treatments-xml/data/CB/66/75/CB6675115E61FFB98CE5BFA0EAC1100D.xml

<document id="5AD4B05F1D37210D708AE89C1B49A16B" ID-CLB-Dataset="148088" ID-DOI="10.11646/zootaxa.5188.4.2" ID-GBIF-Dataset="d2c2237e-f75c-4203-81d0-710f345e32d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7095476" ID-ZooBank="E23EECE7-31F4-4AF1-8215-E070869BB917" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663656221510" checkinUser="plazi" docAuthor="Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy &amp; Araújo, Maíra Xavier" docDate="2022" docId="CB6675115E61FFB98CE5BFA0EAC1100D" docLanguage="en" docName="zootaxa.5188.4.2.pdf" docOrigin="Zootaxa 5188 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wagneromyia Vieira, Cordeiro, Bravo &amp; Araujo 2022, gen. nov." docType="treatment" docVersion="6" lastPageNumber="344" masterDocId="375F0D695E60FFB78C72BE7BEF5D1621" masterDocTitle="A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region" masterLastPageNumber="346" masterPageNumber="330" pageNumber="331" updateTime="1699383553801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0AB4B0196D882ACAF11B1CF4D0C4227A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="221CBF1105AE6156A168B6E9CF3BD6C6">
<mods:title id="B4FA5C1F353C4DE03FE6E58FCF18F580">A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region</mods:title>
</mods:titleInfo>
<mods:name id="6BAC10D5643F1232D9186BFCC3E17DEA" type="personal">
<mods:role id="783BDB5233CB48D26579CB8BE3A85F3A">
<mods:roleTerm id="9EAD577215F3AE1890EA75061784F104">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CBDB09DE19856D271D52EEC8C4881BA8">Vieira, Gabriel Santos</mods:namePart>
<mods:affiliation id="E4AF60631E0A9EEB2449E2F998EF379E">Laboratório de Sistemática de Insetos, Departamento de Ciências Biológicas, Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="2DA8149BE4F1649155F176750D61F99F" type="personal">
<mods:role id="E9019EDC7CE60A1535651E099694B8F9">
<mods:roleTerm id="8FEB50DF54B6C8E03D5464FF17B3FC9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59AB80A8DB3E1591C9BBBD44023582A0">Cordeiro, Danilo Pacheco</mods:namePart>
<mods:nameIdentifier id="998EE59EF250DF689DCF7E8BB427E556" type="ORCID">0000-0002-3085-8628</mods:nameIdentifier>
<mods:affiliation id="DB08EAD0A28ECAEFDF86AF3A1210982E">Instituto Nacional da Mata Atlântica (INMA), Av. José Ruschi, 29650 - 000 Santa Teresa, Espírito Santo, Brasil. d. pacheco. c @ gmail. com; https: // orcid. org / 0000 - 0002 - 3085 - 8628</mods:affiliation>
<mods:nameIdentifier id="896D42E26E8A3CDD6C550F0F0DDC35F8" type="email">d.pacheco.c@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5F107B1E5BD4FD487D1320BDC8940149" type="personal">
<mods:role id="CDCDA49848B056A4A9159CF740BA0719">
<mods:roleTerm id="11366B736D8DD7287AF4CEAE0B948AD1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A0F01DAAF9974692F389D5626BC84266">Bravo, Freddy</mods:namePart>
<mods:nameIdentifier id="47F633ED91F0F5698A20AFE83401703F" type="ORCID">0000-0003-0959-0767</mods:nameIdentifier>
<mods:affiliation id="3BD4E8761B5A462972E03EF1488E17F8">Laboratório de Sistemática de Insetos, Departamento de Ciências Biológicas, Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brazil. &amp; fbravo @ uefs. br; https: // orcid. org / 0000 - 0003 - 0959 - 0767</mods:affiliation>
<mods:nameIdentifier id="2D0548E79E78340936ED24383F2A6241" type="email">fbravo@uefs.br</mods:nameIdentifier>
</mods:name>
<mods:name id="7013F99346E1A63FF3CD802B906BC36D" type="personal">
<mods:role id="0D1E91C775AB91EBE9FA15690A789F7D">
<mods:roleTerm id="B9E411EDA02E481E5CC44676B902F5A4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="807B75A63C25D09327EDB76A42B91498">Araújo, Maíra Xavier</mods:namePart>
<mods:nameIdentifier id="1B48CD324A69DD19D98A1AA0D09C783B" type="ORCID">0000-0003-2447-9901</mods:nameIdentifier>
<mods:affiliation id="DCF19AC429EBB4CF9127D480EFE8B85F">Laboratório de Sistemática de Insetos, Departamento de Ciências Biológicas, Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brazil. &amp; mah. biology @ gmail. com; https: // orcid. org / 0000 - 0003 - 2447 - 9901</mods:affiliation>
<mods:nameIdentifier id="515069963CB371935F38B0038393DFB8" type="email">mah.biology@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A2A8B5DB90FD0D60624CD28FC67A9936">text</mods:typeOfResource>
<mods:relatedItem id="349445EA30C0087869977B3493C6616B" type="host">
<mods:titleInfo id="D485063AF233E48876BBDE7DA72E50AB">
<mods:title id="55C05307A42B4C2F12EC9D504B092202">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6D295BBE395281960DE1EA9FF667D249">
<mods:date id="2753F58B259A99FBE71B70D3B19E91D2">2022</mods:date>
<mods:detail id="61A0C39979741F64D47A6D41B400CA98" type="pubDate">
<mods:number id="457414ED66A1ECD61337450B91ADABD2">2022-09-20</mods:number>
</mods:detail>
<mods:detail id="E2AC38ECC2AA7CC9F2A80C3E7CAE0357" type="volume">
<mods:number id="ACF754E5CB626CB8D270151990CC2C4B">5188</mods:number>
</mods:detail>
<mods:detail id="CDB8F640E9E3ABFB1FFCA2B3DDE081E0" type="issue">
<mods:number id="F281B62670FFA758396E478DE87A143E">4</mods:number>
</mods:detail>
<mods:extent id="7E6E44C15C19C5ECFE5D4DB7A52454CC" unit="page">
<mods:start id="CACE1B6AE3C5B761C104C38D86A90E7A">330</mods:start>
<mods:end id="71F3E65E55C366943CCE6D3C141245C1">346</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="751D6FDAFE1FDF16314FE8999F4D8B6D">journal article</mods:classification>
<mods:identifier id="6CC277B3178B17354323C5F121B7E47E" type="CLB-Dataset">148088</mods:identifier>
<mods:identifier id="D8C16BC2A94C5B71CE3A35DDF7E7A89F" type="DOI">10.11646/zootaxa.5188.4.2</mods:identifier>
<mods:identifier id="142F66C8F1410B90679031433289A00D" type="GBIF-Dataset">d2c2237e-f75c-4203-81d0-710f345e32d2</mods:identifier>
<mods:identifier id="66D5349978DC74AFE65DAE16761C38E7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2C8E5BAB3C1DD70BEF25886EEFDDE33A" type="Zenodo-Dep">7095476</mods:identifier>
<mods:identifier id="1115556659EA1156254E4514BA488257" type="ZooBank">E23EECE7-31F4-4AF1-8215-E070869BB917</mods:identifier>
</mods:mods>
<treatment id="CB6675115E61FFB98CE5BFA0EAC1100D" ID-DOI="http://doi.org/10.5281/zenodo.7105117" ID-GBIF-Taxon="201103510" ID-Zenodo-Dep="7105117" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D" httpUri="http://treatment.plazi.org/id/CB6675115E61FFB98CE5BFA0EAC1100D" lastPageId="14" lastPageNumber="344" pageId="1" pageNumber="331">
<subSubSection id="0BD5978C5E61FFB68CE5BFA0EC3417D4" box="[151,873,474,501]" pageId="1" pageNumber="331" type="nomenclature">
<paragraph id="4370C4075E61FFB68CE5BFA0EC3417D4" blockId="1.[151,873,474,501]" box="[151,873,474,501]" pageId="1" pageNumber="331">
<heading id="1838736B5E61FFB68CE5BFA0EC3417D4" bold="true" box="[151,873,474,501]" fontSize="11" level="1" pageId="1" pageNumber="331" reason="1">
<emphasis id="71BB18155E61FFB68CE5BFA0EC3417D4" bold="true" box="[151,873,474,501]" pageId="1" pageNumber="331">
<taxonomicName id="84CFBF845E61FFB68CE5BFA0EDB317D4" ID-CoL="9J5QP" authority="Vieira, Cordeiro, Bravo &amp; Araujo" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[151,750,474,501]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71BB18155E61FFB68CE5BFA0EE6317D4" bold="true" box="[151,318,475,501]" italics="true" pageId="1" pageNumber="331">Wagneromyia</emphasis>
Vieira, Cordeiro, Bravo &amp; Araújo
</taxonomicName>
, 
<taxonomicNameLabel id="6A88A56E5E61FFB68E8EBFA0EC3417D4" box="[764,873,475,501]" pageId="1" pageNumber="331" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB68CE5BC58EA44141C" box="[151,1305,546,573]" pageId="1" pageNumber="331" type="type_taxon">
<paragraph id="4370C4075E61FFB68CE5BC58EA44141C" blockId="1.[151,1437,546,2013]" box="[151,1305,546,573]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CE5BC58EE68141D" bold="true" box="[151,309,546,573]" pageId="1" pageNumber="331">
<typeStatus id="9C747AA55E61FFB68CE5BC58EF8F141C" box="[151,210,547,573]" pageId="1" pageNumber="331">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="84CFBF845E61FFB68D4EBC58ECFE141C" ID-CoL="9J6XN" authority="Vieira, Cordeiro, Bravo &amp; Araujo" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[316,931,547,573]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E61FFB68D4EBC58ED7D141D" box="[316,544,547,572]" italics="true" pageId="1" pageNumber="331">Wagneromyia aquiri</emphasis>
Vieira, Cordeiro, Bravo &amp; Araújo
</taxonomicName>
, 
<emphasis id="71BB18155E61FFB68FDCBC58EB5A141C" bold="true" box="[942,1031,547,573]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68FDCBC58EB5A141C" box="[942,1031,547,573]" pageId="1" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB68CB5BC3CEC2F14A4" pageId="1" pageNumber="331" type="etymology">
<paragraph id="4370C4075E61FFB68CB5BC3CEC2F14A4" blockId="1.[151,1437,546,2013]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5BC3CEE131440" bold="true" box="[199,334,583,609]" pageId="1" pageNumber="331">Etymology.</emphasis>
The genus name is in honor to Dr. Rüdiger Wagner, dipterologist and specialist in 
<taxonomicName id="84CFBF845E61FFB6897ABC3CEAC51440" box="[1288,1432,583,609]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
, with the Greek suffix 
<emphasis id="71BB18155E61FFB68DFEBC10EE9F14A5" box="[396,450,619,644]" italics="true" pageId="1" pageNumber="331">myia</emphasis>
meaning “fly”. The name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB68CB5BCF5EB2D157C" pageId="1" pageNumber="331" type="diagnosis">
<paragraph id="4370C4075E61FFB68CB5BCF5EB2D157C" blockId="1.[151,1437,546,2013]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5BCF5EE1F1489" bold="true" box="[199,322,654,680]" pageId="1" pageNumber="331">Diagnosis.</emphasis>
Eyes contiguous; antenna with 14 flagellomeres, nodiform (first to 11 
<superScript id="B4BA694F5E61FFB68822BCF5EB0014BD" attach="left" box="[1104,1117,654,668]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
flagellomere), 12 
<superScript id="B4BA694F5E61FFB68951BCF5EA6D14BD" attach="none" box="[1315,1328,654,668]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
, 13 
<superScript id="B4BA694F5E61FFB68928BCF5EA3A14BD" attach="none" box="[1370,1383,654,668]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
, and 14 
<superScript id="B4BA694F5E61FFB68CC1BCC9EF9D14E1" attach="left" box="[179,192,690,704]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
flagellomeres reduced without internodes; setae alveoli of flagellomeres arranged in longitudinal rows; 14th flagellomere with one to three apical spiniform setula; palpus with four segments, fourth segment equal in length or lesser than the first; second segment of the palpus longer than the other segments; first flagellomere with a dorsal patch of sensilla; wing membrane pilose, with pilosity arranged parallel to the veins; gonocoxites contiguous at base; single aedeagus with lateral folds; epandrial appendage with three apical tenacula.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB68CB5BD1DEEFF10B8" pageId="1" pageNumber="331" type="description">
<paragraph id="4370C4075E61FFB68CB5BD1DEB531054" blockId="1.[151,1437,546,2013]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5BD1DEEF815A0" bold="true" box="[199,421,870,897]" pageId="1" pageNumber="331">Description. Male.</emphasis>
Head semicircular, eye bridge with four rows of facets; eyes contiguous; group of setae on the frons not reaching the lower limit of the eye bridge; interocular suture present or absent, when present small and U-inverted shape. antenna with cylindrical scape slightly longer than subspherical pedicel; 14 nodiform flagellomeres; pair of ascoids, Y-shaped, in each flagellomere; small additional sensilla near the ascoid insertion (
<figureCitation id="DBF4D8825E61FFB68CEDBD8CEFBE1230" box="[159,227,1014,1041]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="1" pageNumber="331">Fig. 5</figureCitation>
), on flagellomeres I-X; setae alveoli of flagellomeres arranged in longitudinal rows; flagellomeres 12 
<superScript id="B4BA694F5E61FFB6892BBD8DEA3B1225" attach="none" box="[1369,1382,1014,1028]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
, 13 
<superScript id="B4BA694F5E61FFB689E2BD8DEAC01225" attach="none" box="[1424,1437,1014,1028]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
and 14 
<superScript id="B4BA694F5E61FFB68C93BA61EFB31209" attach="left" box="[225,238,1050,1064]" fontSize="6" pageId="1" pageNumber="331">th</superScript>
reduced, separated from each other, without internodes; 11th, 13th with stout and spiniform setulae and 14th flagellomere with one to three apical spiniform setula; palpus with four segments, first of one with a group of dorsal sensilla near the base (
<figureCitation id="DBF4D8825E61FFB68DEBBA18EE83125C" box="[409,478,1123,1149]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="1" pageNumber="331">Fig. 3</figureCitation>
); length of fourth segment equal to or lesser than the first segment; second segment longer than the other segments. Labrum with convex dorsal margin, slightly expanded (
<figureCitation id="DBF4D8825E61FFB68810BAFCEBF91280" box="[1122,1188,1159,1185]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="1" pageNumber="331">Fig. 3</figureCitation>
); bulbous labella with robust and spiniform setulae on the inner margin and setae on the lateral margin (
<figureCitation id="DBF4D8825E61FFB68851BAD0EB3512E4" box="[1059,1128,1195,1221]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="1" pageNumber="331">Fig. 4</figureCitation>
). Wing membrane pilose in lines parallel to the veins; C vein with two breaks at the base; Sc slightly curved at the apex, ending near the base of R 
<subScript id="DF4BC6425E61FFB68CBBBB7EEF8C1332" attach="left" box="[201,209,1285,1299]" fontSize="6" pageId="1" pageNumber="331">1</subScript>
; costal cell slightly infuscated; medial fork usually incomplete, the base of M 
<subScript id="DF4BC6425E61FFB6884CBB7EEB1B1332" attach="left" box="[1086,1094,1285,1299]" fontSize="6" pageId="1" pageNumber="331">2</subScript>
close to M 
<subScript id="DF4BC6425E61FFB688B7BB7EEB901332" attach="left" box="[1221,1229,1285,1299]" fontSize="6" pageId="1" pageNumber="331">1</subScript>
but always fading before joining M 
<subScript id="DF4BC6425E61FFB68D26BB52EE011316" attach="left" box="[340,348,1321,1335]" fontSize="6" pageId="1" pageNumber="331">1</subScript>
; sudden angulation at the end of R 
<subScript id="DF4BC6425E61FFB68EACBB52EDBB1316" attach="left" box="[734,742,1321,1335]" fontSize="6" pageId="1" pageNumber="331">2</subScript>
. Male terminalia with a narrow hypandrium, like a transversal band; gonocoxites usually dorsally contiguous; gonostylus conical, curved, with sparse setae and with an accessory long subapical seta; single aedeagus, narrow plate-like, with apex sometimes bifurcated and lateral folds; parameres of varied shapes, arising from the gonocoxal apodemes and articulated ventrally to the aedeagus at base; external lateral margin of the parameres generally projecting dorsally to the aedeagus forming a parameral sheath; parameral sheath of varied shape, connected to the gonocoxites, never exceeding the apex of the aedeagus; epandrium wider than long, with concave base and a large pseudospiracular opening; club-shaped epandrial appendages, with a group of setulae at the base, a group of subapical papilla on the dorsal surface and bearing three apical tenacula; hypoproct slightly projected and rounded, with pilose margin; lateral region of the hypoproct sclerotized and densely pilose, with setae a little longer than in the medial region; epiproct pilose and reduced.
</paragraph>
<paragraph id="4370C4075E61FFB68CB5B804EEFF10B8" blockId="1.[151,1437,546,2013]" box="[199,418,1663,1689]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5B804EE7810B8" bold="true" box="[199,293,1663,1689]" pageId="1" pageNumber="331">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB68CB5B8D9EC161150" pageId="1" pageNumber="331" type="distribution">
<paragraph id="4370C4075E61FFB68CB5B8D9ED1D10C0" blockId="1.[151,1437,546,2013]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5B8D9EE3C109D" bold="true" box="[199,353,1698,1724]" pageId="1" pageNumber="331">Distribution.</emphasis>
South and Central America (
<collectingCountry id="3BD884975E61FFB68EE8B8D8EC4E109C" box="[666,787,1699,1725]" name="Costa Rica" pageId="1" pageNumber="331">Costa Rica</collectingCountry>
and 
<collectingRegion id="810B0AE55E61FFB68F31B8D8EC21109C" box="[835,892,1699,1725]" country="Brazil" name="Acre" pageId="1" pageNumber="331">Acre</collectingRegion>
, 
<collectingCountry id="3BD884975E61FFB68FF7B8D8EC93109C" box="[901,974,1699,1725]" name="Brazil" pageId="1" pageNumber="331">Brazil</collectingCountry>
).The current distribution of 
<taxonomicName id="84CFBF845E61FFB68971B8D8EAC1109D" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[1283,1436,1699,1724]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68971B8D8EAC1109D" box="[1283,1436,1699,1724]" italics="true" pageId="1" pageNumber="331">Wagneromyia</emphasis>
</taxonomicName>
is restricted to the Neotropical region.
</paragraph>
<paragraph id="4370C4075E61FFB68CB5B890EC161150" blockId="1.[151,1437,546,2013]" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5B890EEC91125" bold="true" box="[199,404,1770,1797]" pageId="1" pageNumber="331">Included species:</emphasis>
<taxonomicName id="84CFBF845E61FFB68DEEB890EB5B1124" authority="Vieira, Cordeiro, Bravo &amp; Araujo" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[412,1030,1771,1797]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E61FFB68DEEB890EDDC1125" box="[412,641,1771,1796]" italics="true" pageId="1" pageNumber="331">Wagneromyia aquiri</emphasis>
Vieira, Cordeiro, Bravo &amp; Araújo
</taxonomicName>
, 
<emphasis id="71BB18155E61FFB68860B890EB311124" bold="true" box="[1042,1132,1771,1797]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68860B890EB311124" box="[1042,1132,1771,1797]" pageId="1" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84CFBF845E61FFB68808B890EE6B1108" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="bicalcarata" status="comb. nov.">
<emphasis id="71BB18155E61FFB68808B890EAC11124" box="[1146,1436,1771,1797]" italics="true" pageId="1" pageNumber="331">Wagneromyia bicalcarata</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB68CEDB974EE731108" author="Quate, L. W." box="[159,302,1807,1833]" pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68D33B974EE981108" bold="true" box="[321,453,1807,1833]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68D33B974EE981108" box="[321,453,1807,1833]" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84CFBF845E61FFB68DA5B974ECD01108" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[471,909,1807,1833]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="deceptrix" status="comb. nov.">
<emphasis id="71BB18155E61FFB68DA5B974EDBE1108" box="[471,739,1807,1833]" italics="true" pageId="1" pageNumber="331">Wagneromyia deceptrix</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB68E84B974ECD81108" author="Quate, L. W." box="[758,901,1807,1833]" pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68FEAB974EB411108" bold="true" box="[920,1052,1807,1833]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68FEAB974EB411108" box="[920,1052,1807,1833]" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84CFBF845E61FFB6885FB974EF84116C" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="dimorpha" status="comb. nov.">
<emphasis id="71BB18155E61FFB6885FB974EA631108" box="[1069,1342,1807,1833]" italics="true" pageId="1" pageNumber="331">Wagneromyia dimorpha</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB68923B974EF8D116C" author="Quate, L. W." pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68CADB948EE02116C" bold="true" box="[223,351,1843,1869]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68CADB948EE02116C" box="[223,351,1843,1869]" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84CFBF845E61FFB68D1FB948EC4B116C" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[365,790,1843,1869]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="majorina" status="comb. nov.">
<emphasis id="71BB18155E61FFB68D1FB948ED29116D" box="[365,628,1843,1868]" italics="true" pageId="1" pageNumber="331">Wagneromyia majorina</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB68EF1B948EC53116C" author="Quate, L. W." box="[643,782,1843,1869]" pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68F6FB948ECC0116C" bold="true" box="[797,925,1843,1869]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68F6FB948ECC0116C" box="[797,925,1843,1869]" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="84CFBF845E61FFB68FD9B948EA10116C" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[939,1357,1843,1869]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="tripetala" status="comb. nov.">
<emphasis id="71BB18155E61FFB68FD9B948EBF6116C" box="[939,1195,1843,1869]" italics="true" pageId="1" pageNumber="331">Wagneromyia tripetala</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB688C8B948EA18116C" author="Quate, L. W." box="[1210,1349,1843,1869]" pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68926B948EF951150" bold="true" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68926B948EF951150" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="84CFBF845E61FFB68D74B92CED991150" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[262,708,1879,1905]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="tritenacula" status="comb. nov.">
<emphasis id="71BB18155E61FFB68D74B92CED7C1150" box="[262,545,1879,1905]" italics="true" pageId="1" pageNumber="331">Wagneromyia tritenacula</emphasis>
(
<bibRefCitation id="275EB9F65E61FFB68E42B92CEDE61150" author="Quate, L. W." box="[560,699,1879,1905]" pageId="1" pageNumber="331" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<emphasis id="71BB18155E61FFB68EB8B92CEC161150" bold="true" box="[714,843,1879,1905]" pageId="1" pageNumber="331">
<taxonomicNameLabel id="6A88A56E5E61FFB68EB8B92CEC161150" box="[714,843,1879,1905]" pageId="1" pageNumber="331" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E61FFB58CB5B900EE3D1070" lastPageId="2" lastPageNumber="332" pageId="1" pageNumber="331" type="discussion">
<paragraph id="4370C4075E61FFB58CB5B900EB9D173C" blockId="1.[151,1437,546,2013]" lastBlockId="2.[151,1437,151,1617]" lastPageId="2" lastPageNumber="332" pageId="1" pageNumber="331">
<emphasis id="71BB18155E61FFB68CB5B900EE6611B4" bold="true" box="[199,315,1915,1941]" pageId="1" pageNumber="331">Remarks.</emphasis>
Among the 
<taxonomicName id="84CFBF845E61FFB68DB4B900ED1811B4" authorityName="Newman" authorityYear="1834" box="[454,581,1915,1941]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
, the bulbous labellum and symmetrical aedeagus, as found in 
<taxonomicName id="84CFBF845E61FFB6888EB900EAC811B5" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[1276,1429,1915,1940]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB6888EB900EAC811B5" box="[1276,1429,1915,1940]" italics="true" pageId="1" pageNumber="331">Wagneromyia</emphasis>
</taxonomicName>
, are features shared with the genera 
<taxonomicName id="84CFBF845E61FFB68E44B9E4EDF31198" authorityName="Quate" authorityYear="1959" box="[566,686,1951,1977]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68E44B9E4EDF31198" box="[566,686,1951,1977]" italics="true" pageId="1" pageNumber="331">Eurygarka</emphasis>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68ECDB9E4EC341198" authorityName="Vaillant" authorityYear="1974" box="[703,873,1951,1977]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68ECDB9E4EC341198" box="[703,873,1951,1977]" italics="true" pageId="1" pageNumber="331">Feuerborniella</emphasis>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68F0BB9E4EBA01198" authority="Kvifte &amp; Curler, 2018" authorityName="Kvifte &amp; Curler" authorityYear="2018" box="[889,1277,1951,1977]" class="Insecta" family="Psychodidae" genus="Mucomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68F0BB9E4ECB21199" box="[889,1007,1951,1976]" italics="true" pageId="1" pageNumber="331">Mucomyia</emphasis>
<bibRefCitation id="275EB9F65E61FFB68F8BB9E4EBA01198" author="Kvifte, G. M. &amp; Curler, G. R. &amp; Marshall, S. A." box="[1017,1277,1951,1977]" pageId="1" pageNumber="331" pagination="137 - 153" refId="ref8757" refString="Kvifte, G. M., Curler, G. R. &amp; Marshall, S. A. (2018) Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History, 52, 137 - 153. https: // doi. org / 10.1080 / 00222933.2017.1410590" type="journal article" year="2018">Kvifte &amp; Curler, 2018</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68979B9E4EE6611FC" authority="Vaillant, 1973" authorityName="Vaillant" authorityYear="1973" class="Insecta" family="Psychodidae" genus="Perithreticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68979B9E4EAC11198" box="[1291,1436,1951,1977]" italics="true" pageId="1" pageNumber="331">Perithreticus</emphasis>
<bibRefCitation id="275EB9F65E61FFB68CE5B9B8EE6611FC" author="Vaillant, F." box="[151,315,1987,2013]" pageId="1" pageNumber="331" pagination="345 - 379" refId="ref9492" refString="Vaillant, F. (1973) Some new Psychodidae Psychodinae from the United States (Diptera). Annales de la Societe Entomologique de France, New Series, 9, 345 - 379." type="journal article" year="1973">Vaillant, 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68D3BB9B8EC4811FC" authority="Botosaneanu &amp; Vaillant 1970" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[329,789,1987,2013]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68D3BB9B8EEEE11FC" box="[329,435,1987,2013]" italics="true" pageId="1" pageNumber="331">Quatiella</emphasis>
<bibRefCitation id="275EB9F65E61FFB68DCFB9B8EC4811FC" author="Botosaneanu, L. &amp; Vaillant, F." box="[445,789,1987,2013]" pageId="1" pageNumber="331" pagination="173 - 183" refId="ref7854" refString="Botosaneanu, L. &amp; Vaillant, F. (1970) Trois Dipteres Psychodidae nouveaux de Cuba. Travaux du Laboratoire d'hydrobiologie et de pisciculture de l'Universite de Grenoble, 61, 173 - 183." type="journal article" year="1970">Botosaneanu &amp; Vaillant 1970</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68F51B9B8EB7E11FC" authority="Kvifte, 2015" authorityName="Kvifte" authorityYear="2015" box="[803,1059,1986,2013]" class="Insecta" family="Psychodidae" genus="Soeliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68F51B9B8ECD811FC" box="[803,901,1987,2013]" italics="true" pageId="1" pageNumber="331">Soeliella</emphasis>
<bibRefCitation id="275EB9F65E61FFB68FFDB9B8EB7E11FC" author="Kvifte, G. M." box="[911,1059,1986,2013]" pageId="1" pageNumber="331" pagination="115 - 126" refId="ref8577" refString="Kvifte, G. M. (2015) The Afrotropical genera of Psychodini: Redefinition of the tribe, first Afrotropical record of Perithreticus Vaillant, 1973 and description of Soeliella gen. nov. (Diptera: Psychodidae: Psychodinae). Zootaxa, 3986 (1), 115 - 126. https: // doi. org / 10.11646 / zootaxa. 3986.1.6" type="journal article" year="2015">Kvifte, 2015</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E61FFB68840B9B8EA3811FC" authority="Krek, 1999" authorityName="Krek" authorityYear="1999" box="[1074,1381,1987,2013]" class="Insecta" family="Psychodidae" genus="Trichosepedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E61FFB68840B9B8EB8A11FC" box="[1074,1239,1987,2013]" italics="true" pageId="1" pageNumber="331">Trichosepedon</emphasis>
<bibRefCitation id="275EB9F65E61FFB68893B9B8EA3811FC" author="Krek, S." box="[1249,1381,1987,2013]" pageId="1" pageNumber="331" refId="ref8537" refString="Krek, S. (1999) Psychodidae (Diptera, Insecta) Balkanskog Poluotoka. Federacija Bosne i Hercegovine, Ministarstvo Obrazovanja, Nauke, Kulture i Sporta, Studentska Stamparija Univerziteta Sarajevo, Sarajevo, 417 pp." type="book" year="1999">Krek, 1999</bibRefCitation>
</taxonomicName>
, and some species of 
<taxonomicName id="84CFBF845E62FFB58D2ABEECEDE81690" authority="Tonnoir, 1922" authorityName="Tonnoir" authorityYear="1922" box="[344,693,151,177]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D2ABEECED561690" box="[344,523,151,177]" italics="true" pageId="2" pageNumber="332">Trichopsychoda</emphasis>
<bibRefCitation id="275EB9F65E62FFB58E67BEECEDE81690" author="Tonnoir, A. L." box="[533,693,151,177]" pageId="2" pageNumber="332" pagination="49 - 88" refId="ref9346" refString="Tonnoir, A. L. (1922) Synopsis des Especes Europeennes du Genre Psychoda (Dipteres). Annales de la Societe Entomologique de Belgique, 62, 49 - 88." type="journal article" year="1922">Tonnoir, 1922</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="84CFBF845E62FFB58E83BEECECD41690" authorityName="Eaton" authorityYear="1904" box="[753,905,151,177]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E83BEECECD41690" box="[753,905,151,177]" italics="true" pageId="2" pageNumber="332">Philosepedon</emphasis>
</taxonomicName>
. Thus, the species of 
<taxonomicName id="84CFBF845E62FFB588F9BEECEA791691" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[1163,1316,151,176]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB588F9BEECEA791691" box="[1163,1316,151,176]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
are easily differentiated from 
<taxonomicName id="84CFBF845E62FFB58D02BEC0EEB116F4" authorityName="Newman" authorityYear="1834" box="[368,492,187,213]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
genera that include species with laterally-compressed labella and/or an asymmetrical aedeagus, such as 
<taxonomicName id="84CFBF845E62FFB58D10BEA4EE9216D8" box="[354,463,223,249]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D10BEA4EE9216D8" box="[354,463,223,249]" italics="true" pageId="2" pageNumber="332">Psychoda</emphasis>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58DAEBEA4EDB616D8" authority="Quate, 1965" authorityName="Quate" authorityYear="1965" box="[476,747,222,249]" class="Insecta" family="Psychodidae" genus="Epacretron" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58DAEBEA4ED0416D9" box="[476,601,223,248]" italics="true" pageId="2" pageNumber="332">Epacretron</emphasis>
<bibRefCitation id="275EB9F65E62FFB58E12BEA4EDB616D8" author="Quate, L. W." box="[608,747,222,249]" pageId="2" pageNumber="332" pagination="815 - 902" refId="ref9144" refString="Quate, L. W. (1965) A Taxonomic Study of Philippine Psychodidae (Diptera). Pacific Insects, 7, 815 - 902." type="journal article" year="1965">Quate, 1965</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58E87BEA4EB7B16D8" authority="Vaillant, 1963" authorityName="Vaillant" authorityYear="1963" box="[757,1062,223,249]" class="Insecta" family="Psychodidae" genus="Neomaruina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E87BEA4ECDC16D9" box="[757,897,223,248]" italics="true" pageId="2" pageNumber="332">Neomaruina</emphasis>
<bibRefCitation id="275EB9F65E62FFB58FF5BEA4EB7B16D8" author="Vaillant, F." box="[903,1062,223,249]" pageId="2" pageNumber="332" pagination="333 - 343" refId="ref9420" refString="Vaillant, F. (1963) An African psychodid larva with ventral suckers (Diptera, Psychodidae). Annals of the Natal Museum, 15, 333 - 343." type="journal article" year="1963">Vaillant, 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58842BEA4EACA16D8" authority="Vaillant, 1991" authorityName="Vaillant" authorityYear="1991" box="[1072,1431,223,249]" class="Insecta" family="Psychodidae" genus="Rhipidopsychoda" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58842BEA4EBAF16D8" box="[1072,1266,223,249]" italics="true" pageId="2" pageNumber="332">Rhipidopsychoda</emphasis>
<bibRefCitation id="275EB9F65E62FFB5888BBEA4EACA16D8" author="Vaillant, F." box="[1273,1431,223,249]" pageId="2" pageNumber="332" pagination="207 - 209" refId="ref9568" refString="Vaillant, F. (1991) Sur la position generique et tribale de deux especes de Dipteres Psychodidae Psychodinae Classification de 2 especes de Dipteres. Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, 1991, 207 - 209." type="journal article" year="1991">Vaillant, 1991</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58CE5BF78EE99173C" authority="Vaillant, 1972" authorityName="Vaillant" authorityYear="1972" box="[151,452,259,285]" class="Insecta" family="Psychodidae" genus="Nielseniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58CE5BF78EE40173C" box="[151,285,259,285]" italics="true" pageId="2" pageNumber="332">Nielseniella</emphasis>
<bibRefCitation id="275EB9F65E62FFB58D56BF78EE99173C" author="Vaillant, F." box="[292,452,259,285]" pageId="2" pageNumber="332" pagination="79 - 108" refId="ref9452" refString="Vaillant, F. (1972) Psychodidae Psychodinae. In: Linder, E. (Ed.), Die Fliegen der Palearktischen Region. E. Schweizerbartsche Verlangsbuchhandlung, Stuttgart, pp. 79 - 108." type="book chapter" year="1972">Vaillant, 1972</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58DBDBF78ED94173C" authority="Eaton, 1904" authorityName="Eaton" authorityYear="1904" box="[463,713,259,285]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58DBDBF78ED65173C" box="[463,568,259,285]" italics="true" pageId="2" pageNumber="332">Threticus</emphasis>
<bibRefCitation id="275EB9F65E62FFB58E4DBF78ED94173C" author="Eaton, A. E." box="[575,713,259,285]" pageId="2" pageNumber="332" pagination="55 - 59" refId="ref8264" refString="Eaton, A. E. (1904) New genera of European Psychodidae. Entomologist's Monthly Magazine, 15, 55 - 59." type="journal article" year="1904">Eaton, 1904</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="84CFBF845E62FFB58F76BF78EBE0173C" authority="Jezek &amp; Le Pont, 2016" authorityName="Jezek &amp; Le Pont" authorityYear="2016" box="[772,1213,259,285]" class="Insecta" family="Psychodidae" genus="Cookiellocapsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58F76BF78ECEF173C" box="[772,946,259,285]" italics="true" pageId="2" pageNumber="332">Cookiellocapsa</emphasis>
<bibRefCitation id="275EB9F65E62FFB58FCBBF78EBE0173C" author="Jezek, J. &amp; Le Pont, F." box="[953,1213,259,285]" pageId="2" pageNumber="332" pagination="813 - 826" refId="ref8400" refString="Jezek, J. &amp; Le Pont, F. (2016) Psychodidae (Diptera) of New Caledonia: Checklist and description of a new genus and species. Acta Entomologica Musei Nationalis Pragae, 56, 813 - 826." type="journal article" year="2016">Ježek &amp; Le Pont, 2016</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="4370C4075E62FFB58CB5BF5CEE7F17D4" blockId="2.[151,1437,151,1617]" pageId="2" pageNumber="332">
<taxonomicName id="84CFBF845E62FFB58CB5BF5CEE3D1761" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[199,352,295,320]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58CB5BF5CEE3D1761" box="[199,352,295,320]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
and 
<taxonomicName id="84CFBF845E62FFB58DE7BF5CED501760" authorityName="Quate" authorityYear="1959" box="[405,525,295,321]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58DE7BF5CED501760" box="[405,525,295,321]" italics="true" pageId="2" pageNumber="332">Eurygarka</emphasis>
</taxonomicName>
share the presence of contiguous eyes but 
<taxonomicName id="84CFBF845E62FFB58F90BF5CEB071760" authorityName="Quate" authorityYear="1959" box="[994,1114,295,321]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58F90BF5CEB071760" box="[994,1114,295,321]" italics="true" pageId="2" pageNumber="332">Eurygarka</emphasis>
</taxonomicName>
has ascoids with leaf-shaped anterior branches, fourth segment of the palpus greater in length than the other segments, male terminalia with sclerotized and pilose ventral epandrial plate and two tenacula on the epandrial appendages. On the other hand, 
<taxonomicName id="84CFBF845E62FFB58CE5BFE8EED6178C" box="[151,395,403,429]" pageId="2" pageNumber="332">
<emphasis id="71BB18155E62FFB58CE5BFE8EE6D178D" box="[151,304,403,428]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
species
</taxonomicName>
have ascoids with simple posterior branches, fourth segment of the palpus smaller than or equal to the first segment, weakly sclerotized and bare ventral epandrial plate, and three tenacula on the epandrial appendages.
</paragraph>
<paragraph id="4370C4075E62FFB58CB5BF84EDFE14D0" blockId="2.[151,1437,151,1617]" pageId="2" pageNumber="332">
Epandrial appendages with three tenacula, as in 
<taxonomicName id="84CFBF845E62FFB58E8FBF84ECCB1439" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[765,918,511,536]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E8FBF84ECCB1439" box="[765,918,511,536]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
can be found in 
<taxonomicName id="84CFBF845E62FFB58817BF84EB9A1438" authorityName="Kvifte" authorityYear="2015" box="[1125,1223,511,537]" class="Insecta" family="Psychodidae" genus="Soeliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58817BF84EB9A1438" box="[1125,1223,511,537]" italics="true" pageId="2" pageNumber="332">Soeliella</emphasis>
</taxonomicName>
and some species transferred to 
<taxonomicName id="84CFBF845E62FFB58D46BC58EE83141C" authorityName="Vaillant" authorityYear="1974" box="[308,478,547,573]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D46BC58EE83141C" box="[308,478,547,573]" italics="true" pageId="2" pageNumber="332">Feuerborniella</emphasis>
</taxonomicName>
by 
<bibRefCitation id="275EB9F65E62FFB58E74BC58EDA1141C" author="Cordeiro, D. &amp; Bravo, F. &amp; Chagas, C." box="[518,764,546,573]" pageId="2" pageNumber="332" pagination="457 - 472" refId="ref8011" refString="Cordeiro, D., Bravo, F. &amp; Chagas, C. (2015) Four new species of Psychodinae (Diptera: Psychodidae) from the Brazilian semiarid region, with contributions to supraspecific classification of Trichopsychodina and a redefinition of Feuerborniella. Acta Entomologica Musei Nationalis Pragae, 55, 457 - 472. https: // doi. org / 10.5281 / zenodo. 4503883" type="journal article" year="2015">
Cordeiro 
<emphasis id="71BB18155E62FFB58E03BC58EDF4141C" box="[625,681,547,573]" italics="true" pageId="2" pageNumber="332">et al.</emphasis>
(2015)
</bibRefCitation>
, but the new genus can be recognized by the contiguous eye bridge, contrasting with the separate eyes in 
<taxonomicName id="84CFBF845E62FFB58EF0BC3CEC711440" authorityName="Vaillant" authorityYear="1974" box="[642,812,583,609]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58EF0BC3CEC711440" box="[642,812,583,609]" italics="true" pageId="2" pageNumber="332">Feuerborniella</emphasis>
</taxonomicName>
and 
<taxonomicName id="84CFBF845E62FFB58F12BC3CEC9F1440" authorityName="Kvifte" authorityYear="2015" box="[864,962,583,609]" class="Insecta" family="Psychodidae" genus="Soeliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58F12BC3CEC9F1440" box="[864,962,583,609]" italics="true" pageId="2" pageNumber="332">Soeliella</emphasis>
</taxonomicName>
, and by the length of the fourth segment of the palpus, smaller or equal to the first segment in the new genus and greater in length than the other segments in the other two genera. Furthermore, in 
<taxonomicName id="84CFBF845E62FFB58E6CBCF4ED951488" authorityName="Vaillant" authorityYear="1974" box="[542,712,655,681]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E6CBCF4ED951488" box="[542,712,655,681]" italics="true" pageId="2" pageNumber="332">Feuerborniella</emphasis>
</taxonomicName>
the terminal flagellomeres can be fused, while in 
<taxonomicName id="84CFBF845E62FFB58971BCF4EFB514EC" pageId="2" pageNumber="332">
<emphasis id="71BB18155E62FFB58971BCF4EAC11489" box="[1283,1436,655,680]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
species
</taxonomicName>
they are separated, and in 
<taxonomicName id="84CFBF845E62FFB58E6CBCC8EDDD14EC" authorityName="Kvifte" authorityYear="2015" box="[542,640,691,717]" class="Insecta" family="Psychodidae" genus="Soeliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E6CBCC8EDDD14EC" box="[542,640,691,717]" italics="true" pageId="2" pageNumber="332">Soeliella</emphasis>
</taxonomicName>
, the aedeagus is wide and plate-shaped, being narrow and sometimes apically bifurcated in species of 
<taxonomicName id="84CFBF845E62FFB58E70BCACEDC614D1" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[514,667,727,752]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E70BCACEDC614D1" box="[514,667,727,752]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4370C4075E62FFB58CB5BC80EC451230" blockId="2.[151,1437,151,1617]" pageId="2" pageNumber="332">
The species of the genus 
<taxonomicName id="84CFBF845E62FFB58DAABC80ED131535" authorityName="Kvifte &amp; Curler" authorityYear="2018" box="[472,590,763,788]" class="Insecta" family="Psychodidae" genus="Mucomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58DAABC80ED131535" box="[472,590,763,788]" italics="true" pageId="2" pageNumber="332">Mucomyia</emphasis>
</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58E28BC80EDB61534" authorityName="Vaillant" authorityYear="1973" box="[602,747,763,789]" class="Insecta" family="Psychodidae" genus="Perithreticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E28BC80EDB61534" box="[602,747,763,789]" italics="true" pageId="2" pageNumber="332">Perithreticus</emphasis>
</taxonomicName>
, 
<emphasis id="71BB18155E62FFB58E85BC80EB5B1534" box="[759,1030,763,789]" italics="true" pageId="2" pageNumber="332">
<taxonomicName id="84CFBF845E62FFB58E85BC80ECCF1534" authorityName="Eaton" authorityYear="1904" box="[759,914,763,789]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">Philosepedon</taxonomicName>
, 
<taxonomicName id="84CFBF845E62FFB58FEEBC80EB5B1534" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[924,1030,763,789]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">Quatiella</taxonomicName>
</emphasis>
, 
<taxonomicName id="84CFBF845E62FFB58860BC80EB981534" authorityName="Tonnoir" authorityYear="1922" box="[1042,1221,763,789]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58860BC80EB981534" box="[1042,1221,763,789]" italics="true" pageId="2" pageNumber="332">Trichopsychoda</emphasis>
</taxonomicName>
and 
<taxonomicName id="84CFBF845E62FFB58885BC80EAC11534" authorityName="Krek" authorityYear="1999" box="[1271,1436,763,789]" class="Insecta" family="Psychodidae" genus="Trichosepedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58885BC80EAC11534" box="[1271,1436,763,789]" italics="true" pageId="2" pageNumber="332">Trichosepedon</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="84CFBF845E62FFB58D6ABD64EEEC1519" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[280,433,799,824]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D6ABD64EEEC1519" box="[280,433,799,824]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
by the separate eyes, at least in the lower facet rows, and the length of the first segment of the palpus, which is smaller than the others.Also, in 
<taxonomicName id="84CFBF845E62FFB58EAABD38EC13157D" authorityName="Kvifte &amp; Curler" authorityYear="2018" box="[728,846,835,860]" class="Insecta" family="Psychodidae" genus="Mucomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58EAABD38EC13157D" box="[728,846,835,860]" italics="true" pageId="2" pageNumber="332">Mucomyia</emphasis>
</taxonomicName>
the male terminalia has gonostylus without subapical seta, 
<taxonomicName id="84CFBF845E62FFB58CA0BD1CEE3E15A0" authorityName="Vaillant" authorityYear="1973" box="[210,355,871,897]" class="Insecta" family="Psychodidae" genus="Perithreticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58CA0BD1CEE3E15A0" box="[210,355,871,897]" italics="true" pageId="2" pageNumber="332">Perithreticus</emphasis>
</taxonomicName>
has more than five tenacula, 
<taxonomicName id="84CFBF845E62FFB58EC0BD1CEC1715A0" authorityName="Eaton" authorityYear="1904" box="[690,842,871,897]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58EC0BD1CEC1715A0" box="[690,842,871,897]" italics="true" pageId="2" pageNumber="332">Philosepedon</emphasis>
</taxonomicName>
has two tenacula, 
<taxonomicName id="84CFBF845E62FFB5886CBD1CEBD515A0" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[1054,1160,871,897]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB5886CBD1CEBD515A0" box="[1054,1160,871,897]" italics="true" pageId="2" pageNumber="332">Quatiella</emphasis>
</taxonomicName>
has only one tenaculum in the epandrial appendage, in 
<taxonomicName id="84CFBF845E62FFB58D87BDF0EDF51584" authorityName="Tonnoir" authorityYear="1922" box="[501,680,907,933]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D87BDF0EDF51584" box="[501,680,907,933]" italics="true" pageId="2" pageNumber="332">Trichopsychoda</emphasis>
</taxonomicName>
the parameres are absent and in 
<taxonomicName id="84CFBF845E62FFB5886CBDF0EB9E1584" authorityName="Krek" authorityYear="1999" box="[1054,1219,907,933]" class="Insecta" family="Psychodidae" genus="Trichosepedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB5886CBDF0EB9E1584" box="[1054,1219,907,933]" italics="true" pageId="2" pageNumber="332">Trichosepedon</emphasis>
</taxonomicName>
the gonostylus has a longitudinal groove carrying a row of stout setae and the epandrial appendages have two tenacula. In contrast, in males of 
<taxonomicName id="84CFBF845E62FFB58D51BDA8EEE115CD" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[291,444,979,1004]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58D51BDA8EEE115CD" box="[291,444,979,1004]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
the gonostylus has a subapical seta and no distinct longitudinal groove, the epandrial appendage has three tenacula, and parameres are present.
</paragraph>
<paragraph id="4370C4075E62FFB58CB5BA60EC5E1280" blockId="2.[151,1437,151,1617]" pageId="2" pageNumber="332">
<bibRefCitation id="275EB9F65E62FFB58CB5BA60EEB11214" author="Omelkova, M. &amp; Jezek, J." box="[199,492,1051,1077]" pageId="2" pageNumber="332" pagination="29 - 42" refId="ref8926" refString="Omelkova, M. &amp; Jezek, J. (2012) Two new species of Philosepedon Eaton (Diptera, Psychodidae, Psychodinae) from Europe, with comments on subgeneric classification. The Canadian Entomologist, 2012, 29 - 42." type="journal article" year="2012">Omelková &amp; Ježek (2012)</bibRefCitation>
proposed a monospecific subgenus 
<emphasis id="71BB18155E62FFB58F08BA60EB451214" box="[890,1048,1051,1077]" italics="true" pageId="2" pageNumber="332">Philothreticus</emphasis>
<bibRefCitation id="275EB9F65E62FFB5886FBA60EBC41214" author="Krek, S." box="[1053,1177,1051,1077]" pageId="2" pageNumber="332" refId="ref8537" refString="Krek, S. (1999) Psychodidae (Diptera, Insecta) Balkanskog Poluotoka. Federacija Bosne i Hercegovine, Ministarstvo Obrazovanja, Nauke, Kulture i Sporta, Studentska Stamparija Univerziteta Sarajevo, Sarajevo, 417 pp." type="book" year="1999">Krek, 1999</bibRefCitation>
for 
<taxonomicName id="84CFBF845E62FFB588B7BA60EA001214" authorityName="Eaton" authorityYear="1904" box="[1221,1373,1051,1077]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB588B7BA60EA001214" box="[1221,1373,1051,1077]" italics="true" pageId="2" pageNumber="332">Philosepedon</emphasis>
</taxonomicName>
, with a contiguous eye bridge, but with the lower facet rows separated, ascoids with only two branches and asymmetric aedeagal complex, different from the new genus with all the rows of facets touching in the midline, ascoids with three branches, and the aedeagal complex symmetrical.
</paragraph>
<paragraph id="4370C4075E62FFB58CB5BAD0EE3D1070" blockId="2.[151,1437,151,1617]" pageId="2" pageNumber="332">
A feature observed for most species of 
<taxonomicName id="84CFBF845E62FFB58E0FBAD0EC4B12E5" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[637,790,1195,1220]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58E0FBAD0EC4B12E5" box="[637,790,1195,1220]" italics="true" pageId="2" pageNumber="332">Wagneromyia</emphasis>
</taxonomicName>
is the labrum with a convex and slightly expanded dorsal margin. This characteristic has been only observed in this genus until now, but a review of this structure is still needed regarding the other genera of the tribe, as this structure was rarely described and illustrated in previous works. Furthermore, as described for 
<taxonomicName id="84CFBF845E62FFB58E31BB6CEB1D1310" authority="Chagas &amp; Cordeiro, 2015" authorityName="Chagas &amp; Cordeiro" authorityYear="2015" box="[579,1088,1302,1329]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="71BB18155E62FFB58E31BB6CEC491311" box="[579,788,1303,1329]" italics="true" pageId="2" pageNumber="332">Quatiella truncata</emphasis>
Chagas &amp; 
<bibRefCitation id="275EB9F65E62FFB58FE7BB6CEB1D1310" author="Cordeiro, D. &amp; Bravo, F. &amp; Chagas, C." box="[917,1088,1302,1329]" pageId="2" pageNumber="332" pagination="457 - 472" refId="ref8011" refString="Cordeiro, D., Bravo, F. &amp; Chagas, C. (2015) Four new species of Psychodinae (Diptera: Psychodidae) from the Brazilian semiarid region, with contributions to supraspecific classification of Trichopsychodina and a redefinition of Feuerborniella. Acta Entomologica Musei Nationalis Pragae, 55, 457 - 472. https: // doi. org / 10.5281 / zenodo. 4503883" type="journal article" year="2015">Cordeiro, 2015</bibRefCitation>
</taxonomicName>
and species of 
<taxonomicName id="84CFBF845E62FFB58881BB6CEAC01310" authorityName="Vaillant" authorityYear="1974" box="[1267,1437,1303,1329]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E62FFB58881BB6CEAC01310" box="[1267,1437,1303,1329]" italics="true" pageId="2" pageNumber="332">Feuerborniella</emphasis>
</taxonomicName>
auctt (
<bibRefCitation id="275EB9F65E62FFB58CAEBB40EE9C1374" author="Cordeiro, D. &amp; Bravo, F. &amp; Chagas, C." box="[220,449,1338,1365]" pageId="2" pageNumber="332" pagination="457 - 472" refId="ref8011" refString="Cordeiro, D., Bravo, F. &amp; Chagas, C. (2015) Four new species of Psychodinae (Diptera: Psychodidae) from the Brazilian semiarid region, with contributions to supraspecific classification of Trichopsychodina and a redefinition of Feuerborniella. Acta Entomologica Musei Nationalis Pragae, 55, 457 - 472. https: // doi. org / 10.5281 / zenodo. 4503883" type="journal article" year="2015">
Cordeiro 
<emphasis id="71BB18155E62FFB58D3ABB40EEDD1374" box="[328,384,1339,1365]" italics="true" pageId="2" pageNumber="332">et al.</emphasis>
2015
</bibRefCitation>
; 
<bibRefCitation id="275EB9F65E62FFB58DBFBB40EDC31374" author="Pereira, T. P. L. &amp; Bravo, F. &amp; Araujo, M. X. &amp; Cordeiro, D. &amp; Chagas, C. &amp; Delabie, J. H. C." box="[461,670,1338,1365]" pageId="2" pageNumber="332" pagination="18 - 22" refId="ref8970" refString="Pereira, T. P. L., Bravo, F., Araujo, M. X., Cordeiro, D., Chagas, C. &amp; Delabie, J. H. C. (2015) Moth flies (Diptera: Psychodidae) collected in colonies of the fire ant Solenopsis virulens (Smith) (Hymenoptera: Formicidae), with description of two new species. Sociobiology, 62, 18 - 22. https: // doi. org / 10.13102 / sociobiology. v 62 i 1.18 - 22" type="journal article" year="2015">
Pereira 
<emphasis id="71BB18155E62FFB58E51BB40ED061374" box="[547,603,1339,1365]" italics="true" pageId="2" pageNumber="332">et al.</emphasis>
2015
</bibRefCitation>
), the species of the new genus present an extension of the gonocoxal apodemes (= fused parameres), which surround the aedeagus to form a dorsal sheath, but which do not completely cover the aedeagus as in Brachycera (see 
<bibRefCitation id="275EB9F65E62FFB58E16BBF8EC2A13BC" author="Cumming, J. M. &amp; Wood, M. D." box="[612,887,1411,1437]" pageId="2" pageNumber="332" pagination="89 - 133" refId="ref8081" refString="Cumming, J. M. &amp; Wood, M. D. (2017) Adult Morphology and Terminology. In: Kirk-Spriggs, A. H. &amp; Sinclair, B. J. (Eds.), Manual of Afrotropical Diptera Introductory chapters and keys to Diptera families. South African National Biodiversity Institute, Pretoria, pp. 89 - 133." type="book chapter" year="2017">Cumming &amp; Wood 2017</bibRefCitation>
for comments). 
<bibRefCitation id="275EB9F65E62FFB5885CBBF8EB9C13BC" author="Quate, L. W." box="[1070,1217,1411,1437]" pageId="2" pageNumber="332" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate (1996)</bibRefCitation>
cited this structure, referring to it as projections of the anterior gonocoxal apodeme, 
<bibRefCitation id="275EB9F65E62FFB58F1DBBDCEB3D13E0" author="Ibanez-Bernal, S." box="[879,1120,1447,1473]" pageId="2" pageNumber="332" pagination="1 - 8" refId="ref8313" refString="Ibanez-Bernal, S. (2004) A new species of the genus Feuerborniella Vaillant, from Mexico (Diptera: Psychodidae). Zootaxa, 412 (1), 1 - 8. https: // doi. org / 10.11646 / zootaxa. 412.1.1" type="journal article" year="2004">Ibáñez-Bernal (2004)</bibRefCitation>
described it as the aedeagal sheath and 
<bibRefCitation id="275EB9F65E62FFB58D66BBB0EEF013C4" author="Kvifte, G. M." box="[276,429,1483,1509]" pageId="2" pageNumber="332" pagination="1 - 10" refId="ref8644" refString="Kvifte, G. M. (2019) New records of Norwegian Psychodidae, with the first description of the female of Trichosepedon balkanicum (Krek, 1970) comb. nov. Norwegian Journal of Entomology, 66, 1 - 10." type="journal article" year="2019">Kvifte (2019)</bibRefCitation>
named these structures as parameral dorsal bridge. Although still not properly named, this is observed for other genera in illustrations (
<emphasis id="71BB18155E62FFB58EE1BB94EDE61029" box="[659,699,1519,1544]" italics="true" pageId="2" pageNumber="332">e.g.</emphasis>
<bibRefCitation id="275EB9F65E62FFB58EB6BB94ECCC1028" author="Kvifte, G. M. &amp; Andersen, T. &amp; Hagenlund, L. K. &amp; Bello Gonzalez, O. C." box="[708,913,1519,1545]" pageId="2" pageNumber="332" pagination="121 - 127" refId="ref8689" refString="Kvifte, G. M., Andersen, T., Hagenlund, L. K. &amp; Bello Gonzalez, O. C. (2016) Two new Neotropical species of Perithreticus Vaillant 1973 (Diptera: Psychodidae, Psychodinae). Studies on Neotropical Fauna and Environment, 51, 121 - 127. https: // doi. org / 10.1080 / 01650521.2016.1198579" type="journal article" year="2016">
Kvifte 
<emphasis id="71BB18155E62FFB58F66BB94EC131028" box="[788,846,1519,1545]" italics="true" pageId="2" pageNumber="332">et al.</emphasis>
2016
</bibRefCitation>
, figs. 4, 8), however, it has gone unnoticed in works and therefore it is necessary to be revised in 
<taxonomicName id="84CFBF845E62FFB58E92B868EC01100C" authorityName="Newman" authorityYear="1834" box="[736,860,1555,1581]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
species to understand how diagnostic it can be for the tribe’s genera.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E62FFB58CE5B8D8ECF7109C" box="[151,938,1698,1725]" pageId="2" pageNumber="332" type="nomenclature">
<paragraph id="4370C4075E62FFB58CE5B8D8ECF7109C" blockId="2.[151,938,1698,1761]" box="[151,938,1698,1725]" pageId="2" pageNumber="332">
<heading id="1838736B5E62FFB58CE5B8D8ECF7109C" bold="true" box="[151,938,1698,1725]" fontSize="11" level="1" pageId="2" pageNumber="332" reason="1">
<emphasis id="71BB18155E62FFB58CE5B8D8ECF7109C" bold="true" box="[151,938,1698,1725]" pageId="2" pageNumber="332">
<taxonomicName id="84CFBF845E62FFB58CE5B8D8EC62109C" ID-CoL="9J6XN" authority="Vieira, Cordeiro, Bravo &amp; Araujo" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[151,831,1698,1725]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E62FFB58CE5B8D8EED2109C" bold="true" box="[151,399,1699,1725]" italics="true" pageId="2" pageNumber="332">Wagneromyia aquiri</emphasis>
Vieira, Cordeiro, Bravo &amp; Araújo
</taxonomicName>
, 
<taxonomicNameLabel id="6A88A56E5E62FFB58F3EB8D8ECF7109C" box="[844,938,1699,1725]" pageId="2" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E62FFB58CE5B8BCEE5610C0" box="[151,267,1735,1761]" pageId="2" pageNumber="332" type="description">
<paragraph id="4370C4075E62FFB58CE5B8BCEE5610C0" blockId="2.[151,938,1698,1761]" box="[151,267,1735,1761]" pageId="2" pageNumber="332">
(
<figureCitation id="DBF4D8825E62FFB58CEDB8BCEE5F10C0" box="[159,258,1735,1761]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–9" captionStartId-0="3.[152,270,1842,1866]" captionStartId-1="4.[152,270,1896,1920]" captionTargetBox-0="[158,1426,1005,1813]" captionTargetBox-1="[204,1380,198,1856]" captionTargetId-0="figure-411@3.[151,1437,993,1819]" captionTargetId-1="figure-18@4.[197,1390,181,1872]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." captionText-1="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi-0="http://doi.org/10.5281/zenodo.7095480" figureDoi-1="http://doi.org/10.5281/zenodo.7095482" httpUri-0="https://zenodo.org/record/7095480/files/figure.png" httpUri-1="https://zenodo.org/record/7095482/files/figure.png" pageId="2" pageNumber="332">Figs 1–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E62FFB58CE5B973EFA7114B" pageId="2" pageNumber="332" type="materials_examined">
<paragraph id="4370C4075E62FFB58CE5B973EFA7114B" blockId="2.[151,1437,1800,2043]" pageId="2" pageNumber="332">
<emphasis id="71BB18155E62FFB58CE5B973EE141103" bold="true" box="[151,329,1800,1826]" pageId="2" pageNumber="332">Type material:</emphasis>
<materialsCitation id="F3A7CE5A5E62FFB58D21B973ED551102" ID-GBIF-Occurrence="3912952304" box="[339,520,1800,1827]" collectionCode="BR" pageId="2" pageNumber="332" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9C747AA55E62FFB58D21B973EEE81103" box="[339,437,1800,1826]" pageId="2" pageNumber="332" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E62FFB58DCCB972EE8B1102" box="[446,470,1801,1827]" pageId="2" pageNumber="332" type="male">♂</specimenCount>
. 
<collectionCode id="25DE5CC25E62FFB58D91B972ED551102" box="[483,520,1801,1827]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="2" pageNumber="332">BR</collectionCode>
</materialsCitation>
<materialsCitation id="F3A7CE5A5E62FFB58E63B973EC3E1167" ID-GBIF-Occurrence="3912952302" collectingDate="2008-07-07" collectingDateMax="2008-07-16" collectingDateMin="2008-07-07" collectionCode="MZFS" collectorName="Malaise &amp; G. Melo" country="Brazil" county="Km" elevation="170" latitude="67.683334" location="Embrapa" municipality="Rio Branco" pageId="2" pageNumber="332" specimenCount="1" stateProvince="Acre" typeStatus="holotype">
[
<collectingCountry id="3BD884975E62FFB58E6AB973ED391103" box="[536,612,1800,1826]" name="Brazil" pageId="2" pageNumber="332">Brazil</collectingCountry>
], 
<collectingRegion id="810B0AE55E62FFB58E05B973EDED1103" box="[631,688,1800,1826]" country="Brazil" name="Acre" pageId="2" pageNumber="332">Acre</collectingRegion>
, 15 
<collectingCounty id="AA11BC8B5E62FFB58E91B972EC501102" box="[739,781,1801,1827]" pageId="2" pageNumber="332">Km</collectingCounty>
a SE do 
<collectingMunicipality id="A3145E7D5E62FFB58F09B973EB5D1102" box="[891,1024,1800,1827]" pageId="2" pageNumber="332">Rio Branco</collectingMunicipality>
, 
<location id="461092DC5E62FFB5887FB973EB281103" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E62FFB5887FB973EB281103" box="[1037,1141,1800,1826]" country="Brazil" county="Km" latitude="67.683334" municipality="Rio Branco" name="Embrapa" pageId="2" pageNumber="332" stateProvince="Acre">Embrapa</location>
, 
<quantity id="843769E25E62FFB588F1B973EB9E1103" box="[1155,1219,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="332" unit="m" value="170.0">
<elevation id="C8E223345E62FFB588F1B973EB9E1103" box="[1155,1219,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="332" unit="m" value="170.0">170m</elevation>
</quantity>
[altitude], 
<geoCoordinate id="26FBA2C05E62FFB58931B973EAC11103" box="[1347,1436,1800,1826]" degrees="10" direction="south" minutes="01" orientation="latitude" pageId="2" pageNumber="332" precision="925" value="-10.016666">10º01’S</geoCoordinate>
<geoCoordinate id="26FBA2C05E62FFB58CE5B957EFA51167" box="[151,248,1836,1862]" degrees="67" direction="north" minutes="41" orientation="latitude" pageId="2" pageNumber="332" precision="925" value="67.683334">67º41’N</geoCoordinate>
, 
<date id="3771E2C75E62FFB58D76B957EEF61167" box="[260,427,1836,1862]" pageId="2" pageNumber="332" value="2008-07-07" valueMax="2008-07-16" valueMin="2008-07-07">
<collectingDate id="27351B2F5E62FFB58D76B957EEF61167" box="[260,427,1836,1862]" pageId="2" pageNumber="332" value="2008-07-07" valueMax="2008-07-16" valueMin="2008-07-07">7–16.VII.2008</collectingDate>
</date>
, 
<collectorName id="EE3AA1D15E62FFB58DC5B957ED4D1167" box="[439,528,1836,1862]" pageId="2" pageNumber="332">Malaise</collectorName>
[trap] 
<collectorName id="EE3AA1D15E62FFB58E2EB957EDE71167" box="[604,698,1836,1862]" pageId="2" pageNumber="332">G. Melo</collectorName>
[leg.] (
<collectionCode id="25DE5CC25E62FFB58F7EB957EC061167" box="[780,859,1836,1862]" pageId="2" pageNumber="332">MZFS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="F3A7CE5A5E62FFB58F1DB956EFAB114B" ID-GBIF-Occurrence="3912952309" collectionCode="MZFS" pageId="2" pageNumber="332" specimenCount="9" specimenCount-male="9" typeStatus="paratype">
<typeStatus id="9C747AA55E62FFB58F1DB956EC871166" box="[879,986,1837,1863]" pageId="2" pageNumber="332" type="paratype">paratypes</typeStatus>
<specimenCount id="55C90F8E5E62FFB58F90B957EB531166" box="[994,1038,1836,1863]" pageId="2" pageNumber="332" type="male">9 ♂</specimenCount>
, same date and locality as holotype (
<collectionCode id="25DE5CC25E62FFB58CEDB92BEFB3114B" box="[159,238,1872,1898]" pageId="2" pageNumber="332">MZFS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E62FFB58CB5B90FEDF611DB" pageId="2" pageNumber="332" type="etymology">
<paragraph id="4370C4075E62FFB58CB5B90FEDF611DB" blockId="2.[151,1437,1800,2043]" pageId="2" pageNumber="332">
<emphasis id="71BB18155E62FFB58CB5B90FEE1311AF" bold="true" box="[199,334,1908,1934]" pageId="2" pageNumber="332">Etymology.</emphasis>
The specific name, 
<taxonomicName id="84CFBF845E62FFB58E5DB90EED2E11AF" authorityName="Vieira, Cordeiro, Bravo &amp; Araujo" authorityYear="2022" box="[559,627,1909,1934]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="332" phylum="Arthropoda" rank="species" species="aquiri">
<emphasis id="71BB18155E62FFB58E5DB90EED2E11AF" box="[559,627,1909,1934]" italics="true" pageId="2" pageNumber="332">aquiri</emphasis>
</taxonomicName>
, refers to “a’quiri” or “a’kiru”, from the word uwákürü, from the dialect of the Ipurinã tribe, the original inhabitants of the region, from which the toponym 
<collectingRegion id="810B0AE55E62FFB5883EB9E3EBDF1193" box="[1100,1154,1944,1970]" country="Brazil" name="Acre" pageId="2" pageNumber="332">Acre</collectingRegion>
was derived. 
<collectingRegion id="810B0AE55E62FFB58952B9E3EA0B1193" box="[1312,1366,1944,1970]" country="Brazil" name="Acre" pageId="2" pageNumber="332">Acre</collectingRegion>
is the name of the main river of the region and also the name of the Brazilian state where the species was collected. The name should be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E63FFB48CB5BEEDED151744" pageId="3" pageNumber="333" type="diagnosis">
<paragraph id="4370C4075E63FFB48CB5BEEDED151744" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="333">
<emphasis id="71BB18155E63FFB48CB5BEEDEE1F1691" bold="true" box="[199,322,150,176]" pageId="3" pageNumber="333">Diagnosis.</emphasis>
Medial fork incomplete; R 
<subScript id="DF4BC6425E63FFB48E0EBED2EDC81696" attach="left" box="[636,661,169,183]" fontSize="6" pageId="3" pageNumber="333">2+3</subScript>
1.2x length of R 
<subScript id="DF4BC6425E63FFB48F29BED2EC3E1696" attach="left" box="[859,867,169,183]" fontSize="6" pageId="3" pageNumber="333">2</subScript>
; apex of the gonostylus bent, with a small apical accessory seta and a long subapical seta on the ventral side; aedeagus straight, bending ventrolaterally to form a tubular-like structure that narrows to a small, bifurcated apex; parameres well sclerotized, broad at the base and tapering to a hook-shaped apex, curved outwards; parameral sheath subrectangular, with concave posterior margin; epandrial appendages with a group of five subapical papillae on the dorsal surface; hypoproct pilose with straight posterior margin and rounded vertices.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E63FFB48CB5BF15EEFF15E8" pageId="3" pageNumber="333" type="description">
<paragraph id="4370C4075E63FFB48CB5BF15EBD71584" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="333">
<emphasis id="71BB18155E63FFB48CB5BF15EEF817A8" bold="true" box="[199,421,366,393]" pageId="3" pageNumber="333">Description. Male.</emphasis>
Head (
<figureCitation id="DBF4D8825E63FFB48D88BF14ED6117A8" box="[506,572,367,393]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="3" pageNumber="333">Fig. 1</figureCitation>
) as in genus description; eye bridge with a small medial spur on the dorsal margin and five widely spaced supraocular setae; group of setae on frons does not extend to the lower limit of the eye bridge; frontoclypeal suture present, as a sclerotized area; antenna incomplete in all specimens, flagellomeres beyond 10th lost in 
<typeStatus id="9C747AA55E63FFB48D06BFA0EE8717D4" box="[372,474,475,501]" pageId="3" pageNumber="333" type="holotype">holotype</typeStatus>
; scape, pedicel, flagellomeres, and ascoids as in the genus description; palpus formula 1.0:1.6:1.2:1.0 first segment with a dorsolateral group of sensilla in the middle (
<figureCitation id="DBF4D8825E63FFB48854BF84EB361438" box="[1062,1131,511,537]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="3" pageNumber="333">Fig. 3</figureCitation>
); clypeus with four to five prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and three setae on the lateral margin. Wing (
<figureCitation id="DBF4D8825E63FFB48F5BBC3CEC301440" box="[809,877,583,609]" captionStart="FIGURES 1–5" captionStartId="3.[152,270,1842,1866]" captionTargetBox="[158,1426,1005,1813]" captionTargetId="figure-411@3.[151,1437,993,1819]" captionTargetPageId="3" captionText="FIGURES 1–5. Wagneromyia aquiri, sp.nov. 1. Head.2.Wing.3. Mouthparts.4. Labellum (ventral view).5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum." figureDoi="http://doi.org/10.5281/zenodo.7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="3" pageNumber="333">Fig. 2</figureCitation>
) as in genus description; medial fork incomplete; R 
<subScript id="DF4BC6425E63FFB48CD8BC06EF9E14AA" attach="left" box="[170,195,637,651]" fontSize="6" pageId="3" pageNumber="333">2+3</subScript>
1.2x the length of R 
<subScript id="DF4BC6425E63FFB48DC7BC06EEE014AA" attach="left" box="[437,445,637,651]" fontSize="6" pageId="3" pageNumber="333">2</subScript>
. Male terminalia (
<figureCitation id="DBF4D8825E63FFB48EE1BC10EC5C14A4" box="[659,769,619,645]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Figs. 6–9</figureCitation>
) with a narrow band-shaped hypandrium, wider medially; gonocoxites as in genus; gonostylus, with a sudden angulation near the apex, bearing a small apical accessory seta and a long subapical seta on the ventral surface (
<figureCitation id="DBF4D8825E63FFB48EBDBCC8EC6214EC" box="[719,831,691,717]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Figs. 6, 7</figureCitation>
); aedeagus bending ventrally on the sides forming a tubular-like structure with a tapered and bifurcated apex on the ventral view (
<figureCitation id="DBF4D8825E63FFB48F9CBCACEB0A14D0" box="[1006,1111,727,753]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Figs. 6, 7</figureCitation>
); parameres well sclerotized, broad at the base and tapering to a hook-shaped apex, curved outwards; parameres meet each other dorsally to the aedeagus forming a band-shaped parameral sheath with a small concavity at the midline (
<figureCitation id="DBF4D8825E63FFB488F9BD64EB8C1518" box="[1163,1233,799,825]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Fig. 6</figureCitation>
); epandrium as in genus; epandrial appendages as in genus, with a group of five subapical papillae on the dorsal surface and three apical tenacula with half the length of the appendages (
<figureCitation id="DBF4D8825E63FFB48F70BD1CEC1815A0" box="[770,837,871,897]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Fig. 8</figureCitation>
); hypoproct pilose with straight posterior margin and rounded vertices (
<figureCitation id="DBF4D8825E63FFB48D12BDF0EEF91584" box="[352,420,907,933]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Fig. 8</figureCitation>
); epiproct as in genus (
<figureCitation id="DBF4D8825E63FFB48EDBBDF0EDB31584" box="[681,750,907,933]" captionStart="FIGURES 6–9" captionStartId="4.[152,270,1896,1920]" captionTargetBox="[204,1380,198,1856]" captionTargetId="figure-18@4.[197,1390,181,1872]" captionTargetPageId="4" captionText="FIGURES 6–9. Wagneromyia aquiri, sp. nov. 6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="3" pageNumber="333">Fig. 9</figureCitation>
); subepandrial sclerite membranous.
</paragraph>
<paragraph id="4370C4075E63FFB48CB5BDD4EEFF15E8" blockId="3.[151,1437,150,969]" box="[199,418,943,969]" pageId="3" pageNumber="333">
<emphasis id="71BB18155E63FFB48CB5BDD4EE7815E8" bold="true" box="[199,293,943,969]" pageId="3" pageNumber="333">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="17B0948F5E63FFB48CEAB949EB8A114F" ID-DOI="http://doi.org/10.5281/zenodo.7095480" ID-Zenodo-Dep="7095480" httpUri="https://zenodo.org/record/7095480/files/figure.png" pageId="3" pageNumber="333" startId="3.[152,270,1842,1866]" targetBox="[158,1426,1005,1813]" targetPageId="3">
<paragraph id="4370C4075E63FFB48CEAB949EB8A114F" blockId="3.[151,1437,1842,1903]" pageId="3" pageNumber="333">
<emphasis id="71BB18155E63FFB48CEAB949EE63116B" bold="true" box="[152,318,1842,1866]" pageId="3" pageNumber="333">FIGURES 1–5.</emphasis>
<taxonomicName id="84CFBF845E63FFB48D30B948ED52116B" authority="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo, 2022" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" box="[322,527,1843,1866]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E63FFB48D30B948ED52116B" box="[322,527,1843,1866]" italics="true" pageId="3" pageNumber="333">Wagneromyia aquiri</emphasis>
</taxonomicName>
, 
<emphasis id="71BB18155E63FFB48E6BB948ED3A116A" bold="true" box="[537,615,1843,1867]" pageId="3" pageNumber="333">
<taxonomicNameLabel id="6A88A56E5E63FFB48E6BB948ED3A116A" box="[537,615,1843,1867]" pageId="3" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1. Head. 2. Wing. 3. Mouthparts. 4. Labellum (ventral view). 5. Third flagellomere. Abbreviations: asc—ascoid; plp 1 sen—first palpus sensilla; flgm sen—flagellomere sensilla; lbr—labrum.
</paragraph>
</caption>
<caption id="17B0948F5E64FFB38CEAB913EC2711CD" ID-DOI="http://doi.org/10.5281/zenodo.7095482" ID-Zenodo-Dep="7095482" httpUri="https://zenodo.org/record/7095482/files/figure.png" pageId="4" pageNumber="334" startId="4.[152,270,1896,1920]" targetBox="[204,1380,198,1856]" targetPageId="4">
<paragraph id="4370C4075E64FFB38CEAB913EC2711CD" blockId="4.[151,1437,1896,2028]" pageId="4" pageNumber="334">
<emphasis id="71BB18155E64FFB38CEAB913EE1F11A1" bold="true" box="[152,322,1896,1920]" pageId="4" pageNumber="334">FIGURES 6–9.</emphasis>
<taxonomicName id="84CFBF845E64FFB38D38B912ED4711A1" authority="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo, 2022" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" box="[330,538,1897,1920]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E64FFB38D38B912ED4711A1" box="[330,538,1897,1920]" italics="true" pageId="4" pageNumber="334">Wagneromyia aquiri</emphasis>
</taxonomicName>
, 
<emphasis id="71BB18155E64FFB38E5AB913ED2711A1" bold="true" box="[552,634,1896,1920]" pageId="4" pageNumber="334">
<taxonomicNameLabel id="6A88A56E5E64FFB38E5AB913ED2711A1" box="[552,634,1896,1920]" pageId="4" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
6. Gonopods and aedeagal complex (dorsal view). 7. Gonopods and aedeagal complex (ventral view). 8. Epandrium (dorsal view). 9. Epandrium (ventral view). Abbreviations: aed—aedeagus; epand app— epandrial appendages; epiprct—epiproct; goncx apod—gonocoxal apodemes; hypd—hypandrium; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath; sbepand scl—subepandrial sclerite.
</paragraph>
</caption>
<subSubSection id="0BD5978C5E65FFB28CE5BEECEC561690" box="[151,779,151,178]" pageId="5" pageNumber="335" type="nomenclature">
<paragraph id="4370C4075E65FFB28CE5BEECEC561690" blockId="5.[151,779,151,213]" box="[151,779,151,178]" pageId="5" pageNumber="335">
<heading id="1838736B5E65FFB28CE5BEECEC561690" bold="true" box="[151,779,151,178]" fontSize="11" level="1" pageId="5" pageNumber="335" reason="1">
<emphasis id="71BB18155E65FFB28CE5BEECEC561690" bold="true" box="[151,779,151,178]" pageId="5" pageNumber="335">
<taxonomicName id="84CFBF845E65FFB28CE5BEECED231690" ID-CoL="9J67K" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,638,151,178]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="bicalcarata" status="comb. nov.">
<emphasis id="71BB18155E65FFB28CE5BEECEE901693" bold="true" box="[151,461,151,178]" italics="true" pageId="5" pageNumber="335">Wagneromyia bicalcarata</emphasis>
(
<bibRefCitation id="275EB9F65E65FFB28DAEBEECED281690" author="Quate, L. W." box="[476,629,151,177]" pageId="5" pageNumber="335" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E65FFB28EF7BEECEC561690" box="[645,779,151,177]" pageId="5" pageNumber="335" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BEC0EE7A16F4" box="[151,295,187,213]" pageId="5" pageNumber="335" type="description">
<paragraph id="4370C4075E65FFB28CE5BEC0EE7A16F4" blockId="5.[151,779,151,213]" box="[151,295,187,213]" pageId="5" pageNumber="335">
(
<figureCitation id="DBF4D8825E65FFB28CEDBEC0EE4316F4" box="[159,286,187,213]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Figs 10–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BF7BED0B1736" box="[151,598,255,280]" pageId="5" pageNumber="335" type="reference_group">
<paragraph id="4370C4075E65FFB28CE5BF7BED0B1736" blockId="5.[151,598,255,280]" box="[151,598,255,280]" pageId="5" pageNumber="335">
<treatmentCitationGroup id="63DFE3295E65FFB28CE5BF7BED0B1736" box="[151,598,255,280]" pageId="5" pageNumber="335">
<treatmentCitation id="C26EE2165E65FFB28CE5BF7BED0F1736" author="Quate, L. W." box="[151,594,255,280]" page="58" pageId="5" pageNumber="335" year="1996">
<taxonomicName id="84CFBF845E65FFB28CE5BF7BED0F1736" ID-CoL="89WS3" authority="Quate, 1996: 58" authorityName="Quate" authorityPageNumber="58" authorityYear="1996" box="[151,594,255,280]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="bicalcaratus">
<emphasis id="71BB18155E65FFB28CE5BF7BEEFB1739" box="[151,422,256,280]" italics="true" pageId="5" pageNumber="335">Philosepedon bicalcaratus</emphasis>
<bibRefCitation id="275EB9F65E65FFB28DDEBF7BED0F1736" author="Quate, L. W." box="[428,594,255,280]" pageId="5" pageNumber="335" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 58</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BF3EECF417A5" pageId="5" pageNumber="335" type="materials_examined">
<paragraph id="4370C4075E65FFB28CE5BF3EECF417A5" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CE5BF3EEED5177E" bold="true" box="[151,392,325,351]" pageId="5" pageNumber="335">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E65FFB28DE1BF3DECF817A5" ID-GBIF-Occurrence="3912952308" collectionCode="INBIO" collectorName="Lt. Trp &amp; L. W. Quate." country="Costa Rica" location="Pr. Puerto Viejo de Sarapaqui" pageId="5" pageNumber="335" specimenCode="CRI001468679" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="9C747AA55E65FFB28DE1BF3DEEA81741" box="[403,501,326,352]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E65FFB28E73BF3DED4B1741" box="[513,534,326,352]" pageId="5" pageNumber="335" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E65FFB28E58BF3DEDDF1741" box="[554,642,326,352]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 16</figureCitation>
). 
<collectingCountry id="3BD884975E65FFB28EE9BF3DEC191741" box="[667,836,326,352]" name="Costa Rica" pageId="5" pageNumber="335">COSTA RICA</collectingCountry>
, 
<collectingRegion id="810B0AE55E65FFB28F26BF3DECED1741" box="[852,944,326,352]" country="Costa Rica" name="Heredia" pageId="5" pageNumber="335">Heredia</collectingRegion>
, 
<location id="461092DC5E65FFB28FCDBF3DEA421741" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E65FFB28FCDBF3DEA421741" box="[959,1311,326,352]" country="Costa Rica" name="Pr. Puerto Viejo de Sarapaqui" pageId="5" pageNumber="335" stateProvince="Heredia">Pr. Puerto Viejo de Sarapaqui</location>
, Est. Biol. LaSelva, 
<collectorName id="EE3AA1D15E65FFB28D73BF11EE1317A5" box="[257,334,362,388]" pageId="5" pageNumber="335">Lt. Trp</collectorName>
22-25/VII/93, 
<collectorName id="EE3AA1D15E65FFB28D8ABF11EDDB17A5" box="[504,646,362,388]" pageId="5" pageNumber="335">L. W. Quate.</collectorName>
(
<collectionCode id="25DE5CC25E65FFB28EE6BF11EDB917A5" box="[660,740,362,388]" pageId="5" pageNumber="335">INBIO</collectionCode>
<specimenCode id="13696C7C5E65FFB28E99BF11ECC017A5" box="[747,925,362,388]" collectionCode="INBIO" pageId="5" pageNumber="335">CRI001468679</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CB5BFF6EA7A17D1" pageId="5" pageNumber="335" type="diagnosis">
<paragraph id="4370C4075E65FFB28CB5BFF6EA7A17D1" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5BFF6EE1F1786" bold="true" box="[199,322,397,423]" pageId="5" pageNumber="335">Diagnosis.</emphasis>
Medial fork incomplete, M 
<subScript id="DF4BC6425E65FFB28E05BFDBED22178F" attach="left" box="[631,639,416,430]" fontSize="6" pageId="5" pageNumber="335">2</subScript>
not reaching M 
<subScript id="DF4BC6425E65FFB28F43BFDBEC64178F" attach="left" box="[817,825,416,430]" fontSize="6" pageId="5" pageNumber="335">1</subScript>
; gonostylus apically bifurcated, like a chela; aedeagus with slightly enlarged bifid apex; subrectangular parameres apically bifurcated with an external arm like a hook and internal arm rounded; parameral sheath absent; hypoproct with medial lobe subtriangular in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CB5BF82EEFF1211" pageId="5" pageNumber="335" type="description">
<paragraph id="4370C4075E65FFB28CB5BF82EC2C122C" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5BF82ED1C1432" bold="true" box="[199,577,505,532]" pageId="5" pageNumber="335">
Redescription. Male (
<typeStatus id="9C747AA55E65FFB28DBBBF82ED681432" box="[457,565,505,531]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
).
</emphasis>
Head (
<figureCitation id="DBF4D8825E65FFB28EE6BF81EDBB1435" box="[660,742,506,532]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 10</figureCitation>
) as in genus description; eye bridge with small medial dorsal spur and six supraocular setae, the innermost three closest; group of setae on frons extending medially close to the limit of the inferior margin of the eye bridge; frontoclypeal suture as a slightly sclerotized area, incomplete laterally; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and one seta on the lateral margin; antenna (
<figureCitation id="DBF4D8825E65FFB28F81BCF1EB1B1485" box="[1011,1094,650,676]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 11</figureCitation>
) as in genus; palpus formula 1.0:1.5:1.3:0.8, first segment of palpus with a group of sensilla at the slightly inflated base. Wing (
<figureCitation id="DBF4D8825E65FFB28970BCD5EA0814E9" box="[1282,1365,686,712]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 12</figureCitation>
) as in genus; radial fork distal to medial fork; R 
<subScript id="DF4BC6425E65FFB28E01BC9FEDD014D3" attach="left" box="[627,653,740,754]" fontSize="6" pageId="5" pageNumber="335">2+3</subScript>
1.5x the length of R 
<subScript id="DF4BC6425E65FFB28F0ABC9FECDD14D3" attach="left" box="[888,896,740,754]" fontSize="6" pageId="5" pageNumber="335">2</subScript>
; M 
<subScript id="DF4BC6425E65FFB28FD8BC9FECEF14D3" attach="left" box="[938,946,740,754]" fontSize="6" pageId="5" pageNumber="335">2</subScript>
not reaching M 
<subScript id="DF4BC6425E65FFB2881BBC9FEB2C14D3" attach="left" box="[1129,1137,740,754]" fontSize="6" pageId="5" pageNumber="335">1</subScript>
(medial fork incomplete). Male terminalia (
<figureCitation id="DBF4D8825E65FFB28D29BC8DEEBF1530" box="[347,482,758,785]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Figs. 13–15</figureCitation>
) with hypandrium narrow, like a strip; gonocoxites as in genus; gonostylus apically bifurcated (
<figureCitation id="DBF4D8825E65FFB28D65BD61EE341515" box="[279,361,794,820]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 13</figureCitation>
), with two short and strong projections like a chela, and with a long subapical seta on the ventral margin; aedeagus with slightly enlarged bifid apex; parameres subrectangular (
<figureCitation id="DBF4D8825E65FFB28874BD45EBD11579" box="[1030,1164,830,856]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Figs. 13, 14</figureCitation>
), with the apical portion bifurcated into external arm hook-like, positioned dorsally to the aedeagus (
<figureCitation id="DBF4D8825E65FFB28F9CBD19EB1C155D" box="[1006,1089,866,892]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 13</figureCitation>
), and an internal arm rounded, lateral to the aedeagus (
<figureCitation id="DBF4D8825E65FFB28DD7BDFDEEA71581" box="[421,506,902,928]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 14</figureCitation>
); parameres not developed into a parameral sheath; epandrium (
<figureCitation id="DBF4D8825E65FFB288A6BDFDEA7B1580" box="[1236,1318,902,929]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 15</figureCitation>
) damaged and in lateral view in the 
<typeStatus id="9C747AA55E65FFB28DC0BDD1ED4515E5" box="[434,536,938,964]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
; epandrial appendages (
<figureCitation id="DBF4D8825E65FFB28F51BDD1EC2915E4" box="[803,884,938,965]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 15</figureCitation>
) curved in lateral view, base wider than the apex, with a group of three subapical papillae on the dorsal margin and tree apical tenacula with half the length of the appendices; epiproct short; hypoproct with medial lobe (
<figureCitation id="DBF4D8825E65FFB28F62BD89EC3E122C" box="[784,867,1010,1037]" captionStart="FIGURES 10–16" captionStartId="8.[152,270,1943,1967]" captionTargetBox="[179,1400,202,1906]" captionTargetId="figure-18@8.[174,1413,181,1920]" captionTargetPageId="8" captionText="FIGURES 10–16. Wagneromyia bicalcarata. 10. Head. 11.Terminal flagellomeres.12.Wing.13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere." figureDoi="http://doi.org/10.5281/zenodo.7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="5" pageNumber="335">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="4370C4075E65FFB28CB5BA6DEEFF1211" blockId="5.[151,1437,325,1144]" box="[199,418,1046,1072]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5BA6DEE781211" bold="true" box="[199,293,1046,1072]" pageId="5" pageNumber="335">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CB5BA41ECB31259" pageId="5" pageNumber="335" type="discussion">
<paragraph id="4370C4075E65FFB28CB5BA41ECB31259" blockId="5.[151,1437,325,1144]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5BA41EE661275" bold="true" box="[199,315,1082,1108]" pageId="5" pageNumber="335">Remarks.</emphasis>
Although 
<bibRefCitation id="275EB9F65E65FFB28DC3BA41ED181275" author="Quate, L. W." box="[433,581,1082,1108]" pageId="5" pageNumber="335" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate (1996)</bibRefCitation>
described 
<taxonomicName id="84CFBF845E65FFB28ECEBA40EC3F1275" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[700,866,1082,1108]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="bicalcarata">
<emphasis id="71BB18155E65FFB28ECEBA40ED8E1275" box="[700,723,1083,1108]" italics="true" pageId="5" pageNumber="335">W</emphasis>
. 
<emphasis id="71BB18155E65FFB28E93BA41EC3F1275" box="[737,866,1082,1108]" italics="true" pageId="5" pageNumber="335">bicalcarata</emphasis>
</taxonomicName>
with separated gonocoxites, the examination of the 
<typeStatus id="9C747AA55E65FFB28CE5BA25EFA41259" box="[151,249,1118,1144]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
shows that they are contiguous as in the other species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BAB1EDAE12C5" box="[151,755,1225,1252]" pageId="5" pageNumber="335" type="nomenclature">
<paragraph id="4370C4075E65FFB28CE5BAB1EDAE12C5" blockId="5.[151,755,1225,1288]" box="[151,755,1225,1252]" pageId="5" pageNumber="335">
<heading id="1838736B5E65FFB28CE5BAB1EDAE12C5" bold="true" box="[151,755,1225,1252]" fontSize="11" level="1" pageId="5" pageNumber="335" reason="1">
<emphasis id="71BB18155E65FFB28CE5BAB1EDAE12C5" bold="true" box="[151,755,1225,1252]" pageId="5" pageNumber="335">
<taxonomicName id="84CFBF845E65FFB28CE5BAB1ED3812C2" ID-CoL="9J6PH" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,613,1225,1252]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="deceptrix" status="comb. nov.">
<emphasis id="71BB18155E65FFB28CE5BAB1EEE812C5" bold="true" box="[151,437,1225,1252]" italics="true" pageId="5" pageNumber="335">Wagneromyia deceptrix</emphasis>
(
<bibRefCitation id="275EB9F65E65FFB28DB6BAB2ED0012C2" author="Quate, L. W." box="[452,605,1225,1251]" pageId="5" pageNumber="335" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E65FFB28E1FBAB1EDAE12C5" box="[621,755,1226,1252]" pageId="5" pageNumber="335" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BA95EE7A1329" box="[151,295,1262,1288]" pageId="5" pageNumber="335" type="description">
<paragraph id="4370C4075E65FFB28CE5BA95EE7A1329" blockId="5.[151,755,1225,1288]" box="[151,295,1262,1288]" pageId="5" pageNumber="335">
(
<figureCitation id="DBF4D8825E65FFB28CEDBA95EE421329" box="[159,287,1262,1288]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Figs 17–24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BB49ED68136B" box="[151,565,1330,1354]" pageId="5" pageNumber="335" type="reference_group">
<paragraph id="4370C4075E65FFB28CE5BB49ED68136B" blockId="5.[151,565,1330,1354]" box="[151,565,1330,1354]" pageId="5" pageNumber="335">
<treatmentCitationGroup id="63DFE3295E65FFB28CE5BB49ED68136B" box="[151,565,1330,1354]" pageId="5" pageNumber="335">
<treatmentCitation id="C26EE2165E65FFB28CE5BB49ED6C136B" author="Quate, L. W." box="[151,561,1330,1354]" page="58" pageId="5" pageNumber="335" year="1996">
<taxonomicName id="84CFBF845E65FFB28CE5BB49ED6C136B" ID-CoL="4GD2R" authority="Quate, 1996: 58" authorityName="Quate" authorityPageNumber="58" authorityYear="1996" box="[151,561,1330,1354]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="deceptrix">
<emphasis id="71BB18155E65FFB28CE5BB49EED8136B" box="[151,389,1330,1354]" italics="true" pageId="5" pageNumber="335">Philosepedon deceptrix</emphasis>
<bibRefCitation id="275EB9F65E65FFB28DFEBB49ED6C136B" author="Quate, L. W." box="[396,561,1330,1354]" pageId="5" pageNumber="335" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 58</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CE5BB03EC8D1397" pageId="5" pageNumber="335" type="materials_examined">
<paragraph id="4370C4075E65FFB28CE5BB03EC8D1397" blockId="5.[151,1437,1400,2039]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CE5BB03EED913B3" bold="true" box="[151,388,1400,1426]" pageId="5" pageNumber="335">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E65FFB28DFFBB03EC901397" ID-GBIF-Occurrence="3912952310" collectionCode="INBIO" collectorName="L. W. Quate" country="Costa Rica" elevation="1800" location="Vara Blanca" pageId="5" pageNumber="335" specimenCode="CRI001469462" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="9C747AA55E65FFB28DFFBB03EEB213B3" box="[397,495,1400,1426]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E65FFB28D8ABB02ED5013B2" box="[504,525,1401,1427]" pageId="5" pageNumber="335" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E65FFB28E6CBB03ED2913B2" box="[542,628,1400,1427]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 24</figureCitation>
). 
<collectingCountry id="3BD884975E65FFB28EFBBB02EC7213B2" box="[649,815,1401,1427]" name="Costa Rica" pageId="5" pageNumber="335">COSTA RICA</collectingCountry>
, 
<collectingRegion id="810B0AE55E65FFB28F4EBB03ECC813B3" box="[828,917,1400,1426]" country="Costa Rica" name="Heredia" pageId="5" pageNumber="335">Heredia</collectingRegion>
<location id="461092DC5E65FFB28FEFBB02EB7713B3" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E65FFB28FEFBB02EB7713B3" box="[925,1066,1400,1427]" country="Costa Rica" name="Vara Blanca" pageId="5" pageNumber="335" stateProvince="Heredia">Vara Blanca</location>
, 
<quantity id="843769E25E65FFB28844BB02EBDB13B2" box="[1078,1158,1401,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="335" unit="m" value="1800.0">
<elevation id="C8E223345E65FFB28844BB02EBDB13B2" box="[1078,1158,1401,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="335" unit="m" value="1800.0">1800m</elevation>
</quantity>
, 26/VII/ 
<specimenCount id="55C90F8E5E65FFB28895BB03EA7E13B2" box="[1255,1315,1400,1427]" pageId="5" pageNumber="335" type="generic">93 L.</specimenCount>
<collectorName id="EE3AA1D15E65FFB28959BB02EAC513B2" box="[1323,1432,1401,1427]" pageId="5" pageNumber="335">W. Quate</collectorName>
, 
<quantity id="843769E25E65FFB28CE5BBE6EFBA1396" box="[151,231,1437,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="335" unit="m" value="1800.0">
<elevation id="C8E223345E65FFB28CE5BBE6EFBA1396" box="[151,231,1437,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="335" unit="m" value="1800.0">1800m</elevation>
</quantity>
, 
<collectorName id="EE3AA1D15E65FFB28C81BBE6EE611396" box="[243,316,1437,1463]" pageId="5" pageNumber="335">Lt Trp</collectorName>
, 
<collectorName id="EE3AA1D15E65FFB28D3ABBE7EE8F1396" box="[328,466,1436,1463]" pageId="5" pageNumber="335">Euparal, LN</collectorName>
—239.500-519.000 (
<collectionCode id="25DE5CC25E65FFB28ECEBBE6EC511396" box="[700,780,1437,1463]" pageId="5" pageNumber="335">INBIO</collectionCode>
<specimenCode id="13696C7C5E65FFB28F61BBE7EC981397" box="[787,965,1436,1462]" collectionCode="INBIO" pageId="5" pageNumber="335">CRI001469462</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB28CB5BBBBEC321067" pageId="5" pageNumber="335" type="diagnosis">
<paragraph id="4370C4075E65FFB28CB5BBBBEC321067" blockId="5.[151,1437,1400,2039]" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5BBBBEE6713FB" bold="true" box="[199,314,1472,1498]" pageId="5" pageNumber="335">Diagnosis</emphasis>
. 14th flagellomere with two apical spiniform setulae; medial fork complete; subtriangular parameral sheath with a rounded apex; aedeagus straight, with bifid apex, like two ventral lobes; parameral sheath subtriangular with a rounded apex; anterior gonocoxal apodemes with setae on the posteromedial margin, continuous with the ventral setae of the gonocoxite; hypoproct pilose and subovalate.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E65FFB18CB5B82BEEFF1760" lastPageId="6" lastPageNumber="336" pageId="5" pageNumber="335" type="description">
<paragraph id="4370C4075E65FFB18CB5B82BED2C173C" blockId="5.[151,1437,1400,2039]" lastBlockId="6.[151,1436,151,321]" lastPageId="6" lastPageNumber="336" pageId="5" pageNumber="335">
<emphasis id="71BB18155E65FFB28CB5B82BED1F104B" bold="true" box="[199,578,1616,1643]" pageId="5" pageNumber="335">
Redescription. Male (
<typeStatus id="9C747AA55E65FFB28DB8B82BED6B104B" box="[458,566,1616,1642]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
).
</emphasis>
Head (
<figureCitation id="DBF4D8825E65FFB28EE5B82BEDB7104A" box="[663,746,1616,1643]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 17</figureCitation>
) as in genus description; eye bridge with a very short dorsal medial spur and five supraocular setae; group of setae on frons not extending medially; frontoclypeal suture only as a slightly less sclerotized area; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and four setae on the external lateral margin, the most apical one shorter than the others; antenna (
<figureCitation id="DBF4D8825E65FFB28EB4B89BEC4510DA" box="[710,792,1760,1787]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 19</figureCitation>
) as in genus; 14 
<superScript id="B4BA694F5E65FFB28FA6B89BECBC10CF" attach="left" box="[980,993,1760,1774]" fontSize="6" pageId="5" pageNumber="335">th</superScript>
flagellomere with two apical stout and spiniform setulae; palpus formula: 1.0:1.6:1.2:0.8, first segment of palpus with a group of sensilla near the base. Wing (
<figureCitation id="DBF4D8825E65FFB28C96B953EE6B1162" box="[228,310,1832,1859]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 18</figureCitation>
) as in genus description; radial fork distal to medial fork, both complete; R 
<subScript id="DF4BC6425E65FFB288F8B940EBFE1168" attach="left" box="[1162,1187,1851,1865]" fontSize="6" pageId="5" pageNumber="335">2+3</subScript>
1.3x the length of R 
<subScript id="DF4BC6425E65FFB289FFB940EAC81168" attach="left" box="[1421,1429,1851,1865]" fontSize="6" pageId="5" pageNumber="335">2</subScript>
. Male terminalia (
<figureCitation id="DBF4D8825E65FFB28D29B937EEBE1146" box="[347,483,1868,1895]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Figs. 20–23</figureCitation>
) with hypandrium as a narrow band, little sclerotized; gonocoxite with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus (
<figureCitation id="DBF4D8825E65FFB2897BB90BEAC911AA" box="[1289,1428,1904,1931]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Figs. 20, 21</figureCitation>
) straight, with bilobed apex; parameres (
<figureCitation id="DBF4D8825E65FFB28E12B9EFEDE9118E" box="[608,692,1940,1967]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 21</figureCitation>
) with a membranous (non-sclerotized) region between them and the gonocoxal apodemes; parameres with a broad base from where it projects dorsally to the aedeagus forming a subtriangular parameral sheath with a rounded apex (
<figureCitation id="DBF4D8825E65FFB28F60B9A7EC3411D6" box="[786,873,2012,2039]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="5" pageNumber="335">Fig. 20</figureCitation>
); anterior gonocoxal apodemes with setae on the posteromedial margin, continuous with the ventral group of setae of the gonocoxite (
<figureCitation id="DBF4D8825E66FFB18823BEECEBF81690" box="[1105,1189,151,177]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="6" pageNumber="336">Fig. 21</figureCitation>
); epandrium (
<figureCitation id="DBF4D8825E66FFB18930BEECEAC91690" box="[1346,1428,151,177]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="6" pageNumber="336">Fig. 22</figureCitation>
) typical for genus; epandrial appendages (
<figureCitation id="DBF4D8825E66FFB18E2CBEC0EDF316F4" box="[606,686,187,213]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="6" pageNumber="336">Fig. 23</figureCitation>
) as in genus description, with a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct (
<figureCitation id="DBF4D8825E66FFB18817BEA4EBEB16D8" box="[1125,1206,223,249]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="6" pageNumber="336">Fig. 22</figureCitation>
) small and rounded; hypoproct (
<figureCitation id="DBF4D8825E66FFB18D6ABF78EE34173C" box="[280,361,259,285]" captionStart="FIGURES 17–24" captionStartId="9.[152,270,1884,1908]" captionTargetBox="[187,1386,194,1851]" captionTargetId="figure-26@9.[187,1401,181,1860]" captionTargetPageId="9" captionText="FIGURES 17–24. Wagneromyia deceptrix. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="6" pageNumber="336">Fig. 23</figureCitation>
) pilose and subovalate.
</paragraph>
<paragraph id="4370C4075E66FFB18CB5BF5CEEFF1760" blockId="6.[151,1436,151,321]" box="[199,418,295,321]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5BF5CEE781760" bold="true" box="[199,293,295,321]" pageId="6" pageNumber="336">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BFE8EDA7178C" box="[151,762,402,429]" pageId="6" pageNumber="336" type="nomenclature">
<paragraph id="4370C4075E66FFB18CE5BFE8EDA7178C" blockId="6.[151,762,402,465]" box="[151,762,402,429]" pageId="6" pageNumber="336">
<heading id="1838736B5E66FFB18CE5BFE8EDA7178C" bold="true" box="[151,762,402,429]" fontSize="11" level="1" pageId="6" pageNumber="336" reason="1">
<emphasis id="71BB18155E66FFB18CE5BFE8EDA7178C" bold="true" box="[151,762,402,429]" pageId="6" pageNumber="336">
<taxonomicName id="84CFBF845E66FFB18CE5BFE8ED31178D" ID-CoL="9J7FZ" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,620,402,429]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="dimorpha" status="comb. nov.">
<emphasis id="71BB18155E66FFB18CE5BFE8EEE6178C" bold="true" box="[151,443,402,429]" italics="true" pageId="6" pageNumber="336">Wagneromyia dimorpha</emphasis>
(
<bibRefCitation id="275EB9F65E66FFB18DB9BFE9ED39178D" author="Quate, L. W." box="[459,612,402,428]" pageId="6" pageNumber="336" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E66FFB18E01BFE8EDA7178C" box="[627,762,403,429]" pageId="6" pageNumber="336" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BFCCEE7A17F0" box="[151,295,438,465]" pageId="6" pageNumber="336" type="description">
<paragraph id="4370C4075E66FFB18CE5BFCCEE7A17F0" blockId="6.[151,762,402,465]" box="[151,295,438,465]" pageId="6" pageNumber="336">
(
<figureCitation id="DBF4D8825E66FFB18CEDBFCCEE7D17F0" box="[159,288,438,465]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Figs 25–30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BF80ED191432" box="[151,580,507,531]" pageId="6" pageNumber="336" type="reference_group">
<paragraph id="4370C4075E66FFB18CE5BF80ED191432" blockId="6.[151,580,507,531]" box="[151,580,507,531]" pageId="6" pageNumber="336">
<treatmentCitationGroup id="63DFE3295E66FFB18CE5BF80ED191432" box="[151,580,507,531]" pageId="6" pageNumber="336">
<treatmentCitation id="C26EE2165E66FFB18CE5BF80ED621432" author="Quate, L. W." box="[151,575,507,531]" page="58" pageId="6" pageNumber="336" year="1996">
<taxonomicName id="84CFBF845E66FFB18CE5BF80ED621432" ID-CoL="89WSD" authority="Quate, 1996: 58" authorityName="Quate" authorityPageNumber="58" authorityYear="1996" box="[151,575,507,531]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="71BB18155E66FFB18CE5BF80EECE1432" box="[151,403,507,531]" italics="true" pageId="6" pageNumber="336">Philosepedon dimorphus</emphasis>
<bibRefCitation id="275EB9F65E66FFB18DE8BF80ED621432" author="Quate, L. W." box="[410,575,507,531]" pageId="6" pageNumber="336" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 58</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BC3AECD8145E" pageId="6" pageNumber="336" type="materials_examined">
<paragraph id="4370C4075E66FFB18CE5BC3AECD8145E" blockId="6.[151,1437,577,1323]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CE5BC3AEEDE147A" bold="true" box="[151,387,577,603]" pageId="6" pageNumber="336">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E66FFB18DF8BC3AED3B14A1" ID-GBIF-Occurrence="3912952303" collectorName="Lt. Trp &amp; L. W. Quate" country="Costa Rica" elevation="75" location="Puerto Viejo de Sarapaqui" pageId="6" pageNumber="336" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="9C747AA55E66FFB18DF8BC3AEEB1147A" box="[394,492,577,603]" pageId="6" pageNumber="336" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E66FFB18D81BC39ED55147D" box="[499,520,578,604]" pageId="6" pageNumber="336" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E66FFB18E6ABC3AED31147A" box="[536,620,577,603]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 30</figureCitation>
). 
<collectingCountry id="3BD884975E66FFB18E0DBC3AEC78147A" box="[639,805,577,603]" name="Costa Rica" pageId="6" pageNumber="336">COSTA RICA</collectingCountry>
, 
<collectingRegion id="810B0AE55E66FFB18F42BC3AECD1147A" box="[816,908,577,603]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="336">Heredia</collectingRegion>
, 
<location id="461092DC5E66FFB18FE4BC39EBE3147A" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E66FFB18FE4BC39EBE3147A" box="[918,1214,577,604]" country="Costa Rica" name="Puerto Viejo de Sarapaqui" pageId="6" pageNumber="336" stateProvince="Heredia">Puerto Viejo de Sarapaqui</location>
, Est. Biol. LaSelva, 
<collectorName id="EE3AA1D15E66FFB18CE5BC1DEFB814A1" box="[151,229,614,640]" pageId="6" pageNumber="336">Lt. Trp</collectorName>
23/VII/93, 
<collectorName id="EE3AA1D15E66FFB18D1ABC1DEEAE145E" box="[360,499,613,640]" pageId="6" pageNumber="336">L. W. Quate</collectorName>
, 
<quantity id="843769E25E66FFB18D8CBC1EED3B14A1" box="[510,614,613,640]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="336" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="C8E223345E66FFB18D8CBC1EED3B14A1" box="[510,614,613,640]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="336" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
50- 
<specimenCount id="55C90F8E5E66FFB18E51BC1EED3B14A1" box="[547,614,613,640]" pageId="6" pageNumber="336" type="generic">100m</specimenCount>
</elevation>
</quantity>
</materialsCitation>
<materialsCitation id="F3A7CE5A5E66FFB18E14BC1EECD8145E" ID-GBIF-Occurrence="3912952301" box="[614,901,613,640]" collectionCode="INBIO" country="Costa Rica" pageId="6" pageNumber="336" specimenCode="CRI001468641" specimenCount="1" typeStatus="holotype">
. (
<collectionCode id="25DE5CC25E66FFB18E0ABC1EED95145E" box="[632,712,613,639]" pageId="6" pageNumber="336">INBIO</collectionCode>
<specimenCode id="13696C7C5E66FFB18EBDBC1EEC20145E" box="[719,893,613,639]" collectionCode="INBIO" pageId="6" pageNumber="336">CRI001468641</specimenCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CB5BCF2EF80152E" pageId="6" pageNumber="336" type="diagnosis">
<paragraph id="4370C4075E66FFB18CB5BCF2EF80152E" blockId="6.[151,1437,577,1323]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5BCF2EE1F1482" bold="true" box="[199,322,649,675]" pageId="6" pageNumber="336">Diagnosis.</emphasis>
Medial fork incomplete; gonostylus with small subapical spines lined on the ventrolateral margin and a long subapical seta; aedeagus with a straight and tapered bifid apex; parameres long, robust, losing sclerotinization as they approach an outward hook-like apex; parameral sheath poorly sclerotized, as a narrow and membranous stripe.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CB5BD62EEFF130A" pageId="6" pageNumber="336" type="description">
<paragraph id="4370C4075E66FFB18CB5BD62EC4F1326" blockId="6.[151,1437,577,1323]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5BD62EE9F1512" bold="true" box="[199,450,793,819]" pageId="6" pageNumber="336">Redescription. Male.</emphasis>
Head (
<figureCitation id="DBF4D8825E66FFB18E6ABD62ED311515" box="[536,620,793,820]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 25</figureCitation>
) as in genus description; eye bridge with a small medial spur and five supraocular setae, the two or three innermost closest; group of setae on frons extending medially; frontoclypeal suture only as a slightly less sclerotized area; antenna incomplete in 
<typeStatus id="9C747AA55E66FFB18FB4BD1AEB76155A" box="[966,1067,865,891]" pageId="6" pageNumber="336" type="holotype">holotype</typeStatus>
, flagellomeres beyond 7th lost; ascoids lost in 
<typeStatus id="9C747AA55E66FFB18D31BDFEEEF415BE" box="[323,425,901,927]" pageId="6" pageNumber="336" type="holotype">holotype</typeStatus>
; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin and one seta on the external lateral margin; palpus formula 1.0:1.4:1.3:0.9, first segment of palpus as in genus description. Wing (
<figureCitation id="DBF4D8825E66FFB18FC9BDB6EB5215C6" box="[955,1039,973,999]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 26</figureCitation>
) as in genus description; complete radial fork distal to an incomplete medial fork; R 
<subScript id="DF4BC6425E66FFB18EA9BA7FEDA91233" attach="left" box="[731,756,1028,1042]" fontSize="6" pageId="6" pageNumber="336">2+3</subScript>
1.3x the length of R Male terminalia (
<figureCitation id="DBF4D8825E66FFB188BFBD8AEA05122A" box="[1229,1368,1009,1035]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Figs. 27–29</figureCitation>
) with hypandrium as a narrow band; gonocoxites as in genus description; gonostylus with small subapical spines lined on the ventrolateral margin and a long subapical seta; aedeagus straight and bifid on distal half; parameres (
<figureCitation id="DBF4D8825E66FFB1894BBA42EFE91256" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Figs. 27, 28</figureCitation>
) reaching the apex of aedeagus, robust and wide at the base, losing sclerotinization as they approach an outward hook-like apex; parameral sheath (
<figureCitation id="DBF4D8825E66FFB18E57BAFAED2412BA" box="[549,633,1153,1179]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 27</figureCitation>
) poorly sclerotized, like a narrow and membranous stripe; epandrium (
<figureCitation id="DBF4D8825E66FFB18CEDBADEEFAF129E" box="[159,242,1189,1215]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 29</figureCitation>
) as in genus description, but damaged; epandrial appendages with a group of short setulae on the base, a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct (
<figureCitation id="DBF4D8825E66FFB18D70BA96EE091326" box="[258,340,1261,1287]" captionStart="FIGURES 25–30" captionStartId="10.[152,270,1943,1967]" captionTargetBox="[170,1415,199,1911]" captionTargetId="figure-18@10.[163,1424,181,1920]" captionTargetPageId="10" captionText="FIGURES 25–30. Wagneromyia dimorpha. 25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="6" pageNumber="336">Fig. 29</figureCitation>
) short; hypoproct not very pronounced.
</paragraph>
<paragraph id="4370C4075E66FFB18F9FBA7FECA61233" blockId="6.[151,1437,577,1323]" box="[1005,1019,1028,1042]" pageId="6" pageNumber="336">
<subScript id="DF4BC6425E66FFB18F9FBA7FECA61233" attach="left" box="[1005,1019,1028,1042]" fontSize="6" pageId="6" pageNumber="336">2.</subScript>
</paragraph>
<paragraph id="4370C4075E66FFB18CB5BB6AEEFF130A" blockId="6.[151,1437,577,1323]" box="[199,418,1297,1323]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5BB6AEE78130A" bold="true" box="[199,293,1297,1323]" pageId="6" pageNumber="336">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BB06EDAE13B6" box="[151,755,1405,1431]" pageId="6" pageNumber="336" type="nomenclature">
<paragraph id="4370C4075E66FFB18CE5BB06EDAE13B6" blockId="6.[151,755,1405,1467]" box="[151,755,1405,1431]" pageId="6" pageNumber="336">
<heading id="1838736B5E66FFB18CE5BB06EDAE13B6" bold="true" box="[151,755,1405,1431]" fontSize="11" level="1" pageId="6" pageNumber="336" reason="1">
<emphasis id="71BB18155E66FFB18CE5BB06EDAE13B6" bold="true" box="[151,755,1405,1431]" pageId="6" pageNumber="336">
<taxonomicName id="84CFBF845E66FFB18CE5BB06ED3813B6" ID-CoL="9J5VZ" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,613,1405,1431]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="majorina" status="comb. nov.">
<emphasis id="71BB18155E66FFB18CE5BB06EEE813B6" bold="true" box="[151,437,1405,1431]" italics="true" pageId="6" pageNumber="336">Wagneromyia majorina</emphasis>
(
<bibRefCitation id="275EB9F65E66FFB18DB6BB06ED0013B6" author="Quate, L. W." box="[452,605,1405,1431]" pageId="6" pageNumber="336" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E66FFB18E1FBB06EDAE13B6" box="[621,755,1405,1431]" pageId="6" pageNumber="336" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BBDAEE7A139A" box="[151,295,1441,1467]" pageId="6" pageNumber="336" type="description">
<paragraph id="4370C4075E66FFB18CE5BBDAEE7A139A" blockId="6.[151,755,1405,1467]" box="[151,295,1441,1467]" pageId="6" pageNumber="336">
(
<figureCitation id="DBF4D8825E66FFB18CEDBBDAEE42139A" box="[159,287,1441,1467]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Figs 31–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5BB9DED6313DF" box="[151,574,1510,1534]" pageId="6" pageNumber="336" type="reference_group">
<paragraph id="4370C4075E66FFB18CE5BB9DED6313DF" blockId="6.[151,574,1510,1534]" box="[151,574,1510,1534]" pageId="6" pageNumber="336">
<treatmentCitationGroup id="63DFE3295E66FFB18CE5BB9DED6313DF" box="[151,574,1510,1534]" pageId="6" pageNumber="336">
<treatmentCitation id="C26EE2165E66FFB18CE5BB9DED6713DF" author="Quate, L. W." box="[151,570,1510,1534]" page="60" pageId="6" pageNumber="336" year="1996">
<taxonomicName id="84CFBF845E66FFB18CE5BB9DED6713DF" ID-CoL="89WSS" authority="Quate, 1996: 60" authorityName="Quate" authorityPageNumber="60" authorityYear="1996" box="[151,570,1510,1534]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="majorinus">
<emphasis id="71BB18155E66FFB18CE5BB9DEED313DC" box="[151,398,1510,1534]" italics="true" pageId="6" pageNumber="336">Philosepedon majorinus</emphasis>
<bibRefCitation id="275EB9F65E66FFB18DE7BB9DED6713DF" author="Quate, L. W." box="[405,570,1510,1534]" pageId="6" pageNumber="336" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CE5B857ECA4104A" pageId="6" pageNumber="336" type="materials_examined">
<paragraph id="4370C4075E66FFB18CE5B857ECA4104A" blockId="6.[151,1437,1580,2039]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CE5B857EEDF1067" bold="true" box="[151,386,1580,1606]" pageId="6" pageNumber="336">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E66FFB18DFBB857ECA8104A" ID-GBIF-Occurrence="3912952306" collectionCode="INBIO" collectorName="L. W. Quate" country="Costa Rica" elevation="1700" location="Rio Santo Dominigo" municipality="Rafael de Vara Blanca" pageId="6" pageNumber="336" specimenCode="CRI001468528" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="9C747AA55E66FFB18DFBB857EEB61067" box="[393,491,1580,1606]" pageId="6" pageNumber="336" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E66FFB18D80B856ED5A1066" box="[498,519,1581,1607]" pageId="6" pageNumber="336" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E66FFB18E64B857ED371067" box="[534,618,1580,1606]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 37</figureCitation>
). 
<collectingCountry id="3BD884975E66FFB18E0FB857EC7C1067" box="[637,801,1580,1606]" name="Costa Rica" pageId="6" pageNumber="336">COSTA RICA</collectingCountry>
, 
<collectingRegion id="810B0AE55E66FFB18F5EB857ECD51067" box="[812,904,1580,1606]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="336">Heredia</collectingRegion>
, 
<collectorName id="EE3AA1D15E66FFB18FE0B857EBF91067" box="[914,1188,1580,1607]" pageId="6" pageNumber="336">
S 
<collectingMunicipality id="A3145E7D5E66FFB18FDAB857EBF91067" box="[936,1188,1580,1607]" pageId="6" pageNumber="336">Rafael de Vara Blanca</collectingMunicipality>
</collectorName>
, 
<location id="461092DC5E66FFB188DCB857EAC51067" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E66FFB188DCB857EAC51067" box="[1198,1432,1580,1606]" country="Costa Rica" municipality="Rafael de Vara Blanca" name="Rio Santo Dominigo" pageId="6" pageNumber="336" stateProvince="Heredia">Rio Santo Dominigo</location>
, 18-26/VII/ 
<specimenCount id="55C90F8E5E66FFB18D7DB82BEE17104A" box="[271,330,1616,1643]" pageId="6" pageNumber="336" type="generic">93 L.</specimenCount>
<collectorName id="EE3AA1D15E66FFB18D23B82AEEE1104B" box="[337,444,1616,1643]" pageId="6" pageNumber="336">W. Quate</collectorName>
, 
<quantity id="843769E25E66FFB18DB5B82BED4A104B" box="[455,535,1616,1642]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="336" unit="m" value="1700.0">
<elevation id="C8E223345E66FFB18DB5B82BED4A104B" box="[455,535,1616,1642]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="336" unit="m" value="1700.0">1700m</elevation>
</quantity>
, 
<collectorName id="EE3AA1D15E66FFB18E51B82AED31104A" box="[547,620,1617,1643]" pageId="6" pageNumber="336">Lt Trp</collectorName>
, 
<collectorName id="EE3AA1D15E66FFB18E0AB82BED8A104B" box="[632,727,1616,1642]" pageId="6" pageNumber="336">Euparal.</collectorName>
(
<collectionCode id="25DE5CC25E66FFB18E97B82BEC68104B" box="[741,821,1616,1642]" pageId="6" pageNumber="336">INBIO</collectionCode>
<specimenCode id="13696C7C5E66FFB18F49B82BECB3104A" box="[827,1006,1616,1643]" collectionCode="INBIO" pageId="6" pageNumber="336">CRI001468528</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB18CB5B80FECF11093" pageId="6" pageNumber="336" type="diagnosis">
<paragraph id="4370C4075E66FFB18CB5B80FECF11093" blockId="6.[151,1437,1580,2039]" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5B80FEE1F10AF" bold="true" box="[199,322,1652,1678]" pageId="6" pageNumber="336">Diagnosis.</emphasis>
Frontoclypeal suture absent; medial fork complete; aedeagus with bifid and straight apex; parameres broad at the base, not extending posteriorly; parameral sheath bilobed.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E66FFB08CB5B8C7EEFF1760" lastPageId="7" lastPageNumber="337" pageId="6" pageNumber="336" type="description">
<paragraph id="4370C4075E66FFB08CB5B8C7EEBC173C" blockId="6.[151,1437,1580,2039]" lastBlockId="7.[151,1436,151,321]" lastPageId="7" lastPageNumber="337" pageId="6" pageNumber="336">
<emphasis id="71BB18155E66FFB18CB5B8C7EE9C10F7" bold="true" box="[199,449,1724,1750]" pageId="6" pageNumber="336">Redescription. Male.</emphasis>
Head (
<figureCitation id="DBF4D8825E66FFB18E67B8C7ED3510F7" box="[533,616,1724,1750]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 31</figureCitation>
) as in genus description; eye bridge with dorsal medial spur and five to six supraocular setae; group of setae on frons not extending medially to the base of eye bridge; frontoclipeal suture absent; antenna (
<figureCitation id="DBF4D8825E66FFB18D24B97FEEF5113F" box="[342,424,1796,1822]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 32</figureCitation>
) as in genus description; clypeus with four prominent lateral alveoli and slightly concave apical margin; labella with robust spiniform setulae on the inner margin; palpus formula 1.0:1.4:1.2:0.9. Wing (
<figureCitation id="DBF4D8825E66FFB1891DB953EFE91147" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 33</figureCitation>
) as in genus description; radial fork complete, distal to complete medial fork; R 
<subScript id="DF4BC6425E66FFB18837B925EB03114D" attach="left" box="[1093,1118,1886,1900]" fontSize="6" pageId="6" pageNumber="336">2+3</subScript>
1.5x the length of R 
<subScript id="DF4BC6425E66FFB18939B925EA0E114D" attach="left" box="[1355,1363,1886,1900]" fontSize="6" pageId="6" pageNumber="336">2</subScript>
. Male terminalia (
<figureCitation id="DBF4D8825E66FFB18D6AB90BEEC311AB" box="[280,414,1904,1930]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Figs. 34–36</figureCitation>
) with hypandrium reduced to a narrow band wider medially, poorly sclerotized; gonocoxite with setae on the posteromedial dorsal margin, lined up near the base; gonostylus as in genus description; aedeagus (
<figureCitation id="DBF4D8825E66FFB18CEDB9C3EFAE11F3" box="[159,243,1976,2002]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 34</figureCitation>
) with bifid and straight apex; parameres (
<figureCitation id="DBF4D8825E66FFB18EABB9C3EC7011F2" box="[729,813,1976,2003]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="6" pageNumber="336">Fig. 35</figureCitation>
) united to the gonocoxal apodemes by a membranous area; parameres broad at the base, not extending posteriorly, projecting dorsally to the aedeagus to form a bilobed parameral sheath (
<figureCitation id="DBF4D8825E67FFB08D18BEECEE9C1690" box="[362,449,151,177]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="7" pageNumber="337">Fig. 34</figureCitation>
); anterior gonocoxal apodemes without setae; epandrium and appendages (
<figureCitation id="DBF4D8825E67FFB0896DBEECEA2E1690" box="[1311,1395,151,177]" captionStart="FIGURES 31–37" captionStartId="11.[152,270,1936,1960]" captionTargetBox="[160,1426,196,1902]" captionTargetId="figure-26@11.[151,1437,181,1912]" captionTargetPageId="11" captionText="FIGURES 31–37. Wagneromyiamajorina. 31.Head.32.Terminal flagellomeres.33. Wing.34. Gonopods andaedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="7" pageNumber="337">Fig. 36</figureCitation>
) as in genus description; epandrial appendages with a group of three subapical papillae on the dorsal margin and with three apical tenacula longer than half the length of the appendices; epiproct inconspicuous and hypoproct slightly pronounced, wider than long.
</paragraph>
<paragraph id="4370C4075E67FFB08CB5BF5CEEFF1760" blockId="7.[151,1436,151,321]" box="[199,418,295,321]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5BF5CEE781760" bold="true" box="[199,293,295,321]" pageId="7" pageNumber="337">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5BFE8EDB4178C" box="[151,745,402,429]" pageId="7" pageNumber="337" type="nomenclature">
<paragraph id="4370C4075E67FFB08CE5BFE8EDB4178C" blockId="7.[151,745,402,465]" box="[151,745,402,429]" pageId="7" pageNumber="337">
<heading id="1838736B5E67FFB08CE5BFE8EDB4178C" bold="true" box="[151,745,402,429]" fontSize="11" level="1" pageId="7" pageNumber="337" reason="1">
<emphasis id="71BB18155E67FFB08CE5BFE8EDB4178C" bold="true" box="[151,745,402,429]" pageId="7" pageNumber="337">
<taxonomicName id="84CFBF845E67FFB08CE5BFE8ED06178D" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,603,402,429]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="tripetala" status="comb. nov.">
<emphasis id="71BB18155E67FFB08CE5BFE8EEF6178C" bold="true" box="[151,427,402,429]" italics="true" pageId="7" pageNumber="337">Wagneromyia tripetala</emphasis>
(
<bibRefCitation id="275EB9F65E67FFB08DC8BFE9ED0E178D" author="Quate, L. W." box="[442,595,402,428]" pageId="7" pageNumber="337" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E67FFB08E11BFE8EDB4178C" box="[611,745,403,429]" pageId="7" pageNumber="337" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5BFCCEE7A17F0" box="[151,295,439,465]" pageId="7" pageNumber="337" type="description">
<paragraph id="4370C4075E67FFB08CE5BFCCEE7A17F0" blockId="7.[151,745,402,465]" box="[151,295,439,465]" pageId="7" pageNumber="337">
(
<figureCitation id="DBF4D8825E67FFB08CEDBFCCEE4217F0" box="[159,287,439,465]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Figs 38–44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5BF80ED6E1432" box="[151,563,507,531]" pageId="7" pageNumber="337" type="reference_group">
<paragraph id="4370C4075E67FFB08CE5BF80ED6E1432" blockId="7.[151,563,507,531]" box="[151,563,507,531]" pageId="7" pageNumber="337">
<treatmentCitationGroup id="63DFE3295E67FFB08CE5BF80ED6E1432" box="[151,563,507,531]" pageId="7" pageNumber="337">
<treatmentCitation id="C26EE2165E67FFB08CE5BF80ED731432" author="Quate, L. W." box="[151,558,507,531]" page="60" pageId="7" pageNumber="337" year="1996">
<taxonomicName id="84CFBF845E67FFB08CE5BF80ED731432" ID-CoL="4GD4W" authority="Quate, 1996: 60" authorityName="Quate" authorityPageNumber="60" authorityYear="1996" box="[151,558,507,531]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="tripetalis">
<emphasis id="71BB18155E67FFB08CE5BF80EEDE1432" box="[151,387,507,531]" italics="true" pageId="7" pageNumber="337">Philosepedon tripetalis</emphasis>
<bibRefCitation id="275EB9F65E67FFB08DFBBF80ED731432" author="Quate, L. W." box="[393,558,507,531]" pageId="7" pageNumber="337" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5BC3AEC8D14A1" pageId="7" pageNumber="337" type="materials_examined">
<paragraph id="4370C4075E67FFB08CE5BC3AEC8D14A1" blockId="7.[151,1437,577,1431]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CE5BC3AEED9147A" bold="true" box="[151,388,577,603]" pageId="7" pageNumber="337">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E67FFB08DFFBC3AEC9014A1" ID-GBIF-Occurrence="3912952305" collectionCode="INBIO" collectorName="L. W. Quate" country="Costa Rica" elevation="1800" location="Vara Blanca" pageId="7" pageNumber="337" specimenCode="CRI001469454" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="9C747AA55E67FFB08DFFBC3AEEB2147A" box="[397,495,577,603]" pageId="7" pageNumber="337" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E67FFB08D8ABC39ED50147D" box="[504,525,578,604]" pageId="7" pageNumber="337" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E67FFB08E6CBC3AED29147A" box="[542,628,577,603]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 44</figureCitation>
). 
<collectingCountry id="3BD884975E67FFB08EFBBC3AEC72147A" box="[649,815,577,603]" name="Costa Rica" pageId="7" pageNumber="337">COSTA RICA</collectingCountry>
, 
<collectingRegion id="810B0AE55E67FFB08F4EBC3AECC8147A" box="[828,917,577,603]" country="Costa Rica" name="Heredia" pageId="7" pageNumber="337">Heredia</collectingRegion>
<location id="461092DC5E67FFB08FEFBC39EB77147A" LSID="urn:lsid:plazi:treatment:CB6675115E61FFB98CE5BFA0EAC1100D:461092DC5E67FFB08FEFBC39EB77147A" box="[925,1066,577,604]" country="Costa Rica" name="Vara Blanca" pageId="7" pageNumber="337" stateProvince="Heredia">Vara Blanca</location>
, 
<quantity id="843769E25E67FFB08844BC3AEBDB147A" box="[1078,1158,577,603]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="337" unit="m" value="1800.0">
<elevation id="C8E223345E67FFB08844BC3AEBDB147A" box="[1078,1158,577,603]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="337" unit="m" value="1800.0">1800m</elevation>
</quantity>
, 26/VII/ 
<specimenCount id="55C90F8E5E67FFB08895BC3AEA7E147D" box="[1255,1315,577,604]" pageId="7" pageNumber="337" type="generic">93 L.</specimenCount>
<collectorName id="EE3AA1D15E67FFB08959BC39EAC5147A" box="[1323,1432,577,604]" pageId="7" pageNumber="337">W. Quate</collectorName>
, 
<quantity id="843769E25E67FFB08CE5BC1EEFBA145E" box="[151,231,613,639]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="337" unit="m" value="1800.0">
<elevation id="C8E223345E67FFB08CE5BC1EEFBA145E" box="[151,231,613,639]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="337" unit="m" value="1800.0">1800m</elevation>
</quantity>
, 
<collectorName id="EE3AA1D15E67FFB08C81BC1DEE6114A1" box="[243,316,614,640]" pageId="7" pageNumber="337">Lt Trp</collectorName>
, 
<collectorName id="EE3AA1D15E67FFB08D3ABC1EEE8F14A1" box="[328,466,613,640]" pageId="7" pageNumber="337">Euparal, LN</collectorName>
—239.500-519.000 (
<collectionCode id="25DE5CC25E67FFB08ECEBC1EEC51145E" box="[700,780,613,639]" pageId="7" pageNumber="337">INBIO</collectionCode>
<specimenCode id="13696C7C5E67FFB08F61BC1EEC9814A1" box="[787,965,613,640]" collectionCode="INBIO" pageId="7" pageNumber="337">CRI001469454</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CB5BCF2EEE0152E" pageId="7" pageNumber="337" type="diagnosis">
<paragraph id="4370C4075E67FFB08CB5BCF2EEE0152E" blockId="7.[151,1437,577,1431]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5BCF2EE1F1482" bold="true" box="[199,322,649,675]" pageId="7" pageNumber="337">Diagnosis.</emphasis>
Medial fork incomplete; gonocoxites fused at the base; posterior gonocoxal apodemes forming a highly sclerotized ring, dorsal to the aedeagus and the parameral sheath; parameral sheath membranous and rounded; parameres well sclerotized, broad at based tapering to a hook-shaped apex; hypoproct little pronounced, ellipsoid, wider than long.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CB5BD62EEFF13B6" pageId="7" pageNumber="337" type="description">
<paragraph id="4370C4075E67FFB08CB5BD62ED1A1352" blockId="7.[151,1437,577,1431]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5BD62EE9D1512" bold="true" box="[199,448,793,819]" pageId="7" pageNumber="337">Redescription. Male.</emphasis>
Head (
<figureCitation id="DBF4D8825E67FFB08E61BD62ED381512" box="[531,613,793,819]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 38</figureCitation>
) as in genus description; eye bridge without medial dorsal spur and with seven supraocular setae; fourth row of facet nearly separated; group of setae on frons not extending medially toward base of eye bridge; frontoclipeal suture only as a slightly less sclerotized area; clypeus with nine prominent lateral alveoli and slightly concave apical margin; palpus formula 1.0:1.5:1.4:0.9, first segment of palpus with a group of sensilla in the medial region; labella with four robust spiniform setulae on the inner margin, and six setae on the external lateral margin; antenna (
<figureCitation id="DBF4D8825E67FFB08E75BDB6ED0A15C6" box="[519,599,973,999]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 39</figureCitation>
) as in genus description. Wing (
<figureCitation id="DBF4D8825E67FFB08FCEBDB6EB5315C6" box="[956,1038,973,999]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 40</figureCitation>
) as in genus description; radial fork complete, distal to incomplete medial fork; R 
<subScript id="DF4BC6425E67FFB08EF4BA7FEDC21233" attach="left" box="[646,671,1028,1042]" fontSize="6" pageId="7" pageNumber="337">2+3</subScript>
1.4x the length of R 
<subScript id="DF4BC6425E67FFB08F0EBA7FECD91233" attach="left" box="[892,900,1028,1042]" fontSize="6" pageId="7" pageNumber="337">2</subScript>
. Male terminalia (
<figureCitation id="DBF4D8825E67FFB0883FBD8AEB8F122A" box="[1101,1234,1009,1035]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Figs. 41–43</figureCitation>
) with hypandrium as a narrow band, a little wider medially; gonocoxites (
<figureCitation id="DBF4D8825E67FFB08F7ABA6EEC07120E" box="[776,858,1045,1071]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 41</figureCitation>
) fused at the base, with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus narrow, tapering to a single apex; posterior gonocoxal apodeme fused medially, forming a highly sclerotized ring—parameral sclerite, dorsal to the aedeagus and the parameral sheath (
<figureCitation id="DBF4D8825E67FFB08E06BAFAED9512BA" box="[628,712,1153,1179]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 41</figureCitation>
); parameres continuous with the posterior gonocoxal apodemes; parameres (
<figureCitation id="DBF4D8825E67FFB08D69BADEEEFE129E" box="[283,419,1189,1215]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Figs. 41, 42</figureCitation>
) well sclerotized, broad at based tapering to a hook-shaped apex; membranous parameral sheath with a rounded apex (
<figureCitation id="DBF4D8825E67FFB08D9EBAB2ED1912C2" box="[492,580,1225,1251]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 41</figureCitation>
); anterior gonocoxal apodemes (
<figureCitation id="DBF4D8825E67FFB08FB3BAB2EB4A12C2" box="[961,1047,1225,1251]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 42</figureCitation>
) with setae on the posteromedial margin, continuous with the ventral setae of the gonocoxite; epandrium (
<figureCitation id="DBF4D8825E67FFB08FB7BA96EB4B1326" box="[965,1046,1261,1287]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 43</figureCitation>
) as in genus description; epandrial appendages as in genus description with a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct inconspicuous and hypoproct (
<figureCitation id="DBF4D8825E67FFB08837BB4EEBC4136E" box="[1093,1177,1333,1359]" captionStart="FIGURES 38–44" captionStartId="12.[152,270,1907,1931]" captionTargetBox="[197,1401,191,1875]" captionTargetId="figure-18@12.[181,1405,181,1883]" captionTargetPageId="12" captionText="FIGURES 38–44. Wagneromyia tripetala. 38.Head. 39.Terminal flagellomeres.40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="7" pageNumber="337">Fig. 43</figureCitation>
) not very pronounced, ellipsoid, wider than long, both pilose.
</paragraph>
<paragraph id="4370C4075E67FFB08CB5BB06EEFF13B6" blockId="7.[151,1437,577,1431]" box="[199,418,1405,1431]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5BB06EE7813B6" bold="true" box="[199,293,1405,1431]" pageId="7" pageNumber="337">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5BB92EC551022" box="[151,776,1513,1540]" pageId="7" pageNumber="337" type="nomenclature">
<paragraph id="4370C4075E67FFB08CE5BB92EC551022" blockId="7.[151,776,1513,1576]" box="[151,776,1513,1540]" pageId="7" pageNumber="337">
<heading id="1838736B5E67FFB08CE5BB92EC551022" bold="true" box="[151,776,1513,1540]" fontSize="11" level="1" pageId="7" pageNumber="337" reason="1">
<emphasis id="71BB18155E67FFB08CE5BB92EC551022" bold="true" box="[151,776,1513,1540]" pageId="7" pageNumber="337">
<taxonomicName id="84CFBF845E67FFB08CE5BB92ED271022" ID-CoL="9J66P" authority="(Quate, 1996)" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[151,634,1513,1540]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="tritenacula" status="comb. nov.">
<emphasis id="71BB18155E67FFB08CE5BB92EE971025" bold="true" box="[151,458,1513,1540]" italics="true" pageId="7" pageNumber="337">Wagneromyia tritenacula</emphasis>
(
<bibRefCitation id="275EB9F65E67FFB08DABBB92ED2F1022" author="Quate, L. W." box="[473,626,1513,1539]" pageId="7" pageNumber="337" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E67FFB08EF0BB92EC551022" box="[642,776,1513,1539]" pageId="7" pageNumber="337" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5B876EE7A1009" box="[151,295,1549,1576]" pageId="7" pageNumber="337" type="description">
<paragraph id="4370C4075E67FFB08CE5B876EE7A1009" blockId="7.[151,776,1513,1576]" box="[151,295,1549,1576]" pageId="7" pageNumber="337">
(
<figureCitation id="DBF4D8825E67FFB08CEDB876EE421009" box="[159,287,1549,1576]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="7" pageNumber="337">Figs 45–51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5B82EED261051" box="[151,635,1621,1648]" pageId="7" pageNumber="337" type="reference_group">
<paragraph id="4370C4075E67FFB08CE5B82EED261051" blockId="7.[151,635,1621,1648]" box="[151,635,1621,1648]" pageId="7" pageNumber="337">
<treatmentCitationGroup id="63DFE3295E67FFB08CE5B82EED261051" box="[151,635,1621,1648]" pageId="7" pageNumber="337">
<taxonomicName id="84CFBF845E67FFB08CE5B82EED2A1051" ID-CoL="89WTS" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[151,631,1621,1648]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="tritenaculus">
<emphasis id="71BB18155E67FFB08CE5B82EEE72104E" box="[151,303,1621,1647]" italics="true" pageId="7" pageNumber="337">Philosepedon</emphasis>
tritenaculus 
<treatmentCitation id="C26EE2165E67FFB08DB2B82EED2A1051" author="Quate, L. W." box="[448,631,1621,1648]" page="58" pageId="7" pageNumber="337" year="1996">
<bibRefCitation id="275EB9F65E67FFB08DB2B82EED2A1051" author="Quate, L. W." box="[448,631,1621,1648]" pageId="7" pageNumber="337" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate, 1996: 58</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CE5B8E6EC9010FD" pageId="7" pageNumber="337" type="materials_examined">
<paragraph id="4370C4075E67FFB08CE5B8E6EC9010FD" blockId="7.[151,1437,1693,2008]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CE5B8E6EED91096" bold="true" box="[151,388,1693,1719]" pageId="7" pageNumber="337">Material examined:</emphasis>
<materialsCitation id="F3A7CE5A5E67FFB08DF9B8E6EC9710FD" ID-GBIF-Occurrence="3912952307" collectionCode="INBIO" collectorName="L. W. Quate" country="Costa Rica" elevation="700" location="COSTA RICA" pageId="7" pageNumber="337" specimenCode="CRI001469215" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9C747AA55E67FFB08DF9B8E6EEB01096" box="[395,493,1693,1719]" pageId="7" pageNumber="337" type="holotype">holotype</typeStatus>
<specimenCount id="55C90F8E5E67FFB08D87B8E5ED571099" box="[501,522,1694,1720]" pageId="7" pageNumber="337" type="male">♂</specimenCount>
(
<figureCitation id="DBF4D8825E67FFB08E68B8E6ED331099" box="[538,622,1693,1720]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="7" pageNumber="337">Fig. 51</figureCitation>
). 
<collectingCountry id="3BD884975E67FFB08EF1B8E6EC741096" box="[643,809,1693,1719]" name="Costa Rica" pageId="7" pageNumber="337">COSTA RICA</collectingCountry>
, Guanac. Estacion Pitilla, 
<quantity id="843769E25E67FFB08821B8E6EBCB1096" box="[1107,1174,1693,1719]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="337" unit="m" value="700.0">
<specimenCount id="55C90F8E5E67FFB08821B8E6EBCB1096" box="[1107,1174,1693,1719]" pageId="7" pageNumber="337" type="generic">
<elevation id="C8E223345E67FFB08821B8E6EBCB1096" box="[1107,1174,1693,1719]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="337" unit="m" value="700.0">700m</elevation>
</specimenCount>
</quantity>
. 9 
<collectorName id="EE3AA1D15E67FFB088C5B8E5EAC51096" box="[1207,1432,1693,1720]" pageId="7" pageNumber="337">Km S Santa Cecilia</collectorName>
, 10-14/VII/93, 
<collectorName id="EE3AA1D15E67FFB08D4BB8B9EE9E10FA" box="[313,451,1729,1756]" pageId="7" pageNumber="337">L. W. Quate</collectorName>
, 
<collectingMethod id="9A8EBC105E67FFB08DBCB8BAED1810FD" box="[462,581,1729,1756]" pageId="7" pageNumber="337">Light Trap</collectingMethod>
<collectorName id="EE3AA1D15E67FFB08E3EB8BAEDF610FA" box="[588,683,1729,1755]" pageId="7" pageNumber="337">Euparal.</collectorName>
(
<collectionCode id="25DE5CC25E67FFB08ECBB8BAEC5410FA" box="[697,777,1729,1755]" pageId="7" pageNumber="337">INBIO</collectionCode>
<specimenCode id="13696C7C5E67FFB08F7DB8BAEC9C10FD" box="[783,961,1729,1756]" collectionCode="INBIO" pageId="7" pageNumber="337">CRI001469215</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB08CB5B89EEA321166" pageId="7" pageNumber="337" type="diagnosis">
<paragraph id="4370C4075E67FFB08CB5B89EEA321166" blockId="7.[151,1437,1693,2008]" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5B89EEE1F10DE" bold="true" box="[199,322,1765,1791]" pageId="7" pageNumber="337">Diagnosis.</emphasis>
14th flagellomere with three apical stout and spiniform setulae; medial fork complete; aedeagus with lateral folds subapically, forming an abrupt bottleneck anterior to a bifurcated apex; parameres broad at the base, not extending posteriorly; parameral sheath trapezoidal; gonocoxal apodemes with four setae posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E67FFB98CB5B92AEEFF1438" lastPageId="14" lastPageNumber="344" pageId="7" pageNumber="337" type="description">
<paragraph id="4370C4075E67FFB98CB5B92AEA3217D4" blockId="7.[151,1437,1693,2008]" lastBlockId="14.[151,1436,150,537]" lastPageId="14" lastPageNumber="344" pageId="7" pageNumber="337">
<emphasis id="71BB18155E67FFB08CB5B92AEE9D114A" bold="true" box="[199,448,1873,1899]" pageId="7" pageNumber="337">Redescription. Male.</emphasis>
Head (
<figureCitation id="DBF4D8825E67FFB08E61B92AED38114D" box="[531,613,1873,1900]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="7" pageNumber="337">Fig. 45</figureCitation>
) as in genus description; eye bridge without medial dorsal spur and with five to six supraocular setae; group of setae on frons extending medially near the inferior limit of the eye bridge; frontoclipeal suture only as a slightly less sclerotized area; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin and three setae on the external lateral margin; antenna (
<figureCitation id="DBF4D8825E6EFFB98DD6BEECEEA91690" box="[420,500,151,177]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 46</figureCitation>
) as in genus description; palpus formula 1.0:1.5:1.2:0.8, first segment of palpus with a group of sensilla near the base. Wing (
<figureCitation id="DBF4D8825E6EFFB98E21BEC0EDF916F4" box="[595,676,187,213]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 47</figureCitation>
) as in genus description; radial fork complete, distal to the complete medial fork; R 
<subScript id="DF4BC6425E6EFFB98D4EBE8AEE0816DE" attach="left" box="[316,341,241,255]" fontSize="6" pageId="14" pageNumber="344">2+3</subScript>
1.4x the length of R 
<subScript id="DF4BC6425E6EFFB98E4CBE8AED1B16DE" attach="left" box="[574,582,241,255]" fontSize="6" pageId="14" pageNumber="344">2</subScript>
. Male terminalia (
<figureCitation id="DBF4D8825E6EFFB98F65BEA4ECC316D8" box="[791,926,222,249]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Figs. 48–50</figureCitation>
) with hypandrium reduced to a narrow band; gonocoxite with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus (
<figureCitation id="DBF4D8825E6EFFB98D7FBF5CEECF1760" box="[269,402,295,321]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Figs. 48, 49</figureCitation>
) with subapical lateral folds, forming a bottleneck anterior to the bifurcated apex; parameres united to the gonocoxal apodemes by a membranous area; parameres (
<figureCitation id="DBF4D8825E6EFFB98FB7BF30EB441744" box="[965,1049,331,357]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 49</figureCitation>
) broad at the base, not extending posteriorly, projecting dorsally to the aedeagus to form a trapezoidal parameral sheath (
<figureCitation id="DBF4D8825E6EFFB98825BF14EBF717A8" box="[1111,1194,367,393]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 48</figureCitation>
); gonocoxal apodema with four setae posteriorly; epandrium (
<figureCitation id="DBF4D8825E6EFFB98E23BFE8EDFC178C" box="[593,673,402,429]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 50</figureCitation>
) as in genus description; epandrial appendages with a group of short setulae on the base, a group of three subapical papillae on the dorsal margin and three apical tenacula of half the length of the appendices; epiproct inconspicuous and hypoproct (
<figureCitation id="DBF4D8825E6EFFB98F03BFA0EC9E17D4" box="[881,963,474,501]" captionStart="FIGURES 45–51" captionStartId="13.[152,270,1896,1920]" captionTargetBox="[172,1419,162,1865]" captionTargetId="figure-26@13.[159,1428,151,1873]" captionTargetPageId="13" captionText="FIGURES 45–51. Wagneromyia tritenacula. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath." figureDoi="http://doi.org/10.5281/zenodo.7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="14" pageNumber="344">Fig. 50</figureCitation>
) slightly pronounced, wider than long
</paragraph>
<caption id="17B0948F5E68FFBF8CEAB9ECEB4011D6" ID-DOI="http://doi.org/10.5281/zenodo.7095484" ID-Zenodo-Dep="7095484" httpUri="https://zenodo.org/record/7095484/files/figure.png" pageId="8" pageNumber="338" startId="8.[152,270,1943,1967]" targetBox="[179,1400,202,1906]" targetPageId="8">
<paragraph id="4370C4075E68FFBF8CEAB9ECEB4011D6" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="338">
<emphasis id="71BB18155E68FFBF8CEAB9ECEE05118E" bold="true" box="[152,344,1943,1967]" pageId="8" pageNumber="338">FIGURES 10–16.</emphasis>
<taxonomicName id="84CFBF845E68FFBF8D2EB9E3ED3D1191" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[348,608,1944,1968]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="bicalcarata">
<emphasis id="71BB18155E68FFBF8D2EB9E3ED3D1191" box="[348,608,1944,1968]" italics="true" pageId="8" pageNumber="338">Wagneromyia bicalcarata</emphasis>
</taxonomicName>
. 10. Head. 11. Terminal flagellomeres. 12. Wing. 13. Male terminalia (dorsal view). 14. Male terminalia (ventral view). 15. Epandrium and epandrial appendages (lateral view). 16. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere.
</paragraph>
</caption>
<caption id="17B0948F5E69FFBE8CEAB927ECD511C1" ID-DOI="http://doi.org/10.5281/zenodo.7095488" ID-Zenodo-Dep="7095488" httpUri="https://zenodo.org/record/7095488/files/figure.png" pageId="9" pageNumber="339" startId="9.[152,270,1884,1908]" targetBox="[187,1386,194,1851]" targetPageId="9">
<paragraph id="4370C4075E69FFBE8CEAB927ECD511C1" blockId="9.[151,1437,1884,2016]" pageId="9" pageNumber="339">
<emphasis id="71BB18155E69FFBE8CEAB927EE031155" bold="true" box="[152,350,1884,1908]" pageId="9" pageNumber="339">FIGURES 17–24.</emphasis>
<taxonomicName id="84CFBF845E69FFBE8D1AB926ED011155" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[360,604,1884,1908]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="339" phylum="Arthropoda" rank="species" species="deceptrix">
<emphasis id="71BB18155E69FFBE8D1AB926ED011155" box="[360,604,1884,1908]" italics="true" pageId="9" pageNumber="339">Wagneromyia deceptrix</emphasis>
</taxonomicName>
. 17. Head. 18. Wing. 19. Terminal flagellomeres. 20. Gonopods and aedeagal complex (dorsal view). 21. Gonopods and aedeagal complex (ventral view). 22. Epandrium (ventral view). 23. Epandrial appendages (lateral view). 24. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm sh—parameral sheath.
</paragraph>
</caption>
<caption id="17B0948F5E6AFFBD8CEAB9ECEA4211D9" ID-DOI="http://doi.org/10.5281/zenodo.7095492" ID-Zenodo-Dep="7095492" httpUri="https://zenodo.org/record/7095492/files/figure.png" pageId="10" pageNumber="340" startId="10.[152,270,1943,1967]" targetBox="[170,1415,199,1911]" targetPageId="10">
<paragraph id="4370C4075E6AFFBD8CEAB9ECEA4211D9" blockId="10.[151,1437,1943,2040]" pageId="10" pageNumber="340">
<emphasis id="71BB18155E6AFFBD8CEAB9ECEE03118E" bold="true" box="[152,350,1943,1967]" pageId="10" pageNumber="340">FIGURES 25–30.</emphasis>
<emphasis id="71BB18155E6AFFBD8D1AB9E3ED3A1191" box="[360,615,1944,1968]" italics="true" pageId="10" pageNumber="340">
<taxonomicName id="84CFBF845E6AFFBD8D1AB9E3ED3E1191" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[360,611,1944,1968]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="340" phylum="Arthropoda" rank="species" species="dimorpha">Wagneromyia dimorpha</taxonomicName>
.
</emphasis>
25. Head. 26. Wing. 27. Gonopods and aedeagal complex (dorsal view). 28. Gonopods and aedeagal complex (ventral view). 29. Epandrium and epandrial appendages. 30. Holotype labels. Abbreviations: aed—aedeagus; epiprct—epiproct; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath.
</paragraph>
</caption>
<caption id="17B0948F5E6BFFBC8CEAB9EBEA1F11D1" ID-DOI="http://doi.org/10.5281/zenodo.7095496" ID-Zenodo-Dep="7095496" httpUri="https://zenodo.org/record/7095496/files/figure.png" pageId="11" pageNumber="341" startId="11.[152,270,1936,1960]" targetBox="[160,1426,196,1902]" targetPageId="11">
<paragraph id="4370C4075E6BFFBC8CEAB9EBEA1F11D1" blockId="11.[151,1436,1936,2033]" pageId="11" pageNumber="341">
<emphasis id="71BB18155E6BFFBC8CEAB9EBEE051189" bold="true" box="[152,344,1936,1960]" pageId="11" pageNumber="341">FIGURES 31–37.</emphasis>
<taxonomicName id="84CFBF845E6BFFBC8D2EB9EAED1A1189" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[348,583,1937,1960]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="341" phylum="Arthropoda" rank="species" species="majorina">
<emphasis id="71BB18155E6BFFBC8D2EB9EAED1A1189" box="[348,583,1937,1960]" italics="true" pageId="11" pageNumber="341">Wagneromyia majorina</emphasis>
</taxonomicName>
. 31. Head. 32. Terminal flagellomeres. 33. Wing. 34. Gonopods and aedeagal complex (dorsal view). 35. Gonopods and aedeagal complex (ventral view). 36. Epandrium and epandrial appendages. 37. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; pm—paramere; pm sh—parameral sheath.
</paragraph>
</caption>
<caption id="17B0948F5E6CFFBB8CEAB908EDC411D6" ID-DOI="http://doi.org/10.5281/zenodo.7095500" ID-Zenodo-Dep="7095500" httpUri="https://zenodo.org/record/7095500/files/figure.png" pageId="12" pageNumber="342" startId="12.[152,270,1907,1931]" targetBox="[197,1401,191,1875]" targetPageId="12">
<paragraph id="4370C4075E6CFFBB8CEAB908EDC411D6" blockId="12.[151,1437,1907,2039]" pageId="12" pageNumber="342">
<emphasis id="71BB18155E6CFFBB8CEAB908EE0511AA" bold="true" box="[152,344,1907,1931]" pageId="12" pageNumber="342">FIGURES 38–44.</emphasis>
<taxonomicName id="84CFBF845E6CFFBB8D2EB90FED1E11AA" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[348,579,1907,1931]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="342" phylum="Arthropoda" rank="species" species="tripetala">
<emphasis id="71BB18155E6CFFBB8D2EB90FED1E11AA" box="[348,579,1907,1931]" italics="true" pageId="12" pageNumber="342">Wagneromyia tripetala</emphasis>
</taxonomicName>
. 38. Head. 39. Terminal flagellomeres. 40. Wing. 41. Gonopods and aedeagal complex (dorsal view). 42. Gonopods and aedeagal complex (ventral view). 43. Epandrium and epandrial appendages (ventral view). 44. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm—paramere; pm scl—parameral sclerite; pm sh—parameral sheath.
</paragraph>
</caption>
<caption id="17B0948F5E6DFFBA8CEAB913ED5011CD" ID-DOI="http://doi.org/10.5281/zenodo.7095502" ID-Zenodo-Dep="7095502" httpUri="https://zenodo.org/record/7095502/files/figure.png" pageId="13" pageNumber="343" startId="13.[152,270,1896,1920]" targetBox="[172,1419,162,1865]" targetPageId="13">
<paragraph id="4370C4075E6DFFBA8CEAB913ED5011CD" blockId="13.[151,1437,1896,2028]" pageId="13" pageNumber="343">
<emphasis id="71BB18155E6DFFBA8CEAB913EE0011A1" bold="true" box="[152,349,1896,1920]" pageId="13" pageNumber="343">FIGURES 45–51.</emphasis>
<taxonomicName id="84CFBF845E6DFFBA8D14B912ED3411A1" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[358,617,1896,1920]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="343" phylum="Arthropoda" rank="species" species="tritenacula">
<emphasis id="71BB18155E6DFFBA8D14B912ED3411A1" box="[358,617,1896,1920]" italics="true" pageId="13" pageNumber="343">Wagneromyia tritenacula</emphasis>
</taxonomicName>
. 45. Head. 46. Terminal flagellomeres. 47. Wing. 48. Gonopods and aedeagal complex (dorsal view). 49. Gonopods and aedeagal complex (ventral view). 50. Epandrium and epandrial appendages (dorsal view). 51. Holotype labels. Abbreviations: aed—aedeagus; goncx apod—gonocoxal apodemes; hyprct—hypoproct; pm— paramere; pm sh—parameral sheath.
</paragraph>
</caption>
<paragraph id="4370C4075E6EFFB98CB5BF84EEFF1438" blockId="14.[151,1436,150,537]" box="[199,418,511,537]" pageId="14" pageNumber="344">
<emphasis id="71BB18155E6EFFB98CB5BF84EE781438" bold="true" box="[199,293,511,537]" pageId="14" pageNumber="344">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E6EFFB98CE5BC11EBBB14D0" pageId="14" pageNumber="344" type="discussion">
<paragraph id="4370C4075E6EFFB98CE5BC11EC4614A5" blockId="14.[151,795,618,645]" box="[151,795,618,645]" pageId="14" pageNumber="344">
<heading id="1838736B5E6EFFB98CE5BC11EC4614A5" bold="true" box="[151,795,618,645]" fontSize="11" level="1" pageId="14" pageNumber="344" reason="1">
<emphasis id="71BB18155E6EFFB98CE5BC11EC4614A5" bold="true" box="[151,795,618,645]" pageId="14" pageNumber="344">
Identification key for males of 
<taxonomicName id="84CFBF845E6EFFB98E64BC10EC4614A5" box="[534,795,618,645]" pageId="14" pageNumber="344">
<emphasis id="71BB18155E6EFFB98E64BC10EDE014A4" bold="true" box="[534,701,619,645]" italics="true" pageId="14" pageNumber="344">Wagneromyia</emphasis>
species
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BCC8EBBB14D0" blockId="14.[151,1437,691,753]" pageId="14" pageNumber="344">
The species of the new genus are present in the key of 
<bibRefCitation id="275EB9F65E6EFFB98F53BCC8ECE514EC" author="Quate, L. W." box="[801,952,691,717]" pageId="14" pageNumber="344" pagination="1 - 81" refId="ref9172" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de biologia tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate (1996)</bibRefCitation>
to the genus 
<taxonomicName id="84CFBF845E6EFFB98828BCC8EBAF14EC" authorityName="Eaton" authorityYear="1904" box="[1114,1266,691,717]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="71BB18155E6EFFB98828BCC8EBAF14EC" box="[1114,1266,691,717]" italics="true" pageId="14" pageNumber="344">Philosepedon</emphasis>
</taxonomicName>
. A new key is provided below to identify the new described species and the species transferred to the new genus.
</paragraph>
</subSubSection>
<subSubSection id="0BD5978C5E6EFFB98CE5BD67EAC1100D" pageId="14" pageNumber="344" type="key">
<key id="375E93365E6EFFB98CE5BD67EAC1100D" pageId="14" pageNumber="344">
<keyStep id="F83BDCA25E6EFFB98CE5BD67EAC0154D" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BD67EAC11510" blockId="14.[151,1437,796,1580]" box="[151,1436,796,818]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BD67EAC11510" box="[151,1436,796,818]" pageId="14" pageNumber="344">1 Gonostylus ending in a single and narrow apex; each paramere with a non-bifurcated apex........................... 2</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BD42EAC0154D" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BD42EAC0154D" pageId="14" pageNumber="344">
- Gonostylus ending apically bifurcated, like a chela; each paramere apically bifurcated in one external arm hook-like, dorsal to the aedeagus, and an internal arm rounded...................................................... 
<taxonomicName id="84CFBF845E6EFFB9897CBD2CEAC0154D" ID-CoL="9J67K" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1294,1437,854,876]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="bicalcarata">
<emphasis id="71BB18155E6EFFB9897CBD2CEAC0154D" bold="true" box="[1294,1437,854,876]" italics="true" pageId="14" pageNumber="344">W. bicalcarata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F83BDCA25E6EFFB98CE5BD0FEAC115C0" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BD0FEAC11587" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BD0FEAC11587" pageId="14" pageNumber="344">2 Wing with medial fork incomplete; gonocoxites with a group of setae concentrated in the dorsal posteromedial region; parameres well developed, reaching the level of the aedeagus apex...................................................... 3</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BDD4EAC115C0" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BDD4EAC115C0" pageId="14" pageNumber="344">- Wing with medial fork complete; gonocoxites with a group of setae in a row from the apex to near the base; parameres not extending posteriorly ventral to the aedeagus............................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="F83BDCA25E6EFFB98CE5BD92EAC01255" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BD92EAC1123A" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BD92EAC1123A" pageId="14" pageNumber="344">3 Presence of medial dorsal spur on the eye bridge; fourth facet row completely contiguous; gonocoxites contiguous, posterior gonocoxal apodeme not extended medially; parameral sheath wider than long; aedeagus with bifurcated apex........... 4</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BA5FEAC01255" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BA5FEAC01255" pageId="14" pageNumber="344">
- Medial dorsal spur on the eye bridge absent; fourth facet row nearly separated; gonocoxites continuous, with posterior gonocoxal apodemes extended and fused medially; parameral sheath as long and wide at the base; aedeagus with single apex................................................................................................... 
<taxonomicName id="84CFBF845E6EFFB98958BA24EAC01255" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1322,1437,1118,1140]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="tripetala">
<emphasis id="71BB18155E6EFFB98958BA24EAC01255" bold="true" box="[1322,1437,1118,1140]" italics="true" pageId="14" pageNumber="344">W. tripetala</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F83BDCA25E6EFFB98CE5BA07EAC11327" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BA07EAC1128F" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BA07EAC1128F" pageId="14" pageNumber="344">
4 Clypeus with five prominent lateral alveoli; gonostylus bent in the subapical region, bearing a small apical accessory seta and a long subapical seta on the ventral side; parameres greatly sclerotized............................ 
<emphasis id="71BB18155E6EFFB9889CBAE2EAC1128F" bold="true" box="[1262,1436,1177,1198]" pageId="14" pageNumber="344">
<taxonomicName id="84CFBF845E6EFFB9889CBAE2EA14128F" ID-CoL="9J6XN" authority="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo, 2022" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" box="[1262,1353,1177,1198]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="aquiri" status="sp. nov.">
<emphasis id="71BB18155E6EFFB9889CBAE2EA14128F" bold="true" box="[1262,1353,1177,1198]" italics="true" pageId="14" pageNumber="344">W. aquiri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6A88A56E5E6EFFB98922BAE2EAC1128F" box="[1360,1436,1177,1198]" pageId="14" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BACDEAC11327" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BACDEAC11327" pageId="14" pageNumber="344">
<emphasis id="71BB18155E6EFFB98CE5BACDEFC212EA" bold="true" box="[151,159,1206,1227]" pageId="14" pageNumber="344">-</emphasis>
Clypeus with three prominent lateral alveoli; gonostylus slightly curved through its extension, with a row of small subapical spines on the ventrolateral margin and a long subapical seta, almost at the apex; parameres with reduced sclerotization while approaching the apex........................................................................ 
<taxonomicName id="84CFBF845E6EFFB9896FBA8AEAC11327" ID-CoL="9J7FZ" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1309,1436,1264,1286]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="dimorpha">
<emphasis id="71BB18155E6EFFB9896FBA8AEAC11327" bold="true" box="[1309,1436,1264,1286]" italics="true" pageId="14" pageNumber="344">W. dimorpha</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F83BDCA25E6EFFB98CE5BB75EAC01397" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BB75EAC1137E" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BB75EAC1137E" pageId="14" pageNumber="344">5 14th flagellomere with more than one apical spiniform setula; eye bridge without medial dorsal spur; parameral sheath apex with single lobe; anterior gonocoxal apodemes with setae on the posterior region, continuous with the ventrolateral setae of the gonocoxite; length of tenacula equal to half the length of the epandrial appendages................................. 6</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BB1CEAC01397" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BB1CEAC01397" pageId="14" pageNumber="344">
- 14th flagellomere with one apical spiniform setula; eye bridge with medial dorsal spur; parameral sheath apex bilobed; gonocoxal apodemes without setae in the posterior region; tenacula longer than half the length of the epandrial appendages.......................................................................................... 
<taxonomicName id="84CFBF845E6EFFB98950BBDAEAC01397" ID-CoL="9J5VZ" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1314,1437,1441,1462]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="majorina">
<emphasis id="71BB18155E6EFFB98950BBDAEAC01397" bold="true" box="[1314,1437,1441,1462]" italics="true" pageId="14" pageNumber="344">W. majorina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F83BDCA25E6EFFB98CE5BBC5EAC1100D" pageId="14" pageNumber="344">
<paragraph id="4370C4075E6EFFB98CE5BBC5EAC013D0" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BBC5EAC013D0" pageId="14" pageNumber="344">
6 14th flagellomere with two apical spiniform setulae; group of setae on frons not extending medially; apex of parameral sheath with rounded lobe; aedeagus with margins slightly concave before apex............................... 
<taxonomicName id="84CFBF845E6EFFB98950BBA7EAC013D0" ID-CoL="9J6PH" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1314,1437,1499,1521]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="deceptrix">
<emphasis id="71BB18155E6EFFB98950BBA7EAC013D0" bold="true" box="[1314,1437,1499,1521]" italics="true" pageId="14" pageNumber="344">W. deceptrix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4370C4075E6EFFB98CE5BB82EAC1100D" blockId="14.[151,1437,796,1580]" pageId="14" pageNumber="344">
<keyLead id="F83E67325E6EFFB98CE5BB82EAC1100D" pageId="14" pageNumber="344">
- 14th flagellomere with three apical setulae; group of setae on frons extending medially near the base of eye bridge; apex of parameral sheath truncated; aedeagus with sinuous margins like a bottleneck........................... 
<taxonomicName id="84CFBF845E6EFFB98963B86CEAC1100D" ID-CoL="9J66P" authorityName="Vieira &amp; Cordeiro &amp; Bravo &amp; Araújo" authorityYear="2022" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[1297,1436,1558,1580]" class="Insecta" family="Psychodidae" genus="Wagneromyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="tritenacula">
<emphasis id="71BB18155E6EFFB98963B86CEAC1100D" bold="true" box="[1297,1436,1558,1580]" italics="true" pageId="14" pageNumber="344">W. tritenacula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>