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<mods:title id="2C5F876FC25AC87B3E54505E2F5B2EBC">New dendrochirotid sea cucumbers from northern Australia (Echinodermata: Holothuroidea: Dendrochirotida)</mods:title>
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<mods:namePart id="8F36F733A67E2E8FE693AA924466FF37">P. Mark OLoughlin</mods:namePart>
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<treatment id="03871B11FFA53E5DFCE5FD97FB7DFEA1" ID-DOI="http://doi.org/10.5281/zenodo.6138241" ID-GBIF-Taxon="146019603" ID-Zenodo-Dep="6138241" LSID="urn:lsid:plazi:treatment:03871B11FFA53E5DFCE5FD97FB7DFEA1" httpUri="http://treatment.plazi.org/id/03871B11FFA53E5DFCE5FD97FB7DFEA1" lastPageId="15" lastPageNumber="20" pageId="14" pageNumber="19">
<subSubSection id="C334F98CFFA53E5CFCE5FD97FB06FDEC" box="[809,1180,552,574]" pageId="14" pageNumber="19" type="nomenclature">
<paragraph id="8B91AA07FFA53E5CFCE5FD97FB06FDEC" blockId="14.[809,1180,552,574]" box="[809,1180,552,574]" pageId="14" pageNumber="19">
<taxonomicName id="4C2ED184FFA53E5CFCE5FD97FB06FDEC" ID-CoL="9TSG" authority="Ludwig, 1875" authorityName="Ludwig" authorityYear="1875" box="[809,1180,552,574]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="19" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA53E5CFCE5FD97FB97FDEF" bold="true" box="[809,1037,552,573]" italics="true" pageId="14" pageNumber="19">Actinocucumis typica</emphasis>
Ludwig, 1875
</taxonomicName>
</paragraph>
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<paragraph id="8B91AA07FFA53E5CFC83FDECFB22FD48" blockId="14.[809,1441,595,772]" pageId="14" pageNumber="19">
<taxonomicName id="4C2ED184FFA53E5CFC83FDECFB58FDB4" ID-CoL="9TSG" authority="Ludwig, 1875: 91" authorityName="Ludwig" authorityPageNumber="91" authorityYear="1875" box="[847,1218,595,614]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="19" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA53E5CFC83FDECFB82FDB7" box="[847,1048,595,614]" italics="true" pageId="14" pageNumber="19">Actinocucumis typica</emphasis>
Ludwig, 1875: 91
</taxonomicName>
, fig. 24 ad.—
<bibRefCitation id="EFBFD7F6FFA53E5CFA87FDECFCCDFD52" author="Lampert, K." bookContentInfo="311 pp." editor="Semper, C." journalOrPublisher="Wissenschaftliche Resultate 4 (3" pageId="14" pageNumber="19" refId="ref10547" refString="Lampert, K. 1885. Die Seewalzen, Holothurioidea, eine Systematische Monographie mit Bestimmungs und Verbreitungs Tabellen. In Semper, C. (ed.) Reisen im Archipel der Philippinen. Zweiter teil. Wissenschaftliche Resultate 4 (3): 311 pp., 1 pl. C. W. Kreidel's Verlag, Wiesbaden." title="Die Seewalzen, Holothurioidea, eine Systematische Monographie mit Bestimmungs und Verbreitungs Tabellen" type="book" volumeTitle="Reisen im Archipel der Philippinen. Zweiter teil" year="1885">Lampert, 1885</bibRefCitation>
: 177.—
<bibRefCitation id="EFBFD7F6FFA53E5CFC52FDD2FB94FD52" author="Theel, H." box="[926,1038,621,640]" journalOrPublisher="Zoology" pageId="14" pageNumber="19" pagination="1 - 290" part="14" refId="ref11660" refString="Theel, H. 1886. Report on the Holothurioidea dredged by H. M. S. Challenger during the years 1873 - 1876. Report on the scientific results of the voyage of H. M. S. Challenger, Zoology 14 (39): 1 - 290, 16 pls." title="Report on the Holothurioidea dredged by H. M. S. Challenger during the years 1873 - 1876. Report on the scientific results of the voyage of H. M. S. Challenger" type="journal article" year="1886">Théel, 1886</bibRefCitation>
: 84, 125, pl. 12 figs 4, 5.—
<bibRefCitation id="EFBFD7F6FFA53E5CFAD6FDD2FA07FD52" author="Ludwig, H." box="[1306,1437,621,640]" journalOrPublisher="Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="14" pageNumber="19" pagination="805 - 820" part="3" refId="ref10753" refString="Ludwig, H. 1888. Die von Dr. J. Brock im Indischen Archipel gesammelten Holothurien. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 3: 805 - 820, pl. 30." title="Die von Dr. J. Brock im Indischen Archipel gesammelten Holothurien" type="journal article" year="1888">Ludwig, 1888</bibRefCitation>
: 817.—
<bibRefCitation id="EFBFD7F6FFA53E5CFCA8FD37FC53FD49" author="Erwe, W." box="[868,969,648,667]" journalOrPublisher="Jena" pageId="14" pageNumber="19" refId="ref10332" refString="Erwe, W. 1913. Holothurioidea. Pp. 349 - 402. In: Die Fauna Sudwest- Australiens. Ergebenisse der Hamburger sudwest-australischen Forschungsreise 1905 herausgegeben von Prof. Dr. W. Michaelsen und Dr. R. Hartmeyer 4 (9). Fischer, Jena." title="Holothurioidea. Pp. 349 - 402" type="book" volumeTitle="Die Fauna Sudwest- Australiens. Ergebenisse der Hamburger sudwest-australischen Forschungsreise 1905 herausgegeben von Prof. Dr. W. Michaelsen und Dr. R. Hartmeyer 4 (9). Fischer" year="1913">Erwe, 1913</bibRefCitation>
: 364365, pl. 6 fig. 10a, b.
</paragraph>
<paragraph id="8B91AA07FFA53E5CFC83FD1DFAD6FCD6" blockId="14.[809,1441,595,772]" pageId="14" pageNumber="19">
<emphasis id="B95A7615FFA53E5CFC83FD1DFBBBFD66" box="[847,1057,674,693]" italics="true" pageId="14" pageNumber="19">
<taxonomicName id="4C2ED184FFA53E5CFC83FD1DFC4DFD67" ID-CoL="7NMTS" authorityName="Ludwig" authorityYear="1875" box="[847,983,674,693]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="19" phylum="Echinodermata" rank="genus">Actinocucumis</taxonomicName>
typicus
</emphasis>
.—
<bibRefCitation id="EFBFD7F6FFA53E5CFBF1FD1DFB7DFD67" author="Clark, H. L." bookContentInfo="233 pp." box="[1085,1255,674,693]" journalOrPublisher="Carnegie Institution of Washington Publication" pageId="14" pageNumber="19" part="214" refId="ref10062" refString="Clark, H. L. 1921. The Echinoderm fauna of Torres Strait: its composition and its origin. Carnegie Institution of Washington Publication 214. 233 pp., 10 pls." title="The Echinoderm fauna of Torres Strait: its composition and its origin" type="book" year="1921">H. L. Clark, 1921</bibRefCitation>
: 170.—
<bibRefCitation id="EFBFD7F6FFA53E5CFAE2FD1DFA04FD67" author="Clark, H. L." box="[1326,1438,674,693]" journalOrPublisher="Memoirs of the Museum of Comparative Zoology at Harvard College" pageId="14" pageNumber="19" pagination="1 - 596" part="55" refId="ref10097" refString="Clark, H. L. 1938. Echinoderms from Australia. An account of collections made in 1929 and 1932. Memoirs of the Museum of Comparative Zoology at Harvard College 55: 1 - 596, 28 pls, 63 figs." title="Echinoderms from Australia. An account of collections made in 1929 and 1932" type="journal article" year="1938">Clark, 1938</bibRefCitation>
: 479.—
<bibRefCitation id="EFBFD7F6FFA53E5CFCAAFD03FC49FD1D" author="Clark, H. L." box="[870,979,700,719]" journalOrPublisher="Carnegie Institution of Washington Publication" pageId="14" pageNumber="19" pagination="1 - 567" part="566" refId="ref10141" refString="Clark, H. L. 1946. The echinoderm fauna of Australia. Its composition and its origin. Carnegie Institution of Washington Publication 566: 1 - 567." title="The echinoderm fauna of Australia. Its composition and its origin" type="journal article" year="1946">Clark, 1946</bibRefCitation>
: 403.—
<bibRefCitation id="EFBFD7F6FFA53E5CFBD4FD03FA88FD1D" author="Heding, S. G. &amp; Panning, A." bookContentInfo="209 pp." box="[1048,1298,700,719]" journalOrPublisher="Spolia Zoologica Musei Hauniensis 13" pageId="14" pageNumber="19" refId="ref10511" refString="Heding, S. G. and Panning, A. 1954. Phyllophoridae. Eine bearbeitung der polytentaculaten dendrochiroten holothurien des zoologischen museums in Kopenhagen. Spolia Zoologica Musei Hauniensis 13: 209 pp." title="Phyllophoridae. Eine bearbeitung der polytentaculaten dendrochiroten holothurien des zoologischen museums in Kopenhagen" type="journal article" year="1954">Heding and Panning, 1954</bibRefCitation>
: 7274, figs 19, 20 (part).—
<bibRefCitation id="EFBFD7F6FFA53E5CFC5EFD68FB0BFD38" author="Clark, A. M. &amp; Rowe, F. W. E." bookContentInfo="238 pp." box="[914,1169,726,746]" journalOrPublisher="Trustees of the British Museum (Natural History), London. Publication 690" pageId="14" pageNumber="19" pagination="171 - 210" refId="ref10005" refString="Clark, A. M. and Rowe, F. W. E. 1971. Holothurioidea. Pp. 171 - 210. In: Monograph of Shallow Water Indo-West Pacific Echinoderms. Trustees of the British Museum (Natural History), London. Publication 690. 238 pp., 31 pls." title="Holothurioidea. Pp" type="book chapter" volumeTitle="Monograph of Shallow Water Indo-West Pacific Echinoderms" year="1971">A. M. Clark and Rowe, 1971</bibRefCitation>
: fig. 95 e, pl. 30 fig. 4 (part).—
<bibRefCitation id="EFBFD7F6FFA53E5CFCE5FD4EFC76FCD6" author="Liao, Y. &amp; Clark, A. M." bookContentInfo="614 pp." box="[809,1004,753,772]" journalOrPublisher="Science Press, Beijing" pageId="14" pageNumber="19" refId="ref10608" refString="Liao, Y. and Clark, A. M. 1995. The echinoderms of southern China. 614 pp., 23 pls. Science Press, Beijing." title="The echinoderms of southern China" type="book" year="1995">Liao and Clark, 1995</bibRefCitation>
: 481482, fig. 290, pl. 23 fig. 10 (part).
</paragraph>
</subSubSection>
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<collectingCounty id="62F0D28BFFA53E5CFCE5FCA5FCE0FCFF" box="[809,890,794,813]" pageId="14" pageNumber="19">Material</collectingCounty>
examined
</emphasis>
. NE
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,
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:
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, dredged off
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,
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,
<collectorName id="26DBCFD1FFA53E5CFB78FC8AFABAFC9A" box="[1204,1312,821,840]" pageId="14" pageNumber="19">B. J. Smith</collectorName>
,
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<date id="FF908CC7FFA53E5CFAE0FC8AFA07FC9A" box="[1324,1437,821,840]" pageId="14" pageNumber="19" value="1967-09-06">6 Sept 1967</date>
</collectingDate>
,
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F204078 (1); Mackay harbour,
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, Ian Kirwan,
<collectingDate id="EFD4752FFFA53E5CFCE5FCD6FC0BFCAE" box="[809,913,873,892]" pageId="14" pageNumber="19" value="1982-07-14">
<date id="FF908CC7FFA53E5CFCE5FCD6FC0BFCAE" box="[809,913,873,892]" pageId="14" pageNumber="19" value="1982-07-14">14 Jul 1982</date>
</collectingDate>
,
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F206362 (1); near Cairns, Machans Beach,
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, on beach after cyclone,
<collectorName id="26DBCFD1FFA53E5CFBBCFC3BFB48FC45" box="[1136,1234,900,919]" pageId="14" pageNumber="19">B. Collins</collectorName>
,
<collectingDate id="EFD4752FFFA53E5CFB2CFC3BFAC4FC45" box="[1248,1374,900,919]" pageId="14" pageNumber="19" value="1996-12-25">
<date id="FF908CC7FFA53E5CFB2CFC3BFAC4FC45" box="[1248,1374,900,919]" pageId="14" pageNumber="19" value="1996-12-25">25 Dec 1996</date>
</collectingDate>
,
<collectionCode id="ED3F32C2FFA53E5CFAA6FC3BFA3AFC45" box="[1386,1440,900,919]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="14" pageNumber="19">NMV</collectionCode>
F203016 (2); N
<collectingCountry id="F339EA97FFA53E5CFC70FC21FB88FC63" box="[956,1042,926,945]" name="Australia" pageId="14" pageNumber="19">Australia</collectingCountry>
, Joseph Bonaparte Gulf,
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, RV
<collectorName id="26DBCFD1FFA53E5CFC83FC07FC3BFC19" box="[847,929,952,971]" pageId="14" pageNumber="19">
<emphasis id="B95A7615FFA53E5CFC83FC07FC3BFC19" box="[847,929,952,971]" italics="true" pageId="14" pageNumber="19">Solander</emphasis>
</collectorName>
,
<elevation id="00034D34FFA53E5CFC62FC07FC7AFC1E" box="[942,992,952,972]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="19" unit="m" value="48.0">
<quantity id="4CD607E2FFA53E5CFC62FC07FC7AFC1E" box="[942,992,952,972]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="19" unit="m" value="48.0">48 m</quantity>
</elevation>
,
<collectionCode id="ED3F32C2FFA53E5CFC27FC07FBBEFC19" box="[1003,1060,952,971]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12622" name="Australian Institute of Marine Science" pageId="14" pageNumber="19">AIMS</collectionCode>
&amp; GA,
<collectingDate id="EFD4752FFFA53E5CFBBEFC07FB76FC19" box="[1138,1260,952,972]" pageId="14" pageNumber="19" value="2010-08-26">
<date id="FF908CC7FFA53E5CFBBEFC07FB76FC19" box="[1138,1260,952,972]" pageId="14" pageNumber="19" value="2010-08-26">26 Aug 2010</date>
</collectingDate>
,
<collectionCode id="ED3F32C2FFA53E5CFB3AFC06FAB6FC1E" box="[1270,1324,953,972]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="14" pageNumber="19">NMV</collectionCode>
F173265 (1) (GA lot SOL5117 lot 082BS040; UF tissue lot
<specimenCode id="DB88027CFFA53E5CFB26FC6CFAF2FC34" box="[1258,1384,979,998]" collectionCode="MOLAF" pageId="14" pageNumber="19">MOLAF1552</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B46A05AFFA53E5CFAB6FC6CFB92FBE7" ID-GBIF-Occurrence="1851456554" collectingDate="1973-02-01" collectingDateMax="1999-07-24" collectingDateMin="1973-02-01" collectionCode="NMV" collectorName="Eaglehawk I. &amp; Flinders, DA" country="Australia" elevation="13" latitude="-20.67" location="Dampier Archipelago" longLatPrecision="760" longitude="116.46" pageId="14" pageNumber="19" specimenCount="1">
NW
<collectingCountry id="F339EA97FFA53E5CFCE5FC52FCE5FBD2" box="[809,895,1005,1024]" name="Australia" pageId="14" pageNumber="19">Australia</collectingCountry>
,
<location id="8EF1FCDCFFA53E5CFC42FC52FBC3FBD2" LSID="urn:lsid:plazi:treatment:03871B11FFA53E5DFCE5FD97FB7DFEA1:8EF1FCDCFFA53E5CFC42FC52FBC3FBD2" box="[910,1113,1005,1024]" country="Australia" latitude="-20.67" longLatPrecision="760" longitude="116.46" name="Dampier Archipelago" pageId="14" pageNumber="19">Dampier Archipelago</location>
,
<locationDeviation id="9A23B325FFA53E5CFBABFC52FAD4FBD2" box="[1127,1358,1005,1024]" pageId="14" pageNumber="19">
<quantity id="4CD607E2FFA53E5CFBABFC52FB30FBD2" box="[1127,1194,1005,1024]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="19" unit="km" value="1.3">1.3 km</quantity>
E of
<collectorName id="26DBCFD1FFA53E5CFB26FC52FAD4FBD2" box="[1258,1358,1005,1024]" pageId="14" pageNumber="19">Eaglehawk</collectorName>
</locationDeviation>
I., FRV
<collectorName id="26DBCFD1FFA53E5CFCE5FBB8FC3DFBC9" box="[809,935,1031,1051]" pageId="14" pageNumber="19">
<emphasis id="B95A7615FFA53E5CFCE5FBB8FCECFBC8" box="[809,886,1031,1050]" italics="true" pageId="14" pageNumber="19">Flinders</emphasis>
, DA
</collectorName>
<collectingDate id="EFD4752FFFA53E5CFC6BFBB7FC70FBC9" box="[935,1002,1032,1051]" pageId="14" pageNumber="19" value="1973-02-01">
<date id="FF908CC7FFA53E5CFC6BFBB7FC70FBC9" box="[935,1002,1032,1051]" pageId="14" pageNumber="19" value="1973-02-01">2/73/01</date>
</collectingDate>
,
<geoCoordinate id="EE1ACCC0FFA53E5CFC3BFBB8FBA4FBC8" box="[1015,1086,1031,1050]" direction="south" orientation="latitude" pageId="14" pageNumber="19" precision="555" value="-20.67">20.67°S</geoCoordinate>
<geoCoordinate id="EE1ACCC0FFA53E5CFB84FBB8FB01FBC8" box="[1096,1179,1031,1050]" direction="east" orientation="longitude" pageId="14" pageNumber="19" precision="555" value="116.46">116.46°E</geoCoordinate>
, coarse sand,
<elevation id="00034D34FFA53E5CFAE9FBB7FAC3FBC9" box="[1317,1369,1032,1051]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="19" unit="m" value="13.0">
<quantity id="4CD607E2FFA53E5CFAE9FBB7FAC3FBC9" box="[1317,1369,1032,1051]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="19" unit="m" value="13.0">13 m</quantity>
</elevation>
,
<collectingDate id="EFD4752FFFA53E5CFAA9FBB7FCCDFBE7" pageId="14" pageNumber="19" value="1999-07-24">
<date id="FF908CC7FFA53E5CFAA9FBB7FCCDFBE7" pageId="14" pageNumber="19" value="1999-07-24">24 Jul 1999</date>
</collectingDate>
,
<collectionCode id="ED3F32C2FFA53E5CFCACFB9DFC0CFBE7" box="[864,918,1058,1077]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="14" pageNumber="19">NMV</collectionCode>
F209501(1).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C334F98CFFA53E5DFCE5FBF2FE4AFE8A" lastPageId="15" lastPageNumber="20" pageId="14" pageNumber="19" type="diagnosis">
<paragraph id="8B91AA07FFA53E5CFCE5FBF2FAB8FA6C" blockId="14.[809,1440,1100,1905]" pageId="14" pageNumber="19">
<emphasis id="B95A7615FFA53E5CFCE5FBF2FBAEFBB0" box="[809,1076,1101,1122]" italics="true" pageId="14" pageNumber="19">Diagnosis (sensu stricto)</emphasis>
. Dendrochirotid species, uniform brown colour (NMV F204078), or yellowish brown with some red patches and fine black flecking (NMV F173265), or pale brown to cream with fine brown flecking on body and larger brown patches on tube feet (NMV F206362); body pentagonal in section with raised radial ridges; five thick oral valves; about 20 dendritic tentacles, variable in arrangement and size, ventral pair smallest; tube feet confined to radii, small papillae and tube feet also on radii and encroaching inter-radially; tube feet in 46 rows (
<quantity id="4CD607E2FFA53E5CFC61FAEEFC6EFAB5" box="[941,1012,1361,1383]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="19" unit="mm" value="80.0">80 mm</quantity>
long specimen, Ludwig 1875), or in paired rows (smaller
<quantity id="4CD607E2FFA53E5CFC7FFAD1FB85FA56" box="[947,1055,1390,1412]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.4" metricValueMin="3.5" pageId="14" pageNumber="19" unit="mm" value="39.5" valueMax="44.0" valueMin="35.0">3544 mm</quantity>
long specimens, this work); calcareous ring not composite, radial plates with vertical sides, radial and inter-radial plates lacking posterior prolongations.
</paragraph>
<paragraph id="8B91AA07FFA53E5DFC83FA7AFE4AFE8A" blockId="14.[809,1440,1100,1905]" lastBlockId="15.[147,779,232,344]" lastPageId="15" lastPageNumber="20" pageId="14" pageNumber="19">
Body wall ossicles abundant, crowded, small plates and ellipsoids, and rare, thick large plates; bi-perforate plates (figure-8 form, “acorn” plates in Ludwig 1875 and
<bibRefCitation id="EFBFD7F6FFA53E5CFAE6FA40FA07F9C6" author="Clark, H. L." box="[1322,1437,1535,1557]" journalOrPublisher="Memoirs of the Museum of Comparative Zoology at Harvard College" pageId="14" pageNumber="19" pagination="1 - 596" part="55" refId="ref10097" refString="Clark, H. L. 1938. Echinoderms from Australia. An account of collections made in 1929 and 1932. Memoirs of the Museum of Comparative Zoology at Harvard College 55: 1 - 596, 28 pls, 63 figs." title="Echinoderms from Australia. An account of collections made in 1929 and 1932" type="journal article" year="1938">Clark 1938</bibRefCitation>
,
<bibRefCitation id="EFBFD7F6FFA53E5CFCE5F9A3FCC7F9E0" author="Clark, H. L." box="[809,861,1564,1586]" journalOrPublisher="Carnegie Institution of Washington Publication" pageId="14" pageNumber="19" pagination="1 - 567" part="566" refId="ref10141" refString="Clark, H. L. 1946. The echinoderm fauna of Australia. Its composition and its origin. Carnegie Institution of Washington Publication 566: 1 - 567." title="The echinoderm fauna of Australia. Its composition and its origin" type="journal article" year="1946">1946</bibRefCitation>
) predominate, up to about 4050
<emphasis id="B95A7615FFA53E5CFB17F9A2FB73F9E0" box="[1243,1257,1565,1586]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m long with one central and 5 marginal knobs, the apical knob typically projecting; bi-perforate plates inter-grade with abundant but less numerous irregularly ovoid fenestrated ellipsoids, up to about 4055
<emphasis id="B95A7615FFA53E5CFC7DF92EFC25F974" box="[945,959,1681,1702]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m long; large plates thick, irregular in outline, perforation sizes irregular, plates frequently more than 150
<emphasis id="B95A7615FFA53E5CFCE5F974FCADF932" box="[809,823,1739,1760]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m long. Tube feet with endplates and support tables; large endplate diameters about 240
<emphasis id="B95A7615FFA53E5CFBA5F957FBEDF92F" box="[1129,1143,1768,1789]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m, small endplate diameters about 120
<emphasis id="B95A7615FFA53E5CFC5EF8BAFC3AF8C8" box="[914,928,1797,1818]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m; table discs elongate, up to 160
<emphasis id="B95A7615FFA53E5CFB38F8BAFA98F8C8" box="[1268,1282,1797,1818]" italics="true" pageId="14" pageNumber="19">µ</emphasis>
m long, narrow, widened centrally and distally, spires with 4-pillar base, pillars frequently joined to form an arch, arches frequently with single, long, thick, pointed, apical spine. Introvert with abundant tables, spires with four-pillar base and pillars fused with distal narrow arch or long spine or 2 short blunt spines. Papillae with body wall ossicles, rare tables, and lacking endplates. Lacking anal scales.
</paragraph>
</subSubSection>
<subSubSection id="C334F98CFFA43E5DFF5FFED0FB7DFEA1" pageId="15" pageNumber="20" type="discussion">
<paragraph id="8B91AA07FFA43E5DFF5FFED0FC91FD41" blockId="15.[147,779,367,719]" pageId="15" pageNumber="20">
<emphasis id="B95A7615FFA43E5DFF5FFED0FF72FE56" box="[147,232,367,388]" italics="true" pageId="15" pageNumber="20">Remarks</emphasis>
. Because uncertainty remains about the assignment of species to this genus we provide a
<emphasis id="B95A7615FFA43E5DFDC9FE31FD1EFE71" box="[517,644,398,419]" italics="true" pageId="15" pageNumber="20">sensu stricto</emphasis>
diagnosis of
<taxonomicName id="4C2ED184FFA43E5DFF5FFE14FEF2FE13" authorityName="Ludwig" authorityYear="1875" box="[147,360,427,449]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA43E5DFF5FFE14FEF2FE13" box="[147,360,427,449]" italics="true" pageId="15" pageNumber="20">Actinocucumis typica</emphasis>
</taxonomicName>
, based on the morphological characters of the
<typeStatus id="549514A5FFA43E5DFF7AFE75FF7AFE0D" box="[182,224,458,479]" pageId="15" pageNumber="20">type</typeStatus>
species. We examined six northern Australian specimens of
<taxonomicName id="4C2ED184FFA43E5DFF60FE58FEA7FE2E" authorityName="Ludwig" authorityYear="1875" box="[172,317,487,508]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="genus">
<emphasis id="B95A7615FFA43E5DFF60FE58FEA7FE2E" box="[172,317,487,508]" italics="true" pageId="15" pageNumber="20">Actinocucumis</emphasis>
</taxonomicName>
(listed above). We confidently judge that four of these specimens are
<taxonomicName id="4C2ED184FFA43E5DFEB7FDBAFDC3FDC9" authorityName="Ludwig" authorityYear="1875" box="[379,601,517,539]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA43E5DFEB7FDBAFDC3FDC9" box="[379,601,517,539]" italics="true" pageId="15" pageNumber="20">Actinocucumis typica</emphasis>
</taxonomicName>
as all of their morphological characters closely fit the original description and illustrations by Ludwig (1875). These characters are detailed in the generic diagnosis above. With less confidence we judge that the two beach-washed specimens are also
<taxonomicName id="4C2ED184FFA43E5DFDC8FDC1FDC1FD41" authorityName="Ludwig" authorityYear="1875" box="[516,603,638,659]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA43E5DFDC8FDC1FDC1FD41" box="[516,603,638,659]" italics="true" pageId="15" pageNumber="20">A. typica</emphasis>
</taxonomicName>
(NMV F203016).
</paragraph>
<paragraph id="8B91AA07FFA43E5DFF75FD24FB7DFEA1" blockId="15.[147,779,367,719]" lastBlockId="15.[817,1448,232,371]" pageId="15" pageNumber="20">
The ossicles of specimens from the
<collectingCountry id="F339EA97FFA43E5DFDF0FD24FD2AFD63" box="[572,688,667,689]" name="Philippines" pageId="15" pageNumber="20">Philippines</collectingCountry>
that are illustrated by
<bibRefCitation id="EFBFD7F6FFA43E5DFEEEFD06FDD1FD1D" author="Reyes-Leonardo, L. D. &amp; Monzon, R. B. &amp; Navarro, V. C." box="[290,587,697,719]" journalOrPublisher="Natural and Applied Science Bulletin" pageId="15" pageNumber="20" pagination="261 - 284" part="37" refId="ref11223" refString="Reyes-Leonardo, L. D., Monzon, R. B. and Navarro, V. C. 1985. A taxonomic account of shallow water holothurians of Bolinao, Pangasinan. Natural and Applied Science Bulletin 37 (4): 261 - 284." title="A taxonomic account of shallow water holothurians of Bolinao, Pangasinan" type="journal article" year="1985">Reyes-Leonardo et al. (1985)</bibRefCitation>
appear to us to be close to those illustrated for
<taxonomicName id="4C2ED184FFA43E5DFB45FF57FA3BFF2F" authorityName="Clark" authorityYear="1938" box="[1161,1441,232,253]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="species" species="longipedes">
<emphasis id="B95A7615FFA43E5DFB45FF57FA3BFF2F" box="[1161,1441,232,253]" italics="true" pageId="15" pageNumber="20">Actinocucumis longipedes</emphasis>
</taxonomicName>
, although the description refers to the presence of fenestrated ellipsoids and irregular tables with pointed spires. The descriptive reference to “wart-like” podia scattered all over the body” is not characteristic of
<taxonomicName id="4C2ED184FFA43E5DFB49FEE1FB7BFEA1" authorityName="Ludwig" authorityYear="1875" box="[1157,1249,350,371]" class="Holothuroidea" family="Cucumariidae" genus="Actinocucumis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="20" phylum="Echinodermata" rank="species" species="typica">
<emphasis id="B95A7615FFA43E5DFB49FEE1FB7BFEA1" box="[1157,1249,350,371]" italics="true" pageId="15" pageNumber="20">A. typica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>