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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3841.1.1" ID-GBIF-Dataset="07cca84d-e413-47d9-9ea5-5d24be4dce1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="286746" ID-ZooBank="5FAADB3D-2359-453F-8BF5-1C2A33C6D178" checkinTime="1460460685218" checkinUser="plazi" docAuthor="Gerecke, Reinhard" docDate="2014" docId="03AD87B0FF91144BFF46BCDCFE8E803A" docLanguage="en" docName="zt03841p046.pdf" docOrigin="Zootaxa 3841 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@4b05a9b1" docStyleName="zootaxa.2013.journal_article" docTitle="Ljania bipapillata subsp. subterranea Schwoerbel 1964" docType="treatment" docVersion="5" lastPageNumber="35" masterDocId="FF94FFC8FFB31469FFD1BC4BFFE2830B" masterDocTitle="Studies on European species of the water mite family Aturidae Thor (Acari: Hydrachnidia)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="35" updateTime="1677002133583" updateUser="tatiana">
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<mods:title>Studies on European species of the water mite family Aturidae Thor (Acari: Hydrachnidia)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gerecke, Reinhard</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>3841</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3841.1.1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6144809" ID-GBIF-Taxon="127669179" ID-Zenodo-Dep="6144809" LSID="urn:lsid:plazi:treatment:03AD87B0FF91144BFF46BCDCFE8E803A" httpUri="http://treatment.plazi.org/id/03AD87B0FF91144BFF46BCDCFE8E803A" lastPageNumber="35" pageId="34" pageNumber="35">
<subSubSection box="[151,744,151,178]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph blockId="34.[151,744,151,211]" box="[151,744,151,178]" pageId="34" pageNumber="35">
<heading bold="true" box="[151,744,151,178]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<taxonomicName authority="Schwoerbel, 1964" authorityName="Schwoerbel" authorityYear="1964" box="[151,744,151,178]" class="Arachnida" family="Axonopsidae" genus="Ljania" kingdom="Animalia" order="Trombidiformes" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="bipapillata" subSpecies="subterranea">
<emphasis bold="true" box="[151,744,151,178]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[151,519,151,177]" italics="true" pageId="34" pageNumber="35">Ljania bipapillata subterranea</emphasis>
Schwoerbel,
<number box="[685,744,152,178]" pageId="34" pageNumber="35" value="1964.0">1964</number>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,358,186,211]" pageId="34" pageNumber="35" type="reference_group">
<paragraph blockId="34.[151,744,151,211]" box="[151,358,186,211]" pageId="34" pageNumber="35">Subspecies incerta</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="discussion">
<paragraph blockId="34.[151,1437,253,817]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[151,265,253,278]" pageId="34" pageNumber="35">Remarks.</emphasis>
The name
<taxonomicName box="[386,673,253,277]" class="Arachnida" family="Axonopsidae" genus="Ljania" kingdom="Animalia" order="Trombidiformes" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="bipapillata" subSpecies="subterranea">
<emphasis box="[386,673,253,277]" italics="true" pageId="34" pageNumber="35">L. bipapillata subterranea</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation author="Schwoerbel" box="[886,1108,253,278]" pageId="34" pageNumber="35" refString="Schwoerbel, J. (1964 a) Die Bedeutung des Hyporheals fur die benthische Lebensgemeinschaft der Fließgewasser. Verhandlungen des Internationalen Vereins fur theoretische und angewandte Limnologie, 15, 215 - 226." type="journal article" year="1964" yearSuffix="a">
Schwoerbel (
<number box="[1030,1087,253,278]" pageId="34" pageNumber="35" value="1964.0">1964</number>
a)
</bibRefCitation>
without giving any indication about
<typeStatus box="[218,265,290,314]" pageId="34" pageNumber="35">type</typeStatus>
material or a
<typeStatus box="[414,461,290,314]" pageId="34" pageNumber="35">type</typeStatus>
locality (“….diejenigen Populationen, die in das Hyporheal der Bäche eingedrungen sind und hier charakteristische Umbildungen ihres Körpers und ihrer Lebensweise erfahren haben. Diese Populationen sind bei
<bibRefCitation author="Schwoerbel" box="[251,458,361,386]" pageId="34" pageNumber="35" refString="Schwoerbel, J. (1961) Subterrane Wassermilben (Acari: Hydrachnellae, Porohalacaridae und Stygothrombiidae), ihre Okologie und Bedeutung fur die Abgrenzung eines aquatischen Lebensraums zwischen Oberflache und Grundwasser. Archiv fur Hydrobiologie, Supplement 25, 242 - 306." type="book chapter" year="1961" yearSuffix="a">
Schwoerbel
<number box="[391,446,361,386]" pageId="34" pageNumber="35" value="1961.0">1961</number>
a
</bibRefCitation>
und
<number box="[521,576,361,386]" pageId="34" pageNumber="35" value="1964.0">1964</number>
ausführlich gekennzeichnet worden“ “those populations of
<taxonomicName box="[1283,1436,361,385]" class="Arachnida" family="Axonopsidae" genus="Ljania" kingdom="Animalia" order="Trombidiformes" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bipapillata">
<emphasis box="[1283,1436,361,385]" italics="true" pageId="34" pageNumber="35">L. bipapillata</emphasis>
</taxonomicName>
which have immigrated to the hyporheal of streams and here have undergone characteristic transformations of body and life style. These populations were characterized in detail in
<bibRefCitation author="Schwoerbel" box="[862,1066,433,458]" pageId="34" pageNumber="35" refString="Schwoerbel, J. (1961) Subterrane Wassermilben (Acari: Hydrachnellae, Porohalacaridae und Stygothrombiidae), ihre Okologie und Bedeutung fur die Abgrenzung eines aquatischen Lebensraums zwischen Oberflache und Grundwasser. Archiv fur Hydrobiologie, Supplement 25, 242 - 306." type="book chapter" year="1961" yearSuffix="a">
Schwoerbel
<number box="[999,1054,433,458]" pageId="34" pageNumber="35" value="1961.0">1961</number>
a
</bibRefCitation>
and
<number box="[1122,1182,433,458]" pageId="34" pageNumber="35" value="1964.0">1964</number>
”). In the two papers in question we find indications on various populations differing exclusively in mean values and ranges of the idiosoma length/width ratio, and a hypothesis that these differences in idiosoma shape should have developed in parallel to the immigration of the populations in question to the interstitial habitat.
</paragraph>
<paragraph blockId="34.[151,1437,253,817]" pageId="34" pageNumber="35">
There is no chance to select a
<typeStatus box="[536,625,578,602]" pageId="34" pageNumber="35">neotype</typeStatus>
specimen from Schwoerbel's collection or from a specific site in nature. If we would decide to accept the name notwithstanding these negative preconditions, it would surely become a junior synonym of
<taxonomicName authority="The" authorityName="The" box="[389,616,649,674]" class="Arachnida" family="Axonopsidae" genus="Ljania" kingdom="Animalia" order="Trombidiformes" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bipapillata">
<emphasis box="[389,550,649,673]" italics="true" pageId="34" pageNumber="35">L. bipapillata</emphasis>
: The
</taxonomicName>
character state indicated as typical for the taxon is found broadly overlapping in populations from different sizes, as is documented also by a comparison of data from
<bibRefCitation author="Schwoerbel" pageId="34" pageNumber="35" refString="Schwoerbel, J. (1964 a) Die Bedeutung des Hyporheals fur die benthische Lebensgemeinschaft der Fließgewasser. Verhandlungen des Internationalen Vereins fur theoretische und angewandte Limnologie, 15, 215 - 226." type="journal article" year="1964" yearSuffix="a">
Schwoerbel (
<number box="[159,217,721,746]" pageId="34" pageNumber="35" value="1964.0">1964</number>
a)
</bibRefCitation>
and
<bibRefCitation author="Tuzovskij" box="[301,502,721,746]" pageId="34" pageNumber="35" refString="Tuzovskij, P. V. (2012) Water mites of the genus Ljania Thor, 1898 (Acari: Hydrachnidia: Aturidae) in Russia. Zootaxa, 3249, 1 - 17." type="journal article" year="2012">
Tuzovskij (
<number box="[432,494,722,746]" pageId="34" pageNumber="35" value="2012.0">2012</number>
)
</bibRefCitation>
. Schwoerbel had possibly detected in his studies an interesting phenomenon of infraspecific morphological variation that merits attention in future studies, but the taxon has to be considered a
<emphasis box="[151,356,793,817]" italics="true" pageId="34" pageNumber="35">subspecies incerta</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
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