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<document id="5F1AB8B8363FCA5F1AFA765F66AC1EE0" ID="10.11646/zootaxa.4281.1.10SLASH11443" ID-CLB-Dataset="28722" ID-DOI="10.11646/zootaxa.4281.1.10" ID-GBIF-Dataset="a138fb3c-5b32-465d-9fed-9e79e130f803" ID-ISSN="1175-5326" ID-Zenodo-Dep="815973" ID-ZooBank="B5DB709B-4966-4C6F-A7A7-B9639EE0853F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1498119172040" checkinUser="plazi" docAuthor="Gonzalez-Mozo, Laura, Mckamey, Stuart, Ware, Jessica L. &amp; Hamilton, George" docDate="2017" docId="03B287ABAB21E931FF74FF19FA64F8A4" docLanguage="en" docName="zootaxa.4281.1.10SLASH11443.pdf" docOrigin="Zootaxa 4281 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyphotes colombiensis Gonzalez-Mozo &amp; Mckamey &amp; Ware &amp; Hamilton 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="110" masterDocId="FF8BFFD3AB23E933FFE3FFC4FFEEFFC9" masterDocTitle="Two new species of Darnini (Hemiptera: Membracidae) from Colombia and Peru" masterLastPageNumber="114" masterPageNumber="108" pageNumber="110" updateTime="1698448196812" updateUser="ExternalLinkService">
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<mods:title id="5AD48A43D47FF23999ED75F74FB14D4B">Two new species of Darnini (Hemiptera: Membracidae) from Colombia and Peru</mods:title>
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<mods:namePart id="6F92840A0B98BCCBD2CF7F815FA0C2B3">Gonzalez-Mozo, Laura</mods:namePart>
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<mods:namePart id="096E440EDA5780ED5B1037A3715D38D3">Mckamey, Stuart</mods:namePart>
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<mods:namePart id="B89FD4FC1030A59953DEA56509682999">Ware, Jessica L.</mods:namePart>
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<mods:namePart id="31135A67C6386814C045095986E3A8B1">Hamilton, George</mods:namePart>
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<mods:title id="E12226952222AFC39500460CD4B35AFC">Zootaxa</mods:title>
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<mods:date id="4267BF454D569A7C04A9A65765FC9AFA">2017</mods:date>
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<treatment id="03B287ABAB21E931FF74FF19FA64F8A4" ID-DOI="http://doi.org/10.5281/zenodo.6042714" ID-GBIF-Taxon="150808920" ID-Zenodo-Dep="6042714" LSID="urn:lsid:plazi:treatment:03B287ABAB21E931FF74FF19FA64F8A4" httpUri="http://treatment.plazi.org/id/03B287ABAB21E931FF74FF19FA64F8A4" lastPageNumber="110" pageId="2" pageNumber="110">
<subSubSection id="C3016536AB21E931FF74FF19FEC0FED1" pageId="2" pageNumber="110" type="nomenclature">
<paragraph id="8BA436BDAB21E931FF74FF19FD37FF3E" blockId="2.[151,729,221,280]" box="[151,729,221,247]" pageId="2" pageNumber="110">
<heading id="D0EC81D1AB21E931FF74FF19FD37FF3E" bold="true" box="[151,729,221,247]" fontSize="11" level="1" pageId="2" pageNumber="110" reason="1">
<emphasis id="B96FEAAFAB21E931FF74FF19FD37FF3E" bold="true" box="[151,729,221,247]" pageId="2" pageNumber="110">
<taxonomicName id="4C1B4D3EAB21E931FF74FF19FD9DFF3E" authority="Gonzalez-Mozo" authorityName="Gonzalez-Mozo &amp; Mckamey &amp; Ware &amp; Hamilton" authorityYear="2017" box="[151,627,221,247]" class="Insecta" family="Membracidae" genus="Cyphotes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="B96FEAAFAB21E931FF74FF19FE47FF3E" bold="true" box="[151,425,221,247]" italics="true" pageId="2" pageNumber="110">Cyphotes colombiensis</emphasis>
González-Mozo
</taxonomicName>
<taxonomicNameLabel id="A25C57D4AB21E931FD98FF19FD37FF3E" box="[635,729,221,247]" pageId="2" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436BDAB21E931FF74FF3BFEC0FED1" blockId="2.[151,729,221,280]" box="[151,302,255,280]" pageId="2" pageNumber="110">
(
<figureCitation id="13202A38AB21E931FF7DFF3BFEC9FED1" box="[158,295,255,280]" captionStart="FIGURES 2 6" captionStartId="3.[151,264,1940,1963]" captionTargetBox="[198,1452,208,1893]" captionTargetId="figure@3.[167,1452,193,1917]" captionTargetPageId="3" captionText="FIGURES 2 6. Cyphotes colombiensis sp. nov, holotype female (IAvH). 2 4. Anterior, lateral and dorsal view; 5 6. Forewing venation, second female valvulae." httpUri="https://zenodo.org/record/815977/files/figure.png" pageId="2" pageNumber="110">Figures 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF74FE83FAA4FE4D" pageId="2" pageNumber="110" type="diagnosis">
<paragraph id="8BA436BDAB21E931FF74FE83FAA4FE4D" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF74FE83FEFFFEA9" bold="true" box="[151,273,327,352]" pageId="2" pageNumber="110">Diagnosis.</emphasis>
Base of pronotum, in dorsal view, with four lateral longitudinal rugae; two conical middorsal peaks behind humeral angles; one wide black band on pronotum that attains most posterior swelling of pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF24FE4BFE9CFB09" pageId="2" pageNumber="110" type="description">
<paragraph id="8BA436BDAB21E931FF24FE4BFAA4FB9C" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FE4BFEBFFE61" bold="true" box="[199,337,399,424]" pageId="2" pageNumber="110">Description</emphasis>
.
<typeStatus id="54A0881FAB21E931FE81FE4BFE24FE61" box="[354,458,399,424]" pageId="2" pageNumber="110" type="holotype">Holotype</typeStatus>
female. Head brown; pronotum (in dorsal view) with wide black band running from anterior margin to humeral angles, tapering from mid-dorsal portion of pronotum to posterior process, where it expands again, covering most of posterior swelling; dorsolateral margin of pronotum reddish; forewing punctate, semi-opaque between costal margin and R vein, mostly hyaline, punctate basally between veins R, M, and Cu; ventral aspect of head, thorax, legs and abdomen brownish.
<emphasis id="B96FEAAFAB21E931FCBBFDDBFC7AFDF1" box="[856,916,543,568]" italics="true" pageId="2" pageNumber="110">Head</emphasis>
. Subquadrate; densely pubescent; more than twice as long as wide; superior margin arcuate; vertex oblique in lateral view; coronal suture distinctly grooved; frontoclypeus almost tubular, directed forward, anteclypeus directed ventrad; ocelli closer to each other than to eyes; suprantennal ledges oblique.
<emphasis id="B96FEAAFAB21E931FDFDFD49FD63FD6D" box="[542,653,653,676]" italics="true" pageId="2" pageNumber="110">Pronotum</emphasis>
. Rough, pubescent, concealing forewings at rest; metopidium convex in dorsal view; humeral angles subtriangular, projected laterally, notched posterobasally; median carina percurrent, elevated; four longitudinal rugae extended from basal margin of pronotum to tip of humeral angles; two middorsal parallel conic peaks behind humeral angles at each side of medial carina, separated by mediolateral constriction; posterior process with two latero-posterior swellings, slightly compressed anteroposteriorly, strongly expanded laterally behind mediolateral constriction; apex of posterior process acute. In lateral view, medial carina elevated, sinuate; two swellings, one above humeral angles, followed by slight depression, then rising again, forming second swelling, extended from mediolateral constriction to latero-posterior swellings, gradually descending to pronotal apex.
<emphasis id="B96FEAAFAB21E931FF3FFC69FEBEFC0D" box="[220,336,941,964]" italics="true" pageId="2" pageNumber="110">Forewing.</emphasis>
One
<emphasis id="B96FEAAFAB21E931FE71FC69FE73FC0D" box="[402,413,941,964]" italics="true" pageId="2" pageNumber="110">s</emphasis>
, one
<emphasis id="B96FEAAFAB21E931FE3CFC69FDE9FC0D" box="[479,519,941,964]" italics="true" pageId="2" pageNumber="110">r-m</emphasis>
, and two
<emphasis id="B96FEAAFAB21E931FD9DFC69FD58FC0D" box="[638,694,941,964]" italics="true" pageId="2" pageNumber="110">m-cu</emphasis>
crossveins; two discoidal cells; five apical cells.
<emphasis id="B96FEAAFAB21E931FB17FC69FADEFC0D" box="[1268,1328,941,964]" italics="true" pageId="2" pageNumber="110">Legs.</emphasis>
Pro- and mesothoracic femora subcylindrical; ventral aspect striate; two ventral longitudinal rows of cucullate setae; tibiae spatulate, with two ventral proximal rows of cucullate setae. Metathoracic femur without rows of cucullate setae; tibia with three single rows of cucullate setae; first tarsomere with two distal setae. Abdomen strongly sclerotized; conspicuously punctate ventrally.
<emphasis id="B96FEAAFAB21E931FDF7FBF8FD6BFB9C" box="[532,645,1084,1109]" italics="true" pageId="2" pageNumber="110">Genitalia.</emphasis>
Second valvulae blade-shaped, lacking teeth on dorsal surface.
</paragraph>
<paragraph id="8BA436BDAB21E931FF24FB9BFE69FBB1" blockId="2.[151,1436,327,1901]" box="[199,391,1119,1144]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FB9BFEE5FBB1" bold="true" box="[199,267,1119,1144]" pageId="2" pageNumber="110">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA436BDAB21E931FF24FB40FE9CFB09" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FB40FE98FB54" bold="true" box="[199,374,1156,1181]" pageId="2" pageNumber="110">Measurements</emphasis>
(mm): Total length 7.4; head width 2.5; head length 1.5; pronotum length 6.6; width between humeral angles 4.6.
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF24FB08FB8CFAC1" pageId="2" pageNumber="110" type="materials_examined">
<paragraph id="8BA436BDAB21E931FF24FB08FB8CFAC1" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<materialsCitation id="3B733CE0AB21E931FF24FB08FB8CFAC1" ID-GBIF-Occurrence="1950907973" collectingDate="1998-06-22" collectorName="de Iguaque &amp; Golpeteo &amp; D. Diaz &amp; Catalogue No." country="Colombia" elevation="2300" latitude="5.4166665" location="Boyaca" longLatPrecision="1305" longitude="-73.45" pageId="2" pageNumber="110" specimenCount="1" stateProvince="Boyaca" typeStatus="holotype">
<emphasis id="B96FEAAFAB21E931FF24FB08FE54FB2C" bold="true" box="[199,442,1228,1253]" pageId="2" pageNumber="110">Material Examined.</emphasis>
<typeStatus id="54A0881FAB21E931FE27FB08FDB6FB2D" box="[452,600,1228,1252]" pageId="2" pageNumber="110" type="holotype">HOLOTYPE</typeStatus>
female, “
<collectingCountry id="F30C762DAB21E931FD2AFB08FC8AFB2D" box="[713,868,1228,1252]" name="Colombia" pageId="2" pageNumber="110">COLOMBIA</collectingCountry>
,
<collectingRegion id="49DFF85FAB21E931FC91FB08FC27FB2D" box="[882,969,1228,1252]" country="Colombia" name="Boyaca" pageId="2" pageNumber="110">Boyacá</collectingRegion>
, SFF
<collectorName id="26EE536BAB21E931FBF2FB08FB7EFB2C" box="[1041,1168,1228,1253]" pageId="2" pageNumber="110">de Iguaque</collectorName>
<geoCoordinate id="EE2F507AAB21E931FB5BFB08FAE4FB2C" box="[1208,1290,1228,1253]" direction="north" orientation="latitude" pageId="2" pageNumber="110" precision="925" value="5.4166665">5°25N</geoCoordinate>
,
<geoCoordinate id="EE2F507AAB21E931FAFAFB08FA90FB2D" box="[1305,1406,1228,1252]" direction="west" orientation="longitude" pageId="2" pageNumber="110" precision="925" value="-73.45">73°27W</geoCoordinate>
<elevation id="0036D18EAB21E931FF74FB34FF01FAC1" box="[151,239,1264,1288]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="110" unit="m" value="2300.0">
<quantity id="4CE39B58AB21E931FF74FB34FF01FAC1" box="[151,239,1264,1288]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="110" unit="m" value="2300.0">2300 m</quantity>
</elevation>
. │
<collectorName id="26EE536BAB21E931FEF6FB2BFE95FAC1" box="[277,379,1263,1288]" pageId="2" pageNumber="110">Golpeteo</collectorName>
<collectingDate id="EFE1E995AB21E931FE7EFB2BFDF7FAC1" box="[413,537,1263,1288]" pageId="2" pageNumber="110" value="1998-06-22">
<date id="FFA5107DAB21E931FE7EFB2BFDF7FAC1" box="[413,537,1263,1288]" pageId="2" pageNumber="110" value="1998-06-22">22-vi-1998</date>
</collectingDate>
<collectorName id="26EE536BAB21E931FDD8FB35FD7AFAC1" box="[571,660,1264,1288]" pageId="2" pageNumber="110">D. Díaz</collectorName>
” (IavH,
<collectorName id="26EE536BAB21E931FD16FB2BFC79FAC1" box="[757,919,1263,1288]" pageId="2" pageNumber="110">Catalogue No.</collectorName>
IAvH-E-
<date id="FFA5107DAB21E931FBE0FB34FBB9FAC1" box="[1027,1111,1264,1288]" pageId="2" pageNumber="110" value="17-11-14">141117</date>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF24FAD0FDAEFAE5" box="[199,576,1300,1325]" pageId="2" pageNumber="110" type="distribution">
<paragraph id="8BA436BDAB21E931FF24FAD0FDAEFAE5" blockId="2.[151,1436,327,1901]" box="[199,576,1300,1325]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FAD0FE8EFAE4" bold="true" box="[199,352,1300,1325]" pageId="2" pageNumber="110">Distribution.</emphasis>
<collectingCountry id="F30C762DAB21E931FE84FAD0FE34FAE4" box="[359,474,1300,1325]" name="Colombia" pageId="2" pageNumber="110">Colombia</collectingCountry>
:
<collectingRegion id="49DFF85FAB21E931FE07FAD0FDD2FAE5" box="[484,572,1300,1324]" country="Colombia" name="Boyaca" pageId="2" pageNumber="110">Boyacá</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF24FAF3FB5EFA99" box="[199,1200,1335,1361]" pageId="2" pageNumber="110" type="etymology">
<paragraph id="8BA436BDAB21E931FF24FAF3FB5EFA99" blockId="2.[151,1436,327,1901]" box="[199,1200,1335,1361]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FAF3FEA0FA99" bold="true" box="[199,334,1335,1360]" pageId="2" pageNumber="110">Etymology.</emphasis>
This species is named according to the country where the species was found.
</paragraph>
</subSubSection>
<subSubSection id="C3016536AB21E931FF24FA98FA64F8A4" pageId="2" pageNumber="110" type="discussion">
<paragraph id="8BA436BDAB21E931FF24FA98FB91F8C9" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<emphasis id="B96FEAAFAB21E931FF24FA98FEA6FABC" bold="true" box="[199,328,1372,1397]" pageId="2" pageNumber="110">Comments</emphasis>
.
<taxonomicName id="4C1B4D3EAB21E931FEBDFA98FE28FABC" box="[350,454,1372,1397]" class="Insecta" family="Membracidae" genus="Cyphotes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FEBDFA98FE28FABC" box="[350,454,1372,1397]" italics="true" pageId="2" pageNumber="110">Cyphotes</emphasis>
</taxonomicName>
is recognized by the following morphological features: head projected forward, pronotum rugose, with swollen areas (
<bibRefCitation id="EF8A4B4CAB21E931FD9EFABBFCA1FA51" author="Burmeister" box="[637,847,1407,1432]" pageId="2" pageNumber="110" refString="Burmeister, H. (1835) Schnabelkerfe. Rhynchota. In: Handbuch der Entomologie. 2 (1). Gustav Reimer, Berlin, pp. 1 - 396. [overall work published 1832 - 1855]" type="book chapter" year="1835">Burmeister, 1835</bibRefCitation>
;
<bibRefCitation id="EF8A4B4CAB21E931FC84FABBFBC1FA51" author="Silberman" box="[871,1071,1407,1432]" pageId="2" pageNumber="110" refString="Silbermann, G. (1836) Revue Entomologique. Tome IU. A Strasbourg, Au bureau de la revue entomologique, place Saint- Thomas, Paris, 286 pp." type="book" year="1836">Silberman, 1836</bibRefCitation>
,
<bibRefCitation id="EF8A4B4CAB21E931FBA5FABBFB00FA51" author="Goding" box="[1094,1262,1407,1432]" pageId="2" pageNumber="110" refString="Goding, F. (1926) New genera and species of Membracidae. Transactions of the American Entomological Society, 52 (2), 103 - 110." type="journal article" year="1926">Goding, 1926</bibRefCitation>
,
<bibRefCitation id="EF8A4B4CAB21E931FAE6FA45FE96FA74" author="Fonseca" pageId="2" pageNumber="110" refString="Fonseca, J. P. &amp; Diringshofen, R. V. (1969) Contribuicao ao conhecimento dos membracideos neotropicos (Homoptera: Membracidae, VI). Arquivos do Instituto Biologico, 36, 143 - 161." type="journal article" year="1969">Fonseca and Diringshofen, 1969</bibRefCitation>
), and the presence of crossvein
<emphasis id="B96FEAAFAB21E931FD0CFA61FD14FA75" box="[751,762,1445,1468]" italics="true" pageId="2" pageNumber="110">s</emphasis>
(
<bibRefCitation id="EF8A4B4CAB21E931FCEFFA60FC62FA74" author="Deitz" box="[780,908,1444,1469]" pageId="2" pageNumber="110" refString="Deitz, L. L. (1975) Classification of the higher categories of the New World treehoppers (Homoptera: Membracidae). North Carolina Agricultural Experiment Station, Technical Bulletin, 225, 1 - 177." type="journal article" year="1975">Deitz,1975</bibRefCitation>
;
<bibRefCitation id="EF8A4B4CAB21E931FC78FA60FB9FFA74" author="Sakakibara" box="[923,1137,1444,1469]" pageId="2" pageNumber="110" refString="Sakakibara, A. M. (2005 a) The species of Aspona Stal and nomenclatural notes (Hemiptera, Cicadomorpha, Membracidae). Revista Brasileira de Entomologia, 49 (4), 462 - 464." type="journal article" year="2005" yearSuffix="a">Sakakibara, 2005a</bibRefCitation>
;
<bibRefCitation id="EF8A4B4CAB21E931FB9CFA60FB26FA74" author="Sakakibara" box="[1151,1224,1444,1469]" pageId="2" pageNumber="110" refString="Sakakibara, A. M. (2005 b) The genus Hypheodana Metcalf and description of three new species (Hemiptera, Cicadomorpha, Membracidae). Revista Brasileira de Zoologia, 22 (4), 1116 - 1120." type="journal article" year="2005" yearSuffix="b">2005b</bibRefCitation>
;
<bibRefCitation id="EF8A4B4CAB21E931FB34FA60FAFAFA74" author="Sakakibara" box="[1239,1300,1444,1469]" pageId="2" pageNumber="110" refString="Sakakibara, A. M. (2006) A new genus and three new species of treehopper tribe Darnini (Hemiptera, Membracidae). Biociencias, Porto Alegre 14 (2), 189 - 192." type="journal article" year="2006">2006</bibRefCitation>
). All of the above attributes are present in
<taxonomicName id="4C1B4D3EAB21E931FE0DFA0CFD4FFA29" authorityName="Gonzalez-Mozo &amp; Mckamey &amp; Ware &amp; Hamilton" authorityYear="2017" box="[494,673,1479,1504]" class="Insecta" family="Membracidae" genus="Cyphotes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="colombiensis">
<emphasis id="B96FEAAFAB21E931FE0DFA0CFD4FFA29" box="[494,673,1479,1504]" italics="true" pageId="2" pageNumber="110">C. colombiensis</emphasis>
</taxonomicName>
. However, some are also observed in related genera within Darnini, such as
<taxonomicName id="4C1B4D3EAB21E931FF12FA29FE2FF9CC" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[241,449,1516,1541]" class="Insecta" family="Membracidae" genus="Aspona" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FF12FA29FEABF9CD" box="[241,325,1517,1540]" italics="true" pageId="2" pageNumber="110">Aspona</emphasis>
Stal, 1862
</taxonomicName>
;
<taxonomicName id="4C1B4D3EAB21E931FE2EFA28FCEBF9CC" authority="Metcalf, 1954" authorityName="Metcalf" authorityYear="1954" box="[461,773,1516,1541]" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FE2EFA28FDB2F9CC" box="[461,604,1516,1541]" italics="true" pageId="2" pageNumber="110">Hypheodana</emphasis>
Metcalf, 1954
</taxonomicName>
;
<taxonomicName id="4C1B4D3EAB21E931FCF2FA29FBA1F9CC" authority="Sakakibara,2006" authorityName="Sakakibara" authorityYear="2006" box="[785,1103,1516,1541]" class="Insecta" family="Membracidae" genus="Nasuconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FCF2FA29FC67F9CD" box="[785,905,1517,1540]" italics="true" pageId="2" pageNumber="110">Nasuconia</emphasis>
<bibRefCitation id="EF8A4B4CAB21E931FC73FA28FBA1F9CC" author="Sakakibara" box="[912,1103,1516,1541]" pageId="2" pageNumber="110" refString="Sakakibara, A. M. (2006) A new genus and three new species of treehopper tribe Darnini (Hemiptera, Membracidae). Biociencias, Porto Alegre 14 (2), 189 - 192." type="journal article" year="2006">Sakakibara,2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C1B4D3EAB21E931FBB8FA29FA89F9CD" authority="Fonseca, 1934" authorityName="Pinto da Fonseca" authorityYear="1934" box="[1115,1383,1516,1540]" class="Insecta" family="Membracidae" genus="Taunaya" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FBB8FA29FB54F9CD" box="[1115,1210,1517,1540]" italics="true" pageId="2" pageNumber="110">Taunaya</emphasis>
Fonseca, 1934
</taxonomicName>
; and
<emphasis id="B96FEAAFAB21E931FF74F9CBFEA1F9E1" box="[151,335,1551,1576]" italics="true" pageId="2" pageNumber="110">Funkhouseriana</emphasis>
Creão-Duarte, 1999. The head projected forward is observed in the three related genera:
<taxonomicName id="4C1B4D3EAB21E931FAA2F9D4FA7BF9EE" authorityName="Stal" authorityYear="1862" box="[1345,1429,1552,1575]" class="Insecta" family="Membracidae" genus="Aspona" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FAA2F9D4FA7BF9EE" box="[1345,1429,1552,1575]" italics="true" pageId="2" pageNumber="110">Aspona</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1B4D3EAB21E931FF74F9F1FEE1F985" authorityName="Sakakibara" authorityYear="2006" box="[151,271,1589,1612]" class="Insecta" family="Membracidae" genus="Nasuconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FF74F9F1FEE1F985" box="[151,271,1589,1612]" italics="true" pageId="2" pageNumber="110">Nasuconia</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1B4D3EAB21E931FEACF9F0FE30F984" authorityName="Metcalf" authorityYear="1954" box="[335,478,1588,1613]" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FEACF9F0FE30F984" box="[335,478,1588,1613]" italics="true" pageId="2" pageNumber="110">Hypheodana</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1B4D3EAB21E931FE13F9F1FBFEF985" authority="Sakalibara &amp; Lencioni-Neto, 2009" authorityName="Sakalibara &amp; Lencioni-Neto" authorityYear="2009" box="[496,1040,1588,1613]" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="gargionei">
<emphasis id="B96FEAAFAB21E931FE13F9F1FD91F985" box="[496,639,1589,1612]" italics="true" pageId="2" pageNumber="110">H. gargionei</emphasis>
Sakalibara &amp;
<bibRefCitation id="EF8A4B4CAB21E931FCC6F9F0FBFEF985" author="Lencioni-Neto" box="[805,1040,1588,1613]" pageId="2" pageNumber="110" refString="Sakakibara, A. M &amp; Lencioni-Neto, F. (2009) Novos Membracideos (Hemiptera, Membracidae) do Brasil. Papeis Avulsos de Zoologia, 49 (16), 191 - 196." type="journal article" year="2009">Lencioni-Neto, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1B4D3EAB21E931FBA9F9F1FA7FF984" authority="Sakakibara, 2005" authorityName="Sakakibara" authorityYear="2005" box="[1098,1425,1588,1613]" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B96FEAAFAB21E931FBA9F9F1FB51F985" box="[1098,1215,1589,1612]" italics="true" pageId="2" pageNumber="110">H. costata</emphasis>
Sakakibara, 2005
</taxonomicName>
). Swollen areas on the pronotum are known in
<taxonomicName id="4C1B4D3EAB21E931FD5EF99CFC35F9B9" authority="Funkhouseriana" authorityName="Stål" authorityYear="1862" box="[701,987,1623,1648]" class="Insecta" family="Membracidae" genus="Aspona" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FD5EF99CFCFFF9A6" box="[701,785,1624,1647]" italics="true" pageId="2" pageNumber="110">Aspona</emphasis>
,
<emphasis id="B96FEAAFAB21E931FCC7F993FC35F9B9" box="[804,987,1623,1648]" italics="true" pageId="2" pageNumber="110">Funkhouseriana</emphasis>
</taxonomicName>
<emphasis id="B96FEAAFAB21E931FC0BF993FBD2F9B9" box="[1000,1084,1623,1648]" italics="true" pageId="2" pageNumber="110">cerulea</emphasis>
(Funkhouser, 1935),
<taxonomicName id="4C1B4D3EAB21E931FADFF99DFE76F95D" authority="Fonseca, 1934" authorityName="Fonseca" authorityYear="1934" class="Insecta" family="Membracidae" genus="Taunaya" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B96FEAAFAB21E931FADFF99DFF0BF95D" italics="true" pageId="2" pageNumber="110">Taunaya rugosa</emphasis>
Fonseca, 1934
</taxonomicName>
,
<taxonomicName id="4C1B4D3EAB21E931FE44F9B9FC63F95C" authority="Sakakibara, 2006" authorityName="Sakakibara" authorityYear="2006" box="[423,909,1660,1685]" class="Insecta" family="Membracidae" genus="Nasuconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="curculionida">
<emphasis id="B96FEAAFAB21E931FE44F9B9FD56F95C" box="[423,696,1660,1685]" italics="true" pageId="2" pageNumber="110">Nasuconia curculionida</emphasis>
<bibRefCitation id="EF8A4B4CAB21E931FD27F9B8FC63F95C" author="Sakakibara" box="[708,909,1660,1685]" pageId="2" pageNumber="110" refString="Sakakibara, A. M. (2006) A new genus and three new species of treehopper tribe Darnini (Hemiptera, Membracidae). Biociencias, Porto Alegre 14 (2), 189 - 192." type="journal article" year="2006">Sakakibara, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1B4D3EAB21E931FC7FF9B9FB38F95C" authority="Fairmaire, 1846" authorityName="Fairmaire" authorityYear="1846" box="[924,1238,1660,1685]" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B96FEAAFAB21E931FC7FF9B9FC11F95D" box="[924,1023,1661,1684]" italics="true" pageId="2" pageNumber="110">H. ursus</emphasis>
(Fairmaire, 1846)
</taxonomicName>
, and
<taxonomicName id="4C1B4D3EAB21E931FAFAF9B9FE8FF971" authority="Sakakibara, 2005" authorityName="Sakakibara" authorityYear="2005" class="Insecta" family="Membracidae" genus="Hypheodana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="proxima">
<emphasis id="B96FEAAFAB21E931FAFAF9B9FA72F95D" box="[1305,1436,1661,1684]" italics="true" pageId="2" pageNumber="110">H. proxima</emphasis>
Sakakibara, 2005
</taxonomicName>
. The projection and shape of the humeral angles and the forewing venation are also variable across these genera. This suggests that these genera need to be revised based on a morphological assessment of the mentioned features, which should include characters of hind wing venation and genitalia.
</paragraph>
<paragraph id="8BA436BDAB21E931FF24F8C8FA64F8A4" blockId="2.[151,1436,327,1901]" pageId="2" pageNumber="110">
<materialsCitation id="3B733CE0AB21E931FF24F8C8FA64F8A4" ID-GBIF-Occurrence="1950907974" collectorName="Diversidad de Insectos &amp; Instituto de Recursos Biologicos Alexander von Humboldt'" country="Colombia" location="This" municipality="The" pageId="2" pageNumber="110" specimenCount="2" typeStatus="holotype">
<location id="8EC46066AB21E931FF24F8C8FF17F8EC" LSID="urn:lsid:plazi:treatment:03B287ABAB21E931FF74FF19FA64F8A4:8EC46066AB21E931FF24F8C8FF17F8EC" box="[199,249,1804,1829]" country="Colombia" municipality="The" name="This" pageId="2" pageNumber="110">This</location>
is the first record of the genus
<taxonomicName id="4C1B4D3EAB21E931FDBFF8C8FD2AF8EC" box="[604,708,1804,1829]" class="Insecta" family="Membracidae" genus="Cyphotes" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAAFAB21E931FDBFF8C8FD2AF8EC" box="[604,708,1804,1829]" italics="true" pageId="2" pageNumber="110">Cyphotes</emphasis>
</taxonomicName>
from
<collectingCountry id="F30C762DAB21E931FCE9F8C8FC92F8EC" box="[778,892,1804,1829]" name="Colombia" pageId="2" pageNumber="110">Colombia</collectingCountry>
.
<collectingMunicipality id="6BC0ACC7AB21E931FC64F8C8FC5DF8EC" box="[903,947,1804,1829]" pageId="2" pageNumber="110">The</collectingMunicipality>
<typeStatus id="54A0881FAB21E931FC58F8C8FBF3F8EC" box="[955,1053,1804,1829]" pageId="2" pageNumber="110" type="holotype">holotype</typeStatus>
was collected through the project “
<collectorName id="26EE536BAB21E931FF42F8EBFE41F88E" box="[161,431,1839,1864]" pageId="2" pageNumber="110">Diversidad de Insectos</collectorName>
en
<collectingCountry id="F30C762DAB21E931FE00F8EBFDB8F881" box="[483,598,1839,1864]" name="Colombia" pageId="2" pageNumber="110">Colombia</collectingCountry>
”, coordinated by the
<collectorName id="26EE536BAB21E931FC83F8EBFEFCF8A4" pageId="2" pageNumber="110">Instituto de Recursos Biológicos Alexander von Humboldt</collectorName>
(IAvH). The
<typeStatus id="54A0881FAB21E931FE49F890FDE2F8A4" box="[426,524,1876,1901]" pageId="2" pageNumber="110" type="holotype">holotype</typeStatus>
is preserved in alcohol, due to its damaged metopidium and some missing legs.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>