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<document id="88A4F9559748A3C72962EA20620E9B15" ID-CLB-Dataset="29148" ID-DOI="10.11646/zootaxa.4422.2.10" ID-GBIF-Dataset="50565735-7a9a-4154-925a-5d82934ced48" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455589" ID-ZooBank="3683AE27-919D-454B-AD31-CEC3176A11D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539185449720" checkinUser="plazi" docAuthor="Roca-Cusachs, Marcos, Kim, Junggon, Lee, Hodan, Park, Haechul &amp; Jung, Sunghoon" docDate="2018" docId="981E6172D03C0664688FFCF9B559FE7E" docLanguage="en" docName="zootaxa.4422.2.10.pdf" docOrigin="Zootaxa 4422 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraholcostethus irumiae Roca-Cusachs &amp; Kim &amp; Lee &amp; Park &amp; Jung 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="299" masterDocId="6427190AD03D06676818FFA6B609FF99" masterDocTitle="A new species of the genus Paraholcostethus Belousova (Hemiptera: Heteroptera: Pentatomidae: Carpocorini) from Korea" masterLastPageNumber="300" masterPageNumber="296" pageNumber="297" updateTime="1698573689092" updateUser="ExternalLinkService">
<mods:mods id="6C473A0342E5A8C8C6500E7ED1E019E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="97E0B68573FDCE081DEDB3F0C18492B6">
<mods:title id="B2634C3CD6BE4E972BF74B925AA6D85E">A new species of the genus Paraholcostethus Belousova (Hemiptera: Heteroptera: Pentatomidae: Carpocorini) from Korea</mods:title>
</mods:titleInfo>
<mods:name id="7D966C70651146B28C87820CE2140C49" type="personal">
<mods:role id="164A7613F0851AD0E300A9B66EC8E016">
<mods:roleTerm id="120CC05D30874FA42C3016528335677E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3779932DB1415C44D50B945F89DF65C">Roca-Cusachs, Marcos</mods:namePart>
</mods:name>
<mods:name id="299934781B63AFF1E10D6BB20DA06EE1" type="personal">
<mods:role id="F2014BA6EF4507249AC3F9DDB24B1438">
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</mods:role>
<mods:namePart id="62841E611A2580669C15EC08CF060B59">Kim, Junggon</mods:namePart>
</mods:name>
<mods:name id="700BB18AF4136BE88C80482C290EF489" type="personal">
<mods:role id="5DF7CAA68A3182A965011400A89F7AC1">
<mods:roleTerm id="B27FB353E28573C5964EADB12575994F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E7BC75070B0A91FF2937E9F2E648C3E">Lee, Hodan</mods:namePart>
</mods:name>
<mods:name id="BA0221C87F82CA7A45A18F423EF5606B" type="personal">
<mods:role id="C61C0703041700FA4AAF996A1258339B">
<mods:roleTerm id="6059060E8A668462F1396CA58948933F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4888E03F308EF7E1C9CA6ECBF1BDE89C">Park, Haechul</mods:namePart>
</mods:name>
<mods:name id="F3EF5EB95351604502AA80F7C07D297E" type="personal">
<mods:role id="ECB36056A37B09DF93D99763F5F614BC">
<mods:roleTerm id="E757948ED928886F18BFE0FA78E31A29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B7B8A481F2E17E35D75718CAAB6ADB3">Jung, Sunghoon</mods:namePart>
</mods:name>
<mods:typeOfResource id="A849F3B19E1E08CBD49E00015ABF2DD9">text</mods:typeOfResource>
<mods:relatedItem id="F036428043968FB9022D6FF30688B5B7" type="host">
<mods:titleInfo id="63050A3CFFDE430466918C8306E62B43">
<mods:title id="0035E87C3910CE909D169062E2AD9A8B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B73A0D2D203DE9A82F8FD55181197706">
<mods:date id="8E42DF4FEA4B66FB18834BA35F45AC4B">2018</mods:date>
<mods:detail id="A355A4C2AB8C4B4F0C7428703D467FBD" type="volume">
<mods:number id="DF0C860DBA3D337D69FE8F90F0D52502">4422</mods:number>
</mods:detail>
<mods:detail id="2B48C1360973F6C61FEADC952C8C8F52" type="issue">
<mods:number id="2386611CE3285E96EF689441141F79FB">2</mods:number>
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<mods:start id="EE3D7568EBC1C955A2599D60E54CEFE0">296</mods:start>
<mods:end id="AAFD30C73BAE778AECC099B65014E821">300</mods:end>
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<mods:classification id="E8A41674DF59796D16A8F8A21767AA99">journal article</mods:classification>
<mods:identifier id="079D1068D805C340B3AFB9FE4FB0A76C" type="CLB-Dataset">29148</mods:identifier>
<mods:identifier id="AD4EA85C3628F7CF7935CA610937B1EA" type="DOI">10.11646/zootaxa.4422.2.10</mods:identifier>
<mods:identifier id="0791022972C20D0CA712B75E8201D13E" type="GBIF-Dataset">50565735-7a9a-4154-925a-5d82934ced48</mods:identifier>
<mods:identifier id="EC2216B19AA4C8C803D8795FF460A3DE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0EF380F699A4E4E02C1ADCD3F5F36A4C" type="Zenodo-Dep">1455589</mods:identifier>
<mods:identifier id="3D7E013CF8F366A09B90CAAA73DF0DF0" type="ZooBank">3683AE27-919D-454B-AD31-CEC3176A11D4</mods:identifier>
</mods:mods>
<treatment id="981E6172D03C0664688FFCF9B559FE7E" ID-DOI="http://doi.org/10.5281/zenodo.5966116" ID-GBIF-Taxon="148671489" ID-Zenodo-Dep="5966116" LSID="urn:lsid:plazi:treatment:981E6172D03C0664688FFCF9B559FE7E" httpUri="http://treatment.plazi.org/id/981E6172D03C0664688FFCF9B559FE7E" lastPageId="3" lastPageNumber="299" pageId="1" pageNumber="297">
<subSubSection id="58AD83EFD03C0666688FFCF9B5FAFCE3" box="[151,1011,863,890]" pageId="1" pageNumber="297" type="nomenclature">
<paragraph id="1008D064D03C0666688FFCF9B5FAFCE3" blockId="1.[151,1436,863,1996]" box="[151,1011,863,890]" pageId="1" pageNumber="297">
<heading id="4B406708D03C0666688FFCF9B5FAFCE3" bold="true" box="[151,1011,863,890]" fontSize="11" level="1" pageId="1" pageNumber="297" reason="1">
<emphasis id="22C30C76D03C0666688FFCF9B5FAFCE3" bold="true" box="[151,1011,863,890]" pageId="1" pageNumber="297">
<taxonomicName id="D7B7ABE7D03C0666688FFCF9B4E5FCE3" authority="Roca-Cusachs &amp; Jung" authorityName="Roca-Cusachs &amp; Kim &amp; Lee &amp; Park &amp; Jung" authorityYear="2018" box="[151,748,863,890]" class="Insecta" family="Pentatomidae" genus="Paraholcostethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="irumiae" status="sp. nov.">
<emphasis id="22C30C76D03C0666688FFCF9B7C7FCE0" bold="true" box="[151,462,863,889]" italics="true" pageId="1" pageNumber="297">Paraholcostethus irumiae</emphasis>
Roca-Cusachs &amp; Jung
</taxonomicName>
<taxonomicNameLabel id="39F0B10DD03C06666AECFCC6B558FCE3" box="[756,849,864,890]" pageId="1" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="888CCCE1D03C06666B79FCC6B5BAFCE3" box="[865,947,864,890]" captionStart="FIGURE 1" captionStartId="2.[151,244,1937,1959]" captionTargetBox="[228,1377,207,1866]" captionTargetId="figure@2.[184,1406,191,1919]" captionTargetPageId="2" captionText="FIGURE 1. Paraholcostethus irumiae Roсa-Cusaсhs &amp; Jung sp. nov. a, Dorsal habitus; b, Ventral habitus; c–e, Ventral, dorsal and сaudal view of male pygophore; f–h, Lateral, dorsal and сaudal view of aedeagus; i–k, Right paramere; l, Female genital plates; m, Postero-ventral line drawing view of female genital plates. gc, gonoсoxites; la, laterotergites. Sсale bar equals 1 mm." httpUri="https://zenodo.org/record/1455591/files/figure.png" pageId="1" pageNumber="297">Figs. 1</figureCitation>
&amp;
<figureCitation id="888CCCE1D03C06666BC3FCC6B5E2FCE3" box="[987,1003,864,890]" captionStart="FIGURE 2" captionStartId="3.[151,244,996,1018]" captionTargetBox="[151,1379,538,974]" captionTargetId="figure@3.[151,1436,538,975]" captionTargetPageId="3" captionText="FIGURE 2. Internal female genitalia of Paraholcostethus irumiae sp. nov. a, Staсked photograph of spermatheсa; b, line drawing of spermatheсa; c, whole internal genitalia; aaf, anterior annular flange; cs, сapsula seminalis; csp, сapsula seminalis lateral proсesses; paf, posterior annular flange; pi, pars intermedialis; Sсale bar equals 1 mm." httpUri="https://zenodo.org/record/1455593/files/figure.png" pageId="1" pageNumber="297">2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58AD83EFD03C0666688FFC24B35EFBBD" pageId="1" pageNumber="297" type="materials_examined">
<paragraph id="1008D064D03C0666688FFC24B45EFB98" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">
<materialsCitation id="A0DFDA39D03C0666688FFC24B573FC25" ID-GBIF-Occurrence="1915497946" collectingDate="2015-07-09" collectionCode="CNU" collectorName="Deposited" country="South Korea" county="Gojeong-ri" latitude="37.25625" location="Hwaseong-si" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" pageId="1" pageNumber="297" specimenCount="1" stateProvince="Gyeonggi-do" typeStatus="holotype">
<emphasis id="22C30C76D03C0666688FFC24B74FFC00" bold="true" box="[151,326,898,921]" pageId="1" pageNumber="297">
<typeStatus id="CF0C6EC6D03C0666688FFC24B6C6FC00" box="[151,207,898,921]" pageId="1" pageNumber="297">Type</typeStatus>
material:
</emphasis>
<typeStatus id="CF0C6EC6D03C0666694CFC24B7B2FC00" box="[340,443,898,921]" pageId="1" pageNumber="297" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="68A090F4D03C066669D5FC24B425FC00" box="[461,556,898,921]" name="South Korea" pageId="1" pageNumber="297">KOREA</collectingCountry>
.
<specimenCount id="06B11BEDD03C06666A26FC24B463FC00" box="[574,618,898,921]" pageId="1" pageNumber="297" type="generic">1M.</specimenCount>
<collectingDate id="744D0F4CD03C06666A60FC24B505FC00" box="[632,780,898,921]" pageId="1" pageNumber="297" value="2015-07-09">
<date id="6409F6A4D03C06666A60FC24B505FC00" box="[632,780,898,921]" pageId="1" pageNumber="297" value="2015-07-09">09.VII. 2015</date>
</collectingDate>
.
<collectingCounty id="F969A8E8D03C06666B06FC24B59FFC00" box="[798,918,898,921]" pageId="1" pageNumber="297">Gojeong-ri</collectingCounty>
,
<collectingMunicipality id="F06C4A1ED03C06666BBFFC24B25CFC00" box="[935,1109,898,921]" pageId="1" pageNumber="297">Songsan-myeon</collectingMunicipality>
,
<location id="156886BFD03C06666C70FC24B2FFFC00" LSID="urn:lsid:plazi:treatment:981E6172D03C0664688FFCF9B559FE7E:156886BFD03C06666C70FC24B2FFFC00" box="[1128,1270,898,921]" country="South Korea" county="Gojeong-ri" latitude="37.25625" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" name="Hwaseong-si" pageId="1" pageNumber="297" stateProvince="Gyeonggi-do">Hwaseong-si</location>
,
<collectingRegion id="D2731E86D03C06666D10FC24B391FC00" box="[1288,1432,898,921]" country="South Korea" name="Gyeonggi" pageId="1" pageNumber="297">Gyeonggi-do</collectingRegion>
.
<geoCoordinate id="7583B6A3D03C0666688FFC03B721FC25" box="[151,296,933,956]" direction="north" orientation="latitude" pageId="1" pageNumber="297" precision="1" value="37.25625">37°15'22.5&quot;N</geoCoordinate>
<geoCoordinate id="7583B6A3D03C0666692EFC03B7DCFC25" box="[310,469,933,956]" direction="east" orientation="longitude" pageId="1" pageNumber="297" precision="1" value="126.74591">126°44'45.3&quot;E</geoCoordinate>
. Hodan Lee leg.
<collectorName id="BD42B5B2D03C06666ABCFC03B519FC25" box="[676,784,933,956]" pageId="1" pageNumber="297">Deposited</collectorName>
at
<collectionCode id="76A648A1D03C06666B26FC03B573FC25" box="[830,890,933,956]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="1" pageNumber="297">CNU</collectionCode>
</materialsCitation>
. Paratypes: 2M, 3F,
<materialsCitation id="A0DFDA39D03C06666C76FC03B45AFC46" ID-GBIF-Occurrence="1915497944" collectingDate="2010-07-21" collectionCode="CNU" collectorName="Deposited" country="South Korea" county="Gojeong-ri" latitude="37.25625" location="Deposited" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" pageId="1" pageNumber="297" specimenCount="1" stateProvince="Gyeonggi-do" typeStatus="holotype">
same data as in
<typeStatus id="CF0C6EC6D03C06666D2AFC03B390FC25" box="[1330,1433,933,956]" pageId="1" pageNumber="297" type="holotype">Holotype</typeStatus>
.
<location id="156886BFD03C0666688FFC6EB70AFC46" LSID="urn:lsid:plazi:treatment:981E6172D03C0664688FFCF9B559FE7E:156886BFD03C0666688FFC6EB70AFC46" box="[151,259,968,991]" name="Deposited" pageId="1" pageNumber="297">Deposited</location>
at
<collectionCode id="76A648A1D03C06666930FC6EB76DFC46" box="[296,356,968,991]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="1" pageNumber="297">CNU</collectionCode>
; 2F,
<specimenCount id="06B11BEDD03C06666982FC6EB7CFFC46" box="[410,454,968,991]" pageId="1" pageNumber="297" type="generic">1M.</specimenCount>
<collectingDate id="744D0F4CD03C066669D7FC6EB45AFC46" box="[463,595,968,991]" pageId="1" pageNumber="297" value="2010-07-21">
<date id="6409F6A4D03C066669D7FC6EB45AFC46" box="[463,595,968,991]" pageId="1" pageNumber="297" value="2010-07-21">21.VII.2010</date>
</collectingDate>
</materialsCitation>
,
<materialsCitation id="A0DFDA39D03C06666A78FC6EB351FC46" ID-GBIF-Occurrence="1915497945" box="[608,1368,968,991]" collectingDate="2010-07-21" collectionCode="NAAS" collectorName="Deposited" country="South Korea" county="Gojeong-ri" latitude="37.25625" location="Hwaseong-si" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" pageId="1" pageNumber="297" specimenCount="6" stateProvince="Gyeonggi-do" typeStatus="holotype">
same locality as Holotype. Deposited at
<collectionCode id="76A648A1D03C06666C04FC6EB261FC46" box="[1052,1128,968,991]" pageId="1" pageNumber="297">NAAS</collectionCode>
; 1F,
<specimenCount id="06B11BEDD03C06666C87FC6EB2C2FC46" box="[1183,1227,968,991]" pageId="1" pageNumber="297" type="generic">6M.</specimenCount>
<collectingDate id="744D0F4CD03C06666CCBFC6EB351FC46" box="[1235,1368,968,991]" pageId="1" pageNumber="297" value="2010-07-21">
<date id="6409F6A4D03C06666CCBFC6EB351FC46" box="[1235,1368,968,991]" pageId="1" pageNumber="297" value="2010-07-21">21.VII.2010</date>
</collectingDate>
</materialsCitation>
,
<materialsCitation id="A0DFDA39D03C06666D7DFC6EB45EFB98" ID-GBIF-Occurrence="1915497947" collectingDate="2015-07-09" collectionCode="NAAS" collectorName="Deposited" country="South Korea" county="Gojeong-ri" latitude="37.25625" location="Hwaseong-si" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" pageId="1" pageNumber="297" specimenCount="1" stateProvince="Gyeonggi-do" typeStatus="holotype">
same locality as Holotype. Deposited at
<collectionCode id="76A648A1D03C06666A1FFC4CB45AFB98" box="[519,595,1002,1025]" pageId="1" pageNumber="297">NAAS</collectionCode>
;
</materialsCitation>
</paragraph>
<paragraph id="1008D064D03C066668DFFBABB35EFBBD" blockId="1.[151,1436,863,1996]" box="[199,1367,1037,1060]" pageId="1" pageNumber="297">
<materialsCitation id="A0DFDA39D03C066668DFFBABB35EFBBD" ID-GBIF-Occurrence="1915497948" box="[199,1367,1037,1060]" country="South Korea" county="Gojeong-ri" latitude="37.25625" location="Hwaseong-si" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" pageId="1" pageNumber="297" specimenCount="1" stateProvince="Gyeonggi-do" typeStatus="holotype">
<emphasis id="22C30C76D03C066668DFFBABB768FBBD" bold="true" box="[199,353,1037,1060]" pageId="1" pageNumber="297">
<typeStatus id="CF0C6EC6D03C066668DFFBABB6F6FBBD" box="[199,255,1037,1060]" pageId="1" pageNumber="297">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="68A090F4D03C06666970FBABB7A5FBBD" box="[360,428,1037,1060]" name="South Korea" pageId="1" pageNumber="297">Korea</collectingCountry>
,
<collectingCounty id="F969A8E8D03C066669AEFBABB427FBBD" box="[438,558,1037,1060]" pageId="1" pageNumber="297">Gojeong-ri</collectingCounty>
,
<collectingMunicipality id="F06C4A1ED03C06666A20FBABB4EEFBBD" box="[568,743,1037,1060]" pageId="1" pageNumber="297">Songsan-myeon</collectingMunicipality>
,
<location id="156886BFD03C06666AE9FBABB576FBBD" LSID="urn:lsid:plazi:treatment:981E6172D03C0664688FFCF9B559FE7E:156886BFD03C06666AE9FBABB576FBBD" box="[753,895,1037,1060]" country="South Korea" county="Gojeong-ri" latitude="37.25625" longLatPrecision="1" longitude="126.74591" municipality="Songsan-myeon" name="Hwaseong-si" pageId="1" pageNumber="297" stateProvince="Gyeonggi-do">Hwaseong-si</location>
,
<collectingRegion id="D2731E86D03C06666B92FBABB213FBBD" box="[906,1050,1037,1060]" country="South Korea" name="Gyeonggi" pageId="1" pageNumber="297">Gyeonggi-do</collectingRegion>
.
<geoCoordinate id="7583B6A3D03C06666C3CFBABB2BCFBBD" box="[1060,1205,1037,1060]" direction="north" orientation="latitude" pageId="1" pageNumber="297" precision="1" value="37.25625">37°15'22.5&quot;N</geoCoordinate>
<geoCoordinate id="7583B6A3D03C06666CA4FBABB35EFBBD" box="[1212,1367,1037,1060]" direction="east" orientation="longitude" pageId="1" pageNumber="297" precision="1" value="126.74591">126°44'45.3&quot;E</geoCoordinate>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="58AD83EFD03C066668DFFB96B294FB36" pageId="1" pageNumber="297" type="diagnosis">
<paragraph id="1008D064D03C066668DFFB96B294FB36" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">
<emphasis id="22C30C76D03C066668DFFB96B734FBDE" bold="true" box="[199,317,1072,1095]" pageId="1" pageNumber="297">Diagnosis:</emphasis>
Differentiated from
<taxonomicName id="D7B7ABE7D03C06666A36FB96B4BDFBDE" authorityName="Horvath" authorityYear="1897" box="[558,692,1071,1095]" class="Insecta" family="Pentatomidae" genus="Peribalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="22C30C76D03C06666A36FB96B4BDFBDE" box="[558,692,1071,1095]" italics="true" pageId="1" pageNumber="297">P. breviceps</emphasis>
</taxonomicName>
by males having the lateral process of conjunctive processes of aedeagus with a great extension of the sclerotized area, at the apices but not bifurcate. Females having laterotergite IX not surpassing outline of laterotergite VIII, clearly surpassing in
<taxonomicName id="D7B7ABE7D03C06666B5EFBD3B5CCFB15" authorityName="Horvath" authorityYear="1897" box="[838,965,1140,1164]" class="Insecta" family="Pentatomidae" genus="Peribalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="22C30C76D03C06666B5EFBD3B5CCFB15" box="[838,965,1140,1164]" italics="true" pageId="1" pageNumber="297">P. breviceps</emphasis>
</taxonomicName>
; gonocoxite IX concave at base and slightly curved apex. Posterior external margin of gonocoxites slightly angulate, rounded in
<taxonomicName id="D7B7ABE7D03C06666C0FFB3EB29FFB36" authorityName="Horvath" authorityYear="1897" box="[1047,1174,1175,1199]" class="Insecta" family="Pentatomidae" genus="Peribalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="22C30C76D03C06666C0FFB3EB29FFB36" box="[1047,1174,1175,1199]" italics="true" pageId="1" pageNumber="297">P. breviceps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="58AD83EFD03C066468DFFB1CB7A2FEC5" lastPageId="3" lastPageNumber="299" pageId="1" pageNumber="297" type="description">
<paragraph id="1008D064D03C066668DFFB1CB6E6FAA0" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">
<emphasis id="22C30C76D03C066668DFFB1CB75DFB48" bold="true" box="[199,340,1210,1233]" pageId="1" pageNumber="297">Description:</emphasis>
Total body length (8.29–8.76, 8.34–9.68), maximum abdominal width (5.01–5.24, 5.06–5.39). Ventral and dorsal sides of body sandy yellowish, with black punctuation (distance between punctures exceeding puncture diameter). Dorsal surface of abdomen dark brown, ventral surface of abdomen yellowish, with black punctuation (figs. 1a &amp; b).
</paragraph>
<paragraph id="1008D064D03C066668DFFAE3B4E2F990" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">Head: Length (1.68–1.71, 1.84–1.96), width (2.26–2.29, 2.29–2.41). Eyes slightly protruding beyond head profile. Length of head anterior to eyes (0.92–1.03, 0.91–0.96); interocular distance (1.48–1.52, 1.53–1.67). Juga and clypeus of variable proportions, juga never enclosing clypeus. Rostrum just surpassing metacoxae Length of antennal segment I (0.28–0.39, 0.31–0.40), II (0.51–0.59, 0.54–0.56), III (0.74–1.85, 0.75–0.78), IV (0.84–1.05, 0.92–0.96), V (1.12–1.29, 1.03–1.06), segments I, II, and III concolourous with general coloration, segments IV and V bicolored, contrasting; black with base concolourous with general habitus coloration.</paragraph>
<paragraph id="1008D064D03C066668DFF9B3B540F92E" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">Pronotum: Straight sides bordered with whitish-yellow costa, slightly deflexed upwards, pronotal length at median (1.81–2.04, 1.96–2.17), maximum pronotal width (5.06–5.12, 5.02–5.39). Lateral angles widely rounded, almost not projecting beyond hemelytral base. Scutellum slightly longer (3.20–3.37, 3.44–3.63) than wide (3.12–2.19, 3.14–3.34); Scutellum paler at apex. Hemelytra concolourous with body, membrane of hemelytra light brown, slightly darker than dorsal coloration, translucent. Membrane veins scarcely marked.</paragraph>
<paragraph id="1008D064D03C066668DFF964B25CF886" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">Abdomen: dorsally, disc of connexivum yellow, punctures concolourous, no contrast on anterior and posterior margins. Spiracles rounded, black. Dorsal abdominal tergites dark brown, densely scattered with piliferous tubercles. Legs concolourous with ventral surface of thorax; femora with black punctured markings.</paragraph>
<paragraph id="1008D064D03C066668DFF88CB4D4F855" blockId="1.[151,1436,863,1996]" pageId="1" pageNumber="297">
Male genitalia: Pygophore (fig. 1c, d &amp; e) rectangular to trapezoidal; genital cup well exposed dorsally; posterolateral angles of pygophore angulate, blunt, not produced; ventral rim sinuate with a double depression medially; dorsal rim concave, broadly U-shaped. Aedeagus (figs. 1f–h) with 3 pairs of conjunctival processes. Lateral process of aedeagus not bifurcate, sinuous and sclerotized. Paramere narrow, straight; hypophysis flat, with acute apex, triangulate with rounded edges, spherical lobe at base (figs.
<date id="6409F6A4D03C06666A86F813B4B9F855" box="[670,688,1973,1996]" pageId="1" pageNumber="297">1i</date>
–k).
</paragraph>
<caption id="44C880ECD03F0665688FF837B2F0F878" httpUri="https://zenodo.org/record/1455591/files/figure.png" pageId="2" pageNumber="298" startId="2.[151,244,1937,1959]" targetBox="[228,1377,207,1866]" targetPageId="2">
<paragraph id="1008D064D03F0665688FF837B2F0F878" blockId="2.[151,1436,1937,2017]" pageId="2" pageNumber="298">
<emphasis id="22C30C76D03F0665688FF837B707F83E" bold="true" box="[151,270,1937,1959]" pageId="2" pageNumber="298">FIGURE 1.</emphasis>
<taxonomicName id="D7B7ABE7D03F0665690CF837B407F83F" authorityName="Roca-Cusachs &amp; Kim &amp; Lee &amp; Park &amp; Jung" authorityYear="2018" box="[276,526,1937,1958]" class="Insecta" family="Pentatomidae" genus="Paraholcostethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="irumiae">
<emphasis id="22C30C76D03F0665690CF837B407F83F" box="[276,526,1937,1958]" italics="true" pageId="2" pageNumber="298">Paraholcostethus irumiae</emphasis>
</taxonomicName>
Roсa-Cusaсhs &amp; Jung
<emphasis id="22C30C76D03F06656AECF837B55CF83E" bold="true" box="[756,853,1937,1959]" pageId="2" pageNumber="298">sp. nov. a</emphasis>
, Dorsal habitus;
<emphasis id="22C30C76D03F06656BE4F837B200F83E" bold="true" box="[1020,1033,1937,1959]" pageId="2" pageNumber="298">b</emphasis>
, Ventral habitus;
<emphasis id="22C30C76D03F06656CAEF837B2DEF83E" bold="true" box="[1206,1239,1937,1959]" pageId="2" pageNumber="298">c–e</emphasis>
, Ventral, dorsal and сaudal view of male pygophore;
<emphasis id="22C30C76D03F066569C0F808B7F0F85D" bold="true" box="[472,505,1966,1988]" pageId="2" pageNumber="298">f–h</emphasis>
, Lateral, dorsal and сaudal view of aedeagus;
<emphasis id="22C30C76D03F06656BD9F808B5E9F85D" bold="true" box="[961,992,1966,1988]" pageId="2" pageNumber="298">i–k</emphasis>
, Right paramere;
<emphasis id="22C30C76D03F06656C8AF808B290F85D" bold="true" box="[1170,1177,1966,1988]" pageId="2" pageNumber="298">l</emphasis>
, Female genital plates;
<emphasis id="22C30C76D03F06656D9AF808B395F85D" bold="true" box="[1410,1436,1966,1988]" pageId="2" pageNumber="298">m,</emphasis>
Postero-ventral line drawing view of female genital plates.
<emphasis id="22C30C76D03F06656AD6F86DB4EDF878" bold="true" box="[718,740,1995,2017]" pageId="2" pageNumber="298">gc</emphasis>
, gonoсoxites;
<emphasis id="22C30C76D03F06656B6AF86DB58CF878" bold="true" box="[882,901,1995,2017]" pageId="2" pageNumber="298">la</emphasis>
, laterotergites. Sсale bar equals 1 mm.
</paragraph>
</caption>
<paragraph id="1008D064D03E066468DFFF3EB7A2FEC5" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="299">Female genitalia (figs. 1l, m, 2): Laterotergite IX widened at apex, spatulate, short, not surpassing outline of laterotergite VIII; gonocoxite IX wide, trapezoidal, with concave base and slightly curved apex. Posterior internal margins of gonocoxites VIII rounded, external margins slightly angulate. Laterotergite VIII smooth, somewhat triangular. Spermatheca (figs. 2a, b) with small spherical bulb bearing 3 processes, two reaching anterior annular flange (aaf), one slightly surpassing posterior annular flange (paf), long sclerotized duct narrowed toward base, ductus receptaculi short (fig. 2c).</paragraph>
</subSubSection>
<subSubSection id="58AD83EFD03E066468DFFECEB4D0FEE6" box="[199,729,360,383]" pageId="3" pageNumber="299" type="distribution">
<paragraph id="1008D064D03E066468DFFECEB4D0FEE6" blockId="3.[151,1436,152,487]" box="[199,729,360,383]" pageId="3" pageNumber="299">
<emphasis id="22C30C76D03E066468DFFECEB752FEE6" bold="true" box="[199,347,360,383]" pageId="3" pageNumber="299">Distribution:</emphasis>
Known only from the
<typeStatus id="CF0C6EC6D03E06646A56FECEB475FEE6" box="[590,636,360,383]" pageId="3" pageNumber="299">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="58AD83EFD03E066468DFFE2CB559FE7E" pageId="3" pageNumber="299" type="etymology">
<paragraph id="1008D064D03E066468DFFE2CB559FE7E" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="299">
<emphasis id="22C30C76D03E066468DFFE2CB742FE38" bold="true" box="[199,331,394,417]" pageId="3" pageNumber="299">Etymology:</emphasis>
This species is named after the recently born daughter of two members of the laboratory of systematic entomology (CNU), Hodan Lee and Sumin Oh, for her help and contributions. Their daughter’s name, Rumi Lee, is pronounced as Irumi in Korean Language. Noun in genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>
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