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<document ID-DOI="10.5252/geodiversitas2021v43a11" ID-GBIF-Dataset="fc56a00d-cf6b-4fda-aa56-d0492e4781cb" ID-ISSN="1638-9395" ID-Zenodo-Dep="4891107" ID-ZooBank="urn:lsid:zoobank.org:pub:07D1F5A8-98E2-40F1-A716-D25CB975EA4C" checkinTime="1622490877458" checkinUser="felipe" docAuthor="Steurbaut, Etienne &amp; Nolf, Dirk" docDate="2021" docId="1D2F9F2B9A37FFC2FC1FFA86DE90BE43" docLanguage="en" docName="Geodiversitas.43.11.311-363.pdf" docOrigin="Geodiversitas 43 (11)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Trochastrites pyramidalis Steurbaut &amp; Nolf, 2021, n. sp." docType="treatment" docUuid="1A5F4964-D75A-4631-A0A4-E948EF2E29FF" docUuidSource="ZooBank" docVersion="5" lastPageNumber="348" masterDocId="E116E7539A11FFE5FFDBFFFADF3DBA1F" masterDocTitle="The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP" masterLastPageNumber="363" masterPageNumber="311" pageNumber="347" updateTime="1632098410930" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo>
<mods:title>The Mont-des-Récollets section (N France): a key site for the Ypresian-Lutetian transition at mid-latitudes - reassessment of the boundary criterion for the base- Lutetian GSSP</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Steurbaut, Etienne</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium) and Department of Earth, and Environmental Sciences, KU Leuven, Celestijnenlaan 200 E, B- 3001 Leuven (Belgium)</mods:affiliation>
<mods:nameIdentifier type="email">etienne.steurbaut@naturalsciences.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nolf, Dirk</mods:namePart>
<mods:affiliation>Royal Belgian Institute of Natural Sciences, OD Earth and History of Life, Vautierstraat 29, B- 1000 Brussel (Belgium)</mods:affiliation>
</mods:name>
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<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>43</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
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<mods:start>311</mods:start>
<mods:end>363</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2021v43a11</mods:identifier>
<mods:identifier type="GBIF-Dataset">fc56a00d-cf6b-4fda-aa56-d0492e4781cb</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">4891107</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:07D1F5A8-98E2-40F1-A716-D25CB975EA4C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4891206" ID-GBIF-Taxon="182482735" ID-Zenodo-Dep="4891206" LSID="urn:lsid:zoobank.org:act:1A5F4964-D75A-4631-A0A4-E948EF2E29FF" httpUri="http://treatment.plazi.org/id/1D2F9F2B9A37FFC2FC1FFA86DE90BE43" lastPageId="39" lastPageNumber="348" pageId="38" pageNumber="347">
<subSubSection box="[964,1304,1404,1431]" pageId="38" pageNumber="347" type="nomenclature">
<paragraph blockId="38.[964,1304,1404,1431]" box="[964,1304,1404,1431]" pageId="38" pageNumber="347">
<heading box="[964,1304,1404,1431]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="347" reason="2">
<taxonomicName box="[964,1238,1404,1431]" class="Coccolithophyceae" genus="Trochastrites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Discoasterales" pageId="38" pageNumber="347" phylum="Haptophyta" rank="species" species="pyramidalis" status="n. sp.">
<emphasis bold="true" box="[964,1238,1404,1431]" italics="true" pageId="38" pageNumber="347">Trochastrites pyramidalis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1245,1304,1405,1431]" pageId="38" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="description">
<paragraph blockId="38.[1062,1206,1437,1464]" box="[1062,1206,1437,1464]" pageId="38" pageNumber="347">
(
<figureCitation box="[1071,1147,1437,1463]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="38" pageNumber="347">Fig. 20</figureCitation>
T-W)
</paragraph>
<paragraph blockId="38.[813,1456,1504,1521]" box="[813,1456,1504,1521]" pageId="38" pageNumber="347">
<uri box="[813,1456,1504,1521]" pageId="38" pageNumber="347">
urn:lsid:zoobank.org:act:
<uuid box="[1035,1456,1504,1521]" pageId="38" pageNumber="347">1A5F4964-D75A-4631-A0A4-E948EF2E29FF</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="materials_examined">
<paragraph blockId="38.[813,1455,1569,1620]" pageId="38" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3125291308" collectionCode="IRSNB, RBINS" pageId="38" pageNumber="347" specimenCount="1" typeStatus="holotype">
<typeStatus box="[813,919,1570,1593]" pageId="38" pageNumber="347">HOLOTYPE</typeStatus>
. —
<figureCitation box="[967,1056,1569,1594]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="38" pageNumber="347">Fig. 20U</figureCitation>
(
<collectionCode box="[1071,1145,1570,1594]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7161) (negative stored in the collections of the
<collectionCode box="[987,1064,1596,1620]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="38" pageNumber="347" type="Museum">RBINS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="38.[813,1457,1644,1776]" pageId="38" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3125291314" collectionCode="IRSNB" pageId="38" pageNumber="347" specimenCount="1" typeStatus="paratype">
<typeStatus box="[813,920,1645,1668]" pageId="38" pageNumber="347">PARATYPES</typeStatus>
.
<emphasis box="[934,960,1645,1668]" italics="true" pageId="38" pageNumber="347"></emphasis>
<collectorName box="[970,1026,1644,1668]" pageId="38" pageNumber="347">Three</collectorName>
figured specimens from the upper part of the
<collectorName box="[853,1090,1671,1695]" pageId="38" pageNumber="347">Oedelem Sand Member</collectorName>
(top of the Aalter Formation) in the
<collectorName box="[814,947,1698,1722]" pageId="38" pageNumber="347">Oedelem BH</collectorName>
(
<figureCitation box="[962,1078,1698,1722]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="38" pageNumber="347">Fig. 20T, V</figureCitation>
) (
<collectionCode box="[1099,1173,1698,1722]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7160,
<collectionCode box="[1259,1331,1698,1722]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7162) and the
<collectorName box="[851,1085,1725,1749]" pageId="38" pageNumber="347">Vlakte van de Raan BH</collectorName>
(
<figureCitation box="[1098,1193,1725,1749]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="38" pageNumber="347">Fig. 20W</figureCitation>
) (
<collectionCode box="[1213,1287,1725,1749]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="347">IRSNB</collectionCode>
b7163); all from the lower part of NP14.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="diagnosis">
<paragraph blockId="38.[813,1457,1799,1877]" pageId="38" pageNumber="347">
DIAGNOSIS.
<emphasis box="[936,961,1800,1823]" italics="true" pageId="38" pageNumber="347"></emphasis>
Three irregularly bifurcated arms interconnected with a fourth arm to form a pyramidal construction surrounded by a weblike structure.
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="347" type="etymology">
<paragraph blockId="38.[813,1457,1901,1952]" pageId="38" pageNumber="347">DERIVATIO NOMINIS. — Refers to the outline of the solid inner structure of these globular bodies, representing a triangular pyramid.</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="348" pageId="38" pageNumber="347" type="materials_examined">
<paragraph blockId="38.[811,1456,1976,2027]" pageId="38" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3125291324" collectionCode="LOCUS, TYPICUS" collectorName="Grande Carriere', N" country="France" location="Mont-des-Recollets" pageId="38" pageNumber="347" specimenCount="1">
<collectionCode box="[813,955,1976,1999]" pageId="38" pageNumber="347">LOCUS TYPICUS</collectionCode>
. —
<location LSID="urn:lsid:plazi:treatment:1D2F9F2B9A37FFC2FC1FFA86DE90BE43:905978E69A37FFC3FC38F842DBA3BDCF" box="[995,1182,1976,2000]" country="France" name="Mont-des-Recollets" pageId="38" pageNumber="347">Mont-des-Récollets</location>
,
<collectorName box="[1193,1382,1976,2000]" pageId="38" pageNumber="347">Grande Carrière, N</collectorName>
<collectingCountry box="[1386,1452,1976,1999]" name="France" pageId="38" pageNumber="347">France</collectingCountry>
;
<geoCoordinate box="[811,964,2003,2027]" degrees="50" direction="north" minutes="48" orientation="latitude" pageId="38" pageNumber="347" precision="1" seconds="02.74" value="50.800762">50°4802.74”N</geoCoordinate>
,
<geoCoordinate box="[975,1109,2003,2027]" degrees="2" direction="east" minutes="30" orientation="longitude" pageId="38" pageNumber="347" precision="1" seconds="23.06" value="2.5064056">2°3023.06”E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph blockId="39.[132,775,215,268]" pageId="39" pageNumber="348">
STRATUM TYPICUM.
<emphasis box="[331,356,216,239]" italics="true" pageId="39" pageNumber="348"></emphasis>
<materialsCitation ID-GBIF-Occurrence="3125291326" pageId="39" pageNumber="348">
<collectingRegion box="[364,433,215,239]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="39" pageNumber="348">Brussel</collectingRegion>
Sand Formation, base of Unit B3; (sample
<quantity box="[215,404,244,268]" metricMagnitude="2" metricUnit="m" metricValue="1.22185" metricValueMax="1.2221" metricValueMin="1.2216" pageId="39" pageNumber="348" unit="m" value="122.185" valueMax="122.21" valueMin="122.16">122.16 - 122.21 m</quantity>
), lower middle part of NP14
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[132,772,293,318]" pageId="39" pageNumber="348" type="description">
<paragraph blockId="39.[132,772,293,318]" box="[132,772,293,318]" pageId="39" pageNumber="348">
DIMENSIONS. — Diameter =
<quantity box="[419,542,294,318]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="39" pageNumber="348" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10 to 13 µm</quantity>
(
<typeStatus box="[555,645,293,317]" pageId="39" pageNumber="348">holotype</typeStatus>
: d =
<quantity box="[692,760,294,318]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="348" unit="mm" value="12.0">12 µm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="348" type="distribution">
<paragraph blockId="39.[131,776,343,594]" pageId="39" pageNumber="348">
DISTRIBUTION. —
<taxonomicName box="[326,562,343,367]" class="Coccolithophyceae" genus="Trochastrites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Discoasterales" pageId="39" pageNumber="348" phylum="Haptophyta" rank="species" species="pyramidalis" status="n. sp.">
<emphasis box="[326,562,343,367]" italics="true" pageId="39" pageNumber="348">Trochastrites pyramidalis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[568,624,343,366]" pageId="39" pageNumber="348" rank="species">n. sp.</taxonomicNameLabel>
is restricted to the lower part of the
<collectingRegion box="[353,424,371,395]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="39" pageNumber="348">Brussel</collectingRegion>
Sand Formation at the Mont-des- Récollets, with rare records in Unit B1 and Unit B3 (lower NP14). It is also known from Unit BE in the Leuven BH (base of NP14) and from the Aalter-Oedelem area and the offshore Vlakte van de Raan borehole, where it has been identified in the upper part of the Aalter Sand Formation (Unit A4, base of NP14). In this offshore borehole it seems to range up to the base of Unit B4 (sample
<quantity box="[132,217,570,594]" metricMagnitude="1" metricUnit="m" metricValue="7.915" pageId="39" pageNumber="348" unit="m" value="79.15">79.15 m</quantity>
, Steurbaut unpublished).
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="348" type="description">
<paragraph blockId="39.[131,775,653,882]" box="[132,277,653,679]" pageId="39" pageNumber="348">DESCRIPTION</paragraph>
<paragraph blockId="39.[131,775,653,882]" pageId="39" pageNumber="348">
These nannoliths present a pyramidal structure consisting of three irregularly bifurcated arms lying in a plane and interconnected with a fourth arm almost perpendicular to that plane (
<figureCitation box="[207,307,788,814]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="39" pageNumber="348">Fig. 20U</figureCitation>
). This structure is surrounded by a weblike network. The bifurcated part of the arms is quite large compared to the total length of the arms.
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="348" type="discussion">
<paragraph blockId="39.[131,776,921,1116]" box="[132,262,921,947]" pageId="39" pageNumber="348">DISCUSSION</paragraph>
<paragraph blockId="39.[131,776,921,1116]" pageId="39" pageNumber="348">
This taxon differs from
<taxonomicName authorityName="Bouche" authorityYear="1962" baseAuthorityName="Martini" baseAuthorityYear="1958" box="[391,625,955,981]" class="Coccolithophyceae" genus="Trochastrites" higherTaxonomySource="GBIF" kingdom="Chromista" order="Discoasterales" pageId="39" pageNumber="348" phylum="Haptophyta" rank="species" species="hohnensis">
<emphasis box="[391,625,955,981]" italics="true" pageId="39" pageNumber="348">Trochastrites hohnensis</emphasis>
</taxonomicName>
(
<figureCitation box="[641,732,955,981]" captionStart="FIG" captionStartId="42.[132,143,1758,1775]" captionTargetBox="[227,1362,215,1676]" captionTargetId="figure-29@42.[225,795,763,1314]" captionTargetPageId="42" captionText="FIG. 20. — New or less known calcareous nannofossil taxa from the Eocene of the Mont-des-Récollets and other localities in Belgium.Pars 3: A-G, Blackites praeinflatus n. sp.; A, Mont-des-Récollets, 122.26-122.21 m, holotype, l = 23.0 µm (IRSNB b7141); B, paratype, Mont-des-Récollets, 122.16 m, paratype, l = 22.5 µm (IRSNB b7142); C, Mont-des-Récollets, 122.26-122.21 m, paratype, l = 24.0 µm (IRSNB b7143); D, Zaventem, sample 17, paratype, l = 24.0 µm (IRSNB b7144); E, Mont-des-Récollets,122.26-122.21 m, paratype,l = 29.0 µm (IRSNB b7145); F, G, Vossem, otolith sample Nolf, 1974, paratypes (IRSNB b7146, b7147); F, l = 23.05 µm; G, l = 15.2 µm;H-K, Blackites inflatus (Bramlette &amp; Sullivan, 1961);H, Isnes,sample 8, l = 17.5 µm (IRSNB b7148);I, Gobertange,sample Go5, l = 28 µm (IRSNB b7149); J, Isnes, sample 8, l = 18 µm (IRSNB b7150); K, Gobertange, sample Go5, l = 26 µm (IRSNB b7151); L-M, Blackites aff. gladius (Locker, 1967), Isnes, sample 8 (IRSNB b7152, b7153); L, l = 10.46 µm; M, l = 11.76 µm; N-Q, Sphenolithus quadricornutus n. sp.; N, Mont-des-Récollets, 126.50 m, holotype, htot = 9 µm, hpc = hle = 2 µm (IRSNB b7154); O, Balegem, level 6, paratype, hpc + hle = 4 µm (IRSNB b7155); P, Mont-des-Récollets 126.50 m, paratype,htot = 6 µm, hpp + hle = 3 µm (IRSNB b7156); Q, Mont-des-Récollets, 122.85 m, paratype, htot = 9 µm, hpc + hle = 3 µm (IRSNB b7157); R, Sphenolithus perpendicularis Shamrock, 2010, Balegem, level 6, h = 5 µm (IRSNB b7158); S, Trochastrites hohnensis (Martini, 1958). Mont-des-Récollets, 122.85 m, wm = 12 µm (IRSNB b7159); T-W, Trochastrites pyramidalis n. sp.; T, Oedelem BH, 5.50 m, paratype, wm = 10 µm (IRSNB b7160); U, Mont-des-Récollets,122.16 m, holotype, wm = 12 µm (IRSNB b7161); V, Oedelem BH,5.50 m, paratype,wm = 13 µm (IRSNB b7162); W, Vlakte van de Raan BH,80.27 m, paratype, wm = 12 µm (IRSNB b7163)." figureDoi="http://doi.org/10.5281/zenodo.4891144" httpUri="https://zenodo.org/record/4891144/files/figure.png" pageId="39" pageNumber="348">Fig. 20S</figureCitation>
) by its pyramidal outline and the greater length of the bifurcated parts of the arms. In
<taxonomicName authority="(Martini, 1958)" baseAuthorityName="Martini" baseAuthorityYear="1958" box="[344,648,1022,1049]" class="Prymnesiophyceae" family="Coccolithaceae" genus="Discoaster" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coccosphaerales" pageId="39" pageNumber="348" phylum="Haptophyta" rank="species" species="bramlettei">
<emphasis box="[344,473,1022,1048]" italics="true" pageId="39" pageNumber="348">T. bramlettei</emphasis>
(
<bibRefCitation author="MARTINI E." box="[488,640,1022,1049]" pageId="39" pageNumber="348" pagination="353 - 388" refId="ref41605" refString="MARTINI E. 1958. - Discoasteriden und verwandte Formen in N. W. deutschen Eozan (Coccolithophorida). II Teil. Senckenbergiana Lethaea 39 (5 - 6): 353 - 388." type="journal article" year="1958">Martini, 1958</bibRefCitation>
)
</taxonomicName>
the bifurcation of the arms starts much earlier, not far from the central point where the arms meet.
</paragraph>
</subSubSection>
</treatment>
</document>