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<document ID-DOI="10.5252/geodiversitas2021v43a11" ID-GBIF-Dataset="fc56a00d-cf6b-4fda-aa56-d0492e4781cb" ID-ISSN="1638-9395" ID-Zenodo-Dep="4891107" ID-ZooBank="urn:lsid:zoobank.org:pub:07D1F5A8-98E2-40F1-A716-D25CB975EA4C" checkinTime="1622490877458" checkinUser="felipe" docAuthor="Steurbaut, Etienne &amp; Nolf, Dirk" docDate="2021" docId="1D2F9F2B9A3AFFCEFF1FFD0CDDB5BCB0" docLanguage="en" docName="Geodiversitas.43.11.311-363.pdf" docOrigin="Geodiversitas 43 (11)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Blackites gladius Varol 1989" docType="treatment" docVersion="5" lastPageNumber="352" masterDocId="E116E7539A11FFE5FFDBFFFADF3DBA1F" masterDocTitle="The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP" masterLastPageNumber="363" masterPageNumber="311" pageNumber="352" updateTime="1632098410930" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title>The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Steurbaut, Etienne</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium) and Department of Earth, and Environmental Sciences, KU Leuven, Celestijnenlaan 200 E, B- 3001 Leuven (Belgium)</mods:affiliation>
<mods:nameIdentifier type="email">etienne.steurbaut@naturalsciences.be</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nolf, Dirk</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium)</mods:affiliation>
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<mods:title>Geodiversitas</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-05-25</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4891222" ID-GBIF-Taxon="182482762" ID-Zenodo-Dep="4891222" LSID="urn:lsid:plazi:treatment:1D2F9F2B9A3AFFCEFF1FFD0CDDB5BCB0" httpUri="http://treatment.plazi.org/id/1D2F9F2B9A3AFFCEFF1FFD0CDDB5BCB0" lastPageNumber="352" pageId="43" pageNumber="352">
<subSubSection pageId="43" pageNumber="352" type="nomenclature">
<paragraph blockId="43.[196,718,758,817]" pageId="43" pageNumber="352">
<taxonomicName authority="(Locker, 1967) Varol, 1989" authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[196,718,758,785]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis bold="true" box="[196,292,758,785]" italics="true" pageId="43" pageNumber="352">Blackites</emphasis>
aff.
<emphasis bold="true" box="[340,417,758,785]" italics="true" pageId="43" pageNumber="352">gladius</emphasis>
(
<bibRefCitation author="LOCKER S." box="[431,576,758,785]" pageId="43" pageNumber="352" pagination="758 - 768" refId="ref41363" refString="LOCKER S. 1967. - Neue, stratigraphisch wichtige Coccolithophoriden (Flagellata) aus dem norddeutschen Alttertiar. Monatsberichte der Deutschen Akademie der Wissenschaften zu Berlin 9 (9 / 10): 758 - 768." type="journal article" year="1967">Locker, 1967</bibRefCitation>
)
<bibRefCitation author="VAROL O." box="[591,718,758,784]" pageId="43" pageNumber="352" pagination="273 - 320" refId="ref44687" refString="VAROL O. 1989. - Eocene calcareous Nannofossils from Sile (Northwest Turkey). Revista Espanola de of Micropalaeontologia 21 (2): 273 - 320." type="journal article" year="1989">Varol, 1989</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[388,520,790,817]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="43" pageNumber="352">Fig. 20L, M</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="352" type="reference_group">
<paragraph blockId="43.[132,775,847,871]" box="[132,775,847,871]" pageId="43" pageNumber="352">
<treatmentCitationGroup box="[132,775,847,871]" pageId="43" pageNumber="352">
<treatmentCitation author="LOCKER S." box="[132,528,847,871]" page="766" pageId="43" pageNumber="352" year="1967">
<taxonomicName authority="Locker, 1967: 766" authorityName="Locker" authorityPageNumber="766" authorityYear="1967" box="[132,528,847,871]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Rhabdosphaera" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[132,343,847,871]" italics="true" pageId="43" pageNumber="352">Rhabdosphaera gladius</emphasis>
<bibRefCitation author="LOCKER S." box="[348,528,847,871]" pageId="43" pageNumber="352" pagination="758 - 768" refId="ref41363" refString="LOCKER S. 1967. - Neue, stratigraphisch wichtige Coccolithophoriden (Flagellata) aus dem norddeutschen Alttertiar. Monatsberichte der Deutschen Akademie der Wissenschaften zu Berlin 9 (9 / 10): 758 - 768." type="journal article" year="1967">Locker, 1967: 766</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 1, fig. 1; pl. 2, fig. 12.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="43.[132,477,895,919]" box="[132,477,895,919]" pageId="43" pageNumber="352">
<treatmentCitationGroup box="[132,477,895,919]" pageId="43" pageNumber="352">
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[132,285,895,919]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[132,285,895,919]" italics="true" pageId="43" pageNumber="352">Blackites gladius</emphasis>
</taxonomicName>
<treatmentCitation author="VAROL O." box="[311,473,895,919]" page="286" pageId="43" pageNumber="352" year="1989">
<bibRefCitation author="VAROL O." box="[311,473,895,919]" pageId="43" pageNumber="352" pagination="273 - 320" refId="ref44687" refString="VAROL O. 1989. - Eocene calcareous Nannofossils from Sile (Northwest Turkey). Revista Espanola de of Micropalaeontologia 21 (2): 273 - 320." type="journal article" year="1989">Varol 1989: 286</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="352" type="distribution">
<paragraph blockId="43.[131,776,943,1285]" pageId="43" pageNumber="352">
DISTRIBUTION. —
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[322,514,943,967]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[322,404,943,967]" italics="true" pageId="43" pageNumber="352">Blackites</emphasis>
aff.
<emphasis box="[449,514,943,967]" italics="true" pageId="43" pageNumber="352">gladius</emphasis>
</taxonomicName>
is only known from three records, one in
<collectingCountry box="[279,364,969,993]" name="Belgium" pageId="43" pageNumber="352">Belgium</collectingCountry>
, one in the Paris Basin and one in the Aquitaine Basin. It rarely co-occurs with
<taxonomicName baseAuthorityName="Bramlette &amp; Sullivan" baseAuthorityYear="1961" box="[495,655,996,1020]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="inflatus">
<emphasis box="[495,655,996,1020]" italics="true" pageId="43" pageNumber="352">Blackites inflatus</emphasis>
</taxonomicName>
in the Isnes quarry in SW
<collectingCountry box="[271,356,1022,1046]" name="Belgium" pageId="43" pageNumber="352">Belgium</collectingCountry>
, where its presence seems to be restricted to the top of the
<collectingRegion box="[268,337,1049,1073]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="43" pageNumber="352">Brussel</collectingRegion>
Sand Formation. A few specimens have been identified in the Donzacq Marl (upper NP14) at Saint-GeoursdAuribat (
<collectingRegion box="[237,334,1102,1126]" country="France" name="Aquitaine" pageId="43" pageNumber="352">Aquitaine</collectingRegion>
Basin) (
<bibRefCitation author="LIN C. - H. &amp; NOLF D. &amp; STEURBAUT E. &amp; GIRONE A." box="[414,556,1102,1126]" pageId="43" pageNumber="352" pagination="879 - 907" refId="ref41294" refString="LIN C. - H., NOLF D., STEURBAUT E. &amp; GIRONE A. 2017. - Fish otoliths from the Lutetian of the Aquitaine Basin (SW France), a breakthrough in the knowledge of the European Eocene ichthyofauna. Journal of Systematic Palaeontology 15 (11): 879 - 907. https: // doi. org / 10.1080 / 14772019.2016.1246112" type="journal article" year="2017">
Lin
<emphasis box="[454,500,1102,1126]" italics="true" pageId="43" pageNumber="352">et al.</emphasis>
2017
</bibRefCitation>
) and in the upper part of the Glauconie Grossière s.s. at Margival (Paris Basin, sample
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="43" pageNumber="352" unit="in" value="3.0">3 in</quantity>
<bibRefCitation author="STEURBAUT E." box="[158,310,1155,1179]" pageId="43" pageNumber="352" pagination="99 - 115" refId="ref43418" refString="STEURBAUT E. 1988. - New Early and Middle Eocene calcareous nannoplankton events and correlations in middle to high latitudes of the northern hemisphere. Newsletters on Stratigraphy 18 (2): 99 - 115. https: // doi. org / 10.1127 / nos / 18 / 1988 / 99" type="journal article" year="1988">Steurbaut 1988</bibRefCitation>
), where it also co-occurs with
<taxonomicName baseAuthorityName="Bramlette &amp; Sullivan" baseAuthorityYear="1961" box="[607,704,1155,1179]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="inflatus">
<emphasis box="[607,704,1155,1179]" italics="true" pageId="43" pageNumber="352">B. inflatus</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[717,739,1156,1179]" italics="true" pageId="43" pageNumber="352">B.</emphasis>
aff.
<emphasis box="[132,197,1181,1205]" italics="true" pageId="43" pageNumber="352">gladius</emphasis>
</taxonomicName>
seems to indicate late Biochron NP14, also known as Biochon NP14b (
<emphasis box="[272,319,1209,1232]" italics="true" pageId="43" pageNumber="352">sensu</emphasis>
<bibRefCitation author="BERGGREN W. A. &amp; KENT D. V. &amp; SWISSER C. C. &amp; III &amp; AUBRY M. - P." box="[325,521,1208,1232]" pageId="43" pageNumber="352" pagination="129 - 212" refId="ref36889" refString="BERGGREN W. A., KENT D. V., SWISSER C. C. III &amp; AUBRY M. - P. 1995. - A revised Cenozoic geochronology and chronostratigraphy, in BERGGREN W. A., KENT D. V., AUBRY M. - P. &amp; HARDEN- BOL J. (eds), Geochronology, Time Scales and Global Stratigraphic Correlation. Society for Sedimentary Geology (SEPM), Special Publication 54: 129 - 212. https: // doi. org / 10.2110 / pec. 95.04.0129" type="journal article" year="1995">
Berggren
<emphasis box="[419,465,1208,1232]" italics="true" pageId="43" pageNumber="352">et al.</emphasis>
1995
</bibRefCitation>
). It has not been recorded at the Mont-des-Récollets because of major erosion of the top of the
<collectingRegion box="[171,240,1261,1285]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="43" pageNumber="352">Brussel</collectingRegion>
Sand Formation in N
<collectingCountry box="[464,532,1261,1284]" name="France" pageId="43" pageNumber="352">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="352" type="discussion">
<paragraph blockId="43.[130,776,1334,1711]" box="[132,262,1334,1360]" pageId="43" pageNumber="352">DISCUSSION</paragraph>
<paragraph blockId="43.[130,776,1334,1711]" pageId="43" pageNumber="352">
This medium-sized rhabdolith (
<emphasis box="[465,482,1366,1391]" italics="true" pageId="43" pageNumber="352">c.</emphasis>
<quantity box="[487,555,1366,1392]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="43" pageNumber="352" unit="mm" value="11.0">11 µm</quantity>
length) is marked by a very wide basal plate (
<emphasis box="[373,390,1398,1423]" italics="true" pageId="43" pageNumber="352">c.</emphasis>
<quantity box="[394,471,1397,1424]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="43" pageNumber="352" unit="mm" value="4.8">4.8 µm</quantity>
) and a lensiform stem, which maximum width is located in the upper half of the stem. The latter, which presents a basal collar, seems to be thin-walled (
<figureCitation box="[141,231,1493,1520]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="43" pageNumber="352">Fig. 20L</figureCitation>
) and rugose towards the top. These rhabdoliths bear great resemblance with those of
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[471,572,1525,1551]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[471,572,1525,1551]" italics="true" pageId="43" pageNumber="352">B. gladius</emphasis>
</taxonomicName>
(
<bibRefCitation author="VAROL O." box="[588,707,1525,1551]" pageId="43" pageNumber="352" pagination="273 - 320" refId="ref44687" refString="VAROL O. 1989. - Eocene calcareous Nannofossils from Sile (Northwest Turkey). Revista Espanola de of Micropalaeontologia 21 (2): 273 - 320." type="journal article" year="1989">Varol 1989</bibRefCitation>
: pl. 4, fig. 5;
<bibRefCitation author="BOWN P. R." box="[203,330,1557,1584]" pageId="43" pageNumber="352" pagination="21 - 95" refId="ref37380" refString="BOWN P. R. 2005. - Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003 - 4). Journal of Nannoplankton Research 27 (1): 21 - 95." type="journal article" year="2005">Bown 2005</bibRefCitation>
: pl. 21, figs 10-14). However, they differ by certain features of the stem, which is relatively longer and more slender in
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[311,455,1621,1647]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[311,335,1622,1647]" italics="true" pageId="43" pageNumber="352">B.</emphasis>
aff.
<emphasis box="[384,455,1621,1647]" italics="true" pageId="43" pageNumber="352">gladius</emphasis>
</taxonomicName>
, especially in the top part, as well as by the position of its maximum width. The latter is less distal compared to its position in
<taxonomicName authorityName="Varol" authorityYear="1989" baseAuthorityName="Locker" baseAuthorityYear="1967" box="[538,641,1685,1711]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Blackites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="43" pageNumber="352" phylum="Haptophyta" rank="species" species="gladius">
<emphasis box="[538,641,1685,1711]" italics="true" pageId="43" pageNumber="352">B. gladius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>