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<document id="BB2124D9DE0DEB41597C92AD51422483" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638261545392" checkinUser="felipe" docAuthor="Kovařík, František, Yağmur, Ersen Aydin &amp; Fet, Victor" docDate="2019" docId="8F23B8627C28B914FC4EE645340E0835" docLanguage="en" docName="Euscorpius.282.1-30.pdf" docOrigin="Euscorpius 282" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Hottentotta niloticus" docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="731AC01A7C2FB91FFFE8E27A31310E5B" masterDocTitle="Review of Hottentotta described by A. A. Birula, with descriptions of two new species and comments on Birula’s collection (Scorpiones: Buthidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="6" updateTime="1699436433264" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="E652F515F9B74C0559852693974CBA5A">Review of Hottentotta described by A. A. Birula, with descriptions of two new species and comments on Birula’s collection (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="C1D9D0976D1437D4616B7779A62DA65C">Kovařík, František</mods:namePart>
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<mods:namePart id="05D4AC85A1B02F479ED285249C978DBE">Yağmur, Ersen Aydin</mods:namePart>
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<mods:namePart id="FE4B0732C447D9300F37F44509F4EE31">Fet, Victor</mods:namePart>
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<mods:title id="5F4BBDBE287036DFBDAFE085974A4EF4">Euscorpius</mods:title>
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<mods:part id="2EA97B0C708494672B1EA3BD2759C10D">
<mods:date id="9B700E3A5853B5257ADA89E0963BAB1A">2019</mods:date>
<mods:detail id="1E8D76CD83DB45691B0A1E71EC095DE2" type="volume">
<mods:number id="569EE7AE2534BA4CEECC97C3DAF64F98">282</mods:number>
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<mods:identifier id="7D890610FCCAB46B055AFABF5EEC6E1D" type="ISSN">1536-9307</mods:identifier>
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<treatment id="8F23B8627C28B914FC4EE645340E0835" ID-DOI="http://doi.org/10.5281/zenodo.5741313" ID-Zenodo-Dep="5741313" LSID="urn:lsid:plazi:treatment:8F23B8627C28B914FC4EE645340E0835" httpUri="http://treatment.plazi.org/id/8F23B8627C28B914FC4EE645340E0835" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6">
<subSubSection id="4F905AFF7C28B918FC4EE645347D0A03" box="[934,1356,1086,1113]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="073509747C28B918FC4EE645347D0A03" blockId="7.[934,1356,1086,1144]" box="[934,1356,1086,1113]" pageId="7" pageNumber="6">
<heading id="5C7DBE187C28B918FC4EE645347D0A03" box="[934,1356,1086,1113]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="4">
<taxonomicName id="C08A72F77C28B918FC4EE645347D0A03" ID-CoL="3MQXY" authority="(Birula, 1928)" baseAuthorityName="Birula" baseAuthorityYear="1928" box="[934,1356,1086,1113]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C28B918FC4EE64535AF0A02" bold="true" box="[934,1182,1086,1113]" italics="true" pageId="7" pageNumber="6">Hottentotta niloticus</emphasis>
(
<bibRefCitation id="631B74857C28B918FB46E64434750A03" author="BIRULA" box="[1198,1348,1086,1112]" pageId="7" pageNumber="6" pagination="79 - 88" refId="ref10464" refString="BIRULA, A. A. 1928. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner uternommenen Zoologischen Expedition nach dem Anglo-Agyptischen Sudan (Kordofan) 1914. XXV. Skorpione. Denkschriften der Akademie der Wissenschaften in Wien, 101: 79 - 88." type="journal article" year="1928">Birula, 1928</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4F905AFF7C28B918FBEBE61A35DE0A23" box="[1027,1263,1120,1144]" pageId="7" pageNumber="6" type="description">
<paragraph id="073509747C28B918FBEBE61A35DE0A23" blockId="7.[934,1356,1086,1144]" box="[1027,1263,1120,1144]" pageId="7" pageNumber="6">
(
<figureCitation id="9FB115F17C28B918FBE3E61A35BC0A23" box="[1035,1165,1120,1144]" captionStart-0="Figures 23–26" captionStart-1="Figures 27–32" captionStart-2="Figures 33–49" captionStartId-0="8.[113,190,1981,2003]" captionStartId-1="9.[116,193,1970,1992]" captionStartId-2="10.[115,192,1884,1906]" captionTargetBox-0="[121,1454,272,1956]" captionTargetBox-1="[122,1471,246,1948]" captionTargetBox-2="[113,1465,222,1873]" captionTargetId-0="figure-53@8.[113,1474,263,1965]" captionTargetId-1="figure-58@9.[113,1474,234,1957]" captionTargetId-2="figure-140@10.[113,1474,212,1873]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figures 23–26: Hottentotta niloticus. Figures 23–24. Lectotype male, dorsal (23) and ventral (24) views. Figures 25–26. Paralectotype female, dorsal (25) and ventral (26) views. Scale bar = 10 mm." captionText-1="Figures 27–32: Hottentotta niloticus. Figures 27–29. Paralectotype female, metasoma and telson, lateral (27), dorsal (28), and ventral (29) views. Figures 30–32. Lectotype male, metasoma and telson, lateral (30), dorsal (31), and ventral (32) views. Scale bars = 10 mm." captionText-2="Figures 33–49: Hottentotta niloticus. Figures 33–43: Pedipalp segments. Figures 33–34. Paralectotype female, pedipalp chela dorsal (33) and external (34) views. Figures 35–43. Lectotype male, pedipalp chela dorsal (35), external (36) and vental (37) views, patella dorsal (38), external (39) and ventral (40) views, femur and trochanter dorsal (41) and ventral (42), movable finger (43). The trichobothrial pattern is indicated in Figures 35–39, 41. Figures 44, 46, 48–49. Lectotype male, chelicerae, carapace and tergites I–III (44), sternopectinal region and sternites III–IV (46), and distal segments of legs III–IV (48–49), retrolateral view. Figures 45, 47. Paralectotype female, chelicerae, carapace and tergites I–III (45), sternopectinal region and sternite III (47)." pageId="7" pageNumber="6">Figs. 23–49</figureCitation>
,
<tableCitation id="4A083CCF7C28B918FB7FE61A35D40A23" box="[1175,1253,1120,1144]" captionStart="Table 1" captionStartId="7.[116,172,997,1018]" captionText="Table 1. Comparative measurements of Hottentotta buchariensis, H. niloticus and H. penJabensis types.Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4F905AFF7C28B918FCD8E6DA35FD0BC3" pageId="7" pageNumber="6" type="reference_group">
<paragraph id="073509747C28B918FCD8E6DA34420AE3" blockId="7.[816,1475,1184,1336]" box="[816,1395,1184,1208]" pageId="7" pageNumber="6">
<treatmentCitationGroup id="279A2E5A7C28B918FCD8E6DA34420AE3" box="[816,1395,1184,1208]" pageId="7" pageNumber="6">
<treatmentCitation id="862B2F657C28B918FCD8E6DA345E0AE3" author="BIRULA" box="[816,1391,1184,1208]" page="82" pageId="7" pageNumber="6" year="1928">
<taxonomicName id="C08A72F77C28B918FCD8E6DA345E0AE3" authority="Birula, 1928: 82" authorityName="Birula" authorityPageNumber="82" authorityYear="1928" box="[816,1391,1184,1208]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="minax" subGenus="Hottentotta" subSpecies="niloticus">
<emphasis id="35FED5667C28B918FCD8E6DA324B0AE3" box="[816,890,1184,1208]" italics="true" pageId="7" pageNumber="6">Buthus</emphasis>
(
<emphasis id="35FED5667C28B918FC62E6DB35320AE3" box="[906,1027,1185,1208]" italics="true" pageId="7" pageNumber="6">Hottentotta</emphasis>
)
<emphasis id="35FED5667C28B918FBFBE6DB35860AE3" box="[1043,1207,1184,1208]" italics="true" pageId="7" pageNumber="6">minax niloticus</emphasis>
<bibRefCitation id="631B74857C28B918FB55E6DA345E0AE3" author="BIRULA" box="[1213,1391,1184,1208]" pageId="7" pageNumber="6" pagination="79 - 88" refId="ref10464" refString="BIRULA, A. A. 1928. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner uternommenen Zoologischen Expedition nach dem Anglo-Agyptischen Sudan (Kordofan) 1914. XXV. Skorpione. Denkschriften der Akademie der Wissenschaften in Wien, 101: 79 - 88." type="journal article" year="1928">Birula, 1928: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="073509747C28B918FCD8E6BB35A70AA3" blockId="7.[816,1475,1184,1336]" pageId="7" pageNumber="6">
<treatmentCitationGroup id="279A2E5A7C28B918FCD8E6BB35A70AA3" pageId="7" pageNumber="6">
<treatmentCitation id="862B2F657C28B918FCD8E6BB32BB0AA3" author="KOVARIK" page="110" pageId="7" pageNumber="6" year="1998">
<taxonomicName id="C08A72F77C28B918FCD8E6BB32BB0AA3" ID-CoL="3MQXY" authority=": Kovarik, 1998: 110" authorityName="Kovarik" authorityPageNumber="110" authorityYear="1998" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="minax" subGenus="Hottentotta" subSpecies="niloticus">
<emphasis id="35FED5667C28B918FCD8E6BB32980A83" box="[816,937,1217,1240]" italics="true" pageId="7" pageNumber="6">Hottentotta</emphasis>
(
<emphasis id="35FED5667C28B918FC2AE6BB350A0A83" box="[962,1083,1217,1240]" italics="true" pageId="7" pageNumber="6">Hottentotta</emphasis>
)
<emphasis id="35FED5667C28B918FBBCE6BB34300A83" box="[1108,1281,1216,1240]" italics="true" pageId="7" pageNumber="6">minax niloticus</emphasis>
:
<bibRefCitation id="631B74857C28B918FAF0E6BA32BB0AA3" author="KOVARIK" pageId="7" pageNumber="6" refId="ref10766" refString="KOVARIK, F. 1998. StirI [Scorpiones]. Publishing House &quot; Madagaskar &quot;, Jihlava (Czech Republic). 175 pp. (in Czech)." type="book" year="1998">Kovařík, 1998: 110</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Fet &amp; Lowe, 2000: 142.
</treatmentCitationGroup>
</paragraph>
<paragraph id="073509747C28B918FCD8E77B34B50B63" blockId="7.[816,1475,1184,1336]" pageId="7" pageNumber="6">
<treatmentCitationGroup id="279A2E5A7C28B918FCD8E77B34B50B63" pageId="7" pageNumber="6">
<treatmentCitation id="862B2F657C28B918FCD8E77B35DC0B43" author="KOVARIK" box="[816,1261,1280,1304]" page="140" pageId="7" pageNumber="6" year="2003">
<taxonomicName id="C08A72F77C28B918FCD8E77B35DC0B43" ID-CoL="3MQXY" authority=": Kovarik, 2003: 140" authorityName=": Kovarik" authorityPageNumber="140" authorityYear="2003" box="[816,1261,1280,1304]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C28B918FCD8E77B353D0B43" box="[816,1036,1280,1304]" italics="true" pageId="7" pageNumber="6">Hottentotta niloticus</emphasis>
:
<bibRefCitation id="631B74857C28B918FBF2E77A35DC0B43" author="KOVARIK" box="[1050,1261,1280,1304]" pageId="7" pageNumber="6" pagination="133 - 159" refId="ref10797" refString="KOVARIK, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones), with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae, 67: 133 - 159." type="journal article" year="2003">Kovařík, 2003: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="862B2F657C28B918FB10E77A348F0B43" author="KOVARIK" box="[1272,1470,1280,1304]" page="48" pageId="7" pageNumber="6" year="2007">
<bibRefCitation id="631B74857C28B918FB10E77A348F0B43" author="KOVARIK" box="[1272,1470,1280,1304]" pageId="7" pageNumber="6" pagination="1 - 107" refId="ref10897" refString="KOVARIK, F. 2007. A revision of the genus Hottentotta Birula, 1908, with descriptions of four new species (Scorpiones, Buthidae). Euscorpius, 58: 1 - 107." type="journal article" year="2007">Kovařík, 2007: 48</bibRefCitation>
</treatmentCitation>
; Kovařík &amp; Ojanguren, 2013: 170, figs. 1093–1094.
</treatmentCitationGroup>
</paragraph>
<paragraph id="073509747C28B918FCD8E71A35FD0BC3" blockId="7.[816,1474,1376,1432]" pageId="7" pageNumber="6">
TYPELOCALITYANDTYPEREPOSITORY.
<emphasis id="35FED5667C28B918FB47E71A35C90B23" bold="true" box="[1199,1272,1376,1400]" pageId="7" pageNumber="6">
<collectingCountry id="7F9D49E47C28B918FB47E71A35C90B23" box="[1199,1272,1376,1400]" name="Sudan" pageId="7" pageNumber="6">Sudan</collectingCountry>
</emphasis>
:
<emphasis id="35FED5667C28B918FAE9E71A34570B23" box="[1281,1382,1376,1400]" italics="true" pageId="7" pageNumber="6">Kordofan</emphasis>
,Kaducli (now
<emphasis id="35FED5667C28B918FC86E7FA35270BC3" box="[878,1046,1408,1432]" italics="true" pageId="7" pageNumber="6">
<collectingRegion id="C54EC7967C28B918FC86E7FA35270BC3" box="[878,1046,1408,1432]" country="Sudan" name="Janub Kurdufan" pageId="7" pageNumber="6">South Kordofan</collectingRegion>
</emphasis>
, Kaduqli); ZISP.
</paragraph>
</subSubSection>
<subSubSection id="4F905AFF7C28B918FCD8E7BB32FB0803" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="073509747C28B918FCD8E7BB32FB0803" blockId="7.[816,1474,1472,1624]" pageId="7" pageNumber="6">
<materialsCitation id="B7E203297C28B918FCD8E7BB32F70803" collectingDate="1914-03-28" collectingDateMax="1914-03-29" collectingDateMin="1914-03-28" collectionCode="ZISP" collectorName="F. Werner &amp; No." country="Sudan" county="Kordofan" location="Kaduqli" municipality="Kaducli" pageId="7" pageNumber="6" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Janub Kurdufan" typeStatus="lectotype">
TYPE MATERIAL EXAMINED.
<emphasis id="35FED5667C28B918FBB5E7BA35970B83" bold="true" box="[1117,1190,1472,1496]" pageId="7" pageNumber="6">
<collectingCountry id="7F9D49E47C28B918FBB5E7BA35970B83" box="[1117,1190,1472,1496]" name="Sudan" pageId="7" pageNumber="6">Sudan</collectingCountry>
</emphasis>
:
<emphasis id="35FED5667C28B918FB5FE7BA342D0B83" box="[1207,1308,1472,1496]" italics="true" pageId="7" pageNumber="6">
<collectingCounty id="EE5471F87C28B918FB5FE7BA342D0B83" box="[1207,1308,1472,1496]" pageId="7" pageNumber="6">Kordofan</collectingCounty>
</emphasis>
,
<collectingMunicipality id="E751930E7C28B918FAC5E7BA34B00B83" box="[1325,1409,1472,1496]" pageId="7" pageNumber="6">Kaducli</collectingMunicipality>
(now
<emphasis id="35FED5667C28B918FCD8E79A32D20BA3" box="[816,995,1504,1528]" italics="true" pageId="7" pageNumber="6">
<collectingRegion id="C54EC7967C28B918FCD8E79A32D20BA3" box="[816,995,1504,1528]" country="Sudan" name="Janub Kurdufan" pageId="7" pageNumber="6">South Kordofan</collectingRegion>
</emphasis>
,
<location id="02555FAF7C28B918FC13E79A35640BA3" LSID="urn:lsid:plazi:treatment:8F23B8627C28B914FC4EE645340E0835:02555FAF7C28B918FC13E79A35640BA3" box="[1019,1109,1504,1528]" country="Sudan" county="Kordofan" municipality="Kaducli" name="Kaduqli" pageId="7" pageNumber="6" stateProvince="Janub Kurdufan">Kaduqli</location>
),
<date id="73342FB47C28B918FB9AE79A343E0BA3" box="[1138,1295,1504,1528]" pageId="7" pageNumber="6" value="1914-03-28" valueMax="1914-03-29" valueMin="1914-03-28">
<collectingDate id="6370D65C7C28B918FB9AE79A343E0BA3" box="[1138,1295,1504,1528]" pageId="7" pageNumber="6" value="1914-03-28" valueMax="1914-03-29" valueMin="1914-03-28">28-29.III.1914</collectingDate>
</date>
,
<specimenCount id="118CC2FD7C28B918FACDE79A34770BA3" box="[1317,1350,1504,1528]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
(
<typeStatus id="D831B7D67C28B918FAB6E79A34F30BA3" box="[1374,1474,1504,1528]" pageId="7" pageNumber="6" type="lectotype">lectotype</typeStatus>
hereby designated,
<figureCitation id="9FB115F17C28B918FC16E47A354F0843" box="[1022,1150,1536,1560]" captionStart="Figures 23–26" captionStartId="8.[113,190,1981,2003]" captionTargetBox="[121,1454,272,1956]" captionTargetId="figure-53@8.[113,1474,263,1965]" captionTargetPageId="8" captionText="Figures 23–26: Hottentotta niloticus. Figures 23–24. Lectotype male, dorsal (23) and ventral (24) views. Figures 25–26. Paralectotype female, dorsal (25) and ventral (26) views. Scale bar = 10 mm." pageId="7" pageNumber="6">Figs. 23–24</figureCitation>
,
<figureCitation id="9FB115F17C28B918FB60E47A35FC0843" box="[1160,1229,1536,1560]" captionStart="Figures 27–32" captionStartId="9.[116,193,1970,1992]" captionTargetBox="[122,1471,246,1948]" captionTargetId="figure-58@9.[113,1474,234,1957]" captionTargetPageId="9" captionText="Figures 27–32: Hottentotta niloticus. Figures 27–29. Paralectotype female, metasoma and telson, lateral (27), dorsal (28), and ventral (29) views. Figures 30–32. Lectotype male, metasoma and telson, lateral (30), dorsal (31), and ventral (32) views. Scale bars = 10 mm." pageId="7" pageNumber="6">30–32</figureCitation>
,
<figureCitation id="9FB115F17C28B918FB30E47A34A20843" box="[1240,1427,1536,1560]" captionStart="Figures 33–49" captionStartId="10.[115,192,1884,1906]" captionTargetBox="[113,1465,222,1873]" captionTargetId="figure-140@10.[113,1474,212,1873]" captionTargetPageId="10" captionText="Figures 33–49: Hottentotta niloticus. Figures 33–43: Pedipalp segments. Figures 33–34. Paralectotype female, pedipalp chela dorsal (33) and external (34) views. Figures 35–43. Lectotype male, pedipalp chela dorsal (35), external (36) and vental (37) views, patella dorsal (38), external (39) and ventral (40) views, femur and trochanter dorsal (41) and ventral (42), movable finger (43). The trichobothrial pattern is indicated in Figures 35–39, 41. Figures 44, 46, 48–49. Lectotype male, chelicerae, carapace and tergites I–III (44), sternopectinal region and sternites III–IV (46), and distal segments of legs III–IV (48–49), retrolateral view. Figures 45, 47. Paralectotype female, chelicerae, carapace and tergites I–III (45), sternopectinal region and sternite III (47)." pageId="7" pageNumber="6">35–44, 46, 48–49</figureCitation>
)
<specimenCount id="118CC2FD7C28B918FA49E47A34F30843" box="[1441,1474,1536,1560]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
(
<typeStatus id="D831B7D67C28B918FCD0E45A32FD0863" box="[824,972,1568,1592]" pageId="7" pageNumber="6" type="paralectotype">paralectotype</typeStatus>
,
<figureCitation id="9FB115F17C28B918FC32E45A356F0863" box="[986,1118,1568,1592]" captionStart-0="Figures 23–26" captionStart-1="Figures 27–32" captionStartId-0="8.[113,190,1981,2003]" captionStartId-1="9.[116,193,1970,1992]" captionTargetBox-0="[121,1454,272,1956]" captionTargetBox-1="[122,1471,246,1948]" captionTargetId-0="figure-53@8.[113,1474,263,1965]" captionTargetId-1="figure-58@9.[113,1474,234,1957]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figures 23–26: Hottentotta niloticus. Figures 23–24. Lectotype male, dorsal (23) and ventral (24) views. Figures 25–26. Paralectotype female, dorsal (25) and ventral (26) views. Scale bar = 10 mm." captionText-1="Figures 27–32: Hottentotta niloticus. Figures 27–29. Paralectotype female, metasoma and telson, lateral (27), dorsal (28), and ventral (29) views. Figures 30–32. Lectotype male, metasoma and telson, lateral (30), dorsal (31), and ventral (32) views. Scale bars = 10 mm." pageId="7" pageNumber="6">Figs. 25–29</figureCitation>
,
<figureCitation id="9FB115F17C28B918FB84E45A34380863" box="[1132,1289,1568,1592]" captionStart="Figures 33–49" captionStartId="10.[115,192,1884,1906]" captionTargetBox="[113,1465,222,1873]" captionTargetId="figure-140@10.[113,1474,212,1873]" captionTargetPageId="10" captionText="Figures 33–49: Hottentotta niloticus. Figures 33–43: Pedipalp segments. Figures 33–34. Paralectotype female, pedipalp chela dorsal (33) and external (34) views. Figures 35–43. Lectotype male, pedipalp chela dorsal (35), external (36) and vental (37) views, patella dorsal (38), external (39) and ventral (40) views, femur and trochanter dorsal (41) and ventral (42), movable finger (43). The trichobothrial pattern is indicated in Figures 35–39, 41. Figures 44, 46, 48–49. Lectotype male, chelicerae, carapace and tergites I–III (44), sternopectinal region and sternites III–IV (46), and distal segments of legs III–IV (48–49), retrolateral view. Figures 45, 47. Paralectotype female, chelicerae, carapace and tergites I–III (45), sternopectinal region and sternite III (47)." pageId="7" pageNumber="6">33–34, 45, 47</figureCitation>
), leg.
<collectorName id="AA7F6CA27C28B918FAB9E45B348E0863" box="[1361,1471,1569,1592]" pageId="7" pageNumber="6">F. Werner</collectorName>
,
<collectionCode id="619B91B17C28B918FCD8E43A32570803" box="[816,870,1600,1624]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="6">ZISP</collectionCode>
<collectorName id="AA7F6CA27C28B918FC85E43B32A40803" box="[877,917,1601,1624]" pageId="7" pageNumber="6">No.</collectorName>
564
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4F905AFF7C28B914FCD8E4FA30E90974" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="073509747C28B914FCD8E4FA30E90974" blockId="7.[816,1475,1664,2009]" lastBlockId="11.[113,771,1399,1839]" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6">
DIAGNOSIS. Total length
<quantity id="C072A4917C28B918FBACE4FA358E08C3" box="[1092,1215,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="7" pageNumber="6" unit="mm" value="57.5" valueMax="70.0" valueMin="45.0">45–70 mm</quantity>
. Trichobothrium
<emphasis id="35FED5667C28B918FA69E4FA34AD08C3" box="[1409,1436,1664,1688]" italics="true" pageId="7" pageNumber="6">db</emphasis>
on fixed finger of pedipalp situated between trichobothria
<emphasis id="35FED5667C28B918FA47E4DB34F308E3" box="[1455,1474,1697,1720]" italics="true" pageId="7" pageNumber="6">et</emphasis>
and
<emphasis id="35FED5667C28B918FCB7E4BB324C0883" box="[863,893,1729,1752]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, may be on level with
<emphasis id="35FED5667C28B918FB9FE4BB35A40883" box="[1143,1173,1729,1752]" italics="true" pageId="7" pageNumber="6">est</emphasis>
. Manus of pedipalp usually of same width in both sexes, but males have fingers twisted whereas females have them straight. Pectinal teeth number 19–28. Chelicerae yellow, without reticulation, only tips of teeth on cheliceral fingers are black. Pedipalps sparsely hirsute. Metasoma bears only a few setae. Color usually uniformly yellowish brown, only ventral carinae of metasoma black; mesosoma and carapace may be black in some specimens. Femur of pedipalps with 5 carinae that may be incomplete. Patella with 8 carinae, of which some are smooth, without granules and obsolete. Chela lacks carinae. Movable fingers of pedipalps with 12–13 rows of granules and 5 terminal granules. Seventh sternite with 4 well marked carinae. First to third metasomal segments with 10 carinae; fourth with 8 or 10 carinae; fifth segment with 5 carinae. Lateral carinae may not be discernible in some males. All carinae granulated, dorsal carinae bear larger terminal granules. Metasoma strongly granulated, accessoric rows of granules present on dorsal surfaces of segments as well as on ventral surface of fifth segment. First metasomal segment of adults always wider than long; second metasomal segment usually also wider than long but may be longer than wide. Second to fourth metasomal segment width ratio less than 1.2.
</paragraph>
</subSubSection>
<caption id="53F559FC7C27B917FF99E5C733FC09B4" pageId="8" pageNumber="7" startId="8.[113,190,1981,2003]" targetBox="[121,1454,272,1956]" targetPageId="8">
<paragraph id="073509747C27B917FF99E5C733FC09B4" blockId="8.[113,1472,1981,2032]" pageId="8" pageNumber="7">
<emphasis id="35FED5667C27B917FF99E5C730350989" bold="true" box="[113,260,1981,2003]" pageId="8" pageNumber="7">Figures 23–26</emphasis>
:
<taxonomicName id="C08A72F77C27B917FEFCE5C730EC0989" authorityName=": Kovarik" authorityYear="2003" box="[276,477,1981,2002]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C27B917FEFCE5C730EC0989" box="[276,477,1981,2002]" italics="true" pageId="8" pageNumber="7">Hottentotta niloticus</emphasis>
</taxonomicName>
.
<emphasis id="35FED5667C27B917FE04E5C7334E0989" bold="true" box="[492,639,1981,2003]" pageId="8" pageNumber="7">Figures 23–24</emphasis>
. Lectotype male, dorsal (23) and ventral (24) views.
<emphasis id="35FED5667C27B917FB73E5C7343B0989" bold="true" box="[1179,1290,1981,2003]" pageId="8" pageNumber="7">Figures 25</emphasis>
–2
<emphasis id="35FED5667C27B917FACAE5C7341F0989" bold="true" box="[1314,1326,1981,2002]" pageId="8" pageNumber="7">6</emphasis>
. Paralectotype female, dorsal (25) and ventral (26) views. Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="53F559FC7C26B916FF9CE5C8346C09BE" pageId="9" pageNumber="8" startId="9.[116,193,1970,1992]" targetBox="[122,1471,246,1948]" targetPageId="9">
<paragraph id="073509747C26B916FF9CE5C8346C09BE" blockId="9.[116,1474,1970,2022]" pageId="9" pageNumber="8">
<emphasis id="35FED5667C26B916FF9CE5C830350993" bold="true" box="[116,260,1970,1992]" pageId="9" pageNumber="8">Figures 27–32</emphasis>
:
<taxonomicName id="C08A72F77C26B916FEFAE5C930EB0993" authorityName=": Kovarik" authorityYear="2003" box="[274,474,1971,1992]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C26B916FEFAE5C930EB0993" box="[274,474,1971,1992]" italics="true" pageId="9" pageNumber="8">Hottentotta niloticus</emphasis>
</taxonomicName>
.
<emphasis id="35FED5667C26B916FE0FE5C833460993" bold="true" box="[487,631,1970,1992]" pageId="9" pageNumber="8">Figures 27–29</emphasis>
. Paralectotype female, metasoma and telson, lateral (27), dorsal (28), and ventral (29) views.
<emphasis id="35FED5667C26B916FF50E5B5307609BF" bold="true" box="[184,327,1999,2021]" pageId="9" pageNumber="8">Figures 30–32</emphasis>
. Lectotype male, metasoma and telson, lateral (30), dorsal (31), and ventral (32) views. Scale bars = 10 mm.
</paragraph>
</caption>
<caption id="53F559FC7C25B915FF9BE52633EE0658" pageId="10" pageNumber="9" startId="10.[115,192,1884,1906]" targetBox="[113,1465,222,1873]" targetPageId="10">
<paragraph id="073509747C25B915FF9BE52633EE0658" blockId="10.[115,1473,1884,2051]" pageId="10" pageNumber="9">
<emphasis id="35FED5667C25B915FF9BE5263032092A" bold="true" box="[115,259,1884,1906]" pageId="10" pageNumber="9">Figures 33–49</emphasis>
:
<taxonomicName id="C08A72F77C25B915FEFAE52730EB092A" authorityName=": Kovarik" authorityYear="2003" box="[274,474,1884,1906]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C25B915FEFAE52730EB092A" box="[274,474,1884,1906]" italics="true" pageId="10" pageNumber="9">Hottentotta niloticus</emphasis>
</taxonomicName>
.
<emphasis id="35FED5667C25B915FE0FE5263349092A" bold="true" box="[487,632,1884,1906]" pageId="10" pageNumber="9">Figures 33–43</emphasis>
: Pedipalp segments.
<emphasis id="35FED5667C25B915FCA1E52632EB092A" bold="true" box="[841,986,1884,1906]" pageId="10" pageNumber="9">Figures 33–34</emphasis>
. Paralectotype female, pedipalp chela dorsal (33) and external (34) views.
<emphasis id="35FED5667C25B915FE8BE50330C209D5" bold="true" box="[355,499,1913,1935]" pageId="10" pageNumber="9">Figures 35–43</emphasis>
. Lectotype male, pedipalp chela dorsal (35), external (36) and vental (37) views, patella dorsal (38), external (39) and ventral (40) views, femur and trochanter dorsal (41) and ventral (42), movable finger (43). The trichobothrial pattern is indicated in Figures 35–39, 41.
<emphasis id="35FED5667C25B915FE4DE5C933210993" bold="true" box="[421,528,1971,1993]" pageId="10" pageNumber="9">Figures 44</emphasis>
,
<emphasis id="35FED5667C25B915FDF4E5C933050993" bold="true" box="[540,564,1971,1992]" pageId="10" pageNumber="9">46</emphasis>
,
<emphasis id="35FED5667C25B915FDA8E5C9334D0993" bold="true" box="[576,636,1971,1992]" pageId="10" pageNumber="9">48–49</emphasis>
. Lectotype male, chelicerae, carapace and tergites I–III (44), sternopectinal region and sternites III–IV (46), and distal segments of legs III–IV (48–49), retrolateral view.
<emphasis id="35FED5667C25B915FC66E5B532C809BF" bold="true" box="[910,1017,1999,2021]" pageId="10" pageNumber="9">Figures 45</emphasis>
,
<emphasis id="35FED5667C25B915FBEDE5B5352C09BF" bold="true" box="[1029,1053,1999,2020]" pageId="10" pageNumber="9">47</emphasis>
. Paralectotype female, chelicerae, carapace and tergites I–III (45), sternopectinal region and sternite III (47).
</paragraph>
</caption>
<caption id="53F559FC7C24B914FF9CE74335840B14" box="[116,1205,1337,1360]" pageId="11" pageNumber="10" startId="11.[116,193,1337,1359]" targetBox="[127,1448,252,1295]" targetPageId="11">
<paragraph id="073509747C24B914FF9CE74335840B14" blockId="11.[116,1205,1337,1360]" box="[116,1205,1337,1360]" pageId="11" pageNumber="10">
<emphasis id="35FED5667C24B914FF9CE74330320B15" bold="true" box="[116,259,1337,1359]" pageId="11" pageNumber="10">Figures 50–51</emphasis>
.
<taxonomicName id="C08A72F77C24B914FEE7E74030C20B14" box="[271,499,1338,1359]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="penjabensis">
<emphasis id="35FED5667C24B914FEE7E74030C20B14" box="[271,499,1338,1359]" italics="true" pageId="11" pageNumber="10">Hottentotta penJabensis</emphasis>
</taxonomicName>
, holotype female, dorsal (50) and ventral (51) views. Scale bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="4F905AFF7C24B914FF99E52D340E0835" pageId="11" pageNumber="10" type="discussion">
<paragraph id="073509747C24B914FF99E52D340E0835" blockId="11.[113,771,1879,2031]" lastBlockId="11.[816,1475,1399,1647]" pageId="11" pageNumber="10">
COMMENTS. Kovařík in Kovařík &amp; Ojanguren (2013: 168) assumed that
<taxonomicName id="C08A72F77C24B914FEE2E50D30BB09D4" authorityName=": Kovarik" authorityYear="2003" box="[266,394,1911,1935]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C24B914FEE2E50D30BB09D4" box="[266,394,1911,1935]" italics="true" pageId="11" pageNumber="10">H. niloticus</emphasis>
</taxonomicName>
could be a valid species different from
<taxonomicName id="C08A72F77C24B914FF5AE5ED302B09F5" baseAuthorityName="L.Koch" baseAuthorityYear="1975" box="[178,282,1943,1966]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="35FED5667C24B914FF5AE5ED302B09F5" box="[178,282,1943,1966]" italics="true" pageId="11" pageNumber="10">H. minax</emphasis>
</taxonomicName>
and that studying the types can solve the problem. Studying the types of
<taxonomicName id="C08A72F77C24B914FE56E5CD332E0995" baseAuthorityName="L.Koch" baseAuthorityYear="1975" box="[446,543,1975,1998]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="35FED5667C24B914FE56E5CD332E0995" box="[446,543,1975,1998]" italics="true" pageId="11" pageNumber="10">H. minax</emphasis>
</taxonomicName>
(male
<typeStatus id="D831B7D67C24B914FD80E5CD33FB0994" box="[616,714,1975,1999]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
from
<collectingCountry id="7F9D49E47C24B914FF99E5AD318309B4" box="[113,178,2007,2031]" name="Egypt" pageId="11" pageNumber="10">Egypt</collectingCountry>
) and types of
<taxonomicName id="C08A72F77C24B914FEA1E5AD30F409B4" authorityName=": Kovarik" authorityYear="2003" box="[329,453,2007,2031]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="35FED5667C24B914FEA1E5AD30F409B4" box="[329,453,2007,2031]" italics="true" pageId="11" pageNumber="10">H. niloticus</emphasis>
</taxonomicName>
(
<typeStatus id="D831B7D67C24B914FE3AE5AD330709B4" box="[466,566,2007,2031]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="D831B7D67C24B914FD81E5AD323209B4" box="[617,771,2007,2031]" pageId="11" pageNumber="10" type="paralectotype">paralectotypes</typeStatus>
from
<collectingCountry id="7F9D49E47C24B914FC83E70D329E0BD4" box="[875,943,1399,1423]" name="Sudan" pageId="11" pageNumber="10">Sudan</collectingCountry>
) show agreement in the following key characters: trichobothrial pattern, pedipalp finger dentation, pectinal tooth count and lamellar structure, proportions, setation, carination and sculpture of pedipalps, carapace, tergites, sternites, and metasoma, shape of the telson, as well as armature of chelicerae and pedipalp fingers. The logical conclusion is that
<taxonomicName id="C08A72F77C24B914FCD8E44D347A0814" authority="Birula, 1928" authorityName="Birula" authorityYear="1928" box="[816,1355,1591,1615]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="minax" subGenus="Hottentotta" subSpecies="niloticus">
<emphasis id="35FED5667C24B914FCD8E44D324B0814" box="[816,890,1591,1615]" italics="true" pageId="11" pageNumber="10">Buthus</emphasis>
(
<emphasis id="35FED5667C24B914FC64E44D35340815" box="[908,1029,1591,1614]" italics="true" pageId="11" pageNumber="10">Hottentotta</emphasis>
)
<emphasis id="35FED5667C24B914FBFFE44D358D0814" box="[1047,1212,1591,1615]" italics="true" pageId="11" pageNumber="10">minax niloticus</emphasis>
<bibRefCitation id="631B74857C24B914FB2CE44D347A0814" author="BIRULA" box="[1220,1355,1591,1615]" pageId="11" pageNumber="10" pagination="79 - 88" refId="ref10464" refString="BIRULA, A. A. 1928. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner uternommenen Zoologischen Expedition nach dem Anglo-Agyptischen Sudan (Kordofan) 1914. XXV. Skorpione. Denkschriften der Akademie der Wissenschaften in Wien, 101: 79 - 88." type="journal article" year="1928">Birula, 1928</bibRefCitation>
</taxonomicName>
is a junior synonym of
<taxonomicName id="C08A72F77C24B914FC5EE42D35DA0834" authority="L. Koch, 1875" authorityName="L. Koch" authorityYear="1875" box="[950,1259,1623,1647]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="35FED5667C24B914FC5EE42D35760835" box="[950,1095,1623,1647]" italics="true" pageId="11" pageNumber="10">Buthus minax</emphasis>
L. Koch, 1875
</taxonomicName>
,
<taxonomicNameLabel id="2ECD681D7C24B914FB1EE42D340E0835" box="[1270,1343,1623,1647]" pageId="11" pageNumber="10">
<emphasis id="35FED5667C24B914FB1EE42D342C0834" bold="true" box="[1270,1309,1623,1647]" pageId="11" pageNumber="10">syn</emphasis>
.
<emphasis id="35FED5667C24B914FAC2E42D34080834" bold="true" box="[1322,1337,1623,1647]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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