageId="27" pageNumber="26">138)</emphasis>
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Metaleg.
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<emphasis id="B970EADEFFCCAE4DFF02340DFF2E4918" bold="true" box="[192,244,1832,1858]" pageId="27" pageNumber="26">139)</emphasis>
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Median lobe, as it appears in holotype.
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<emphasis id="B970EADEFFCCAE4DFD79340DFD354918" bold="true" box="[699,751,1832,1858]" pageId="27" pageNumber="26">140)</emphasis>
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Median lobe, lateral view, as it appears in the holotype.
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<emphasis id="B970EADEFFCCAE4DFAAC340DFA7B4918" bold="true" box="[1390,1441,1832,1858]" pageId="27" pageNumber="26">141)</emphasis>
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Basal plate.
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<emphasis id="B970EADEFFCCAE4DFE84346FFEA3493E" bold="true" box="[326,377,1866,1892]" pageId="27" pageNumber="26">142)</emphasis>
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Median lobe, hypothetical reconstruction.
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<emphasis id="B970EADEFFCCAE4DFCA1346FFC4C493E" bold="true" box="[867,918,1866,1892]" pageId="27" pageNumber="26">143)</emphasis>
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Median lobe, lateral view, hypothetical reconstruction.
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</paragraph>
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</caption>
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<paragraph id="8BBB36CCFFCBAE4AFEC232C1FDA84CDF" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">Pronotum strongly convex, 1.75× as wide as long; lateral margins in dorsal view arcuate converging anteriorly. Lateral lobe deplanate, anterolateral and lateral margin distinctly bordered; both lateral angles rounded creating an almost rectangular lateral lobe that is slightly explanate; surface alutaceous and minutely punctured with short recumbent sparse setae; interspaces smooth. Scutellar shield large and triangular with slightly convex margins and alutaceous on the surface.</paragraph>
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<paragraph id="8BBB36CCFFCBAE4AFEC231A8FD0E4D74" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">Elytra evenly convex in lateral aspect, combined width wider than long (1.17× as wide as long). Suture distinctly bordered from scutellar shield all the way to the base. Lateral portions strongly deplanate, almost vertical. Epipleural fold narrowly visible and without setae. Punctures of elytral disc larger than on pronotum, generally distinct and evenly distributed with overall surface alutaceous. Apical margins separately arcuate and pointed at apex with a sinuate margin. Hindwings present.</paragraph>
|
||
<paragraph id="8BBB36CCFFCBAE4AFEC23013FC2A4A66" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">Prosternum strongly carinate in middle. Mesoventrite asetose, depressed and slightly shorter than metaventrite. Metaventrite longer than mesoventrite and 2.6× wider than long. Metaventrite slightly convex, uniformly punctate and setose; setae long and numerous on lateral half becoming smaller and less numerous in the center. Inter-metacoxal distance wide, 2× as wide as inter-mesocoxal distance. Inter-mesocoxal distance narrow with the apical process of metaventrite coming to a rounded point and separating the coxae. Abdominal ventrites covered with long, brownish setae; anterior process of abdominal ventrite I wide, truncate; femoral lines strongly expressed beyond anterior margin of abdominal ventrite I. Abdominal ventrite I longer than abdominal ventrites II-IV combined. Pygidium broadly rounded at apex and covered in setae.</paragraph>
|
||
<paragraph id="8BBB36CCFFCBAE4AFEC23761FE3A4AFB" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">
|
||
Femora wider than tibiae but only slightly expanded. Tibiae with curved margin and expanding slightly towards apex. Tarsomeres lobed, claws simple. Terminal tarsomere longest and longer than tarsomeres II
|
||
<emphasis id="B970EADEFFCBAE4AFAB73740FA594A25" bold="true" box="[1397,1411,1125,1151]" pageId="28" pageNumber="27">–</emphasis>
|
||
III combined (
|
||
<figureCitation id="133F2A49FFCBAE4AFEFC37A2FE0E4AFB" box="[318,468,1159,1185]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 136–138</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBB36CCFFCBAE4AFEC2378CFE094B72" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFEC2378CFE574A99" box="[256,397,1193,1219]" italics="true" pageId="28" pageNumber="27">Median lobe:</emphasis>
|
||
Sides parallel and curving into a rounded point, apical curve slightly convex (Fig. 139,142). In profile, slightly curved from middle (
|
||
<figureCitation id="133F2A49FFCBAE4AFDA237EFFD2E4ABF" box="[608,756,1226,1253]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 140, 143</figureCitation>
|
||
). Median plate on surface elevated.
|
||
<emphasis id="B970EADEFFCBAE4AFB4537EFFADC4ABE" box="[1159,1286,1226,1252]" italics="true" pageId="28" pageNumber="27">Basal plate:</emphasis>
|
||
Sides parallel and curving into a sharp point with crenulate margins along the pointed apex and long setae along the curved apical margin (
|
||
<figureCitation id="133F2A49FFCBAE4AFEAA362BFE1C4B72" box="[360,454,1294,1320]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 141</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBB36CCFFCBAE4AFEC23615FD8B4B36" blockId="28.[191,1443,214,1388]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFEC23615FE814B10" bold="true" box="[256,347,1328,1354]" pageId="28" pageNumber="27">Female.</emphasis>
|
||
Similar to male but with abdominal ventrite V longer than in male with a more tapered shape to the lateral margins of the pygidium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E6547FFCBAE4AFF02365EFD734BCC" box="[192,681,1403,1430]" pageId="28" pageNumber="27" type="distribution">
|
||
<paragraph id="8BBB36CCFFCBAE4AFF02365EFD734BCC" blockId="28.[192,681,1403,1430]" box="[192,681,1403,1430]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFF02365EFE814BCF" bold="true" box="[192,347,1403,1429]" pageId="28" pageNumber="27">Distribution.</emphasis>
|
||
<collectingCountry id="F313765CFFCBAE4AFEA3365EFE6E4BCF" box="[353,436,1403,1429]" name="Mexico" pageId="28" pageNumber="27">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="49C0F82EFFCBAE4AFE03365EFD464BCC" box="[449,668,1403,1430]" country="Mexico" name="Baja California Sur" pageId="28" pageNumber="27">Baja California Sur</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E6547FFCBAE4AFF023680FD364BE5" box="[192,748,1444,1471]" pageId="28" pageNumber="27" type="biology_ecology">
|
||
<paragraph id="8BBB36CCFFCBAE4AFF023680FD364BE5" blockId="28.[192,748,1444,1471]" box="[192,748,1444,1471]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFF023680FED04BE5" bold="true" box="[192,266,1445,1471]" pageId="28" pageNumber="27">Hosts.</emphasis>
|
||
<taxonomicName id="4C044D4FFFCBAE4AFED23680FE434BE5" box="[272,409,1445,1471]" class="Insecta" family="Diaspididae" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="family">Diaspididae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C044D4FFFCBAE4AFE603683FDA94BE5" baseAuthorityName="Bouche" baseAuthorityYear="1833" box="[418,627,1445,1471]" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="echinocacti">
|
||
<emphasis id="B970EADEFFCBAE4AFE603683FDA94BE5" box="[418,627,1445,1471]" italics="true" pageId="28" pageNumber="27">Diaspis echinocacti</emphasis>
|
||
</taxonomicName>
|
||
(Bouch
|
||
<emphasis id="B970EADEFFCBAE4AFD093681FD0D4BE5" bold="true" box="[715,727,1444,1471]" pageId="28" pageNumber="27">é</emphasis>
|
||
)?
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E6547FFCBAE4AFF0236EAFA4148AD" pageId="28" pageNumber="27" type="materials_examined">
|
||
<paragraph id="8BBB36CCFFCBAE4AFF0236EAFA4148AD" blockId="28.[191,1444,1487,1785]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFF0236EAFE074BB3" bold="true" box="[192,477,1487,1513]" pageId="28" pageNumber="27">Type material examined.</emphasis>
|
||
The
|
||
<typeStatus id="54BF886EFFCBAE4AFDD036EAFDAF4BB3" box="[530,629,1487,1513]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
, deposited in the
|
||
<collectionCode id="ED15AE09FFCBAE4AFCF436EAFC594BB3" box="[822,899,1487,1513]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="28" pageNumber="27" type="University or college">UCRC</collectionCode>
|
||
, is a disarticulated male specimen glued to a card with the following labels:
|
||
<collectingRegion id="49C0F82EFFCBAE4AFE1F36D4FD714851" box="[477,683,1521,1547]" inferredBySuffix="true" pageId="28" pageNumber="27">Mex. Baja Cal. Sur</collectingRegion>
|
||
, Las Barracas,
|
||
<date id="FFBA100CFFCBAE4AFC8D36D4FC6D4851" box="[847,951,1521,1547]" pageId="28" pageNumber="27" value="1986-05-02">2-V-1986</date>
|
||
(printed) [white rectangular label] / Coll. P. DeBach (printed) [white rectangular label] / Ex. cactus scale on
|
||
<taxonomicName id="4C044D4FFFCBAE4AFC443537FBF74876" box="[902,1069,1554,1580]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="28" pageNumber="27" phylum="Tracheophyta" rank="species" species="cholla">Opuntia cholla</taxonomicName>
|
||
(handwritten) [white rectangular label] /
|
||
<typeStatus id="54BF886EFFCBAE4AFEDA3511FE724814" box="[280,424,1588,1614]" pageId="28" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C044D4FFFCBAE4AFE713511FD464814" authorityName="Smith" authorityYear="2022" box="[435,668,1588,1614]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="skelleyi">
|
||
<emphasis id="B970EADEFFCBAE4AFE713511FD464814" box="[435,668,1588,1614]" italics="true" pageId="28" pageNumber="27">Cybocephalus skelleyi</emphasis>
|
||
</taxonomicName>
|
||
<collectionCode id="ED15AE09FFCBAE4AFD603510FD694815" box="[674,691,1589,1615]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="27">T</collectionCode>
|
||
.
|
||
<collectionCode id="ED15AE09FFCBAE4AFD7D3510FD094815" box="[703,723,1589,1615]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="27">R</collectionCode>
|
||
. Smith Det: Trevor Smith (printed) [red rectangular label]. The
|
||
<typeStatus id="54BF886EFFCBAE4AFF003573FEC6482A" box="[194,284,1622,1648]" pageId="28" pageNumber="27" type="allotype">allotype</typeStatus>
|
||
, deposited in the
|
||
<collectionCode id="ED15AE09FFCBAE4AFE193573FDF2482A" box="[475,552,1622,1648]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="28" pageNumber="27" type="University or college">UCRC</collectionCode>
|
||
, is a female specimen glued to a card with the following labels:
|
||
<collectingRegion id="49C0F82EFFCBAE4AFB173573FA44482A" box="[1237,1438,1622,1648]" inferredBySuffix="true" pageId="28" pageNumber="27">Mex. Baja Cal. Sur</collectingRegion>
|
||
, Las Barracas,
|
||
<date id="FFBA100CFFCBAE4AFE9B355DFE1B48C8" box="[345,449,1656,1682]" pageId="28" pageNumber="27" value="1986-05-02">2-V-1986</date>
|
||
(printed) [white rectangular label] / Coll. P. DeBach (printed) [white rectangular label] / Ex. cactus scale on
|
||
<taxonomicName id="4C044D4FFFCBAE4AFE5535BCFDE448E9" box="[407,574,1689,1715]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="28" pageNumber="27" phylum="Tracheophyta" rank="species" species="cholla">Opuntia cholla</taxonomicName>
|
||
(handwritten) [white rectangular label] /
|
||
<typeStatus id="54BF886EFFCBAE4AFBCC35BFFB4C48EE" box="[1038,1174,1690,1716]" pageId="28" pageNumber="27" type="allotype">ALLOTYPE</typeStatus>
|
||
<taxonomicName id="4C044D4FFFCBAE4AFB5F35BCFA5F48E9" authorityName="Smith" authorityYear="2022" box="[1181,1413,1689,1715]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="skelleyi">
|
||
<emphasis id="B970EADEFFCBAE4AFB5F35BCFA5F48E9" box="[1181,1413,1689,1715]" italics="true" pageId="28" pageNumber="27">Cybocephalus skelleyi</emphasis>
|
||
</taxonomicName>
|
||
<collectionCode id="ED15AE09FFCBAE4AFA4E35BFFA7A48EE" box="[1420,1440,1690,1716]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="27">T</collectionCode>
|
||
.
|
||
<collectionCode id="ED15AE09FFCBAE4AFF023599FF0E488C" box="[192,212,1724,1750]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="27">R</collectionCode>
|
||
. Smith Det: Trevor Smith (printed) [blue rectangular label].
|
||
<typeStatus id="54BF886EFFCBAE4AFCB53599FC3F488C" box="[887,997,1724,1750]" pageId="28" pageNumber="27" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectingCountry id="F313765CFFCBAE4AFC2D359EFB87488F" box="[1007,1117,1723,1749]" name="Mexico" pageId="28" pageNumber="27">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C0F82EFFCBAE4AFBAA359EFB66488F" box="[1128,1212,1723,1749]" country="Mexico" name="Mexico" pageId="28" pageNumber="27">Mexico</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C0F82EFFCBAE4AFB05359EFA46488F" box="[1223,1436,1723,1749]" country="Mexico" name="Baja California Sur" pageId="28" pageNumber="27">Baja California Sur</collectingRegion>
|
||
, Las Barracas;
|
||
<date id="FFBA100CFFCBAE4AFE9B35F8FE1948AD" box="[345,451,1757,1783]" pageId="28" pageNumber="27" value="1986-05-02">2-V-1986</date>
|
||
; Coll. P. DeBach; Ex. cactus scale on
|
||
<taxonomicName id="4C044D4FFFCBAE4AFC9535F8FC2748AD" box="[855,1021,1757,1783]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="28" pageNumber="27" phylum="Tracheophyta" rank="species" species="cholla">Opuntia cholla</taxonomicName>
|
||
(
|
||
<specimenCount id="9D02FD45FFCBAE4AFBC935F8FBF748A3" box="[1035,1069,1757,1785]" pageId="28" pageNumber="27" type="female">4♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED15AE09FFCBAE4AFBF835F8FB5D48AD" box="[1082,1159,1757,1783]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="28" pageNumber="27" type="University or college">UCRC</collectionCode>
|
||
;
|
||
<specimenCount id="9D02FD45FFCBAE4AFB5335F8FB6948A3" box="[1169,1203,1757,1785]" pageId="28" pageNumber="27" type="female">2♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED15AE09FFCBAE4AFB7D35F8FAD948AD" box="[1215,1283,1757,1783]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="28" pageNumber="27" type="Museum">FSCA</collectionCode>
|
||
;
|
||
<specimenCount id="9D02FD45FFCBAE4AFACF35F8FAEA48A3" box="[1293,1328,1757,1785]" pageId="28" pageNumber="27" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED15AE09FFCBAE4AFAFE35F8FA4A48AD" box="[1340,1424,1757,1783]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="27" type="Museum">USNM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E6547FFCBAE4BFF023422FB7A4F0E" lastPageId="29" lastPageNumber="28" pageId="28" pageNumber="27" type="discussion">
|
||
<paragraph id="8BBB36CCFFCBAE4AFF023422FAD349DC" blockId="28.[191,1440,1799,1927]" pageId="28" pageNumber="27">
|
||
<emphasis id="B970EADEFFCBAE4AFF023422FEF5497B" bold="true" box="[192,303,1799,1825]" pageId="28" pageNumber="27">Remarks.</emphasis>
|
||
The aedeagus of the
|
||
<typeStatus id="54BF886EFFCBAE4AFDD43422FDA0497B" box="[534,634,1799,1825]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
, the only known male, was slightly damaged. Therefore, the author illustrated the median lobe as it actually appears and has been preserved (
|
||
<figureCitation id="133F2A49FFCBAE4AFC78340DFB924919" box="[954,1096,1832,1859]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 139-140</figureCitation>
|
||
) but also provided illustrations of the median lobe as it most likely looks undamaged (
|
||
<figureCitation id="133F2A49FFCBAE4AFCFF346FFC08493E" box="[829,978,1866,1892]" captionStart="Figures 135–143" captionStartId="27.[192,279,1799,1825]" captionTargetBox="[230,1402,316,1741]" captionTargetId="figure-60@27.[192,1440,280,1770]" captionTargetPageId="27" captionText="Figures 135–143. Cybocephalus skelleyi T. R. Smith. 135) Antenna. 136) Proleg. 137) Mesoleg. 138) Metaleg. 139) Median lobe, as it appears in holotype. 140) Median lobe, lateral view, as it appears in the holotype. 141) Basal plate. 142) Median lobe, hypothetical reconstruction. 143) Median lobe, lateral view, hypothetical reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7300650" httpUri="https://zenodo.org/record/7300650/files/figure.png" pageId="28" pageNumber="27">Fig. 142-143</figureCitation>
|
||
). These hypothetical reconstructions are based on the author’s extensive experience with the anatomy and morphology of this beetle family.
|
||
</paragraph>
|
||
<paragraph id="8BBB36CCFFCAAE4BFEC233F3FB7A4F0E" blockId="29.[192,1440,214,341]" pageId="29" pageNumber="28">
|
||
This species was collected feeding on scale insects labeled as “cactus scale” infesting
|
||
<taxonomicName id="4C044D4FFFCAAE4BFB0633F3FAB04EAA" box="[1220,1386,214,240]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="29" pageNumber="27" phylum="Tracheophyta" rank="species" species="cholla">opuntia cholla</taxonomicName>
|
||
. The common name
|
||
<taxonomicName id="4C044D4FFFCAAE4BFEB233D2FDD54F4B" box="[368,527,247,273]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="29" pageNumber="27" phylum="Tracheophyta" rank="species" species="cholla">opuntia cholla</taxonomicName>
|
||
refers to a group of over 20 species in the genus
|
||
<taxonomicName id="4C044D4FFFCAAE4BFBEE33DDFB654F4B" authority="Mill" authorityName="Mill" box="[1068,1215,247,274]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="29" pageNumber="28" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B970EADEFFCAAE4BFBEE33DDFB524F48" box="[1068,1160,248,274]" italics="true" pageId="29" pageNumber="28">Opuntia</emphasis>
|
||
Mill
|
||
</taxonomicName>
|
||
..
|
||
<taxonomicName id="4C044D4FFFCAAE4BFB1233DDFA7A4F4B" baseAuthorityName="Bouche" baseAuthorityYear="1833" box="[1232,1440,247,273]" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="echinocacti">
|
||
<emphasis id="B970EADEFFCAAE4BFB1233DDFA7A4F4B" box="[1232,1440,247,273]" italics="true" pageId="29" pageNumber="28">Diaspis echinocacti</emphasis>
|
||
</taxonomicName>
|
||
(Bouch
|
||
<emphasis id="B970EADEFFCAAE4BFED0323DFEC44F69" bold="true" box="[274,286,280,307]" pageId="29" pageNumber="28">é</emphasis>
|
||
) is commonly referred to as the cactus scale and is most likely what the collector was referring to.
|
||
<taxonomicName id="4C044D4FFFCAAE4BFA8C323FFEE34F0E" baseAuthorityName="Bouche" baseAuthorityYear="1833" class="Insecta" family="Diaspididae" genus="Diaspis" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="echinocacti">
|
||
<emphasis id="B970EADEFFCAAE4BFA8C323FFEE34F0E" italics="true" pageId="29" pageNumber="28">Diaspis echinocacti</emphasis>
|
||
</taxonomicName>
|
||
is a known host of
|
||
<taxonomicName id="4C044D4FFFCAAE4BFDD3321EFD744F0E" box="[529,686,314,341]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B970EADEFFCAAE4BFDD3321EFD744F0E" box="[529,686,314,341]" italics="true" pageId="29" pageNumber="28">C. californicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B970EADEFFCAAE4BFD26321EFCA64F0E" box="[740,892,314,341]" italics="true" pageId="29" pageNumber="28">
|
||
<taxonomicName id="4C044D4FFFCAAE4BFD26321EFCA34F0E" authorityName="T. R. Smith in T. R. Smith & Cave" authorityYear="2007" box="[740,889,314,341]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="championi">C. championi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
also occurring in
|
||
<collectingCountry id="F313765CFFCAAE4BFB85321FFB464F0E" box="[1095,1180,314,340]" name="Mexico" pageId="29" pageNumber="28">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |