201 lines
34 KiB
XML
201 lines
34 KiB
XML
<document id="62A0E5FDDE1A2BD51AD6CD5EB88F3708" ID-DOI="10.11646/zootaxa.4040.1.1" ID-GBIF-Dataset="3b927167-5686-4666-861f-75b65e82fcdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="289973" ID-ZooBank="51219593-CC2D-4077-B9FA-5E2F3255FC24" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333548483" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, Taiti, Stefano & Sfenthourakis, Spyros" docDate="2015" docId="03B287A20E3B4037FF06FE621C0CFEAC" docLanguage="en" docName="zt04040p016.pdf" docOrigin="Zootaxa 4040 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Levantoniscus bicostulatus Cardoso, Taiti & Sfenthourakis, 2015, n. sp." docType="treatment" docVersion="6" lastPageNumber="4" masterDocId="FF8BFFDA0E394034FF91FFD51D0EFFB1" masterDocTitle="A new genus of Trachelipodidae Strouhal, 1953 (Crustacea, Isopoda, Oniscidea) from the eastern Mediterranean" masterLastPageNumber="16" masterPageNumber="1" pageNumber="3" updateTime="1698618859390" updateUser="plazi">
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<mods:title id="9B6E233B93D20C8A46903EBC2C2A04E8">A new genus of Trachelipodidae Strouhal, 1953 (Crustacea, Isopoda, Oniscidea) from the eastern Mediterranean</mods:title>
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<mods:name id="A290106959B1DF63C3EBC754632FAC18" type="personal">
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<mods:namePart id="4BF56E9A83483A294C5C051287734442">Cardoso, Giovanna Monticelli</mods:namePart>
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<mods:namePart id="3A19CBBCB2A55588BAF9DD8273A9F8E8">Taiti, Stefano</mods:namePart>
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<mods:roleTerm id="D4B3C437B09B66B1D778507D8360FB79">Author</mods:roleTerm>
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<mods:namePart id="A0426C67AE73776555427F71E1A55963">Sfenthourakis, Spyros</mods:namePart>
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<mods:typeOfResource id="97EBB592ABC4A7B0670A94DC7B32E81B">text</mods:typeOfResource>
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<mods:title id="B0DFF9628B7003C3226097195C27E2DD">Zootaxa</mods:title>
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<mods:part id="0611E2F9A31402C7D26163B6F14D7E60">
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<mods:date id="C0B546974AE00173716188531DC35D64">2015</mods:date>
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<mods:detail id="210621D0F6AE7CA5EB288F4D9F5FFC61" type="volume">
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<mods:number id="D850DF1713836C56A455FA4CCA7590B8">4040</mods:number>
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<mods:classification id="D996EB936B65B284B96289FA29FCAFDB">journal article</mods:classification>
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<mods:identifier id="6D819BA1421ADE517BA65193AAA18A5F" type="DOI">10.11646/zootaxa.4040.1.1</mods:identifier>
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<mods:identifier id="F5675345873BD3255746461F50B8E27F" type="GBIF-Dataset">3b927167-5686-4666-861f-75b65e82fcdb</mods:identifier>
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<treatment id="03B287A20E3B4037FF06FE621C0CFEAC" ID-DOI="http://doi.org/10.5281/zenodo.6118234" ID-GBIF-Taxon="127694048" ID-Zenodo-Dep="6118234" LSID="urn:lsid:plazi:treatment:03B287A20E3B4037FF06FE621C0CFEAC" httpUri="http://treatment.plazi.org/id/03B287A20E3B4037FF06FE621C0CFEAC" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="C301653F0E3B4036FF06FE621C12FE42" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BA436B40E3B4036FF06FE621F29FE63" blockId="2.[151,551,439,499]" box="[151,551,439,466]" pageId="2" pageNumber="3">
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<heading id="D0EC81D80E3B4036FF06FE621F29FE63" bold="true" box="[151,551,439,466]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="B96FEAA60E3B4036FF06FE621F29FE63" bold="true" box="[151,551,439,466]" pageId="2" pageNumber="3">
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<taxonomicName id="4C1B4D370E3B4036FF06FE621CD3FE60" ID-CoL="6Q7H9" box="[151,477,439,465]" class="Malacostraca" family="Trachelipodidae" genus="Levantoniscus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicostulatus" status="sp. nov.">
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<emphasis id="B96FEAA60E3B4036FF06FE621CD3FE60" bold="true" box="[151,477,439,465]" italics="true" pageId="2" pageNumber="3">Levantoniscus bicostulatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C57DD0E3B4036FE75FE6D1F29FE63" box="[484,551,440,466]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA436B40E3B4036FF06FE0F1C12FE42" blockId="2.[151,551,439,499]" box="[151,284,474,499]" pageId="2" pageNumber="3">
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<figureCitation id="13202A310E3B4036FF06FE0F1C12FE42" box="[151,284,474,499]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,249,1821,1843]" captionStartId-1="4.[151,249,1851,1873]" captionStartId-2="5.[151,249,1828,1850]" captionStartId-3="6.[151,249,1173,1195]" captionStartId-4="7.[151,249,1944,1966]" captionStartId-5="8.[151,249,1247,1269]" captionTargetBox-0="[173,1402,366,1772]" captionTargetBox-1="[156,1419,205,1820]" captionTargetBox-2="[172,1414,207,1790]" captionTargetBox-3="[184,1395,200,1145]" captionTargetBox-4="[218,1369,214,1868]" captionTargetBox-5="[164,1416,187,1211]" captionTargetId-0="figure@3.[160,1426,363,1787]" captionTargetId-1="figure@4.[151,1436,193,1829]" captionTargetId-2="figure@5.[151,1436,193,1807]" captionTargetId-3="figure@6.[173,1411,193,1155]" captionTargetId-4="figure@7.[199,1386,193,1888]" captionTargetId-5="figure@8.[151,1436,177,1226]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." captionText-1="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." captionText-2="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." captionText-3="FIGURE 4. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pleopod 3 exopodite, ventral view, B, pleopod 3 exopodite, dorsal view; C, pleopod 4 exopodite, ventral view, D, pleopod 4 exopodite, dorsal view; E, pleopod 5 exopodite, ventral view, F, pleopod 5 exopodite, dorsal view." captionText-4="FIGURE 5. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, cephalon and pereonite 1, dorsal view; C, pereonites 6 and 7, pleon, telson and uropods, dorsal view; D, pereonites 2 and 3, details of the cuticular structure; E, tubercle on pereonite 7, medial surface." captionText-5="FIGURE 6. Levantoniscus bicostulatus n. sp. Paratype ♀: A, pleopod 1 exopodite; B, pleopod 2 exopodite, detail of respiratory surface; C, pleopod 4 exopodite, dorsal view; D, detail of respiratory surface on pleopod 4 exopodite." httpUri-0="https://zenodo.org/record/289974/files/figure.png" httpUri-1="https://zenodo.org/record/289975/files/figure.png" httpUri-2="https://zenodo.org/record/289976/files/figure.png" httpUri-3="https://zenodo.org/record/289977/files/figure.png" httpUri-4="https://zenodo.org/record/289978/files/figure.png" httpUri-5="https://zenodo.org/record/289979/files/figure.png" pageId="2" pageNumber="3">Figures 1–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C301653F0E3B4036FF06FDF71E10FD5E" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8BA436B40E3B4036FF06FDF71E10FD5E" blockId="2.[151,1437,546,2011]" pageId="2" pageNumber="3">
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<emphasis id="B96FEAA60E3B4036FF06FDF71C8AFD8A" bold="true" box="[151,388,546,571]" pageId="2" pageNumber="3">Material examined.</emphasis>
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<collectingCountry id="F30C76240E3B4036FE1FFDF61CEDFD8A" box="[398,483,547,571]" name="Cyprus" pageId="2" pageNumber="3">Cyprus</collectingCountry>
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:
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<specimenCount id="9D1DFD3D0E3B4036FE60FDF61F11FD8B" box="[497,543,547,571]" pageId="2" pageNumber="3" type="generic">1 ♂</specimenCount>
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<typeStatus id="54A088160E3B4036FDB8FDF71F9FFD8A" box="[553,657,546,571]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="ED0AAE710E3B4036FD32FDF61FFDFD8A" box="[675,755,547,571]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="3">MZUF</collectionCode>
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9609), Kampos Potamos, northern slopes of Troodos Mt.,
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<geoCoordinate id="EE2F50730E3B4036FF06FD931C49FDEE" box="[151,327,582,607]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="35.07911">35°04’44.80’’N</geoCoordinate>
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<geoCoordinate id="EE2F50730E3B4036FEDCFD931CF3FDEE" box="[333,509,582,607]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="32.735043">32°44’06.15’’E</geoCoordinate>
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,
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<quantity id="4CE39B510E3B4036FD99FD921F5EFDEF" box="[520,592,583,607]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="3" unit="m" value="430.0">430 m</quantity>
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,
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<date id="FFA510740E3B4036FDCDFD921FD3FDEE" box="[604,733,583,607]" pageId="2" pageNumber="3" value="2012-11-29">29.XI.2012</date>
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, leg. S. Sfenthourakis; 5 ♀
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<typeStatus id="54A088160E3B4036FB9FFD921975FDEE" box="[1038,1147,583,607]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
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(
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<collectionCode id="ED0AAE710E3B4036FB18FD9219D7FDEE" box="[1161,1241,583,607]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="3">MZUF</collectionCode>
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9609), same data as
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<typeStatus id="54A088160E3B4036FF29FDBF1C2AFD32" box="[184,292,618,643]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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;
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<specimenCount id="9D1DFD3D0E3B4036FEA3FDBF1C6EFD33" box="[306,352,618,643]" pageId="2" pageNumber="3" type="generic">6 ♂</specimenCount>
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, 6 ♀
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<typeStatus id="54A088160E3B4036FE34FDBE1F1CFD32" box="[421,530,619,643]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
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(
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<collectionCode id="ED0AAE710E3B4036FDB2FDBE1F7DFD32" box="[547,627,619,643]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="3">MZUF</collectionCode>
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9610), same locality,
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<date id="FFA510740E3B4036FCE4FDBF1EF8FD32" box="[885,1014,618,643]" pageId="2" pageNumber="3" value="2015-03-25">25.III.2015</date>
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, leg. S. Sfenthourakis; 1 ♀
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<typeStatus id="54A088160E3B4036FAABFDBE1892FD32" box="[1338,1436,619,643]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
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(
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<collectionCode id="ED0AAE710E3B4036FF0FFD5A1DE0FD16" box="[158,238,655,679]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="3">MZUF</collectionCode>
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9611), Chalefka, Pentadaktylos Mt.,
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<geoCoordinate id="EE2F50730E3B4036FD56FD5B1E7BFD16" box="[711,885,654,679]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="35.285843">35º17’09.03’’N</geoCoordinate>
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<geoCoordinate id="EE2F50730E3B4036FC16FD5A193BFD16" box="[903,1077,655,679]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="33.535633">33º32’08.28’’E</geoCoordinate>
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,
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<quantity id="4CE39B510E3B4036FBDAFD5B1991FD17" box="[1099,1183,654,679]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="3" unit="m" value="520.0">520 m</quantity>
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,
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<date id="FFA510740E3B4036FB27FD5B183DFD16" box="[1206,1331,654,679]" pageId="2" pageNumber="3" value="2012-12-05">5.XII.2012</date>
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||
, leg. S. Sfenthourakis;
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<specimenCount id="9D1DFD3D0E3B4036FED0FD661C62FD7B" box="[321,364,691,715]" pageId="2" pageNumber="3" type="generic">2 ♂</specimenCount>
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, 3 ♀
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||
<typeStatus id="54A088160E3B4036FE39FD661F1BFD7A" box="[424,533,691,715]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED0AAE710E3B4036FDB2FD661F7DFD7A" box="[547,627,691,715]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="3">MZUF</collectionCode>
|
||
9612), Akanthou, northern slopes of Pentadaktylos Mt.,
|
||
<geoCoordinate id="EE2F50730E3B4036FB7FFD671892FD7A" box="[1262,1436,690,715]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="35.36937">35º22’09.73’’N</geoCoordinate>
|
||
<geoCoordinate id="EE2F50730E3B4036FF06FD031C4AFD5E" box="[151,324,726,751]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="33.755028">33º45’18.10’’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CE39B510E3B4036FEDEFD021C96FD5F" box="[335,408,727,751]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="120.0">120 m</quantity>
|
||
,
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||
<date id="FFA510740E3B4036FE35FD031F2FFD5E" box="[420,545,726,751]" pageId="2" pageNumber="3" value="2012-12-05">5.XII.2012</date>
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||
, leg. S. Sfenthourakis.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C301653F0E3B4036FF56FD2F1E2AF822" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="8BA436B40E3B4036FF56FD2F1E9AF90A" blockId="2.[151,1437,546,2011]" pageId="2" pageNumber="3">
|
||
<emphasis id="B96FEAA60E3B4036FF56FD2F1C57FCA2" bold="true" box="[199,345,762,787]" pageId="2" pageNumber="3">Description.</emphasis>
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Maximum size: ♂: 4.0 x
|
||
<quantity id="4CE39B510E3B4036FDE7FD2E1FC0FCA3" box="[630,718,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="3" unit="mm" value="2.7">2.7 mm</quantity>
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, ♀: 4.2 x 2.0 mm. Colour in ethanol: head with brown lobes and pale vertex; pereonites brown in median part, two pale stripes between tubercles and epimera; pereon and pleon epimera brown; tergites of pleonite 4 and 5 pale; antennae, pereopods, pleopods, telson and uropods pale. Body with distinct dorsal ornamentation (
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||
<figureCitation id="13202A310E3B4036FDB5FCB31F69FCCE" box="[548,615,870,895]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
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A, B, 5A–C, E) disposed as follows: two quadrangular lamellar tubercles on posterior part of cephalon and an oblique line of four low rounded tubercles on each side next to eyes; two quadrangular lamellar tubercles on each pereonite, and two on pleonite 1; two, progressively more pointed tubercles on pleonites 2–4; no tubercles on pleonite 5 and telson. Dorsum with cuticular structure consisting of hexagonal or pentagonal cells and small wart-like protuberances (
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<figureCitation id="13202A310E3B4036FC30FC231EE7FBBE" box="[929,1001,1014,1039]" captionStart="FIGURE 5" captionStartId="7.[151,249,1944,1966]" captionTargetBox="[218,1369,214,1868]" captionTargetId="figure@7.[199,1386,193,1888]" captionTargetPageId="7" captionText="FIGURE 5. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, cephalon and pereonite 1, dorsal view; C, pereonites 6 and 7, pleon, telson and uropods, dorsal view; D, pereonites 2 and 3, details of the cuticular structure; E, tubercle on pereonite 7, medial surface." httpUri="https://zenodo.org/record/289978/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
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D), whole surface covered with long piliform setae (
|
||
<figureCitation id="13202A310E3B4036FED2FBCF1C8DFB82" box="[323,387,1050,1075]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
C), which are longer and denser on tubercles; noduli laterales (
|
||
<figureCitation id="13202A310E3B4036FBDEFBCF1986FB82" box="[1103,1160,1050,1075]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Fig 1</figureCitation>
|
||
A, B) on pereonites 1–4 inserted on outer surface of tubercles near tubercle posterior corner, and on pereonites 5 to 7 on postero-lateral corner of pereonites, progressively closer to posterior margin. Cephalon (
|
||
<figureCitation id="13202A310E3B4036FC4AFBB7192EFBCA" box="[987,1056,1122,1147]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
||
D, 5B) with quadrangular frontal lateral lobes, obliquely directed outwards, and rectangular median lobe more protruding than lateral lobes and directed upwards. Eyes small with five ommatidia. Epimera (
|
||
<figureCitation id="13202A310E3B4036FCDCFB7F1E9FFB72" box="[845,913,1194,1219]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
|
||
A, B, 5A) of pereonite 1–3 quadrangular with posterior margins slightly concave; epimera of pereonites 4–7 progressively more acute and backwards directed. Telson (
|
||
<figureCitation id="13202A310E3B4036FF63FB271C3BFABA" box="[242,309,1266,1291]" captionStart="FIGURE 5" captionStartId="7.[151,249,1944,1966]" captionTargetBox="[218,1369,214,1868]" captionTargetId="figure@7.[199,1386,193,1888]" captionTargetPageId="7" captionText="FIGURE 5. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, cephalon and pereonite 1, dorsal view; C, pereonites 6 and 7, pleon, telson and uropods, dorsal view; D, pereonites 2 and 3, details of the cuticular structure; E, tubercle on pereonite 7, medial surface." httpUri="https://zenodo.org/record/289978/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
|
||
C) almost twice as broad as long, distal part with rounded corners and very slightly concave distal margin. Antennula (
|
||
<figureCitation id="13202A310E3B4036FEE8FAC31CB6FA9E" box="[377,440,1302,1327]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
E) with second article shorter than first and third; distal articles with two apical and five subapical aesthetascs. Antenna (
|
||
<figureCitation id="13202A310E3B4036FD93FAEF1F4FFAE2" box="[514,577,1338,1363]" captionStart="FIGURE 1" captionStartId="3.[151,249,1821,1843]" captionTargetBox="[173,1402,366,1772]" captionTargetId="figure@3.[160,1426,363,1787]" captionTargetPageId="3" captionText="FIGURE 1. Levantoniscus bicostulatus n. sp. Paratype ♀: A, adult specimen, dorsal view; B, adult specimen, lateral view; C, dorsal scale-seta; D, cephalon, frontal view. Paratype ♂: E, antennula; F, antenna." httpUri="https://zenodo.org/record/289974/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
F) short, reaching posterior margin of pereonite 1; flagellum slightly shorter than fifth segment of peduncle, first flagellar article 3 times shorter than second, second flagellar article with row of three aesthetascs. Right mandible (
|
||
<figureCitation id="13202A310E3B4036FDC7FA571F93FA2A" box="[598,669,1410,1435]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
A) with 1+1 and left mandible (
|
||
<figureCitation id="13202A310E3B4036FBB1FA571967FA2A" box="[1056,1129,1410,1435]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
B) with 2+1 free penicils. Maxillula (
|
||
<figureCitation id="13202A310E3B4036FE86FA731C53FA0E" box="[279,349,1446,1471]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
C) outer branch with 4+6 teeth; inner branch with distinct triangular posterior point and two subequal penicils. Maxilla (
|
||
<figureCitation id="13202A310E3B4036FE47FA1F1F15FA52" box="[470,539,1482,1507]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
D) with inner lobe 1.5 times larger than outer lobe, covered with thick apical setae; three large setae between two lobes. Maxilliped (
|
||
<figureCitation id="13202A310E3B4036FCB7FA3B1E62F9B6" box="[806,876,1518,1543]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
E) endite with broad and rounded distal margin bearing three triangular teeth and long seta; pulp short with proximal article bearing two thick setae. Pereopod 1 (
|
||
<figureCitation id="13202A310E3B4036FF0EF9E31DEBF9FE" box="[159,229,1590,1615]" captionStart="FIGURE 3" captionStartId="5.[151,249,1828,1850]" captionTargetBox="[172,1414,207,1790]" captionTargetId="figure@5.[151,1436,193,1807]" captionTargetPageId="5" captionText="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." httpUri="https://zenodo.org/record/289976/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
A) carpus with antennal brush consisting of large area of short setae obliquely directed. Pleopods with uncovered lungs largely exposed on exopodites 1 and 2 (
|
||
<figureCitation id="13202A310E3B4036FC87F98F1E57F9C2" box="[790,857,1626,1651]" captionStart="FIGURE 6" captionStartId="8.[151,249,1247,1269]" captionTargetBox="[164,1416,187,1211]" captionTargetId="figure@8.[151,1436,177,1226]" captionTargetPageId="8" captionText="FIGURE 6. Levantoniscus bicostulatus n. sp. Paratype ♀: A, pleopod 1 exopodite; B, pleopod 2 exopodite, detail of respiratory surface; C, pleopod 4 exopodite, dorsal view; D, detail of respiratory surface on pleopod 4 exopodite." httpUri="https://zenodo.org/record/289979/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
|
||
A, B), and in invaginations on exopodites 3–5 (
|
||
<figureCitation id="13202A310E3B4036FAFEF98F1DA8F926" captionStart="FIGURE 6" captionStartId="8.[151,249,1247,1269]" captionTargetBox="[164,1416,187,1211]" captionTargetId="figure@8.[151,1436,177,1226]" captionTargetPageId="8" captionText="FIGURE 6. Levantoniscus bicostulatus n. sp. Paratype ♀: A, pleopod 1 exopodite; B, pleopod 2 exopodite, detail of respiratory surface; C, pleopod 4 exopodite, dorsal view; D, detail of respiratory surface on pleopod 4 exopodite." httpUri="https://zenodo.org/record/289979/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
|
||
C, D). Uropod (
|
||
<figureCitation id="13202A310E3B4036FEF7F9AB1CA0F926" box="[358,430,1662,1687]" captionStart="FIGURE 2" captionStartId="4.[151,249,1851,1873]" captionTargetBox="[156,1419,205,1820]" captionTargetId="figure@4.[151,1436,193,1829]" captionTargetPageId="4" captionText="FIGURE 2. Levantoniscus bicostulatus n. sp. Paratype ♀: A, right mandible; B, left mandible; C, maxillula; D, maxilla; E, maxilliped; F, uropod." httpUri="https://zenodo.org/record/289975/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
F) protopodite with concave outer margin, not grooved; exopodite ventrally grooved, slightly longer than endopodite and reaching distal margin of telson.
|
||
</paragraph>
|
||
<paragraph id="8BA436B40E3B4036FF56F9131E2AF822" blockId="2.[151,1437,546,2011]" pageId="2" pageNumber="3">
|
||
Male. Pereopod 1 (
|
||
<figureCitation id="13202A310E3B4036FE0DF9131CD3F96E" box="[412,477,1734,1759]" captionStart="FIGURE 3" captionStartId="5.[151,249,1828,1850]" captionTargetBox="[172,1414,207,1790]" captionTargetId="figure@5.[151,1436,193,1807]" captionTargetPageId="5" captionText="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." httpUri="https://zenodo.org/record/289976/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
A) merus and carpus with line of strong setae on sternal margin. Pereopod 7 (
|
||
<figureCitation id="13202A310E3B4036FAAFF9131871F96E" box="[1342,1407,1734,1759]" captionStart="FIGURE 3" captionStartId="5.[151,249,1828,1850]" captionTargetBox="[172,1414,207,1790]" captionTargetId="figure@5.[151,1436,193,1807]" captionTargetPageId="5" captionText="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." httpUri="https://zenodo.org/record/289976/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
B) with enlarged basis, ischium with straight sternal margin, merus and carpus with slightly concave sternal margins. Pleopod 1 (
|
||
<figureCitation id="13202A310E3B4036FE89F8DB1C55F896" box="[280,347,1806,1831]" captionStart="FIGURE 3" captionStartId="5.[151,249,1828,1850]" captionTargetBox="[172,1414,207,1790]" captionTargetId="figure@5.[151,1436,193,1807]" captionTargetPageId="5" captionText="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." httpUri="https://zenodo.org/record/289976/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
C, D) exopodite small, subrectangular, about 2.5 times as wide as long; endopodite with thickset distal part and pointed apex. Pleopod 2 (
|
||
<figureCitation id="13202A310E3B4036FDC7F8E71F99F8FA" box="[598,663,1842,1867]" captionStart="FIGURE 3" captionStartId="5.[151,249,1828,1850]" captionTargetBox="[172,1414,207,1790]" captionTargetId="figure@5.[151,1436,193,1807]" captionTargetPageId="5" captionText="FIGURE 3. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pereopod 1; B, pereopod 7; C, pleopod 1, ventral view; D, pleopod 1 exopodite, dorsal view; E, pleopod 2, ventral view; F, pleopod 2 exopodite, dorsal view." httpUri="https://zenodo.org/record/289976/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
E, F) exopodite triangular distinctly shorter than endopodite. Pleopod 3 (
|
||
<figureCitation id="13202A310E3B4036FF28F8831C0CF8DE" box="[185,258,1878,1903]" captionStart="FIGURE 4" captionStartId="6.[151,249,1173,1195]" captionTargetBox="[184,1395,200,1145]" captionTargetId="figure@6.[173,1411,193,1155]" captionTargetPageId="6" captionText="FIGURE 4. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pleopod 3 exopodite, ventral view, B, pleopod 3 exopodite, dorsal view; C, pleopod 4 exopodite, ventral view, D, pleopod 4 exopodite, dorsal view; E, pleopod 5 exopodite, ventral view, F, pleopod 5 exopodite, dorsal view." httpUri="https://zenodo.org/record/289977/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
A, B) and 4 (
|
||
<figureCitation id="13202A310E3B4036FE39F8831CFFF8DE" box="[424,497,1878,1903]" captionStart="FIGURE 4" captionStartId="6.[151,249,1173,1195]" captionTargetBox="[184,1395,200,1145]" captionTargetId="figure@6.[173,1411,193,1155]" captionTargetPageId="6" captionText="FIGURE 4. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pleopod 3 exopodite, ventral view, B, pleopod 3 exopodite, dorsal view; C, pleopod 4 exopodite, ventral view, D, pleopod 4 exopodite, dorsal view; E, pleopod 5 exopodite, ventral view, F, pleopod 5 exopodite, dorsal view." httpUri="https://zenodo.org/record/289977/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
C, D) trapezoidal with distinctly concave distal margin; pleopod 5 (
|
||
<figureCitation id="13202A310E3B4036FA86F883186EF8DE" box="[1303,1376,1878,1903]" captionStart="FIGURE 4" captionStartId="6.[151,249,1173,1195]" captionTargetBox="[184,1395,200,1145]" captionTargetId="figure@6.[173,1411,193,1155]" captionTargetPageId="6" captionText="FIGURE 4. Levantoniscus bicostulatus n. sp. Holotype ♂: A, pleopod 3 exopodite, ventral view, B, pleopod 3 exopodite, dorsal view; C, pleopod 4 exopodite, ventral view, D, pleopod 4 exopodite, dorsal view; E, pleopod 5 exopodite, ventral view, F, pleopod 5 exopodite, dorsal view." httpUri="https://zenodo.org/record/289977/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
E, F) trapezoidal with distal margin oblique and almost straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301653F0E3B4036FF56F84B1FE0F86A" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="8BA436B40E3B4036FF56F84B1FE0F86A" blockId="2.[151,1437,546,2011]" pageId="2" pageNumber="3">
|
||
<emphasis id="B96FEAA60E3B4036FF56F84B1C40F806" bold="true" box="[199,334,1950,1975]" pageId="2" pageNumber="3">Etymology.</emphasis>
|
||
Latin:
|
||
<emphasis id="B96FEAA60E3B4036FE33F84B1CB6F806" box="[418,440,1950,1975]" italics="true" pageId="2" pageNumber="3">bi</emphasis>
|
||
= double +
|
||
<emphasis id="B96FEAA60E3B4036FDD7F84B1FB6F806" box="[582,696,1950,1975]" italics="true" pageId="2" pageNumber="3">costulatus</emphasis>
|
||
= with ribs. The name of the species refers to the double line of lamellar tubercles on the cephalon, pereon and pleon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301653F0E3A4037FF56FF421C0CFEAC" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="8BA436B40E3A4037FF56FF421C0CFEAC" blockId="3.[151,1436,151,285]" pageId="3" pageNumber="4">
|
||
<emphasis id="B96FEAA60E3A4037FF56FF421C35FF01" bold="true" box="[199,315,151,176]" pageId="3" pageNumber="4">Remarks.</emphasis>
|
||
The species has been found only at the close vicinity of running water, under leaves and stones hidden inside the wet litter layer of hygrophilic plants. Taking into account its dorsal ornamentation, the species should be considered rather as an endogean species, coming to the surface only in locations with very high humidity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |