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<document ID-DOI="10.5252/geodiversitas2021v43a11" ID-GBIF-Dataset="fc56a00d-cf6b-4fda-aa56-d0492e4781cb" ID-ISSN="1638-9395" ID-Zenodo-Dep="4891107" ID-ZooBank="urn:lsid:zoobank.org:pub:07D1F5A8-98E2-40F1-A716-D25CB975EA4C" checkinTime="1622490877458" checkinUser="felipe" docAuthor="Steurbaut, Etienne &amp; Nolf, Dirk" docDate="2021" docId="1D2F9F2B9A37FFC3FEE3FD2FDA93B82B" docLanguage="en" docName="Geodiversitas.43.11.311-363.pdf" docOrigin="Geodiversitas 43 (11)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Trochoaster nodosus Steurbaut &amp; Nolf, 2021, n. sp." docType="treatment" docUuid="BEE661BF-859D-4C27-AC92-BF41326F94A1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="347" masterDocId="E116E7539A11FFE5FFDBFFFADF3DBA1F" masterDocTitle="The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP" masterLastPageNumber="363" masterPageNumber="311" pageNumber="347" updateTime="1632098410930" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Steurbaut, Etienne</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium) and Department of Earth, and Environmental Sciences, KU Leuven, Celestijnenlaan 200 E, B- 3001 Leuven (Belgium)</mods:affiliation>
<mods:nameIdentifier type="email">etienne.steurbaut@naturalsciences.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nolf, Dirk</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium)</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>43</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>311</mods:start>
<mods:end>363</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2021v43a11</mods:identifier>
<mods:identifier type="GBIF-Dataset">fc56a00d-cf6b-4fda-aa56-d0492e4781cb</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">4891107</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:07D1F5A8-98E2-40F1-A716-D25CB975EA4C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4891200" ID-GBIF-Taxon="182482748" ID-Zenodo-Dep="4891200" LSID="urn:lsid:zoobank.org:act:BEE661BF-859D-4C27-AC92-BF41326F94A1" httpUri="http://treatment.plazi.org/id/1D2F9F2B9A37FFC3FEE3FD2FDA93B82B" lastPageNumber="347" pageId="38" pageNumber="347">
<subSubSection box="[312,596,725,752]" pageId="38" pageNumber="347" type="nomenclature">
<paragraph blockId="38.[312,596,725,752]" box="[312,596,725,752]" pageId="38" pageNumber="347">
<heading box="[312,596,725,752]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="347" reason="2">
<taxonomicName box="[312,530,725,752]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="nodosus" status="n. sp.">
<emphasis bold="true" box="[312,530,725,752]" italics="true" pageId="38" pageNumber="347">Trochoaster nodosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[537,596,726,752]" pageId="38" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="description">
<paragraph blockId="38.[381,526,757,784]" box="[381,526,757,784]" pageId="38" pageNumber="347">
(
<figureCitation box="[390,466,757,783]" captionStart="FIG" captionStartId="34.[132,142,1690,1707]" captionTargetBox="[242,1344,215,1682]" captionTargetId="figure-29@34.[241,794,765,1321]" captionTargetPageId="34" captionText="FIG. 18. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium;a, b, c, views of the same specimen, in transmitted light or cross-polarized light or vice versa, and in different positions with respect to the polarisation directions. Numbers (e.g. IRSNB bXXXX) refer to the collections of the Royal Belgian Institute of Natural Sciences (Brussels).Pars 1: A-D, Braarudosphaera stylifera Troelsen &amp; Quadros,1971,Mont-des-Récollets,122.85 m (IRSNB b7093,b7094,b7096);A, wm = 13 µm;B, wm = 12 µm;D, wm = 14.5 µm;C, Mont-des-Récollets,124.10 m (IRSNB b7095),wm = 13 µm;E, Lanternithus minutus Stradner, 1962, Mont-des-Récollets, 122.85 m, l = 5.6 µm (IRSNB b7097); F, G, Martiniaster cecellanoriae n. sp.: F, Chaumont-en-Vexin, sample 11, 0.55 m above base of the Lutetian, paratype,d = 13 µm (IRSNB b7098);G, Leuven BH,23.70 m depth, holotype,d = 14 µm (IRSNB b7099);H, Discoaster praebifax Wei &amp; Wise,1989, Mont-des-Récollets,116.45 m, d = 7 µm (IRSNB b7100);I-M, Discoaster sublodoensis Bramlette &amp; Sullivan,1961,Mont-des-Récollets,116.45 m (IRSNB b7101,b7102); J, d = 15.2 µm; I, d = 16 µm; K, Mont-des-Récollets, 118.35 m, d = 11.6 µm (IRSNB b7103); L, Vossem, otolith sample Nolf 1974, d = 7 µm (IRSNB b7104); M, Gisors, Bassin de Paris,sample 3,d = 14 µm (IRSNB b7105);N, Martiniaster fragilis (Martini,1961), Mont-des-Récollets,122.85 m, wm = 12 µm (IRSNB b7106);O-R, Discoaster wemmelensis Achuthan &amp; Stradner,1969;O, Mont-des-Récollets, 122.85 m, d = 5.6 µm (IRSNB b7107); P-R, Vossem,otolith sample Nolf 1974 (IRSNB b7108, b7109, b7110);P, d = 5.3 µm; Q, d = 4.4 µm; R, d = 4.3 µm;S-V, Blackites minusculus n. sp.; S, Mont-des-Récollets,117.46 m, paratype, l = 8.8 µm (IRSNB b7111); T-V, Vossem, otolith sample Nolf 1974; T, U, paratypes (IRSNB b7112, b7113); T, I = 8.2 µm;U, I = 7.2 µm;V, holotype,I = 8.56 µm (IRSNB b7114); W-Y, Trochoaster nodosus n. sp.; W, Chaumont-en-Vexin,sample 16, 0.80 m above base of the Lutetian,holotype, d = 16 µm (IRSNB b7115);X, Vilvoorde BH, 5.80 m depth,paratype,d = 15 µm (IRSNB b7116); Y, Oedelem BH, 11.70 m depth, paratype, d = 15 µm (IRSNB b7117); Z-AA, Sphenolithus recolletensis n. sp.; Z, Mont-des-Récollets, 116.45 m, paratype, htot = 20 µm (IRSNB b7118);AA, Mont-des-Récollets, 117.46 m, holotype (a, c, parallel to the polarization directions;b, at 45°), htot = 11.4 µm (IRSNB b7119)." figureDoi="http://doi.org/10.5281/zenodo.4891140" httpUri="https://zenodo.org/record/4891140/files/figure.png" pageId="38" pageNumber="347">Fig. 18</figureCitation>
W-Y)
</paragraph>
<paragraph blockId="38.[132,775,824,841]" box="[132,775,824,841]" pageId="38" pageNumber="347">
<uri box="[132,775,824,841]" pageId="38" pageNumber="347">
urn:lsid:zoobank.org:act:
<uuid box="[350,775,824,841]" pageId="38" pageNumber="347">BEE661BF-859D-4C27-AC92-BF41326F94A1</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="materials_examined">
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3125291332" collectionCode="IRSNB, RBINS" pageId="38" pageNumber="347" specimenCount="1" typeStatus="holotype">
<typeStatus box="[132,239,883,906]" pageId="38" pageNumber="347">HOLOTYPE</typeStatus>
. —
<figureCitation box="[285,379,883,907]" captionStart="FIG" captionStartId="34.[132,142,1690,1707]" captionTargetBox="[242,1344,215,1682]" captionTargetId="figure-29@34.[241,794,765,1321]" captionTargetPageId="34" captionText="FIG. 18. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium;a, b, c, views of the same specimen, in transmitted light or cross-polarized light or vice versa, and in different positions with respect to the polarisation directions. Numbers (e.g. IRSNB bXXXX) refer to the collections of the Royal Belgian Institute of Natural Sciences (Brussels).Pars 1: A-D, Braarudosphaera stylifera Troelsen &amp; Quadros,1971,Mont-des-Récollets,122.85 m (IRSNB b7093,b7094,b7096);A, wm = 13 µm;B, wm = 12 µm;D, wm = 14.5 µm;C, Mont-des-Récollets,124.10 m (IRSNB b7095),wm = 13 µm;E, Lanternithus minutus Stradner, 1962, Mont-des-Récollets, 122.85 m, l = 5.6 µm (IRSNB b7097); F, G, Martiniaster cecellanoriae n. sp.: F, Chaumont-en-Vexin, sample 11, 0.55 m above base of the Lutetian, paratype,d = 13 µm (IRSNB b7098);G, Leuven BH,23.70 m depth, holotype,d = 14 µm (IRSNB b7099);H, Discoaster praebifax Wei &amp; Wise,1989, Mont-des-Récollets,116.45 m, d = 7 µm (IRSNB b7100);I-M, Discoaster sublodoensis Bramlette &amp; Sullivan,1961,Mont-des-Récollets,116.45 m (IRSNB b7101,b7102); J, d = 15.2 µm; I, d = 16 µm; K, Mont-des-Récollets, 118.35 m, d = 11.6 µm (IRSNB b7103); L, Vossem, otolith sample Nolf 1974, d = 7 µm (IRSNB b7104); M, Gisors, Bassin de Paris,sample 3,d = 14 µm (IRSNB b7105);N, Martiniaster fragilis (Martini,1961), Mont-des-Récollets,122.85 m, wm = 12 µm (IRSNB b7106);O-R, Discoaster wemmelensis Achuthan &amp; Stradner,1969;O, Mont-des-Récollets, 122.85 m, d = 5.6 µm (IRSNB b7107); P-R, Vossem,otolith sample Nolf 1974 (IRSNB b7108, b7109, b7110);P, d = 5.3 µm; Q, d = 4.4 µm; R, d = 4.3 µm;S-V, Blackites minusculus n. sp.; S, Mont-des-Récollets,117.46 m, paratype, l = 8.8 µm (IRSNB b7111); T-V, Vossem, otolith sample Nolf 1974; T, U, paratypes (IRSNB b7112, b7113); T, I = 8.2 µm;U, I = 7.2 µm;V, holotype,I = 8.56 µm (IRSNB b7114); W-Y, Trochoaster nodosus n. sp.; W, Chaumont-en-Vexin,sample 16, 0.80 m above base of the Lutetian,holotype, d = 16 µm (IRSNB b7115);X, Vilvoorde BH, 5.80 m depth,paratype,d = 15 µm (IRSNB b7116); Y, Oedelem BH, 11.70 m depth, paratype, d = 15 µm (IRSNB b7117); Z-AA, Sphenolithus recolletensis n. sp.; Z, Mont-des-Récollets, 116.45 m, paratype, htot = 20 µm (IRSNB b7118);AA, Mont-des-Récollets, 117.46 m, holotype (a, c, parallel to the polarization directions;b, at 45°), htot = 11.4 µm (IRSNB b7119)." figureDoi="http://doi.org/10.5281/zenodo.4891140" httpUri="https://zenodo.org/record/4891140/files/figure.png" pageId="38" pageNumber="347">Fig. 18W</figureCitation>
(
<collectionCode box="[394,468,883,907]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7115) (negative stored in the collections of the
<collectionCode box="[306,383,910,934]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="38" pageNumber="347" type="Museum">RBINS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3125291317" collectionCode="IRSNB" collectorName="Oedelem BH" elevation="6" pageId="38" pageNumber="347" specimenCount="1" typeStatus="paratype">
<typeStatus box="[132,236,951,974]" pageId="38" pageNumber="347">PARATYPES</typeStatus>
.
<emphasis box="[248,273,952,975]" italics="true" pageId="38" pageNumber="347">—</emphasis>
2 figured specimens; 1 from the Vilvoorde borehole at
<elevation box="[157,295,977,1001]" metricMagnitude="0" metricUnit="m" metricValue="5.8" pageId="38" pageNumber="347" unit="m" value="5.8">
<quantity box="[157,230,977,1001]" metricMagnitude="0" metricUnit="m" metricValue="5.8" pageId="38" pageNumber="347" unit="m" value="5.8">5.80 m</quantity>
depth
</elevation>
(d =
<quantity box="[353,420,977,1002]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="347" unit="mm" value="15.0">15 µm</quantity>
) (
<figureCitation box="[445,534,977,1002]" captionStart="FIG" captionStartId="34.[132,142,1690,1707]" captionTargetBox="[242,1344,215,1682]" captionTargetId="figure-29@34.[241,794,765,1321]" captionTargetPageId="34" captionText="FIG. 18. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium;a, b, c, views of the same specimen, in transmitted light or cross-polarized light or vice versa, and in different positions with respect to the polarisation directions. Numbers (e.g. IRSNB bXXXX) refer to the collections of the Royal Belgian Institute of Natural Sciences (Brussels).Pars 1: A-D, Braarudosphaera stylifera Troelsen &amp; Quadros,1971,Mont-des-Récollets,122.85 m (IRSNB b7093,b7094,b7096);A, wm = 13 µm;B, wm = 12 µm;D, wm = 14.5 µm;C, Mont-des-Récollets,124.10 m (IRSNB b7095),wm = 13 µm;E, Lanternithus minutus Stradner, 1962, Mont-des-Récollets, 122.85 m, l = 5.6 µm (IRSNB b7097); F, G, Martiniaster cecellanoriae n. sp.: F, Chaumont-en-Vexin, sample 11, 0.55 m above base of the Lutetian, paratype,d = 13 µm (IRSNB b7098);G, Leuven BH,23.70 m depth, holotype,d = 14 µm (IRSNB b7099);H, Discoaster praebifax Wei &amp; Wise,1989, Mont-des-Récollets,116.45 m, d = 7 µm (IRSNB b7100);I-M, Discoaster sublodoensis Bramlette &amp; Sullivan,1961,Mont-des-Récollets,116.45 m (IRSNB b7101,b7102); J, d = 15.2 µm; I, d = 16 µm; K, Mont-des-Récollets, 118.35 m, d = 11.6 µm (IRSNB b7103); L, Vossem, otolith sample Nolf 1974, d = 7 µm (IRSNB b7104); M, Gisors, Bassin de Paris,sample 3,d = 14 µm (IRSNB b7105);N, Martiniaster fragilis (Martini,1961), Mont-des-Récollets,122.85 m, wm = 12 µm (IRSNB b7106);O-R, Discoaster wemmelensis Achuthan &amp; Stradner,1969;O, Mont-des-Récollets, 122.85 m, d = 5.6 µm (IRSNB b7107); P-R, Vossem,otolith sample Nolf 1974 (IRSNB b7108, b7109, b7110);P, d = 5.3 µm; Q, d = 4.4 µm; R, d = 4.3 µm;S-V, Blackites minusculus n. sp.; S, Mont-des-Récollets,117.46 m, paratype, l = 8.8 µm (IRSNB b7111); T-V, Vossem, otolith sample Nolf 1974; T, U, paratypes (IRSNB b7112, b7113); T, I = 8.2 µm;U, I = 7.2 µm;V, holotype,I = 8.56 µm (IRSNB b7114); W-Y, Trochoaster nodosus n. sp.; W, Chaumont-en-Vexin,sample 16, 0.80 m above base of the Lutetian,holotype, d = 16 µm (IRSNB b7115);X, Vilvoorde BH, 5.80 m depth,paratype,d = 15 µm (IRSNB b7116); Y, Oedelem BH, 11.70 m depth, paratype, d = 15 µm (IRSNB b7117); Z-AA, Sphenolithus recolletensis n. sp.; Z, Mont-des-Récollets, 116.45 m, paratype, htot = 20 µm (IRSNB b7118);AA, Mont-des-Récollets, 117.46 m, holotype (a, c, parallel to the polarization directions;b, at 45°), htot = 11.4 µm (IRSNB b7119)." figureDoi="http://doi.org/10.5281/zenodo.4891140" httpUri="https://zenodo.org/record/4891140/files/figure.png" pageId="38" pageNumber="347">Fig. 18X</figureCitation>
) (
<collectionCode box="[555,629,978,1002]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7116) and 1 from the middle part of the Oedelem Sand Member (Aalter Sand Formation) in the
<collectorName box="[312,443,1030,1054]" pageId="38" pageNumber="347">Oedelem BH</collectorName>
(
<quantity box="[456,541,1030,1054]" metricMagnitude="1" metricUnit="m" metricValue="1.17" pageId="38" pageNumber="347" unit="m" value="11.7">
<elevation box="[456,541,1030,1054]" metricMagnitude="1" metricUnit="m" metricValue="1.17" pageId="38" pageNumber="347" unit="m" value="11.7">11.70 m</elevation>
</quantity>
) (d =
<quantity box="[599,664,1030,1054]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="347" unit="mm" value="15.0">15 µm</quantity>
) (
<figureCitation box="[684,769,1030,1054]" captionStart="FIG" captionStartId="34.[132,142,1690,1707]" captionTargetBox="[242,1344,215,1682]" captionTargetId="figure-29@34.[241,794,765,1321]" captionTargetPageId="34" captionText="FIG. 18. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium;a, b, c, views of the same specimen, in transmitted light or cross-polarized light or vice versa, and in different positions with respect to the polarisation directions. Numbers (e.g. IRSNB bXXXX) refer to the collections of the Royal Belgian Institute of Natural Sciences (Brussels).Pars 1: A-D, Braarudosphaera stylifera Troelsen &amp; Quadros,1971,Mont-des-Récollets,122.85 m (IRSNB b7093,b7094,b7096);A, wm = 13 µm;B, wm = 12 µm;D, wm = 14.5 µm;C, Mont-des-Récollets,124.10 m (IRSNB b7095),wm = 13 µm;E, Lanternithus minutus Stradner, 1962, Mont-des-Récollets, 122.85 m, l = 5.6 µm (IRSNB b7097); F, G, Martiniaster cecellanoriae n. sp.: F, Chaumont-en-Vexin, sample 11, 0.55 m above base of the Lutetian, paratype,d = 13 µm (IRSNB b7098);G, Leuven BH,23.70 m depth, holotype,d = 14 µm (IRSNB b7099);H, Discoaster praebifax Wei &amp; Wise,1989, Mont-des-Récollets,116.45 m, d = 7 µm (IRSNB b7100);I-M, Discoaster sublodoensis Bramlette &amp; Sullivan,1961,Mont-des-Récollets,116.45 m (IRSNB b7101,b7102); J, d = 15.2 µm; I, d = 16 µm; K, Mont-des-Récollets, 118.35 m, d = 11.6 µm (IRSNB b7103); L, Vossem, otolith sample Nolf 1974, d = 7 µm (IRSNB b7104); M, Gisors, Bassin de Paris,sample 3,d = 14 µm (IRSNB b7105);N, Martiniaster fragilis (Martini,1961), Mont-des-Récollets,122.85 m, wm = 12 µm (IRSNB b7106);O-R, Discoaster wemmelensis Achuthan &amp; Stradner,1969;O, Mont-des-Récollets, 122.85 m, d = 5.6 µm (IRSNB b7107); P-R, Vossem,otolith sample Nolf 1974 (IRSNB b7108, b7109, b7110);P, d = 5.3 µm; Q, d = 4.4 µm; R, d = 4.3 µm;S-V, Blackites minusculus n. sp.; S, Mont-des-Récollets,117.46 m, paratype, l = 8.8 µm (IRSNB b7111); T-V, Vossem, otolith sample Nolf 1974; T, U, paratypes (IRSNB b7112, b7113); T, I = 8.2 µm;U, I = 7.2 µm;V, holotype,I = 8.56 µm (IRSNB b7114); W-Y, Trochoaster nodosus n. sp.; W, Chaumont-en-Vexin,sample 16, 0.80 m above base of the Lutetian,holotype, d = 16 µm (IRSNB b7115);X, Vilvoorde BH, 5.80 m depth,paratype,d = 15 µm (IRSNB b7116); Y, Oedelem BH, 11.70 m depth, paratype, d = 15 µm (IRSNB b7117); Z-AA, Sphenolithus recolletensis n. sp.; Z, Mont-des-Récollets, 116.45 m, paratype, htot = 20 µm (IRSNB b7118);AA, Mont-des-Récollets, 117.46 m, holotype (a, c, parallel to the polarization directions;b, at 45°), htot = 11.4 µm (IRSNB b7119)." figureDoi="http://doi.org/10.5281/zenodo.4891140" httpUri="https://zenodo.org/record/4891140/files/figure.png" pageId="38" pageNumber="347">Fig. 18Y</figureCitation>
) (
<collectionCode box="[139,213,1057,1081]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7117); both base of NP14.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="diagnosis">
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
DIAGNOSIS.
<emphasis box="[253,278,1099,1122]" italics="true" pageId="38" pageNumber="347">—</emphasis>
Six-rayed body, enterily covered with bumps, and marked by strong median ridges along the rays, which are connected with a central circular ring.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="etymology">
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">DERIVATIO NOMINIS. — The species name refers to the presence of numerous small bumps popping up all over this robust six-rayed holococcolith.</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="materials_examined">
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
LOCUS TYPICUS. —
<materialsCitation ID-GBIF-Occurrence="3125291320" pageId="38" pageNumber="347">
Quarry at Chaumont-en-Vexin, sample 16, Paris Basin;
<emphasis box="[198,214,1315,1338]" italics="true" pageId="38" pageNumber="347">c.</emphasis>
<geoCoordinate box="[219,373,1314,1338]" degrees="49" direction="north" minutes="15" orientation="latitude" pageId="38" pageNumber="347" precision="1" seconds="33.59" value="49.25933">49°15’33.59”N</geoCoordinate>
,
<geoCoordinate box="[383,517,1314,1338]" degrees="1" direction="east" minutes="52" orientation="longitude" pageId="38" pageNumber="347" precision="1" seconds="44.20" value="1.8789444">1°52’44.20”E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
STRATUM TYPICUM.
<emphasis box="[327,352,1356,1379]" italics="true" pageId="38" pageNumber="347">—</emphasis>
‘
<materialsCitation ID-GBIF-Occurrence="3125291309" pageId="38" pageNumber="347">Chaumont-en-Vexin sands’; base of NP14; basal Lutetian.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="description">
<paragraph blockId="38.[131,777,883,1754]" pageId="38" pageNumber="347">
DIMENSIONS. — Diameter =
<quantity box="[419,542,1423,1448]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="38" pageNumber="347" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15 to 16 µm</quantity>
(
<typeStatus box="[555,645,1423,1447]" pageId="38" pageNumber="347">holotype</typeStatus>
: d =
<quantity box="[692,760,1423,1448]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="347" unit="mm" value="16.0">16 µm</quantity>
). DISTRIBUTION. —
<taxonomicName box="[318,500,1465,1489]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="nodosus" status="n. sp.">
<emphasis box="[318,500,1465,1489]" italics="true" pageId="38" pageNumber="347">Trochoaster nodosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[507,560,1465,1488]" pageId="38" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
has been identified in many outcrop and borehole sections in the Belgian Basin and the Paris Basin. At the Mont-des-Récollets it has been recorded at the base of Unit A
<quantity box="[279,318,1544,1568]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="38" pageNumber="347" unit="in" value="4.0">4 in</quantity>
the upper part of the Aalter Sand Formation. It has been found in the same position (Unit A4) in the Oedelem BH and the Vlakte van de Raan BH. It has been encountered in the Vilvoorde BH at the base of the
<collectingRegion box="[484,553,1624,1648]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="38" pageNumber="347">Brussel</collectingRegion>
Sand Formation (Unit BA) and is present throughout the ‘Chaumont-en-Vexin sands‘ at Chaumont-en-Vexin. The LO of
<taxonomicName box="[452,548,1677,1701]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="nodosus" status="n. sp.">
<emphasis box="[452,548,1677,1701]" italics="true" pageId="38" pageNumber="347">T. nodosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[554,607,1677,1700]" pageId="38" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
is also part of the famous calcareous nannofossil turnover, marking the base of NP14 and the base of the historical Lutetian stratotype.
</paragraph>
<paragraph blockId="38.[130,776,1810,2028]" box="[132,277,1810,1836]" pageId="38" pageNumber="347">DESCRIPTION</paragraph>
<paragraph blockId="38.[130,776,1810,2028]" lastBlockId="38.[811,1457,215,274]" pageId="38" pageNumber="347">
This
<taxonomicName box="[185,381,1841,1867]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="undetermined">
<emphasis box="[185,302,1841,1867]" italics="true" pageId="38" pageNumber="347">Trochoaster</emphasis>
species
</taxonomicName>
is characterised by six symmetrically positioned protruding rays with strong median ridges along the rays. The ridges are connected with a central circular ring, which surrounds a triradiate central bridge. The entire body, also the areas between the centrally positioned ridges of the arms, is covered with numerous bumps. These flat interray areas show no furher ornamentation. Specimens of this taxon do not exhibit birefringence under crossed nicols.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="discussion">
<paragraph blockId="38.[811,1455,312,564]" box="[813,942,312,338]" pageId="38" pageNumber="347">DISCUSSION</paragraph>
<paragraph blockId="38.[811,1455,312,564]" pageId="38" pageNumber="347">
This taxon differs from all known
<taxonomicName authorityName="Klumpp" authorityYear="1953" box="[1175,1290,344,370]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="genus">
<emphasis box="[1175,1290,344,370]" italics="true" pageId="38" pageNumber="347">Trochoaster</emphasis>
</taxonomicName>
taxa by its rugged and knobby outline and the configuration of the central ring, where the median ridges along the rays meet. It presents a vague resemblance with
<taxonomicName box="[1083,1197,441,466]" class="Coccolithophyceae" family="Lithostromationaceae" genus="Trochoaster" higherTaxonomySource="GBIF" kingdom="Chromista" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="martinii">
<emphasis box="[1083,1197,441,466]" italics="true" pageId="38" pageNumber="347">T. martinii</emphasis>
</taxonomicName>
from the Early Miocene of
<collectingCountry box="[837,936,473,499]" name="Germany" pageId="38" pageNumber="347">Germany</collectingCountry>
because of the presence of six protruding rays.The latter is less knobby and presents some fundamental differences in configuration of the central structure and the interray areas.
</paragraph>
</subSubSection>
</treatment>
</document>
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