220 lines
22 KiB
XML
220 lines
22 KiB
XML
<document id="0DDF5E93684ED36C4153ACA3C7A07CBE" ID-CLB-Dataset="34534" ID-DOI="http://dx.doi.org/10.3897/zookeys.661.11224" ID-GBIF-Dataset="a52331ef-0444-4ca9-83cc-d0187677e7a4" ID-PMC="PMC5539368" ID-Pensoft-Pub="1313-2970-661-91" ID-PubMed="28769604" ID-ZooBank="7CBF7FB1EFEA4DC18F645BF862694AA0" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-661-91" ModsDocOrigin="ZooKeys 661" ModsDocTitle="Molecular phylogeny of Atractus (Serpentes, Dipsadidae), with emphasis on Ecuadorian species and the description of three new taxa" checkinTime="1489657683944" checkinUser="pensoft" docAuthor="Arteaga, Alejandro, Mebert, Konrad, Valencia, Jorge H., Cisneros-Heredia, Diego F., Penafiel, Nicolas, Reyes-Puig, Carolina, Vieira-Fernandes, Jose L. & Guayasamin, Juan M." docDate="2017" docId="1DA5E4AF8E2ABD4A028D99E76E6F74E5" docLanguage="en" docName="ZooKeys 661: 91-123" docOrigin="ZooKeys 661" docSource="http://dx.doi.org/10.3897/zookeys.661.11224" docTitle="Atractus pyroni Arteaga, Mebert, Valencia, Cisneros-Heredia, Penafiel, Reyes-Puig, Vieira-Fernandes & Guayasamin, 2017, sp. n." docType="treatment" docUuid="36145E29-02B6-4C66-A097-44EFC1BC3A92" docUuidSource="ZooBank" docVersion="7" lastPageNumber="106" masterDocId="812DFFA6EF6EFFB9AD1EFFB9570BFF9A" masterDocTitle="Molecular phylogeny of Atractus (Serpentes, Dipsadidae), with emphasis on Ecuadorian species and the description of three new taxa" masterLastPageNumber="123" masterPageNumber="91" pageNumber="105" updateTime="1701375718023" updateUser="ExternalLinkService">
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<mods:titleInfo id="5F59E2D5F36EF27353C8134B9F2947B8">
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<mods:title id="09BBDC0E1D8BE39787ECF4D9DA0C32D0">Molecular phylogeny of Atractus (Serpentes, Dipsadidae), with emphasis on Ecuadorian species and the description of three new taxa</mods:title>
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<mods:namePart id="B0FC74647D2D2F26A77DC00AC9DDAC92">Arteaga, Alejandro</mods:namePart>
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<mods:namePart id="16434AEDCBC9FC9B939BA40EC033A3D7">Mebert, Konrad</mods:namePart>
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<mods:namePart id="8E2621EF8AAEA0463E9BC73732E0CFA4">Valencia, Jorge H.</mods:namePart>
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<mods:name id="3A97808436467C711B7D9FDBF1EA71DD" type="personal">
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<mods:namePart id="AAA84B1A2C6BCA6DBBCB29C12E5DF34B">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:roleTerm id="63B3B2E387B08F5B56C73E37C0057453">Author</mods:roleTerm>
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<mods:namePart id="E4E8D44CA2D6CA26923D953D81EC34E1">Penafiel, Nicolas</mods:namePart>
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<mods:name id="28C84E001CB36B8F12E0F675F8D7D0FB" type="personal">
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<mods:namePart id="07D9E162160F4533E09E88FBA4B838BE">Reyes-Puig, Carolina</mods:namePart>
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<mods:namePart id="DBEFB2FC447FE4D9D203A373D3FA81F3">Vieira-Fernandes, Jose L.</mods:namePart>
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<mods:namePart id="93D3DA4A0BDD77BDE3C108B1EA1997C2">Guayasamin, Juan M.</mods:namePart>
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<mods:typeOfResource id="1BF5D52C0FCBB09D641D556A59936094">text</mods:typeOfResource>
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<mods:title id="7A1519D79B2C08BB7C3C45BC6ECE701F">ZooKeys</mods:title>
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<mods:part id="4B934520409FF3EA07F517DF98086A8A">
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<mods:date id="E128672653F9E1953DACA0709E9296B7">2017</mods:date>
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<mods:detail id="8B29130E9F3248448A946E2C0F391D6F" type="volume">
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<mods:number id="5FDE734FCDC16AC352B6310460439F24">661</mods:number>
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<mods:start id="48D72928F924422BD1D9F32FFC217ADB">91</mods:start>
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<mods:end id="C592E3813E07F3413D0EEBDF36CFA946">123</mods:end>
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<mods:url id="0961C3F97FE16CDA937B04085FD1B37A">http://dx.doi.org/10.3897/zookeys.661.11224</mods:url>
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<mods:classification id="5DC2A3642E248DB048830776B3872182">journal article</mods:classification>
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<mods:identifier id="701BF884D729AF98AA67CF0BDAB84527" type="DOI">http://dx.doi.org/10.3897/zookeys.661.11224</mods:identifier>
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<mods:identifier id="EECD442328AE502A510D087E9820E82D" type="Pensoft-Pub">1313-2970-661-91</mods:identifier>
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<mods:identifier id="0F71DF86BB016CC0DDB0F65D2BBC66F0" type="ZooBank">7CBF7FB1EFEA4DC18F645BF862694AA0</mods:identifier>
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<treatment id="1DA5E4AF8E2ABD4A028D99E76E6F74E5" ID-GBIF-Taxon="127883985" LSID="urn:lsid:zoobank.org:act:36145E29-02B6-4C66-A097-44EFC1BC3A92" httpUri="http://treatment.plazi.org/id/1DA5E4AF8E2ABD4A028D99E76E6F74E5" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105">
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<subSubSection id="DDEB110429F46F124B34CC5F8DA5699A" pageId="14" pageNumber="105" type="nomenclature">
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<paragraph id="09DDC3806086DD42C948A1546408F73C" pageId="14" pageNumber="105">
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<taxonomicName id="32F8E5F3C8EC40CB22B3E2BE2F79380F" LSID="http://zoobank.org/36145E29-02B6-4C66-A097-44EFC1BC3A92" class="Squamata" family="Dipsadidae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" order="Squamata" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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<taxonomicNameLabel id="4C3AF9ED014CBA020D4D0C758107C86E" pageId="14" pageNumber="105">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="68FA62E2733698A5AE3F02E08C00B14D" pageId="14" pageNumber="105" type="proposed standard english name">
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<paragraph id="6DFB85EBA4D8406A65742EFC6CBD7C73" pageId="14" pageNumber="105">Proposed standard English name.</paragraph>
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<paragraph id="12885ACA55E38423F4210C8B1A807B13" pageId="14" pageNumber="105">
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<normalizedToken id="5F2BAF6777B720F928F5C07BED2A8D69" originalValue="Pyron’s">Pyron's</normalizedToken>
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Groundsnake
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</paragraph>
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</subSubSection>
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<subSubSection id="79B4F35DF6AA9F5C6CEAE6A2C1779AC5" pageId="14" pageNumber="105" type="proposed standard spanish name">
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<paragraph id="30B8820A63D9603B89823C24C7D97C9E" pageId="14" pageNumber="105">Proposed standard Spanish name.</paragraph>
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<paragraph id="C2150C8A3F35E014E98F79278C82CB78" pageId="14" pageNumber="105">Tierrera de Pyron</paragraph>
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</subSubSection>
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<subSubSection id="967DAC192E56BB9813D306304816E24E" pageId="14" pageNumber="105" type="holotype">
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<paragraph id="1C7A7C0E8871FBAD5299EE61B9A513CA" pageId="14" pageNumber="105">Holotype.</paragraph>
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<paragraph id="6FE097D03C171FE7C4689D7C87D423EF" pageId="14" pageNumber="105">
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MZUTI 5107 (Fig. 7), adult male collected by
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<normalizedToken id="E1078AF93F2E467E9AF0354C32D67F29" originalValue="José">Jose</normalizedToken>
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L. Vieira-Fernandes and Carlos
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<normalizedToken id="9308D1DE3DB1A879D4E2874F39DC735A" originalValue="Durán">Duran</normalizedToken>
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on May 23, 2016 between Balzapamba and
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<normalizedToken id="B1ACEA1192C9C4ED832F535B5E7B0575" originalValue="Bilován">Bilovan</normalizedToken>
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, province of
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<normalizedToken id="7823DB7DDB2370182EFD8015154DD003" originalValue="Bolívar">Bolivar</normalizedToken>
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, Ecuador (S1.83601, W79.13322; 2026 m).
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</paragraph>
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<caption id="F9FEC938F609FBA639F076DB7F8DC4E8" pageId="14" pageNumber="105">
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<paragraph id="C395C84ED20256DABA4A463A192E27FC" pageId="14" pageNumber="105">
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Figure 7. Adult female holotype of
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<taxonomicName id="71902D701E8D8365C674CB76ED41F3A8" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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. MZUTI 5107. Scale = 1 cm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="1A731DC75C016095989435A1695C0D26" pageId="14" pageNumber="105" type="diagnosis">
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<paragraph id="9B168A655748D14BA872731F6DA07D64" pageId="14" pageNumber="105">Diagnosis.</paragraph>
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<paragraph id="AC6E1C522D06B1E941F865E2AA721FE9" pageId="14" pageNumber="105">
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<taxonomicName id="409ECED703DE734A6F8BDC599F208848" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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is placed in the genus
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<taxonomicName id="3D379C223F976EB46C6B59442479A2D0" class="Insecta" family="Coreidae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus" order="Hemiptera" pageId="14" pageNumber="105" phylum="Arthropoda" rank="genus">Atractus</taxonomicName>
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as diagnosed by
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<bibRefCitation id="D23175710AE675076EDBD7FBEB39899D" author="Savage, JM" journalOrPublisher="Miscellaneous Publications, Museum of Zoology, University of Michigan" pageId="21" pageNumber="112" pagination="1 - 86" title="A revision of the Ecuadorian snakes of the colubrid genus Atractus." volume="112" year="1960">Savage (1960)</bibRefCitation>
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, based on phylogenetic (Fig. 1) and morphological (Table 3) evidence. It is included in the
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<taxonomicName id="5ADF4B4E7F072BF38BA69403D3AF5A6B" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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group due to its 15/15/15 dorsal scale rows and its phylogenetic position (Fig. 1). The species is diagnosed based on the following combination of characters: (1) 15/15/15 smooth dorsals; (2) one postocular; (3) loreal long; (4) temporals 1+2; (5) six supralabials, third and fourth contacting orbit; (6) five infralabials, first four contacting chinshields (7) eight maxillary teeth; (8) 3 gular scale rows; (9) 2 preventrals; (10) 143 ventrals; (11) 16 subcaudals; (12) dorsal ground color dark brown with a series of light golden brown paravertebral scales running along the entire dorsum (Fig. 7); (13) venter dark brown with scattered scales of a lighter color; (14) 443 mm SVL; (15) 34 mm TL.
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</paragraph>
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</subSubSection>
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<subSubSection id="E53E008DB44110B5B5C1AB3334B8BB43" pageId="14" pageNumber="105" type="comparisons">
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<paragraph id="F318C28202AB1033106E815BB07EEC5B" pageId="14" pageNumber="105">Comparisons.</paragraph>
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<paragraph id="821A35C29A66746021C172AB4ED8196C" pageId="14" pageNumber="105">
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<taxonomicName id="4CB9DDC241B688CEC4AF9CAB01691E11" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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is compared to members of the
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<taxonomicName id="1148B7757DCEADDC90CE7802078ED0B6" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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species group:
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<taxonomicName id="676E8A80EEDA77ED58832656EBE6E453" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus carrioni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="carrioni">Atractus carrioni</taxonomicName>
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,
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<taxonomicName id="CA5C9D8BA26E12241584125691061BE9" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus lehmanni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="lehmanni">Atractus lehmanni</taxonomicName>
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, and
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<taxonomicName id="EE2F7940431D2E6C92BF3B9725163680" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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(Fig. 2). From
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<taxonomicName id="C55C224DCFA0732555E68025197F0FC9" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus carrioni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="carrioni">Atractus carrioni</taxonomicName>
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, it differs by having a loreal. From
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<taxonomicName id="8C34402516575DDAAF9F275A8773524C" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus lehmanni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="lehmanni">Atractus lehmanni</taxonomicName>
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and
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<taxonomicName id="0D9B1B88583235FB855CF8FFC45EB618" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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, it differs in size and color pattern.
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<taxonomicName id="93D39FCAB604EA9AA7D06BA7293906F2" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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is 443 mm in SVL; whereas
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<taxonomicName id="3876C9E44B2FF62CC90DB5564FC36D12" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus lehmanni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="lehmanni">Atractus lehmanni</taxonomicName>
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is 262-321 in SVL, and
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<taxonomicName id="8692304318BDA34E6545C969489BB359" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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is 230-396. Both
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<taxonomicName id="A1E58E6223D94D8E312F8EC08D428613" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus lehmanni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="lehmanni">Atractus lehmanni</taxonomicName>
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and
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<taxonomicName id="66437F226AAC7939E10ED932B0FBE53A" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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have uniform dorsal ground color, whereas
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<taxonomicName id="3217D0E9929EDD2C5B61F13F1397453D" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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has a distinct dorsal bicolored pattern (Fig. 7). Finally, in life,
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<taxonomicName id="4F151E8472FFB9A8AA30B507191D194A" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
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is darker than the remaining members of the
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<taxonomicName id="2C0297CADF0E172C3E039D88A95CC21F" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus roulei" pageId="14" pageNumber="105" phylum="Chordata" rank="species" species="roulei">Atractus roulei</taxonomicName>
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species group and has a ventral pattern that, instead of having fine speckles, has conspicuous scattered blotches of a contrasting color.
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</paragraph>
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</subSubSection>
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<subSubSection id="DAAC250E5265C8545578331FFA5F1C5B" pageId="14" pageNumber="105" type="color pattern">
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<paragraph id="1896169A41041CD1E852D27994064672" pageId="14" pageNumber="105">Color pattern.</paragraph>
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<paragraph id="B5531B425F63EAC67AA9B1B508050E61" pageId="14" pageNumber="105">The dorsal ground color is blackish with a dark vertebral (mid-dorsal) scale row flanked by a dark yellow scale row on either side (the 7th dorsal scale row), irregularly adjoined by one to few additional yellow scales on the 6th dorsal scale row, rendering an appearance of an irregularly edged mid-dorsal striped pattern (Fig. 7). The dorsal and lateral surfaces of the head are dark grayish brown and the labials are dark mustard yellow. All ventral surfaces are glossy grayish black except for the throat and some scattered blotches, which are dark mustard yellow.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="84FD504462D049B79C85BC48E8C22958" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105" type="description">
|
||
<paragraph id="6B6F06088D1968FB22F7850106C6CAC4" pageId="14" pageNumber="105">Description of holotype.</paragraph>
|
||
<paragraph id="D15FA80292763A97136D4308AF116366" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105">
|
||
Adult female, SVL 443 mm, tail length 34 mm (7.7% SVL); body diameter 11.6 mm; head length 14.4 mm (3.3% SVL); head width 9.8 mm
|
||
<pageBreakToken id="CC389EFA1A7EDACE565331AB60EF8D85" pageId="15" pageNumber="106" start="start">(</pageBreakToken>
|
||
2.2% SVL); interocular distance 5.1 mm; head slightly distinct from body;
|
||
<normalizedToken id="752D1DE9FCDF0EA636859F9D81C1F578" originalValue="snout–orbit">snout-orbit</normalizedToken>
|
||
distance 5.7 mm; rostral 2.8 mm wide, about two times broader than high; internasals 1.5 mm wide; internasal suture sinistral relative to prefrontal suture; prefrontals 2.8 mm wide; frontal 3.5 mm wide, with a curvilinear triangle shape in dorsal view; parietals 4.0 mm wide, about twice as long as wide; nasal divided; loreal 3.7 mm long, about 3 times longer than high; eye diameter 1.8 mm; pupil round; supraoculars 2.1 mm wide; one postocular; temporals 1+2, upper posterior temporal elongate, about five times longer than high, and twice as long as first temporal; six supralabials, 3
|
||
<normalizedToken id="7A701A4568B9E4AE172002E703E3D81C" originalValue="rd–">rd-</normalizedToken>
|
||
4th contacting orbit; symphisial 2.4 mm wide, separated from chin shields by first pair of infralabials; five infralabials, 1
|
||
<normalizedToken id="7E415837496D382D6E2792FDD54B637B" originalValue="st–">st-</normalizedToken>
|
||
4th contacting chin shields; anterior chin shields about three times as long as broad, posterior chin shields absent; three series of gular scales; dorsal scales 15/15/15 rows, smooth without apical pits; preventrals 2; ventrals 143; anal plate single; paired subcaudals 16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31B78EB4A522254389EDD41FE44EB74F" pageId="15" pageNumber="106" type="natural history">
|
||
<paragraph id="10508FF8DCA528CB8E83DE1EE9A25C5A" pageId="15" pageNumber="106">Natural history.</paragraph>
|
||
<paragraph id="4555480C9D9C6ADDA4C5C2679C40DF42" pageId="15" pageNumber="106">
|
||
The only known specimen of
|
||
<taxonomicName id="29DBFE813DF32BAC37792829D0475477" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="15" pageNumber="106" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
|
||
was found dead on a dirt road surrounded by silvopastures and remnants of native montane cloudforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A0B5E7E10E3AA50587D842F16C283FE0" pageId="15" pageNumber="106" type="distribution">
|
||
<paragraph id="BCA787B37463000CBF3931FFDE143C3F" pageId="15" pageNumber="106">Distribution.</paragraph>
|
||
<paragraph id="71E157D16229257C355FA65454510393" pageId="15" pageNumber="106">
|
||
Known only from the type locality, between Balzapamba and
|
||
<normalizedToken id="79ACE30C9A2C5BD97EDE4C2088D2BE47" originalValue="Bilován">Bilovan</normalizedToken>
|
||
, in the Ecuadorian province of
|
||
<normalizedToken id="54CB80436954DBA9F0F7A7A35DE2FE28" originalValue="Bolívar">Bolivar</normalizedToken>
|
||
at 2026 m (Fig. 7).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8411BFE0FAA8D8CA19AA30B8E6D1581" pageId="15" pageNumber="106" type="etymology">
|
||
<paragraph id="6F48FA0F77E1EDB8D878AA6C2C75A2E5" pageId="15" pageNumber="106">Etymology.</paragraph>
|
||
<paragraph id="C52026DFDBCB65F473928E777CFE45BF" pageId="15" pageNumber="106">
|
||
Named after R. Alexander Pyron, one of the most prolific contemporary herpetologists, in recognition of his invaluable contribution to systematics and evolution of the
|
||
<normalizedToken id="05569FFC0B16A58E9FDB9FEA2DF6CB26" originalValue="world’s">world's</normalizedToken>
|
||
reptiles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B56FFAA89591A78B459AB14D17ACC33D" pageId="15" pageNumber="106" type="conservation status">
|
||
<paragraph id="18E70DC344B0C69633B276806D3FA8EB" pageId="15" pageNumber="106">Conservation status.</paragraph>
|
||
<paragraph id="FE9C3EC59C2EA7244D05E648B626D607" pageId="15" pageNumber="106">
|
||
We consider
|
||
<taxonomicName id="4821C3085BC636FF39A2C396E89541FB" class="Squamata" family="Colubridae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus pyroni" pageId="15" pageNumber="106" phylum="Chordata" rank="species" species="pyroni">Atractus pyroni</taxonomicName>
|
||
to be to be Data Deficient following IUCN because there is inadequate information to make a direct, or indirect, assessment of its extinction risk based on its scarce distribution data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |