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222 lines
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<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="5F6B14D73F4F589AB6421AE0A0780643" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) brunneus Karsch 1881, stat. rev." docType="treatment" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
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<mods:title id="AE08C6C4C24037B0256472C51F6BE2ED">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart id="3A4FB83EA53BF4FA7E5A2F8A063D8AE5">Gueorguiev, Borislav</mods:namePart>
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<mods:affiliation id="C8136322D35812CDE4001C75ACC27DEB">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier id="F8107243209EBAEC1865C3DBED66AC6C" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart id="BD757C24BF1F75A8BE98F78BE61B449E">Wrase, David W.</mods:namePart>
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<mods:nameIdentifier id="9D4E5E814AE4DB81F9C27A72F5B2A4F9" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation id="59EAC62830E8CC82B351E71E2F1A0A71">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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<mods:namePart id="203497A42A42878C49F9F309B843C98F">Assmann, Thorsten</mods:namePart>
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<mods:affiliation id="205ECA69AD6A2A57E195B258AB5B79FD">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:namePart id="2E1EAC7475C34AF135058DE8E6358765">Muilwijk, Jan</mods:namePart>
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<mods:affiliation id="CE35E7B7992B3EDEE7D2DEFCB5FF8624">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="E732C2B51F391014F1069EA35F145EB1">Machard, Patrice</mods:namePart>
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<mods:affiliation id="110DD42FDC8556B4A5B6DBE776C627B0">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:date id="C45EA83A41ABAAD164010BFEAE733C61">2022</mods:date>
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<treatment id="5F6B14D73F4F589AB6421AE0A0780643" ID-GBIF-Taxon="195868755" LSID="urn:lsid:plazi:treatment:5F6B14D73F4F589AB6421AE0A0780643" httpUri="http://treatment.plazi.org/id/5F6B14D73F4F589AB6421AE0A0780643" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
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<subSubSection id="5C1F6AA48A42AA488E6B81637B69E1B1" pageId="0" pageNumber="71" type="nomenclature">
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<paragraph id="268BDEA1A9976E129CA70E3029677BBB" pageId="0" pageNumber="71">
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15.
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<taxonomicName id="0F1DE4A4ECC1EA906A530EE2EBBD2C30" ID-CoL="4JQH4" LSID="5F6B14D7-3F4F-589A-B642-1AE0A0780643" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" status="stat. rev." subGenus="Eremoderus">Platyderus (Eremoderus) brunneus Karsch, 1881</taxonomicName>
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<taxonomicNameLabel id="E387B294865FA8F410D52D6B3B137DD9" pageId="0" pageNumber="71">stat. rev.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D6469DC7516A83171CE53D495AF0ACFA" pageId="0" pageNumber="71" type="description">
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<paragraph id="87197DC0089602A5086464917D294666" pageId="0" pageNumber="71">
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<figureCitation id="22ED3D4ACD4025612E209F4DDC60B0BB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, F</figureCitation>
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<figureCitation id="1392E83F4C7610D41D06B31BE58384E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche & Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5H</figureCitation>
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<figureCitation id="6B8D9602E175523E7241198679C543FE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A, B</figureCitation>
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<figureCitation id="A7CD08F76E23A2A153A603B2F3DFBC85" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I, J</figureCitation>
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<figureCitation id="F11D97EB658399622857F668668D5ADA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C, D</figureCitation>
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<figureCitation id="C9296947179883CF8E7A95DFB6DF70CA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B, C</figureCitation>
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<figureCitation id="1825677A5F16A3BA6E05869BF65AB701" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche & Saulcy, 1855), male paralectotype, " Syria ". Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J, K</figureCitation>
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<figureCitation id="465BF25C2FAA4D280D90B02A5E4BBE71" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A, B</figureCitation>
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<figureCitation id="7A1E5569D2E1995B0F3536D150F8E9FF" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E7D6571A98FC2BB624E35183F094732C" pageId="0" pageNumber="71" type="diagnosis">
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<paragraph id="5F1CC9BE6239D33C1CB7B9AE2F2EEE8D" pageId="0" pageNumber="71">Diagnosis.</paragraph>
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<paragraph id="E528E28AAC55A9B19F3CD2B0B5F3BF41" pageId="0" pageNumber="71">
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A species of
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<taxonomicName id="3A0B3FDBBF62AF989AA8CF04A57DEABB" authority="" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Platyderus (Eremoderus)</taxonomicName>
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, with mesotarsomeres and metatarsomeres dorsally more or less flattened and grooved (Fig.
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<figureCitation id="F189906866308B41FCCDE6E236F13FE1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6A, B</figureCitation>
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), cordate pronotum, with sides from widest point to base markedly (nominotypical subspecies) or barely concave (ssp.
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<taxonomicName id="1844B684789F1770582FC86DC4BB59DB" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
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<emphasis id="173D670E818C979C9E35F24A01462E48" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
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|
</taxonomicName>
|
|
) and posterior angles obtuse, with tips laterally more (nominotypical subspecies) or less protruding (ssp.
|
|
<taxonomicName id="1EB42EC04FCCAF21C617A9AD709F109A" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
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<emphasis id="ACB522282CAA2718AAD80B5E515BAF68" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
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</taxonomicName>
|
|
), and median lobe of aedeagus with basal bulb more slanting to shaft (Fig.
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|
<figureCitation id="670362D686C533AE36BF9D8787CB6D3E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
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).
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</paragraph>
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|
</subSubSection>
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<subSubSection id="98033BABF677886AF007E7BC6B6C60DF" pageId="0" pageNumber="71" type="redescription">
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<paragraph id="793431F73E953B37CDA254382CCC5837" pageId="0" pageNumber="71">Redescription.</paragraph>
|
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<paragraph id="DE63A72FDFAB6A3B0ED46B6A7B6DC57F" pageId="0" pageNumber="71">
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<emphasis id="C050192CB099947A88FA16A0AEB67020" bold="true" pageId="0" pageNumber="71">
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<emphasis id="1DCDB8A7E1B2ADF53D5E8FFAF85BDE5E" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
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.
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</emphasis>
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Moderately to large-sized species (BL: 7.60-9.30 mm; BW: 2.70-3.40 mm), with elongate and moderately convex body (Fig.
|
|
<figureCitation id="93B0BE401E7212622090EDF61B90F38B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3D, F</figureCitation>
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).
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<emphasis id="774B584CA10DC8DCA09F55FD26F09B58" bold="true" pageId="0" pageNumber="71">
|
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<emphasis id="3C6C5FDC6A61997008CD3969C2520D2F" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
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.
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</emphasis>
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See Table
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<tableCitation id="551B9FA5D8160B6B5A9520F2DCDDECB4" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
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.
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<emphasis id="868D5BC981FD81C6951E727EEB194ECA" bold="true" pageId="0" pageNumber="71">
|
|
<emphasis id="E40DF5A69EED5116AD724DAF6FDD74D1" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
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.
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</emphasis>
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Body, palpi, antennae and most of legs yellow-brown to reddish- brown, with head usually darker than pronotum and elytra and femora lighter than tibiae and tarsomeres. Integument slightly shiny (less shiny than in
|
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<taxonomicName id="D6EBAD4CDDAB6027760D6F88BE2F4840" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
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<emphasis id="3DCFFB2830AAD8DCAB1F13C13E48E1E1" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
|
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</taxonomicName>
|
|
).
|
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<emphasis id="26C71D2E823F036C3696DDCB8260FF4D" bold="true" pageId="0" pageNumber="71">
|
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<emphasis id="94A9FB322F70624E4B61F6BFFD8F8B78" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
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.
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</emphasis>
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Pronotum with evident microreticulation, sculpticells regularly isodiametric. Elytral intervals, scutellum and lateral gutterwith distinct isodiametric sculpticells; basal margin with reduced microreticulation. Ventral surface with impressed slight transverse sculpticells or with sculpticells scarcely-visible. Head impunctate, usually smooth, clypeus and areas laterad of frontal furrows sometimes with fine longitudinal wrinkles. Surface of pronotum impunctate, largely smooth, only basal area medially behind posterior transverse impression longitudinally wrinkled, rarely apical part medially in front of anterior transverse impression also wrinkled; basal foveae and adjacent lateral areas smooth, smoother than those in
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<taxonomicName id="93067E9758C78D3FB0060C7E2A8E3D82" class="Insecta" kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
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<emphasis id="F772415986A89AB38E8FB0567B2050E9" italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
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</taxonomicName>
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. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled in whole, 2-5 finely wrinkled at sides.
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<emphasis id="A694D7205E9DA9E1AAA078CAEE20F48F" bold="true" pageId="0" pageNumber="71">
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<emphasis id="BB901E9997C726A210B022C738D67E2B" italics="true" pageId="0" pageNumber="71">Head</emphasis>
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.
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</emphasis>
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|
About one-third as wide as pronotum (PW/HW= 1.28-1.41). Eye long, moderately convex. Labrum subrectangular, as long as clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced to disappeared at sides. Frontal furrows small, rather shallow, punctiform. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, nearly reaching level of posterior supraorbital pore.
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<emphasis id="D660F8505963C7A9E450B0BE623AE6B8" bold="true" pageId="0" pageNumber="71">
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<emphasis id="D1C19D698F9BD5F2E3D3A72446860273" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
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.
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</emphasis>
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|
Pronotum one-tenth to one-fifth wider than long (PW/PL= 1.10-1.21), with widest point at second quarter. Anterior transverse impression absent, posterior transverse impression short, barely distinct. Sides convex medially and anteriorly, slightly concave to straight in base; anterior bead present laterally, lacking or present in medial eight to tenth; basal bead present laterally, reduced to absent in medial fifth to half. Metepisternum slightly longer than wide or nearly as long as wide, MA/MI= 0.87-1.00, with greater values occurring in specimens of ssp.
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|
<taxonomicName id="718E2F93D1620D0B0A685AAEC2E34218" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
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<emphasis id="22BFAD0FC1817899DC5F2FF6C5BDE9F1" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<emphasis id="1F6D93E01AB9581E98D4568121EC7A7D" bold="true" pageId="0" pageNumber="71">
|
|
<emphasis id="F87A5AB1CFE85A9505374EC543715D1B" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
|
|
.
|
|
</emphasis>
|
|
Oblong, about one-and-a-half times to one and two thirds as long as wide (EL/EW= 1.52-1.72), two-and-a-half times or little more as long as pronotum (EL/PL= 2.46-2.72), and one and a third to a one-and-a-half times as wide as pronotum (EW/PW= 1.35-1.48), with widest point at medial third. Parascutellar striole and striae 1-7 very shallow to indistinct, stria 8 evident, clearly deeper than other striae (nominotypical form) or parascutellar striole and striae 1-7 distinct, slightly impressed, stria 8 somewhat deeper than other striae (ssp.
|
|
<taxonomicName id="841648D170DB928744CE0696646452DA" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
|
|
<emphasis id="B5D85A9A346D5C26F444AC575E3DFC7E" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
|
|
</taxonomicName>
|
|
); parascutellar striole, if present, not joining stria 1; striae 1-8 superficially to moderately punctate; bases of striae 1-7 more or less reduced, not reaching basal bead (nominotypical form) or distinct, reaching basal bead (ssp.
|
|
<taxonomicName id="84D1B0B001C53C30D0D5EC4CB5517B42" authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
|
|
<emphasis id="7C8EA7B4B52B33AF6168B340878D1904" italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
|
|
</taxonomicName>
|
|
). Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 or 17 punctures on each side.
|
|
<emphasis id="666AE40683FF3220DDF0B8BBB322BE75" bold="true" pageId="0" pageNumber="71">
|
|
<emphasis id="C4F976109EE4AACAC702091644D2C006" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
|
|
.
|
|
</emphasis>
|
|
Posterior side of profemur with one or two setae in basal third and one or two in medial third. Mesofemur with 4-7 setiferous punctures on anterior side ventrally (Fig.
|
|
<figureCitation id="88C70F923A479EBDA55F380EF51EE947" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche & Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5H</figureCitation>
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). Anterior side of metafemur ventrally with a few long setae, one in basal third and one to three in apical half.
|
|
<emphasis id="48081BB624194F0B12BD2B14BC71CF07" bold="true" pageId="0" pageNumber="71">
|
|
<emphasis id="3304463B9913792161028E2CE029C28B" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
|
|
.
|
|
</emphasis>
|
|
Urite IX suboval, with proximal margin slightly asymmetrical, insignificantly turned to left (Fig.
|
|
<figureCitation id="05863C0D3C7837AC7C312BC36435A7B1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">7I, J</figureCitation>
|
|
). Median lobe of aedeagus in lateral view slender, with narrow basal bulb, long shaft and short, straight apex (Fig.
|
|
<figureCitation id="BC23E6A36E8F699F1CBB78A7568DCA85" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
|
|
); median lobe in ventral view straight, 3.4-3.5 times longer than wide (Fig.
|
|
<figureCitation id="3AC01AFA6CE42E929551AA39C633B5B3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
|
|
); apical lamella (dorsal view) short, scarcely asymmetrical, rounded at tip, with right side straight and left side slightly concave. Internal sac in lateral view (Fig.
|
|
<figureCitation id="872021723E9DEE0B10889D3202A41572" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9C, D</figureCitation>
|
|
) with ventral sclerite long, broadened and rounded distally; same in ventral view (Fig.
|
|
<figureCitation id="176D2FFDAE74CD6883F78B81B3DCC4CC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11B, C</figureCitation>
|
|
), with dorsal sclerite large, reticulate, with left-sided and right-sided protuberances distinct, and ventral sclerite narrow and distally very slightly curved to left. Right paramere as on Fig.
|
|
<figureCitation id="F9F3EC427B086E6DF794101D76693D6B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche & Saulcy, 1855), male paralectotype, " Syria ". Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12J, K</figureCitation>
|
|
.
|
|
<emphasis id="48944FC475AF4EC301803FD9EA23C43F" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
|
|
(Fig.
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<figureCitation id="4D06A1C52F6E34F1A92D102F529D962D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15A, B</figureCitation>
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). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
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</paragraph>
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</subSubSection>
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<subSubSection id="124F985D828777688D6AC58C96990E4F" pageId="0" pageNumber="71" type="notes">
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<paragraph id="20726AD7087D775FF2660CEC0690ABD7" pageId="0" pageNumber="71">Notes.</paragraph>
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<paragraph id="CAA8B757FBB16B605453BEBD097E3F41" pageId="0" pageNumber="71">
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Due to minor morphological differences, the populations from the lower course of Nile River and Southwest Israel are treated as a distinct subspecies of
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<taxonomicName id="B0FD5534214588707C17CBF6CE53C1D3" class="Insecta" kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="0A02F5D014E827B373C676F678FC6FC2" italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
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</taxonomicName>
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(see Fig.
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<figureCitation id="2EA668B60E1A8797CDA95F189A97F0B7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
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). Moreover, no verifiable records of the species are so far known from the vast area between Misrata District (Libya) and Cairo Governorate (Egypt).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |