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<document ID-DOI="10.11865/zs.2021402" ID-GBIF-Dataset="322fcc8b-1bb1-4dbe-9607-d803273e26eb" ID-ISSN="2095-6827" ID-Zenodo-Dep="7175787" ID-ZooBank="E29E0326-F45F-4749-B930-0DD9062FCCBC" checkinTime="1665294850988" checkinUser="felipe" docAuthor="Li, Haitao, Jin, Daochao &amp; Guo, Jianjun" docDate="2021" docId="0387972EFFE4FFA0FF1AFE7474642A72" docLanguage="en" docName="ZoolSyst.46.4.281288.pdf" docOrigin="Zoological Systematics 46 (4)" docSource="http://zoobank.org/e29e0326-f45f-4749-b930-0dd9062fccbc" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Stygothrombium garzensis Li &amp; Guo 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="287" masterDocId="FFBEEF56FFE6FFA7FF92FFBE77492B55" masterDocTitle="A new species of the genus Stygothrombium, representing a newly recorded superfamily Stygothrombioidea Mullen &amp; Vercammen-Grandjean, 1980 (Acari: Stygothrombiae) from China" masterLastPageNumber="288" masterPageNumber="281" pageNumber="283" updateTime="1665312989738" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A new species of the genus Stygothrombium, representing a newly recorded superfamily Stygothrombioidea Mullen &amp; Vercammen-Grandjean, 1980 (Acari: Stygothrombiae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Haitao</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jin, Daochao</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guo, Jianjun</mods:namePart>
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<mods:title>Zoological Systematics</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:identifier type="DOI">10.11865/zs.2021402</mods:identifier>
<mods:identifier type="GBIF-Dataset">322fcc8b-1bb1-4dbe-9607-d803273e26eb</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172154" ID-GBIF-Taxon="202185823" ID-Zenodo-Dep="7172154" LSID="urn:lsid:plazi:treatment:0387972EFFE4FFA0FF1AFE7474642A72" httpUri="http://treatment.plazi.org/id/0387972EFFE4FFA0FF1AFE7474642A72" lastPageId="7" lastPageNumber="287" pageId="2" pageNumber="283">
<subSubSection pageId="2" pageNumber="283" type="nomenclature">
<paragraph blockId="2.[136,777,458,482]" box="[136,777,458,482]" pageId="2" pageNumber="283">
<heading box="[136,777,458,482]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="283" reason="2">
<emphasis bold="true" box="[136,644,458,482]" pageId="2" pageNumber="283">
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[136,546,458,482]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis bold="true" box="[136,429,458,482]" italics="true" pageId="2" pageNumber="283">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<taxonomicNameLabel box="[559,644,458,482]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[674,768,458,482]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="2.[155,218,2014,2036]" captionStartId-1="3.[155,218,2027,2049]" captionStartId-2="4.[155,218,2024,2046]" captionStartId-3="5.[155,218,2024,2046]" captionStartId-4="6.[155,218,1791,1813]" captionTargetBox-0="[189,1399,820,1984]" captionTargetBox-1="[199,1388,723,1984]" captionTargetBox-2="[194,1392,711,1987]" captionTargetBox-3="[200,1386,726,1961]" captionTargetBox-4="[189,1398,473,1758]" captionTargetId-0="figure-188@2.[189,1399,820,1984]" captionTargetId-1="figure-267@3.[189,1399,710,1995]" captionTargetId-2="figure-299@4.[189,1399,707,1992]" captionTargetId-3="figure-259@5.[189,1398,721,1976]" captionTargetId-4="figure-225@6.[189,1398,473,1758]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Figure 1. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Ventral view. B. Gll-1. C. Gll-2. D. I-L-4, stomatoid lyrifissures. Scale bars: A=500μm; BD=10μm." captionText-1="Figure 2. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Dorsal view. B. Ventral view. Scale bars =500μm." captionText-2="Figure 3. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar=200μm." captionText-3="Figure 4. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Anterior coxal group (Cx-I+Cx-II). B. Posterior coxal group (Cx-III+Cx-IV), genital field. Scale bar=200μm." captionText-4="Figure 5. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. I-L-16. B. II-L-16. C. III-L-16. D. IV-L-16. Scale bar=200μm." figureDoi-0="http://doi.org/10.5281/zenodo.7175789" figureDoi-1="http://doi.org/10.5281/zenodo.7175791" figureDoi-2="http://doi.org/10.5281/zenodo.7175793" figureDoi-3="http://doi.org/10.5281/zenodo.7175795" figureDoi-4="http://doi.org/10.5281/zenodo.7175797" httpUri-0="https://zenodo.org/record/7175789/files/figure.png" httpUri-1="https://zenodo.org/record/7175791/files/figure.png" httpUri-2="https://zenodo.org/record/7175793/files/figure.png" httpUri-3="https://zenodo.org/record/7175795/files/figure.png" httpUri-4="https://zenodo.org/record/7175797/files/figure.png" pageId="2" pageNumber="283">Figs 15</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="2.[136,1452,514,781]" box="[189,475,514,538]" pageId="2" pageNumber="283">Habitat. Interstitial waters.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="287" pageId="2" pageNumber="283" type="materials_examined">
<paragraph blockId="2.[136,1452,514,781]" pageId="2" pageNumber="283">
<materialsCitation collectingDate="2020-08-30" collectorName="Boyan Li &amp; Slides No." country="China" elevation="3523" location="Material" municipality="Chaqingsongduo National Nature Reserve" pageId="2" pageNumber="283" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<location LSID="urn:lsid:plazi:treatment:0387972EFFE4FFA0FF1AFE7474642A72:8EF170E3FFE4FFA5FF2FFD9A765E2969" box="[189,279,548,572]" country="China" municipality="Chaqingsongduo National Nature Reserve" name="Material" pageId="2" pageNumber="283" stateProvince="Sichuan">Material</location>
examined.
<typeStatus box="[419,518,548,572]" pageId="2" pageNumber="283">Holotype</typeStatus>
female,
<collectingMunicipality box="[627,1105,548,572]" pageId="2" pageNumber="283">Chaqingsongduo National Nature Reserve</collectingMunicipality>
,
<collectingRegion box="[1123,1212,548,572]" country="China" name="Sichuan" pageId="2" pageNumber="283">Sichuan</collectingRegion>
,
<collectingCountry box="[1230,1293,548,572]" name="China" pageId="2" pageNumber="283">China</collectingCountry>
(31°0088N, 99°2471E, elev.
<quantity box="[333,415,583,607]" metricMagnitude="3" metricUnit="m" metricValue="3.523" pageId="2" pageNumber="283" unit="m" value="3523.0">
<elevation box="[333,415,583,607]" metricMagnitude="3" metricUnit="m" metricValue="3.523" pageId="2" pageNumber="283" unit="m" value="3523.0">3523 m</elevation>
</quantity>
), water depth
<quantity box="[570,676,583,607]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="283" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">3040 cm</quantity>
, located at the hillside, running water with organic detritus, dead wood and leaves on the bottom, collected by
<collectorName box="[552,655,618,642]" pageId="2" pageNumber="283">Boyan Li</collectorName>
,
<date box="[666,806,618,642]" pageId="2" pageNumber="283" value="2020-08-30">
<collectingDate box="[666,806,618,642]" pageId="2" pageNumber="283" value="2020-08-30">30.VIII.2020</collectingDate>
</date>
,
<collectorName box="[817,929,618,642]" pageId="2" pageNumber="283">Slides No.</collectorName>
SC-ST-2020080101.
</materialsCitation>
</paragraph>
<paragraph blockId="2.[136,1452,514,781]" pageId="2" pageNumber="283">
Etymology. “
<emphasis box="[331,390,653,676]" italics="true" pageId="2" pageNumber="283">garze</emphasis>
-” is derived from the name of the Garze Tibetan Autonomous Prefecture,
<collectingRegion box="[1181,1371,652,676]" country="China" name="Sichuan" pageId="2" pageNumber="283">Sichuan Province</collectingRegion>
,
<collectingCountry box="[1381,1447,652,676]" name="China" pageId="2" pageNumber="283">China</collectingCountry>
, where the specimen was collected.
</paragraph>
<paragraph blockId="2.[136,1452,514,781]" lastBlockId="3.[136,1452,236,677]" lastPageId="3" lastPageNumber="284" pageId="2" pageNumber="283">
Diagnosis. Soft and papillate idiosoma vermiform; eyes completely absent; infracapitulum base approximately 3.5 times as long as rostrum; P-A with two stout subventrodistal harpagones, above harpago seta thicker than other one; on one side of P-B, tibia with one four-pronged odontus claws bulging above tarsus;
<emphasis box="[986,1008,237,260]" italics="true" pageId="3" pageNumber="284">se</emphasis>
about half length of posterior portion of prodorsal plate; one seta and two rather long setae at lateral margin of Cx-I; stalked Ac-13 arranging in an almost straight line,
<emphasis box="[193,235,306,330]" italics="true" pageId="3" pageNumber="284">glv-</emphasis>
2 between Ac-3 and inner posterior angle of Cx-IV, located at one-third of interval near Ac-3 approximately; empodium considerably smaller than lateral claws.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175789" ID-Zenodo-Dep="7175789" httpUri="https://zenodo.org/record/7175789/files/figure.png" pageId="2" pageNumber="283" startId="2.[155,218,2014,2036]" targetBox="[189,1399,820,1984]" targetPageId="2">
<paragraph blockId="2.[155,1433,2014,2068]" pageId="2" pageNumber="283">
Figure 1.
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[257,626,2014,2036]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[257,516,2014,2036]" italics="true" pageId="2" pageNumber="283">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[645,726,2015,2036]" pageId="2" pageNumber="283">
<taxonomicNameLabel box="[645,726,2015,2036]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (SC-ST-20200801). A. Ventral view. B.
<emphasis box="[1276,1307,2014,2035]" italics="true" pageId="2" pageNumber="283">Gll</emphasis>
-1. C.
<emphasis box="[1376,1407,2014,2035]" italics="true" pageId="2" pageNumber="283">Gll</emphasis>
-2. D. I-L-4, stomatoid lyrifissures. Scale bars: A = 500 μm; BD = 10 μm.
</paragraph>
</caption>
<paragraph blockId="3.[136,1452,236,677]" pageId="3" pageNumber="284">
Description. Female (SC-ST-2020080101). Soft and papillate idiosoma vermiform (
<figureCitation box="[1093,1180,375,399]" captionStart="Figure 1" captionStartId="2.[155,218,2014,2036]" captionTargetBox="[189,1399,820,1984]" captionTargetId="figure-188@2.[189,1399,820,1984]" captionTargetPageId="2" captionText="Figure 1. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Ventral view. B. Gll-1. C. Gll-2. D. I-L-4, stomatoid lyrifissures. Scale bars: A=500μm; BD=10μm." figureDoi="http://doi.org/10.5281/zenodo.7175789" httpUri="https://zenodo.org/record/7175789/files/figure.png" pageId="3" pageNumber="284">Fig. 1A</figureCitation>
); eyes completely absent (
<figureCitation box="[144,231,410,434]" captionStart="Figure 2" captionStartId="3.[155,218,2027,2049]" captionTargetBox="[199,1388,723,1984]" captionTargetId="figure-267@3.[189,1399,710,1995]" captionTargetPageId="3" captionText="Figure 2. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Dorsal view. B. Ventral view. Scale bars =500μm." figureDoi="http://doi.org/10.5281/zenodo.7175791" httpUri="https://zenodo.org/record/7175791/files/figure.png" pageId="3" pageNumber="284">Fig. 2A</figureCitation>
); with eight pairs of
<emphasis box="[456,490,410,434]" italics="true" pageId="3" pageNumber="284">gld</emphasis>
and seven pairs of
<emphasis box="[701,729,410,434]" italics="true" pageId="3" pageNumber="284">gll</emphasis>
(
<emphasis box="[745,788,410,434]" italics="true" pageId="3" pageNumber="284">gld-</emphasis>
18,
<emphasis box="[842,879,410,434]" italics="true" pageId="3" pageNumber="284">gll-</emphasis>
<quantity box="[879,948,410,434]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.778" metricValueMin="0.254" pageId="3" pageNumber="284" unit="in" value="4.0" valueMax="7.0" valueMin="1.0">17 in</quantity>
<figureCitation box="[955,1041,410,434]" captionStart="Figure 2" captionStartId="3.[155,218,2027,2049]" captionTargetBox="[199,1388,723,1984]" captionTargetId="figure-267@3.[189,1399,710,1995]" captionTargetPageId="3" captionText="Figure 2. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Dorsal view. B. Ventral view. Scale bars =500μm." figureDoi="http://doi.org/10.5281/zenodo.7175791" httpUri="https://zenodo.org/record/7175791/files/figure.png" pageId="3" pageNumber="284">Fig. 2A</figureCitation>
), six pairs of
<emphasis box="[1188,1221,410,434]" italics="true" pageId="3" pageNumber="284">glv</emphasis>
(
<emphasis box="[1237,1278,410,434]" italics="true" pageId="3" pageNumber="284">glv-</emphasis>
<quantity box="[1279,1347,410,434]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="15.24" metricValueMin="2.54" pageId="3" pageNumber="284" unit="in" value="3.5" valueMax="6.0" valueMin="1.0">16 in</quantity>
<figureCitation box="[1354,1438,410,434]" captionStart="Figure 2" captionStartId="3.[155,218,2027,2049]" captionTargetBox="[199,1388,723,1984]" captionTargetId="figure-267@3.[189,1399,710,1995]" captionTargetPageId="3" captionText="Figure 2. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Dorsal view. B. Ventral view. Scale bars =500μm." figureDoi="http://doi.org/10.5281/zenodo.7175791" httpUri="https://zenodo.org/record/7175791/files/figure.png" pageId="3" pageNumber="284">Fig. 2B</figureCitation>
); idiosomal glandularia (or modified stomatoid lyrifissures) without sclerite platelets but surrounded by unpapillate cuticle, a long seta and short seta associated with glandularia except
<emphasis box="[773,801,479,503]" italics="true" pageId="3" pageNumber="284">gll</emphasis>
-1 with only a short seta (
<figureCitation box="[1078,1198,479,503]" captionStart="Figure 1" captionStartId="2.[155,218,2014,2036]" captionTargetBox="[189,1399,820,1984]" captionTargetId="figure-188@2.[189,1399,820,1984]" captionTargetPageId="2" captionText="Figure 1. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Ventral view. B. Gll-1. C. Gll-2. D. I-L-4, stomatoid lyrifissures. Scale bars: A=500μm; BD=10μm." figureDoi="http://doi.org/10.5281/zenodo.7175789" httpUri="https://zenodo.org/record/7175789/files/figure.png" pageId="3" pageNumber="284">Figs 1BC</figureCitation>
); excretory pore placed near posterior end of ventral surface and between
<emphasis box="[669,701,514,538]" italics="true" pageId="3" pageNumber="284">glv</emphasis>
-5 (
<figureCitation box="[739,823,514,538]" captionStart="Figure 2" captionStartId="3.[155,218,2027,2049]" captionTargetBox="[199,1388,723,1984]" captionTargetId="figure-267@3.[189,1399,710,1995]" captionTargetPageId="3" captionText="Figure 2. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Dorsal view. B. Ventral view. Scale bars =500μm." figureDoi="http://doi.org/10.5281/zenodo.7175791" httpUri="https://zenodo.org/record/7175791/files/figure.png" pageId="3" pageNumber="284">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph blockId="3.[136,1452,236,677]" lastBlockId="4.[136,1452,236,642]" lastPageId="4" lastPageNumber="285" pageId="3" pageNumber="284">
Gnathosoma retractable into idiosoma; infracapitulum base approximately 3.5 times as long as rostrum; two pairs of setae on rostrum, posterior one longer than anterior one (
<figureCitation box="[754,841,583,607]" captionStart="Figure 3" captionStartId="4.[155,218,2024,2046]" captionTargetBox="[194,1392,711,1987]" captionTargetId="figure-299@4.[189,1399,707,1992]" captionTargetPageId="4" captionText="Figure 3. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175793" httpUri="https://zenodo.org/record/7175793/files/figure.png" pageId="3" pageNumber="284">Fig. 3A</figureCitation>
); chelicera two-segmented, basal segment expanded and long (
<figureCitation box="[198,280,618,642]" captionStart="Figure 3" captionStartId="4.[155,218,2024,2046]" captionTargetBox="[194,1392,711,1987]" captionTargetId="figure-299@4.[189,1399,707,1992]" captionTargetPageId="4" captionText="Figure 3. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175793" httpUri="https://zenodo.org/record/7175793/files/figure.png" pageId="3" pageNumber="284">Fig. 3B</figureCitation>
); fused pedipalp two-segmented: first segment including trochanter + femur + genu (P-A); P-A with numerous dorsal setae, a thin ventrodistal seta, two stout subventrodistal harpagones (above harpago seta thicker than other one) and two ventroproximal setae (one normal seta and one long curved seta); tibia + tarsus fused into short second segment (P-B); on one side of P-B, tibia with one four-pronged odontus claws bulging above tarsus, one stout gladius seta, one thick and short lancea seta, one parodontus, and two normal setae on tarsus; P-B with six normal setae on another side, single solenidion on dorsum and alantoid seta with rounded tip at terminus (
<figureCitation box="[876,998,340,365]" captionStart="Figure 3" captionStartId="4.[155,218,2024,2046]" captionTargetBox="[194,1392,711,1987]" captionTargetId="figure-299@4.[189,1399,707,1992]" captionTargetPageId="4" captionText="Figure 3. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175793" httpUri="https://zenodo.org/record/7175793/files/figure.png" pageId="4" pageNumber="285">Figs 3CD</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175791" ID-Zenodo-Dep="7175791" httpUri="https://zenodo.org/record/7175791/files/figure.png" pageId="3" pageNumber="284" startId="3.[155,218,2027,2049]" targetBox="[199,1388,723,1984]" targetPageId="3">
<paragraph blockId="3.[155,1432,2027,2081]" pageId="3" pageNumber="284">
Figure 2.
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[248,604,2027,2049]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[248,503,2027,2049]" italics="true" pageId="3" pageNumber="284">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[614,691,2028,2049]" pageId="3" pageNumber="284">
<taxonomicNameLabel box="[614,691,2028,2049]" pageId="3" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (SC-ST-20200801). A. Dorsal view. B. Ventral view. Scale bars = 500 μm.
</paragraph>
</caption>
<paragraph blockId="4.[136,1452,236,642]" pageId="4" pageNumber="285">
Prodorsal plate bearing
<emphasis box="[446,465,376,399]" italics="true" pageId="4" pageNumber="285">vi</emphasis>
,
<emphasis box="[479,497,376,399]" italics="true" pageId="4" pageNumber="285">si</emphasis>
,
<emphasis box="[511,535,376,399]" italics="true" pageId="4" pageNumber="285">ve</emphasis>
and
<emphasis box="[588,610,376,399]" italics="true" pageId="4" pageNumber="285">se</emphasis>
; long and narrow posterior portion of prodorsal plate approximately six times as long as short and wide anterior portion;
<emphasis box="[592,614,411,434]" italics="true" pageId="4" pageNumber="285">se</emphasis>
about half length of posterior portion, longer than
<emphasis box="[1157,1176,411,434]" italics="true" pageId="4" pageNumber="285">vi</emphasis>
,
<emphasis box="[1189,1207,411,434]" italics="true" pageId="4" pageNumber="285">si</emphasis>
and
<emphasis box="[1259,1283,411,434]" italics="true" pageId="4" pageNumber="285">ve</emphasis>
(
<figureCitation box="[1297,1377,410,434]" captionStart="Figure 3" captionStartId="4.[155,218,2024,2046]" captionTargetBox="[194,1392,711,1987]" captionTargetId="figure-299@4.[189,1399,707,1992]" captionTargetPageId="4" captionText="Figure 3. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801).A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175793" httpUri="https://zenodo.org/record/7175793/files/figure.png" pageId="4" pageNumber="285">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph blockId="4.[136,1452,236,642]" pageId="4" pageNumber="285">
Coxal plates in four groups; two ACG not fused but close, two PCG widely separated; Cx-I trapezoidal, with numerous setae near inner apical angles and three setae (one seta and two rather long setae) at lateral margin; Cx-II trapezoidal, with one seta near outer posterior angles (
<figureCitation box="[528,614,514,538]" captionStart="Figure 4" captionStartId="5.[155,218,2024,2046]" captionTargetBox="[200,1386,726,1961]" captionTargetId="figure-259@5.[189,1398,721,1976]" captionTargetPageId="5" captionText="Figure 4. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Anterior coxal group (Cx-I+Cx-II). B. Posterior coxal group (Cx-III+Cx-IV), genital field. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175795" httpUri="https://zenodo.org/record/7175795/files/figure.png" pageId="4" pageNumber="285">Fig. 4A</figureCitation>
); Cx-III and Cx-IV nearly triangular, with some setae respectively (
<figureCitation box="[1336,1420,514,538]" captionStart="Figure 4" captionStartId="5.[155,218,2024,2046]" captionTargetBox="[200,1386,726,1961]" captionTargetId="figure-259@5.[189,1398,721,1976]" captionTargetPageId="5" captionText="Figure 4. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Anterior coxal group (Cx-I+Cx-II). B. Posterior coxal group (Cx-III+Cx-IV), genital field. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175795" httpUri="https://zenodo.org/record/7175795/files/figure.png" pageId="4" pageNumber="285">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph blockId="4.[136,1452,236,642]" pageId="4" pageNumber="285">
Genital field with about twenty tiny setae; stalked Ac-13 arranging in an almost straight line, interval from Ac-1 to Ac-2 and Ac-2 to Ac-3 almost equal; gonopore surrounded by three or four setae on each side;
<emphasis box="[1138,1179,583,607]" italics="true" pageId="4" pageNumber="285">glv-</emphasis>
2 between Ac-3 and inner posterior angle of Cx-IV, located at one-third of interval near Ac-3 approximately (
<figureCitation box="[1019,1104,618,642]" captionStart="Figure 4" captionStartId="5.[155,218,2024,2046]" captionTargetBox="[200,1386,726,1961]" captionTargetId="figure-259@5.[189,1398,721,1976]" captionTargetPageId="5" captionText="Figure 4. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Anterior coxal group (Cx-I+Cx-II). B. Posterior coxal group (Cx-III+Cx-IV), genital field. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175795" httpUri="https://zenodo.org/record/7175795/files/figure.png" pageId="4" pageNumber="285">Fig. 4B</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175793" ID-Zenodo-Dep="7175793" httpUri="https://zenodo.org/record/7175793/files/figure.png" pageId="4" pageNumber="285" startId="4.[155,218,2024,2046]" targetBox="[194,1392,711,1987]" targetPageId="4">
<paragraph blockId="4.[155,1433,2024,2078]" pageId="4" pageNumber="285">
Figure 3.
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[246,599,2024,2046]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[246,500,2024,2046]" italics="true" pageId="4" pageNumber="285">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[607,683,2024,2045]" pageId="4" pageNumber="285">
<taxonomicNameLabel box="[607,683,2024,2045]" pageId="4" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (SC-ST-20200801). A. Infracapitulum, dorsal view. B. Chelicera. C. Pedipalp, one lateral view. D. Pedipalp, another lateral view. E. Prodorsal plate. Scale bar= 200 μm.
</paragraph>
</caption>
<paragraph blockId="5.[136,1452,236,642]" pageId="5" pageNumber="286">
Legs without swimming setae but numerous setae present; I-L thicker than other pairs of legs, I-L-35 with stomatoid lyrifissures on one side (
<figureCitation box="[400,524,271,295]" captionStart="Figure 5" captionStartId="6.[155,218,1791,1813]" captionTargetBox="[189,1398,473,1758]" captionTargetId="figure-225@6.[189,1398,473,1758]" captionTargetPageId="6" captionText="Figure 5. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. I-L-16. B. II-L-16. C. III-L-16. D. IV-L-16. Scale bar=200μm." figureDoi="http://doi.org/10.5281/zenodo.7175797" httpUri="https://zenodo.org/record/7175797/files/figure.png" pageId="5" pageNumber="286">Figs 5AD</figureCitation>
); tarsus of IIV-L with two pectinate lateral claws and one smooth middle empodium, empodium considerably smaller than lateral claws (
<figureCitation box="[687,771,306,330]" captionStart="Figure 1" captionStartId="2.[155,218,2014,2036]" captionTargetBox="[189,1399,820,1984]" captionTargetId="figure-188@2.[189,1399,820,1984]" captionTargetPageId="2" captionText="Figure 1. Stygothrombium garzensis Li &amp; Guo, sp. nov., ♀, holotype (SC-ST-20200801). A. Ventral view. B. Gll-1. C. Gll-2. D. I-L-4, stomatoid lyrifissures. Scale bars: A=500μm; BD=10μm." figureDoi="http://doi.org/10.5281/zenodo.7175789" httpUri="https://zenodo.org/record/7175789/files/figure.png" pageId="5" pageNumber="286">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph blockId="5.[136,1452,236,642]" box="[189,370,340,364]" pageId="5" pageNumber="286">Male. Unknown.</paragraph>
<paragraph blockId="5.[136,1452,236,642]" pageId="5" pageNumber="286">
Measurements (
<typeStatus box="[360,459,375,399]" pageId="5" pageNumber="286">holotype</typeStatus>
). Idiosoma L 3857 (from peak of protrusible integument to bottom), W 1114; excretory pore L 108; Infracapitulum L 783, base L 607, rostrum 176; Chelicera base segment L 477, claw L 102; P-A dL 253, P-B dL 97, alantoid seta L 60, above harpago seta L 64, nether harpago seta L 66; prodorsal plate L 478, anterior portion L 73, posterior portion L 405; ACG L 274; PCG L 346; gonopore L 119, Ac-1 L 76, Ac-2 L 66, Ac-3 L 80; Legs segments dL: -L-1 100, - L-2 164, -L-3 173, -L-4 182, -L-5 189, -L-6 159, claw dL 88; II-L-1 88, II-L-2 134, II-L-3 136, II-L-4 166, II-L-5 190, II-L-6 140, claw dL 84; III-L-1 123, III-L-2 134, III-L-3 139, III-L-4 186, III-L-5 205, III-L-6 153, claw dL 78; IV-L-1 101, IV-L-2 173, IV-L-3 213, IV-L-4 259, IV-L-5 280, IV-L-6 191, claw dL 81.
</paragraph>
<paragraph blockId="5.[136,1452,236,642]" lastBlockId="6.[136,1452,236,434]" lastPageId="6" lastPageNumber="287" pageId="5" pageNumber="286">
Remarks. The present new species is similar to
<taxonomicName authority="Nagasawa &amp; Abe, 2014" authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[711,1149,618,642]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[711,882,618,642]" italics="true" pageId="5" pageNumber="286">S. monotrichum</emphasis>
Nagasawa &amp; Abé, 2014
</taxonomicName>
from
<collectingCountry box="[1218,1279,618,641]" name="Japan" pageId="5" pageNumber="286">Japan</collectingCountry>
in quantity and location of idiosomal glandularia and pedipalp structures. The new species differs from the latter in the following aspects: (1) idiosoma L
<quantity box="[298,378,271,295]" metricMagnitude="1" metricUnit="m" metricValue="9.79678" pageId="6" pageNumber="287" unit="in" value="3857.0">3857 in</quantity>
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[384,621,271,295]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[384,511,271,295]" italics="true" pageId="6" pageNumber="287">S. garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[631,716,271,295]" pageId="6" pageNumber="287">
<taxonomicNameLabel box="[631,716,271,295]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but
<quantity box="[768,915,271,295]" metricMagnitude="1" metricUnit="m" metricValue="4.3053" metricValueMax="4.6482" metricValueMin="3.9624" pageId="6" pageNumber="287" unit="in" value="1695.0" valueMax="1830.0" valueMin="1560.0">15601830 in</quantity>
<taxonomicName authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[920,1088,271,295]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[920,1088,271,295]" italics="true" pageId="6" pageNumber="287">S. monotrichum</emphasis>
</taxonomicName>
; (2)
<emphasis box="[1139,1172,271,295]" italics="true" pageId="6" pageNumber="287">glv</emphasis>
-1 between ACG and PCG in
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[163,402,306,330]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[163,291,306,330]" italics="true" pageId="6" pageNumber="287">S. garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[413,498,306,330]" pageId="6" pageNumber="287">
<taxonomicNameLabel box="[413,498,306,330]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while absent in
<taxonomicName authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[678,846,306,330]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[678,846,306,330]" italics="true" pageId="6" pageNumber="287">S. monotrichum</emphasis>
</taxonomicName>
; (3)
<emphasis box="[897,938,306,330]" italics="true" pageId="6" pageNumber="287">glv-</emphasis>
2 between Ac-3 and inner posterior angle of CxIV in
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[197,436,340,365]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[197,325,341,364]" italics="true" pageId="6" pageNumber="287">S. garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[446,531,341,365]" pageId="6" pageNumber="287">
<taxonomicNameLabel box="[446,531,341,365]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but located posteromedially to the posterior acetabula in
<taxonomicName authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[1146,1314,340,364]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[1146,1314,340,364]" italics="true" pageId="6" pageNumber="287">S. monotrichum</emphasis>
</taxonomicName>
; (4) setae on ACG and pedipalp much more multiple than
<taxonomicName authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[617,786,375,399]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[617,786,375,399]" italics="true" pageId="6" pageNumber="287">S. monotrichum</emphasis>
</taxonomicName>
; (5) P-A dL
<quantity box="[921,989,375,399]" metricMagnitude="0" metricUnit="m" metricValue="6.4262" pageId="6" pageNumber="287" unit="in" value="253.0">253 in</quantity>
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[995,1235,375,399]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[995,1122,375,399]" italics="true" pageId="6" pageNumber="287">S. garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[1245,1331,375,399]" pageId="6" pageNumber="287">
<taxonomicNameLabel box="[1245,1331,375,399]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while P-A dL
<quantity box="[172,226,410,434]" metricMagnitude="0" metricUnit="m" metricValue="2.2352" pageId="6" pageNumber="287" unit="in" value="88.0">88 in</quantity>
<taxonomicName authorityName="Nagasawa &amp; Abe" authorityYear="2014" box="[232,401,410,434]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="monotrichum">
<emphasis box="[232,401,410,434]" italics="true" pageId="6" pageNumber="287">S. monotrichum</emphasis>
</taxonomicName>
. (Nagasawa &amp; Abé 2014)
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175795" ID-Zenodo-Dep="7175795" httpUri="https://zenodo.org/record/7175795/files/figure.png" pageId="5" pageNumber="286" startId="5.[155,218,2024,2046]" targetBox="[200,1386,726,1961]" targetPageId="5">
<paragraph blockId="5.[155,1433,2024,2078]" pageId="5" pageNumber="286">
Figure 4.
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[255,620,2024,2046]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[255,512,2024,2046]" italics="true" pageId="5" pageNumber="286">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[636,716,2024,2045]" pageId="5" pageNumber="286">
<taxonomicNameLabel box="[636,716,2024,2045]" pageId="5" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (SC-ST-20200801). A. Anterior coxal group (Cx-I+Cx-II). B. Posterior coxal group (Cx-III+Cx-IV), genital field. Scale bar =200 μm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175797" ID-Zenodo-Dep="7175797" httpUri="https://zenodo.org/record/7175797/files/figure.png" pageId="6" pageNumber="287" startId="6.[155,218,1791,1813]" targetBox="[189,1398,473,1758]" targetPageId="6">
<paragraph blockId="6.[155,1433,1790,1844]" pageId="6" pageNumber="287">
Figure 5.
<taxonomicName authority="Li &amp; Guo" authorityName="Li &amp; Guo" authorityYear="2021" box="[255,621,1791,1813]" class="Arachnida" family="Stygothrombiidae" genus="Stygothrombium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="garzensis" status="sp. nov.">
<emphasis box="[255,512,1791,1812]" italics="true" pageId="6" pageNumber="287">Stygothrombium garzensis</emphasis>
Li &amp; Guo
</taxonomicName>
,
<emphasis bold="true" box="[636,716,1791,1812]" pageId="6" pageNumber="287">
<taxonomicNameLabel box="[636,716,1791,1812]" pageId="6" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (SC-ST-20200801). A. I-L-16. B. II-L-16. C. III-L-16. D. IV-L-16. Scale bar= 200 μm.
</paragraph>
</caption>
<paragraph blockId="6.[136,1452,1912,2005]" pageId="6" pageNumber="287">
<emphasis bold="true" box="[136,232,1912,1936]" inLineHeading="true" pageId="6" pageNumber="287" reason="1">Funding</emphasis>
This research was supported by National Natural Science Foundation of
<collectingCountry box="[1031,1095,1912,1936]" name="China" pageId="6" pageNumber="287">China</collectingCountry>
(31772421, 31750002),
<collectingRegion box="[1360,1452,1912,1936]" country="China" name="Guizhou" pageId="6" pageNumber="287">Guizhou</collectingRegion>
Science and Technology Project (Qiankehe Pingtai Rencai [2017] 5788) and National Key
<collectionCode box="[1160,1178,1947,1970]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="287">R</collectionCode>
&amp; D Program of
<collectingCountry box="[1388,1452,1946,1970]" name="China" pageId="6" pageNumber="287">China</collectingCountry>
(2017YFD0201000).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287" type="description">
<paragraph blockId="6.[136,1451,2057,2082]" lastBlockId="7.[136,1452,236,295]" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287">
<emphasis bold="true" box="[136,357,2057,2081]" inLineHeading="true" pageId="6" pageNumber="287" reason="1">Acknowledgements</emphasis>
<materialsCitation collectionCode="R" collectorName="University, P." country="China" county="Institute of Entomology" lastPageId="7" lastPageNumber="288" location="We" municipality="Institute of Entomology" pageId="6" pageNumber="287" specimenCount="1" stateProvince="Guizhou">
<location LSID="urn:lsid:plazi:treatment:0387972EFFE4FFA0FF1AFE7474642A72:8EF170E3FFE0FFA1FEEEF7B476D62374" box="[380,415,2058,2081]" country="China" county="Institute of Entomology" municipality="Institute of Entomology" name="We" pageId="6" pageNumber="287" stateProvince="Guizhou">We</location>
are grateful to
<location LSID="urn:lsid:plazi:treatment:0387972EFFE4FFA0FF1AFE7474642A72:8EF170E3FFE0FFA1FDADF7B475EA2374" box="[575,675,2057,2082]" country="China" county="Institute of Entomology" municipality="Institute of Entomology" name="Boyan Li" pageId="6" pageNumber="287" stateProvince="Guizhou">Boyan Li</location>
(
<collectingMunicipality box="[688,944,2057,2081]" pageId="6" pageNumber="287">Institute of Entomology</collectingMunicipality>
,
<collectingRegion box="[954,1046,2057,2081]" country="China" name="Guizhou" pageId="6" pageNumber="287">Guizhou</collectingRegion>
<collectorName box="[1052,1195,2057,2081]" pageId="6" pageNumber="287">University, P.</collectorName>
<collectionCode box="[1200,1221,2058,2081]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="287">R</collectionCode>
.
<collectingCountry box="[1230,1296,2057,2081]" name="China" pageId="6" pageNumber="287">China</collectingCountry>
) for collecting the specimen and processing the photos. Furthermore, we are thankful to Zhuhui Ding (
<collectingCounty box="[1087,1348,236,260]" pageId="7" pageNumber="288">Institute of Entomology</collectingCounty>
,
<collectingRegion box="[1360,1452,236,260]" country="China" name="Guizhou" pageId="7" pageNumber="288">Guizhou</collectingRegion>
<collectorName box="[136,280,271,295]" pageId="7" pageNumber="288">University, P.</collectorName>
<collectionCode box="[286,307,272,295]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="288">R</collectionCode>
.
<collectingCountry box="[317,382,271,295]" name="China" pageId="7" pageNumber="288">China</collectingCountry>
) for helping to take photos of specimen
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>