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<document id="E5457862A8C915FC52DCA00B2063F5C0" ID-DOI="10.4314/met.v32i1.2" ID-ISSN="2307-5031" ID-Zenodo-Dep="14118512" IM.bibliography_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1731461235859" checkinUser="felipe" docAuthor="Libert, Michel" docDate="2021" docId="03B287CAFF8F253FFF270E3CFB8689CE" docLanguage="en" docName="Metamorphosis.32.13-14.pdf" docOrigin="Metamorphosis 32 (1)" docSource="http://dx.doi.org/10.4314/met.v32i1.2" docStyle="DocumentStyle:73CBAE06DE6A380ACC54A94377F5FCBA.1:Metamorphosis.2021-.journal_article" docStyleId="73CBAE06DE6A380ACC54A94377F5FCBA" docStyleName="Metamorphosis.2021-.journal_article" docStyleVersion="1" docTitle="Liptena augusta Suffert 1904" docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="FF8BFFB2FF8F253FFFA30A6DFFC28C55" masterDocTitle="The types of Liptena augusta Suffert, 1904 and Liptena subundularis (Staudinger, 1892) (Papilionoidea: Lycaenidae: Poritiinae)" masterLastPageNumber="14" masterPageNumber="13" pageNumber="13" updateTime="1732200405856" updateUser="julia" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="2B9A3A729B2163104738F8F4B37E82E9">The types of Liptena augusta Suffert, 1904 and Liptena subundularis (Staudinger, 1892) (Papilionoidea: Lycaenidae: Poritiinae)</mods:title>
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<mods:namePart id="0D83E47F35640B832F49E44DD021E360">Libert, Michel</mods:namePart>
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<treatment id="03B287CAFF8F253FFF270E3CFB8689CE" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE" httpUri="http://treatment.plazi.org/id/03B287CAFF8F253FFF270E3CFB8689CE" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection id="C3016557FF8F253FFF270E3CFE08883C" box="[132,458,1105,1129]" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="8BA436DCFF8F253FFF270E3CFE08883C" blockId="0.[132,458,1105,1129]" box="[132,458,1105,1129]" pageId="0" pageNumber="13">
<heading id="D0EC81B0FF8F253FFF270E3CFE08883C" box="[132,458,1105,1129]" fontSize="10" level="2" pageId="0" pageNumber="13" reason="8">
<taxonomicName id="4C1B4D5FFF8F253FFF270E3CFE08883C" authority="Suffert, 1904" authorityName="Suffert" authorityYear="1904" box="[132,458,1105,1129]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFF270E3CFEF4883C" bold="true" box="[132,310,1105,1129]" italics="true" pageId="0" pageNumber="13">Liptena augusta</emphasis>
<bibRefCitation id="EF8A4B2DFF8F253FFE9E0E3CFE08883C" author="SUFFERT E." box="[317,458,1105,1129]" pageId="0" pageNumber="13" pagination="12 - 107" refId="ref1670" refString="SUFFERT E., 1904. Neue afrikanische Tagfalter aus dem kon. zool. Museum, Berlin, und meiner Sammlung. Deutsche Entomologische Zeitschrift, Iris, 17: 12 - 107." type="journal article" year="1904">Suffert, 1904</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016557FF8F253FFF270EE5FD8D8B63" pageId="0" pageNumber="13" type="discussion">
<paragraph id="8BA436DCFF8F253FFF270EE5FEE888A9" blockId="0.[132,775,1160,1276]" pageId="0" pageNumber="13">
The description of
<taxonomicName id="4C1B4D5FFF8F253FFEF90EE4FE0C88F5" authorityName="Suffert" authorityYear="1904" box="[346,462,1161,1184]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFEF90EE4FE0C88F5" box="[346,462,1161,1184]" italics="true" pageId="0" pageNumber="13">L. augusta</emphasis>
</taxonomicName>
is based on
<specimenCount id="9D1DFD55FF8F253FFDC20EE5FCC588F5" box="[609,775,1160,1184]" count="4" pageId="0" pageNumber="13" type="generic">four specimens</specimenCount>
collected in
<collectingCountry id="F30C764CFF8F253FFEBF0ECAFE4F88EA" box="[284,397,1191,1215]" name="Cameroon" pageId="0" pageNumber="13">Cameroon</collectingCountry>
,
<specimenCount id="9D1DFD55FF8F253FFE000ECAFDCD88EA" box="[419,527,1191,1215]" count="1" pageId="0" pageNumber="13" type="male">one male</specimenCount>
from Lolodorf (
<emphasis id="B96FEACEFF8F253FFD7C0ECAFF1F8888" italics="true" pageId="0" pageNumber="13">leg. Conradt</emphasis>
) and
<specimenCount id="9D1DFD55FF8F253FFE880EA8FE018888" box="[299,451,1221,1245]" count="3" pageId="0" pageNumber="13" type="female">three females</specimenCount>
from Bipindi (
<emphasis id="B96FEACEFF8F253FFDDE0EA8FD3C8888" box="[637,766,1221,1245]" italics="true" pageId="0" pageNumber="13">leg. Zenker</emphasis>
) (
<bibRefCitation id="EF8A4B2DFF8F253FFF280E89FEDC88A9" author="SUFFERT E." box="[139,286,1252,1276]" pageId="0" pageNumber="13" pagination="12 - 107" refId="ref1670" refString="SUFFERT E., 1904. Neue afrikanische Tagfalter aus dem kon. zool. Museum, Berlin, und meiner Sammlung. Deutsche Entomologische Zeitschrift, Iris, 17: 12 - 107." type="journal article" year="1904">Suffert, 1904</bibRefCitation>
).
</paragraph>
<paragraph id="8BA436DCFF8F253FFF270F71FD8D89B9" blockId="0.[132,774,1308,1516]" pageId="0" pageNumber="13">
<bibRefCitation id="EF8A4B2DFF8F253FFF270F71FE1B8961" author="STEMPFFER H. &amp; BENNETT N. H. &amp; MAY S. J." box="[132,473,1308,1332]" pageId="0" pageNumber="13" pagination="109 - 181" refId="ref1614" refString="STEMPFFER H., BENNETT N. H. &amp; MAY S. J., 1974. A Revision of some groups of Liptena Westwood (Lepidoptera: Lycaenidae). Bulletin of the British Museum (Natural History) (Entomology) 30 (2): 109 - 181; 6 pl." type="journal article" year="1974">
Stempffer
<emphasis id="B96FEACEFF8F253FFEA30F70FEFB8961" box="[256,313,1308,1332]" italics="true" pageId="0" pageNumber="13">et al</emphasis>
. (1974: 138)
</bibRefCitation>
considered that the
<specimenCount id="9D1DFD55FF8F253FFD790F71FF268906" count="4" pageId="0" pageNumber="13" type="generic" typeStatus="syntype">four syntypes</specimenCount>
, which had been deposited in the Berlin museum, were &quot;destroyed in the last war&quot;, and designated a male
<typeStatus id="54A0887EFF8F253FFF270F15FF1A89C5" box="[132,216,1400,1424]" pageId="0" pageNumber="13" type="neotype">neotype</typeStatus>
from
<location id="8EC46007FF8F253FFEB50F1AFE8889DA" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE:8EC46007FF8F253FFEB50F1AFE8889DA" box="[278,330,1399,1423]" county="Article" municipality="Code of Zoological Nomenclature" name="Bitje" pageId="0" pageNumber="13">Bitje</location>
(illustrated pl. 3,
<location id="8EC46007FF8F253FFDA10F1AFDF289DA" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE:8EC46007FF8F253FFDA10F1AFDF289DA" box="[514,560,1399,1423]" county="Article" municipality="Code of Zoological Nomenclature" name="Figs" pageId="0" pageNumber="13">Figs</location>
41 &amp; 44).
<location id="8EC46007FF8F253FFD3D0F15FCC089DA" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE:8EC46007FF8F253FFD3D0F15FCC089DA" box="[670,770,1400,1423]" county="Article" municipality="Code of Zoological Nomenclature" name="However" pageId="0" pageNumber="13">However</location>
, a visit to the
<location id="8EC46007FF8F253FFEBD0FFBFEA389FB" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE:8EC46007FF8F253FFEBD0FFBFEA389FB" box="[286,353,1430,1454]" county="Article" municipality="Code of Zoological Nomenclature" name="Berlin" pageId="0" pageNumber="13">Berlin</location>
museum made it possible to find the complete type-series, which invalidates this
<typeStatus id="54A0887EFF8F253FFDCB0FDBFD7E899B" box="[616,700,1462,1486]" pageId="0" pageNumber="13" type="neotype">neotype</typeStatus>
(
<collectingMunicipality id="6BC0ACA6FF8F253FFD6F0FD8FE7789B9" pageId="0" pageNumber="13">Code of Zoological Nomenclature</collectingMunicipality>
,
<collectingCounty id="62C54E50FF8F253FFE1C0FB9FDCB89B9" box="[447,521,1492,1516]" pageId="0" pageNumber="13">Article</collectingCounty>
75.8).
</paragraph>
<paragraph id="8BA436DCFF8F253FFF270C66FD8D8B63" blockId="0.[132,775,1547,1847]" pageId="0" pageNumber="13">
Moreover, the
<specimenCount id="9D1DFD55FF8F253FFE8C0C66FE048A76" box="[303,454,1547,1571]" count="4" pageId="0" pageNumber="13" type="generic" typeStatus="syntype">four syntypes</specimenCount>
do not belong to the same species, and a
<typeStatus id="54A0887EFF8F253FFE820C44FE408A14" box="[289,386,1577,1601]" pageId="0" pageNumber="13" type="lectotype">lectotype</typeStatus>
must be designated. The male from Lolodorf is most probably the specimen illustrated by
<bibRefCitation id="EF8A4B2DFF8F253FFD660C24FF078A2A" author="DRUCE H. H." pageId="0" pageNumber="13" refId="ref1391" refString="DRUCE H. H., 1910. Illustrations of African Lycaenidae, being photographic representations of type specimens contained in the Imperial Zoological Museum at Berlin. 34 pp., 8 pls. London." type="book" year="1910">Druce (1910</bibRefCitation>
, pl. III,
<figureCitation id="13202A59FF8F253FFE8A0C0AFE0B8A2A" box="[297,457,1639,1663]" captionStart="Figure 2" captionStartId="1.[380,448,924,945]" captionTargetBox="[368,1250,462,889]" captionTargetId="figure-674@1.[368,1250,462,889]" captionTargetPageId="1" captionText="Figure 2 Male genitalia of ♂ lectotype of Liptena augusta (prép. Libert, 120-030) A: right lateral view; B: ventral view of sternites 7 &amp; 8; C: dorsal view of uncus. D: reproduction of Fig. 4 in Stempffer 1957." figureDoi="http://doi.org/10.5281/zenodo.14118518" httpUri="https://zenodo.org/record/14118518/files/figure.png" pageId="0" pageNumber="13">Figs 2 and 2a</figureCitation>
); in his commentary, Druce synonymysed
<taxonomicName id="4C1B4D5FFF8F253FFE850CEBFEB98AC8" authorityName="Suffert" authorityYear="1904" box="[294,379,1670,1693]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFE850CEBFEB98AC8" box="[294,379,1670,1693]" italics="true" pageId="0" pageNumber="13">augusta</emphasis>
</taxonomicName>
with
<emphasis id="B96FEACEFF8F253FFE6B0CE8FDE38AC8" box="[456,545,1669,1693]" italics="true" pageId="0" pageNumber="13">alluaudi</emphasis>
Mabille, 1890, but
<bibRefCitation id="EF8A4B2DFF8F253FFF270CC9FEB08AE9" author="STEMPFFER H." box="[132,370,1700,1724]" pageId="0" pageNumber="13" pagination="209 - 227" refId="ref1578" refString="STEMPFFER H., 1957. Contribution a l'etude des Lycaenidae de la faune ethiopienne. Bulletin de l'Institut Francais d'Afrique Noire, serie. A, 19 (1): 209 - 227." type="journal article" year="1957">Stempffer (1957: 212)</bibRefCitation>
showed that the two species are quite distinct and reinstated
<taxonomicName id="4C1B4D5FFF8F253FFEDA0CA9FE298A8E" authorityName="Suffert" authorityYear="1904" box="[377,491,1732,1755]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFEDA0CA9FE298A8E" box="[377,491,1732,1755]" italics="true" pageId="0" pageNumber="13">L. augusta</emphasis>
</taxonomicName>
(his reasoning is repeated in
<bibRefCitation id="EF8A4B2DFF8F253FFF010C8CFE0C8AAC" author="STEMPFFER H. &amp; BENNETT N. H. &amp; MAY S. J." box="[162,462,1761,1785]" pageId="0" pageNumber="13" pagination="109 - 181" refId="ref1614" refString="STEMPFFER H., BENNETT N. H. &amp; MAY S. J., 1974. A Revision of some groups of Liptena Westwood (Lepidoptera: Lycaenidae). Bulletin of the British Museum (Natural History) (Entomology) 30 (2): 109 - 181; 6 pl." type="journal article" year="1974">
Stempffer
<emphasis id="B96FEACEFF8F253FFEB60C8FFE8F8AAC" box="[277,333,1761,1785]" italics="true" pageId="0" pageNumber="13">et al.</emphasis>
, 1974: 136
</bibRefCitation>
). The male from Lolodorf is designated as a
<typeStatus id="54A0887EFF8F253FFEED0D6DFE6D8B4D" box="[334,431,1792,1816]" pageId="0" pageNumber="13" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B96FEACEFF8F253FFE690D6CFD708B4C" bold="true" box="[458,690,1793,1817]" pageId="0" pageNumber="13">present designation</emphasis>
); it is illustrated, with the original labels (
<figureCitation id="13202A59FF8F253FFDA30D72FDFC8B62" box="[512,574,1823,1847]" captionStart="Figure 1" captionStartId="1.[377,445,412,433]" captionTargetBox="[370,1245,130,377]" captionTargetId="figure-671@1.[370,1245,130,377]" captionTargetPageId="1" captionText="Figure 1 Male lectotype of Liptena augusta with labels." figureDoi="http://doi.org/10.5281/zenodo.14118514" httpUri="https://zenodo.org/record/14118514/files/figure.png" pageId="0" pageNumber="13">
Fig.
<docIssue id="CF784933FF8F253FFD910D72FDFC8B62" box="[562,574,1823,1847]" pageId="0" pageNumber="13">1</docIssue>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016557FF8F253FFC8E0839FBC08EFC" pageId="0" pageNumber="13" type="materials_examined">
<paragraph id="8BA436DCFF8F253FFC8E0839FBC08EFC" blockId="0.[813,1455,596,681]" pageId="0" pageNumber="13">
<materialsCitation id="3B733C81FF8F253FFC8E0839FBC08EFC" collectingDate="1895-07-29" collectorName="S. Conradt" country="S Cameroun" location="Lolodorf" pageId="0" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B96FEACEFF8F253FFC8E0839FC5C8E39" bold="true" box="[813,926,596,620]" pageId="0" pageNumber="13">
<typeStatus id="54A0887EFF8F253FFC8E0839FC5C8E39" box="[813,926,596,620]" pageId="0" pageNumber="13" type="lectotype">Lectotype</typeStatus>
</emphasis>
:
<specimenCount id="9D1DFD55FF8F253FFC630839FC148E3E" box="[960,982,596,619]" count="1" pageId="0" pageNumber="13" type="male"></specimenCount>
,
<location id="8EC46007FF8F253FFC560839FB948E39" LSID="urn:lsid:plazi:treatment:03B287CAFF8F253FFF270E3CFB8689CE:8EC46007FF8F253FFC560839FB948E39" box="[1013,1110,596,620]" country="S Cameroun" name="Lolodorf" pageId="0" pageNumber="13">Lolodorf</location>
,
<collectingCountry id="F30C764CFF8F253FFBD60839FAD28E39" box="[1141,1296,596,620]" pageId="0" pageNumber="13">S Cameroun</collectingCountry>
,
<date id="FFA5101CFF8F253FFA8D0839FA698E39" box="[1326,1451,596,620]" pageId="0" pageNumber="13" value="1895-07-29">
<collectingDate id="EFE1E9F4FF8F253FFA8D0839FA698E39" box="[1326,1451,596,620]" pageId="0" pageNumber="13" value="1895-07-29">29.vii.1895</collectingDate>
</date>
, (
<emphasis id="B96FEACEFF8F253FFC95081EFC688EDE" box="[822,938,627,651]" italics="true" pageId="0" pageNumber="13">
<collectorName id="26EE530AFF8F253FFC95081EFC688EDE" box="[822,938,627,651]" pageId="0" pageNumber="13">S. Conradt</collectorName>
</emphasis>
); genitalia Libert, 120-030; Museum für Naturkunde, Berlin.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3016557FF8F253FFC8E08A4FB8689CE" pageId="0" pageNumber="13" type="discussion">
<paragraph id="8BA436DCFF8F253FFC8E08A4FB198FCC" blockId="0.[813,1456,713,1135]" pageId="0" pageNumber="13">
Thanks to Viola Richter, of the Berlin Museum, this male could be dissected, and its genitalia (
<figureCitation id="13202A59FF8F253FFB70088AFAED8EAA" box="[1235,1327,743,767]" captionStart="Figure 2" captionStartId="1.[380,448,924,945]" captionTargetBox="[368,1250,462,889]" captionTargetId="figure-674@1.[368,1250,462,889]" captionTargetPageId="1" captionText="Figure 2 Male genitalia of ♂ lectotype of Liptena augusta (prép. Libert, 120-030) A: right lateral view; B: ventral view of sternites 7 &amp; 8; C: dorsal view of uncus. D: reproduction of Fig. 4 in Stempffer 1957." figureDoi="http://doi.org/10.5281/zenodo.14118518" httpUri="https://zenodo.org/record/14118518/files/figure.png" pageId="0" pageNumber="13">Figs 2A</figureCitation>
to 2C) are identical to those illustrated by
<bibRefCitation id="EF8A4B2DFF8F253FFB2E096BFAFC8F4B" author="STEMPFFER H." box="[1165,1342,774,798]" pageId="0" pageNumber="13" pagination="209 - 227" refId="ref1578" refString="STEMPFFER H., 1957. Contribution a l'etude des Lycaenidae de la faune ethiopienne. Bulletin de l'Institut Francais d'Afrique Noire, serie. A, 19 (1): 209 - 227." type="journal article" year="1957">Stempffer, 1957</bibRefCitation>
as Fig. 4. The same figure is used by Stempffer
<emphasis id="B96FEACEFF8F253FFB660948FAC08F69" box="[1221,1282,804,828]" italics="true" pageId="0" pageNumber="13">et. al.</emphasis>
, 1974 (Fig. 18), which suggests that the male dissected by Stempffer is the one designated as a
<typeStatus id="54A0887EFF8F253FFBB0090EFBA58F2E" box="[1043,1127,867,891]" pageId="0" pageNumber="13" type="neotype">neotype</typeStatus>
in the same publication; this figure is also reproduced here (
<figureCitation id="13202A59FF8F253FFBDB09ECFB0C8FCC" box="[1144,1230,897,921]" captionStart="Figure 2" captionStartId="1.[380,448,924,945]" captionTargetBox="[368,1250,462,889]" captionTargetId="figure-674@1.[368,1250,462,889]" captionTargetPageId="1" captionText="Figure 2 Male genitalia of ♂ lectotype of Liptena augusta (prép. Libert, 120-030) A: right lateral view; B: ventral view of sternites 7 &amp; 8; C: dorsal view of uncus. D: reproduction of Fig. 4 in Stempffer 1957." figureDoi="http://doi.org/10.5281/zenodo.14118518" httpUri="https://zenodo.org/record/14118518/files/figure.png" pageId="0" pageNumber="13">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8BA436DCFF8F253FFC8E09F2FBA9883A" blockId="0.[813,1456,713,1135]" pageId="0" pageNumber="13">
The designation of the male from Lolodorf as a
<typeStatus id="54A0887EFF8F253FFAEE09F2FA6D8FE2" box="[1357,1455,927,951]" pageId="0" pageNumber="13" type="lectotype">lectotype</typeStatus>
preserves the stability of the nomenclature. It would have been quite different if the choice had been one of the females from Bipindi, since these are actually females of
<taxonomicName id="4C1B4D5FFF8F253FFA390991FB568867" authority="(Staudinger, 1892)" baseAuthorityName="Staudinger" baseAuthorityYear="1892" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="subundularis">
<emphasis id="B96FEACEFF8F253FFA390991FC7A8867" italics="true" pageId="0" pageNumber="13">L. subundularis</emphasis>
(Staudinger, 1892)
</taxonomicName>
, and it would have been necessary to find a replacement name for
<taxonomicName id="4C1B4D5FFF8F253FFB5A0E54FA8F8805" authorityName="Suffert" authorityYear="1904" box="[1273,1357,1081,1104]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFB5A0E54FA8F8805" box="[1273,1357,1081,1104]" italics="true" pageId="0" pageNumber="13">augusta</emphasis>
</taxonomicName>
, all with significant risks of confusion.
</paragraph>
<paragraph id="8BA436DCFF8F253FFC8E0EE3FB8689CE" blockId="0.[813,1456,1166,1436]" pageId="0" pageNumber="13">
This is also why it is propitious that the decision by
<bibRefCitation id="EF8A4B2DFF8F253FFAF00EE3FC7A8890" author="SCHULTZE A." pageId="0" pageNumber="13" pagination="1113 - 1242" refId="ref1539" refString="SCHULTZE A., 1923, in SCHULTZE A. &amp; AURIVILLIUS C., 1923 - Lepidoptera. III. Teil. Ergebnisse der Zweiten Deutschen Zentral-Afrika Expedition, 1 (17): 1113 - 1242." type="journal article" year="1923">Schultze (1923: 1179)</bibRefCitation>
to synonymyse
<taxonomicName id="4C1B4D5FFF8F253FFBC40EC3FB7E8890" authorityName="Suffert" authorityYear="1904" box="[1127,1212,1198,1221]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFBC40EC3FB7E8890" box="[1127,1212,1198,1221]" italics="true" pageId="0" pageNumber="13">augusta</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1B4D5FFF8F253FFB5A0EC0FA468890" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[1273,1412,1197,1221]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="subundularis">
<emphasis id="B96FEACEFF8F253FFB5A0EC0FA468890" box="[1273,1412,1197,1221]" italics="true" pageId="0" pageNumber="13">subundularis</emphasis>
</taxonomicName>
has fallen into disuse, which is perfectly justified since Schultze overlooked the fact that the male and the females of the type-series did not belong to the same species. It is however preferable to emphasise it [the same error is found in Stempffer
<emphasis id="B96FEACEFF8F253FFC020F2AFC18890B" box="[929,986,1350,1374]" italics="true" pageId="0" pageNumber="13">et. al</emphasis>
. when they contest Schultze's decision (p. 156), but they could not avoid it as they had not seen the type-series of
<taxonomicName id="4C1B4D5FFF8F253FFC670FE8FBF789C9" authorityName="Suffert" authorityYear="1904" box="[964,1077,1413,1436]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="augusta">
<emphasis id="B96FEACEFF8F253FFC670FE8FBF789C9" box="[964,1077,1413,1436]" italics="true" pageId="0" pageNumber="13">L. augusta</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>