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<document id="2783BE2F54F2513B816090783A69A147" ID-DOI="10.11646/zootaxa.5227.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7523269" ID-ZooBank="2556E988-3FBB-4185-9C54-981CAD99041C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673389024490" checkinUser="plazi" docAuthor="Kronestedt, Torbjörn" docDate="2023" docId="03B2878CFF93FF93FF61FDB7BF57FC58" docLanguage="en" docName="zootaxa.5227.5.2.pdf" docOrigin="Zootaxa 5227 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wadicosa mabweana Kronestedt 2023, comb. nov." docType="treatment" docVersion="3" lastPageNumber="539" masterDocId="FF8BFFF4FF95FF9BFFF6FFDDBB04FFB5" masterDocTitle="Species of Wadicosa (Araneae, Lycosidae): transfer of four species from Africa currently placed in Pardosa" masterLastPageNumber="548" masterPageNumber="531" pageNumber="537" updateTime="1699446229162" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AE26AE19E0C564EBB125E62E9C841487">Species of Wadicosa (Araneae, Lycosidae): transfer of four species from Africa currently placed in Pardosa</mods:title>
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<mods:date id="8A7383FC3F967E2143E4A19AFC1D7F51">2023</mods:date>
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<taxonomicName id="4C1B4D19FF93FF9DFF61FDB7B951FD31" authority="(Roewer, 1959)" authorityName="Kronestedt" authorityYear="2023" baseAuthorityName="Roewer" baseAuthorityYear="1959" box="[151,597,618,645]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="mabweana" status="comb. nov.">
<emphasis id="B96FEA88FF93FF9DFF61FDB7BA90FD30" bold="true" box="[151,404,618,645]" italics="true" pageId="6" pageNumber="537">Wadicosa mabweana</emphasis>
(
<bibRefCitation id="EF8A4B6BFF93FF9DFE55FDB7B949FD31" author="Roewer, C. F." box="[419,589,618,644]" pageId="6" pageNumber="537" pagination="1 - 518" refId="ref11081" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II (Lycosidae). Exploration du Parc National de l'Upemba, Mission G. F. De Witte, 55, 1 - 518." type="journal article" year="1959">Roewer, 1959</bibRefCitation>
)
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<subSubSection id="C3016511FF93FF9DFF61FD52B891FD1C" box="[151,917,654,681]" pageId="6" pageNumber="537" type="description">
<paragraph id="8BA4369AFF93FF9DFF61FD52B891FD1C" blockId="6.[151,917,618,681]" box="[151,917,654,681]" pageId="6" pageNumber="537">
<figureCitation id="13202A1FFF93FF9DFF61FD52BBF7FD1C" box="[151,243,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1751]" captionTargetBox="[159,1419,416,1689]" captionTargetId="figure-117@3.[159,1429,416,1703]" captionTargetPageId="3" captionText="FIGURE 1. A–C. Wadicosa manubriata (Simon), male. A. Right palp in retrolateral view, from Simon (1898: fig. 356). B–C. Holotype, right palp in retrolateral view (B), and left palp in antero-ventral view (C). Photographs by C. Hervé, MNHN. D–E. W. sordidecolorata (Strand), epigynes, from Strand (1908): D pl. 8, fig. 16 (“Lycosa sordide-colorata”) and E pl. 8, fig.9 (“Lycosa darolii”). F. W. enucleata (Roewer), epigyne, from Roewer (1959, fig. 62b). G–I. W. manubriata (Simon), from Roewer 1959, fig. 63a–c), G. Right palp in retrolateral view. H. Left palp in ventral view. I. Epigyne. (G and H obviously drawn from microslide SMF 13616 (RII 4914/252) in which the embolus in ventral view is distorted similarly to that shown in H.) K. W. mabweana (Roewer), epigyne, from Roewer (1959, fig. 54a). L–M. W. sordidecolorata (Strand), from Roewer (1959, fig. 30a–c). L. Right male palp. M. “Lamella characteristica”= tegular apophysis. N. Epigyne. O. Pardosa darolii (Strand). from Roewer (1959, fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.7523271" httpUri="https://zenodo.org/record/7523271/files/figure.png" pageId="6" pageNumber="537">Figs 1K</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFF09FD52BA3BFD1C" box="[255,319,655,681]" captionStart="FIGURE 2" captionStartId="4.[151,250,1888,1912]" captionTargetBox="[193,1395,179,1862]" captionTargetId="figure-18@4.[192,1395,179,1862]" captionTargetPageId="4" captionText="FIGURE 2. Habitus, dorsal view.A–C. Wadicosa enucleata (Roewer).A ♁ from Tanzania: Mkomazi, B ♁ from Nigeria: Ilorin. C ♀ from Tanzania: Kyela. D–F. W. mabweana (Roewer). D ♁ and F ♀ from DR Congo: Mikembo, E ♁ from Namibia:Andara. G–I. W. manubriata (Simon). G ♁ from Namibia: Brukkaros Crater. H ♁ from South Africa: Dendron. I ♀ from Zimbabwe: Umguza River. Scale bar: 2 mm (A–I)." figureDoi="http://doi.org/10.5281/zenodo.7523273" httpUri="https://zenodo.org/record/7523273/files/figure.png" pageId="6" pageNumber="537">2D–F</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFEBCFD52BA88FD1C" box="[330,396,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[163,1419,185,1757]" captionTargetId="figure-15@5.[151,1436,179,1772]" captionTargetPageId="5" captionText="FIGURE 3. Right male palp, ventral view (A–C) and retrolateral view (D–F). A, D. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E. W. mabweana (Roewer) from DR Congo: Mikembo. C, F. W. manubriata (Simon) from Namibia: Brukkaros Crater. Scale bar: 0.5 mm (A–F)." figureDoi="http://doi.org/10.5281/zenodo.7523275" httpUri="https://zenodo.org/record/7523275/files/figure.png" pageId="6" pageNumber="537">3B, E</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFE6EFD52BAF9FD1C" box="[408,509,655,681]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="6" pageNumber="537">4B, E, H</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFDFEFD53B948FD1C" box="[520,588,654,681]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[163,1427,1037,1869]" captionTargetId="figure-375@8.[151,1436,1026,1874]" captionTargetPageId="8" captionText="FIGURE 5. Epigyne, ventral view. A. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, C. W. mabweana (Roewer), from DR Congo: Mikembo. D. W. manubriata (Simon), from Zimbabwe: Umguza River. E. W. sordidecolorata (Strand), from Cameroon: Koum. Scale bar: 0.2 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.7523279" httpUri="https://zenodo.org/record/7523279/files/figure.png" pageId="6" pageNumber="537">5B–C</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFDA1FD52B999FD1C" box="[599,669,655,681]" captionStart="FIGURE 6" captionStartId="9.[151,250,878,902]" captionTargetBox="[151,1436,179,852]" captionTargetId="figure-15@9.[151,1436,179,852]" captionTargetPageId="9" captionText="FIGURE 6. Epigyne, ventral view.A–B. Wadicosa enucleata (Roewer), A holotype, B from Zimbabwe: Batoka Gorge. C–D. W. mabweana (Roewer), C lectotype, D from Namibia:Andara. E–F. W. manubriata (Simon), E from Namibia: Windhoek, F from Namibia: Brukkaros Crater. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7523281" httpUri="https://zenodo.org/record/7523281/files/figure.png" pageId="6" pageNumber="537">6C–D</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFD5EFD52B809FD1C" box="[680,781,655,681]" captionStart="FIGURE 7" captionStartId="10.[151,250,1840,1864]" captionTargetBox="[151,1435,846,1814]" captionTargetId="figure-325@10.[151,1435,846,1814]" captionTargetPageId="10" captionText="FIGURE 7. Epigyne in ventral (A, B, C) and female genitalia in dorsal (D–F, details in G–I) view.A, D, G. Wadicosa enucleata (Roewer) from Zimbabwe: Batoka Gorge. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon), C from Namibia: Windhoek, F from South Africa: Paarl (F). Scale bars: 300 µm (A–F), 50 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523285" httpUri="https://zenodo.org/record/7523285/files/figure.png" pageId="6" pageNumber="537">7B, E, H</figureCitation>
, 
<figureCitation id="13202A1FFF93FF9DFCEEFD52B833FD1C" box="[792,823,655,681]" captionStart="FIGURE 10" captionStartId="15.[151,250,1891,1915]" captionTargetBox="[151,1436,479,1865]" captionTargetId="figure-76@15.[151,1436,479,1865]" captionTargetPageId="15" captionText="FIGURE 10. Collection localities for Wadicosa enucleata (Roewer), W. mabweana (Roewer), W. manubriata (Simon) and W. sordidecolorata (Strand)." figureDoi="http://doi.org/10.5281/zenodo.7523291" httpUri="https://zenodo.org/record/7523291/files/figure.png" pageId="6" pageNumber="537">10</figureCitation>
; 
<tableCitation id="C6990321FF93FF9DFCB5FD52B891FD1C" box="[835,917,655,681]" captionStart="TABLE 1" captionStartId="14.[152,244,151,175]" captionText="TABLE 1. Leg measurements (in mm) of Wadicosa enucleata, W. mabweana, W. manubriata and W. sordidecolorata." pageId="6" pageNumber="537">Table 1</tableCitation>
</paragraph>
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<treatmentCitationGroup id="AB0B11B4FF93FF9DFF61FD13B9CEFD53" box="[151,714,718,742]" pageId="6" pageNumber="537">
<treatmentCitation id="0ABA108BFF93FF9DFF61FD13B92CFD53" author="Roewer, C. F." box="[151,552,718,742]" page="112" pageId="6" pageNumber="537" year="1959">
<taxonomicName id="4C1B4D19FF93FF9DFF61FD13B92CFD53" ID-CoL="4DJV4" authority="Roewer, 1959: 112" authorityName="Roewer" authorityPageNumber="112" authorityYear="1959" box="[151,552,718,742]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="mabweana">
<emphasis id="B96FEA88FF93FF9DFF61FD13BA64FD53" box="[151,352,718,742]" italics="true" pageId="6" pageNumber="537">Pardosa mabweana</emphasis>
<bibRefCitation id="EF8A4B6BFF93FF9DFE91FD13B92CFD53" author="Roewer, C. F." box="[359,552,718,742]" pageId="6" pageNumber="537" pagination="1 - 518" refId="ref11081" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II (Lycosidae). Exploration du Parc National de l'Upemba, Mission G. F. De Witte, 55, 1 - 518." type="journal article" year="1959">Roewer, 1959: 112</bibRefCitation>
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, fig. 54a, b (
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).
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<emphasis id="B96FEA88FF93FF9DFF61FCD2BA4CFC9D" bold="true" box="[151,328,782,809]" pageId="6" pageNumber="537">Type material.</emphasis>
<materialsCitation id="3B733CC7FF93FF9DFEA7FCD2BE98FCF8" collectingDate="1948-09-27" collectionCode="MRAC" country="Democratic Republic of the Congo" county="Mission De Witte" elevation="1750" latitude="-8.783334" location="Haut-Katanga" longLatPrecision="1300" longitude="26.933332" municipality="Buye-Bala" pageId="6" pageNumber="537" specimenCode="MRAC 139460" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
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<typeStatus id="54A08838FF93FF9DFEA7FCD2BABBFC9C" box="[337,447,783,809]" pageId="6" pageNumber="537" type="lectotype">Lectotype</typeStatus>
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from 
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, 
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<location id="8EC46041FF93FF9DFD3AFCD2B875FC9D" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFD3AFCD2B875FC9D" box="[716,881,783,808]" country="Democratic Republic of the Congo" county="Mission De Witte" latitude="-8.783334" longLatPrecision="1300" longitude="26.933332" municipality="Buye-Bala" name="Haut-Katanga" pageId="6" pageNumber="537">Haut-Katanga</location>
,
</emphasis>
<location id="8EC46041FF93FF9DFC88FCD2BF83FC9C" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFC88FCD2BF83FC9C" box="[894,1159,783,809]" country="Democratic Republic of the Congo" county="Mission De Witte" latitude="-8.783334" longLatPrecision="1300" longitude="26.933332" municipality="Buye-Bala" name="Upemba National Park" pageId="6" pageNumber="537">Upemba National Park</location>
, 
<collectingMunicipality id="6BC0ACE0FF93FF9DFB63FCD2BE14FC9C" box="[1173,1296,783,809]" pageId="6" pageNumber="537">Buye-Bala</collectingMunicipality>
, [
<emphasis id="B96FEA88FF93FF9DFAD0FCD2BE44FC9D" box="[1318,1344,783,808]" italics="true" pageId="6" pageNumber="537">ca</emphasis>
<geoCoordinate id="EE2F505DFF93FF9DFABCFCCDBE93FC9E" box="[1354,1431,784,811]" degrees="8" direction="south" minutes="47" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-8.783334">8°47′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFF61FCE9BBF3FCFA" box="[151,247,820,847]" degrees="26" direction="east" minutes="56" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="26.933332">26°56′E</geoCoordinate>
], alt. 
<quantity id="4CE39B7FFF93FF9DFEC8FCEFBA9FFCF8" box="[318,411,818,845]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="537" unit="m" value="1750.0">
<elevation id="0036D1A9FF93FF9DFEC8FCEFBA9FFCF8" box="[318,411,818,845]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="537" unit="m" value="1750.0">1750 m</elevation>
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, 
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<collectingDate id="EFE1E9B2FF93FF9DFE5BFCEEB94FFCF8" box="[429,587,819,845]" pageId="6" pageNumber="537" value="1948-09-27">27 Sep. 1948</collectingDate>
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(
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, 
<specimenCode id="DBBD9EE1FF93FF9DFCB0FCEEB8F9FCF8" box="[838,1021,819,845]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 139460</specimenCode>
), examined and here designated.—
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</paragraph>
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<materialsCitation id="3B733CC7FF93FF9DFF61FC8BB9E2FC20" collectingDate="1948-09-27" collectionCode="MRAC, SMF" country="Democratic Republic of the Congo" county="Upemba Park" elevation="1750" latitude="-8.783334" location="Haut-Katanga" longLatPrecision="1300" longitude="26.933332" municipality="Mission De Witte" pageId="6" pageNumber="537" specimenCode="MRAC 139461, SMF 9911343" specimenCount="4" specimenCount-female="4" typeStatus="paralectotype">
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.
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<collectingCountry id="F30C760AFF93FF9DFEB8FC8ABAE0FCC4" box="[334,484,855,881]" name="Democratic Republic of the Congo" pageId="6" pageNumber="537">DR CONGO</collectingCountry>
. 
<emphasis id="B96FEA88FF93FF9DFE19FC8AB99EFCC5" box="[495,666,855,880]" italics="true" pageId="6" pageNumber="537">
<location id="8EC46041FF93FF9DFE19FC8AB990FCC5" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFE19FC8AB990FCC5" box="[495,660,855,880]" country="Democratic Republic of the Congo" county="Upemba Park" municipality="Mission De Witte" name="Haut-Katanga" pageId="6" pageNumber="537">Haut-Katanga</location>
:
</emphasis>
data as lectotype, 
<specimenCount id="9D1DFD13FF93FF9DFC9BFC8AB88BFCC4" box="[877,911,855,881]" pageId="6" pageNumber="537" type="female">2♀</specimenCount>
(
<collectingMunicipality id="6BC0ACE0FF93FF9DFC69FC8ABF6EFCC4" box="[927,1130,855,881]" pageId="6" pageNumber="537">Mission De Witte</collectingMunicipality>
, 
<specimenCode id="DBBD9EE1FF93FF9DFB80FC8ABE2EFCC4" box="[1142,1322,855,881]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 139461</specimenCode>
); 
<collectingCounty id="62C54E16FF93FF9DFACBFC8ABBC9FC20" pageId="6" pageNumber="537">Upemba Park</collectingCounty>
, 
<specimenCount id="9D1DFD13FF93FF9DFF2FFCA6BBF9FC20" box="[217,253,891,917]" pageId="6" pageNumber="537" type="female">2♀</specimenCount>
, RII/11343/679 (
<specimenCode id="DBBD9EE1FF93FF9DFE4AFCA6B967FC20" box="[444,611,891,917]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="537">SMF 9911343</specimenCode>
), examined
</materialsCitation>
.
</paragraph>
<paragraph id="8BA4369AFF93FF9DFF31FC43B9A8FAF0" blockId="6.[151,1437,782,2033]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31FC43BAFBFC0D" bold="true" box="[199,511,926,952]" pageId="6" pageNumber="537">Other material examined.</emphasis>
<materialsCitation id="3B733CC7FF93FF9DFDFEFC42B9B9FC68" collectionCode="MHNG" collectorName="A. Monard" country="Angola" latitude="-15.416667" location="Kangela" longLatPrecision="1285" longitude="15.733334" municipality="Cankela" pageId="6" pageNumber="537" specimenCount="9" specimenCount-female="9" stateProvince="Huila">
<collectingCountry id="F30C760AFF93FF9DFDFEFC42B986FC0C" box="[520,642,927,953]" name="Angola" pageId="6" pageNumber="537">ANGOLA</collectingCountry>
. 
<emphasis id="B96FEA88FF93FF9DFD78FC42B9D3FC0C" box="[654,727,927,953]" italics="true" pageId="6" pageNumber="537">
<collectingRegion id="49DFF878FF93FF9DFD78FC42B9D5FC0C" box="[654,721,927,953]" country="Angola" name="Huila" pageId="6" pageNumber="537">Huila</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC0ACE0FF93FF9DFD16FC42B838FC0C" box="[736,828,927,953]" pageId="6" pageNumber="537">Cankela</collectingMunicipality>
(= 
<location id="8EC46041FF93FF9DFC90FC42B8C2FC0C" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFC90FC42B8C2FC0C" box="[870,966,927,953]" country="Angola" latitude="-15.416667" longLatPrecision="1285" longitude="15.733334" municipality="Cankela" name="Kangela" pageId="6" pageNumber="537" stateProvince="Huila">Kangela</location>
, [
<geoCoordinate id="EE2F505DFF93FF9DFC2DFC7DBF3CFC0E" box="[987,1080,928,955]" degrees="15" direction="south" minutes="25" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-15.416667">15°25′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFBB3FC7DBFA2FC0E" box="[1093,1190,928,955]" degrees="15" direction="east" minutes="44" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="15.733334">15°44′E</geoCoordinate>
]), 
<collectorName id="26EE534CFF93FF9DFB37FC42BE45FC0C" box="[1217,1345,927,953]" pageId="6" pageNumber="537">A. Monard</collectorName>
, 1♁ 
<specimenCount id="9D1DFD13FF93FF9DFA8FFC42BE99FC0C" box="[1401,1437,927,953]" pageId="6" pageNumber="537" type="female">9♀</specimenCount>
(
<collectionCode id="ED0AAE5FFF93FF9DFF69FC1EBBFCFC68" box="[159,248,963,989]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="537" type="Museum">MHNG</collectionCode>
) (
<bibRefCitation id="EF8A4B6BFF93FF9DFEFAFC1EBA99FC68" author="Lessert, R. de" box="[268,413,963,989]" pageId="6" pageNumber="537" pagination="207 - 306" refId="ref10894" refString="Lessert, R. de (1936) Araignees de l'Afrique orientale portugaise recueillies par MM. P. Lesne et H. - B. Cott. Revue Suisse de Zoologie, 43 (9), 207 - 306. https: // doi. org / 10.5962 / bhl. part. 144393" type="journal article" year="1936">Lessert 1936</bibRefCitation>
<emphasis id="B96FEA88FF93FF9DFE54FC1EB9B2FC68" box="[418,694,963,989]" italics="true" pageId="6" pageNumber="537">
sub 
<taxonomicName id="4C1B4D19FF93FF9DFE39FC1EB9B2FC68" authorityName="Simon" authorityYear="1898" box="[463,694,963,989]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="manubriata">Pardosa manubriata</taxonomicName>
</emphasis>
)
</materialsCitation>
.— 
<materialsCitation id="3B733CC7FF93FF9DFD29FC1EB8E5FBB4" collectingDate="2001-04-13" collectionCode="MRAC" country="Democratic Republic of the Congo" county="Haut-Katanga" latitude="-8.933333" location="Ntumbwa river" longLatPrecision="1300" longitude="27.2" municipality="Lusinga" pageId="6" pageNumber="537" specimenCode="MRAC 211722" specimenCount="2" specimenCount-female="2" stateProvince="Huila">
<collectingCountry id="F30C760AFF93FF9DFD29FC1EB871FC68" box="[735,885,963,989]" name="Democratic Republic of the Congo" pageId="6" pageNumber="537">DR CONGO</collectingCountry>
. 
<emphasis id="B96FEA88FF93FF9DFC88FC1EBF2DFC69" box="[894,1065,963,988]" italics="true" pageId="6" pageNumber="537">
<collectingCounty id="62C54E16FF93FF9DFC88FC1EBF27FC69" box="[894,1059,963,988]" pageId="6" pageNumber="537">Haut-Katanga</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BC0ACE0FF93FF9DFBD9FC1EBF88FC68" box="[1071,1164,963,989]" pageId="6" pageNumber="537">Lusinga</collectingMunicipality>
, 
<location id="8EC46041FF93FF9DFB60FC1EBE45FC68" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFB60FC1EBE45FC68" box="[1174,1345,963,989]" country="Democratic Republic of the Congo" county="Haut-Katanga" latitude="-8.933333" longLatPrecision="1300" longitude="27.2" municipality="Lusinga" name="Ntumbwa river" pageId="6" pageNumber="537" stateProvince="Huila">Ntumbwa river</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFABCFC19BE93FC6A" box="[1354,1431,964,991]" degrees="8" direction="south" minutes="56" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-8.933333">8°56′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFF61FC35BBF0FBB6" box="[151,244,1000,1027]" degrees="27" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="27.2">27°12′E</geoCoordinate>
, 9 &amp; 
<date id="FFA5105AFF93FF9DFECBFC3ABADDFBB4" box="[317,473,999,1025]" pageId="6" pageNumber="537" value="2001-04-13">
<collectingDate id="EFE1E9B2FF93FF9DFECBFC3ABADDFBB4" box="[317,473,999,1025]" pageId="6" pageNumber="537" value="2001-04-13">13 Apr. 2001</collectingDate>
</date>
, PNU staff, 
<specimenCount id="9D1DFD13FF93FF9DFD98FC3AB994FBB4" box="[622,656,999,1025]" pageId="6" pageNumber="537" type="female">2♀</specimenCount>
(
<specimenCode id="DBBD9EE1FF93FF9DFD55FC3AB851FBB4" box="[675,853,999,1025]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 211722</specimenCode>
&amp; 211725)
</materialsCitation>
; 
<materialsCitation id="3B733CC7FF93FF9DFC19FC3AB8BAFBFC" collectingDate="2010-07-18" collectingDateMax="2010-08-08" collectingDateMin="2010-07-18" collectionCode="NHRS" collectorName="M. Hasson" country="Democratic Republic of the Congo" latitude="-11.466666" location="Mikembo" longLatPrecision="1295" longitude="27.65" pageId="6" pageNumber="537" specimenCode="MRAC 232512" specimenCount="51" specimenCount-female="51" stateProvince="Huila">
<location id="8EC46041FF93FF9DFC19FC3ABF5BFBB4" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFC19FC3ABF5BFBB4" box="[1007,1119,999,1025]" country="Democratic Republic of the Congo" latitude="-11.466666" longLatPrecision="1295" longitude="27.65" name="Mikembo" pageId="6" pageNumber="537" stateProvince="Huila">Mikembo</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFB98FC35BFC2FBB6" box="[1134,1222,1000,1027]" degrees="11" direction="south" minutes="28" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-11.466666">11º28′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFB23FC35BE34FBB6" box="[1237,1328,1000,1027]" degrees="27" direction="east" minutes="39" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="27.65">27º39′E</geoCoordinate>
, miombo woodland, 
<collectingMethod id="525A4E8DFF93FF9DFEE5FBD6BA97FB90" box="[275,403,1035,1061]" pageId="6" pageNumber="537">pitfall traps</collectingMethod>
and hand catch, 
<date id="FFA5105AFF93FF9DFDA6FBD6B831FB90" box="[592,821,1035,1061]" pageId="6" pageNumber="537" value="2010-07-18" valueMax="2010-08-08" valueMin="2010-07-18">
<collectingDate id="EFE1E9B2FF93FF9DFDA6FBD6B831FB90" box="[592,821,1035,1061]" pageId="6" pageNumber="537" value="2010-07-18" valueMax="2010-08-08" valueMin="2010-07-18">18 Jul.–8 Aug. 2010</collectingDate>
</date>
, 
<collectorName id="26EE534CFF93FF9DFCC9FBD6B8BFFB90" box="[831,955,1035,1061]" pageId="6" pageNumber="537">M. Hasson</collectorName>
, 11♁ 
<specimenCount id="9D1DFD13FF93FF9DFC0BFBD7BF2AFB90" box="[1021,1070,1034,1061]" pageId="6" pageNumber="537" type="female">35♀</specimenCount>
(
<specimenCode id="DBBD9EE1FF93FF9DFBCAFBD6BFE8FB90" box="[1084,1260,1034,1061]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 232512</specimenCode>
[2♁ 
<specimenCount id="9D1DFD13FF93FF9DFAD0FBD6BE4CFB90" box="[1318,1352,1035,1061]" pageId="6" pageNumber="537" type="female">2♀</specimenCount>
now in 
<collectionCode id="ED0AAE5FFF93FF9DFF61FBF2BBE3FBFC" box="[151,231,1071,1097]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="6" pageNumber="537">NHRS</collectionCode>
], 232518, 232532 [together with 3♁ 
<specimenCount id="9D1DFD13FF93FF9DFD7EFBF2B9BDFBFC" box="[648,697,1071,1097]" pageId="6" pageNumber="537" type="female">14♀</specimenCount>
of 
<taxonomicName id="4C1B4D19FF93FF9DFD29FBF2B842FBFC" baseAuthorityName="Lawrence" baseAuthorityYear="1927" box="[735,838,1071,1097]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="oncka">
<emphasis id="B96FEA88FF93FF9DFD29FBF2B842FBFC" box="[735,838,1071,1097]" italics="true" pageId="6" pageNumber="537">W. oncka</emphasis>
</taxonomicName>
], 232533)
</materialsCitation>
. 
<materialsCitation id="3B733CC7FF93FF9DFC3FFBF2B98EFBD8" collectingDate="1937-02" collectionCode="MRAC" collectorName="E. Dartevelle" country="Democratic Republic of the Congo" latitude="-5.8166666" location="Matadi" longLatPrecision="1305" longitude="13.466666" pageId="6" pageNumber="537" specimenCode="MRAC 1219" specimenCount="1" stateProvince="Bas-Congo">
<emphasis id="B96FEA88FF93FF9DFC3FFBF2BF78FBFC" box="[969,1148,1071,1097]" italics="true" pageId="6" pageNumber="537">
<collectingRegion id="49DFF878FF93FF9DFC3FFBF2BF72FBFC" box="[969,1142,1071,1097]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="6" pageNumber="537">Kongo-Central</collectingRegion>
:
</emphasis>
<location id="8EC46041FF93FF9DFB72FBF2BFD2FBFC" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFB72FBF2BFD2FBFC" box="[1156,1238,1071,1097]" country="Democratic Republic of the Congo" latitude="-5.8166666" longLatPrecision="1305" longitude="13.466666" name="Matadi" pageId="6" pageNumber="537" stateProvince="Bas-Congo">Matadi</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFB17FBEDBE2AFBFE" box="[1249,1326,1072,1099]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-5.8166666">5°49′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFACDFBEDBE93FBFE" box="[1339,1431,1072,1099]" degrees="13" direction="east" minutes="28" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="13.466666">13°28′E</geoCoordinate>
, 
<date id="FFA5105AFF93FF9DFF61FB8EBA08FBD8" box="[151,268,1107,1133]" pageId="6" pageNumber="537" value="1937-02">
<collectingDate id="EFE1E9B2FF93FF9DFF61FB8EBA08FBD8" box="[151,268,1107,1133]" pageId="6" pageNumber="537" value="1937-02">Feb. 1937</collectingDate>
</date>
, 
<collectorName id="26EE534CFF93FF9DFEEFFB8EBAB6FBD8" box="[281,434,1107,1133]" pageId="6" pageNumber="537">E. Dartevelle</collectorName>
, 1♁ (
<specimenCode id="DBBD9EE1FF93FF9DFE07FB8EB987FBD8" box="[497,643,1107,1133]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 1219</specimenCode>
)
</materialsCitation>
.— 
<materialsCitation id="3B733CC7FF93FF9DFD5DFB8EBF3EFB24" collectingDate="1989-07-20" collectionCode="MRAC" collectorName="M. Alderweireldt" country="Equatorial Guinea" latitude="1.15" location="Rio Muni" longLatPrecision="1308" longitude="9.55" municipality="Between Cogo" pageId="6" pageNumber="537" specimenCode="MRAC 170101" specimenCount="4" specimenCount-female="4" stateProvince="Kongo-Central">
<collectingCountry id="F30C760AFF93FF9DFD5DFB8EB8DEFBD8" box="[683,986,1107,1133]" name="Equatorial Guinea" pageId="6" pageNumber="537">EQUATORIAL GUINEA</collectingCountry>
. 
<collectingMunicipality id="6BC0ACE0FF93FF9DFC10FB8EBF8AFBD8" box="[998,1166,1107,1133]" pageId="6" pageNumber="537">Between Cogo</collectingMunicipality>
and 
<location id="8EC46041FF93FF9DFB3EFB8EBE3CFBD8" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFB3EFB8EBE3CFBD8" box="[1224,1336,1107,1133]" country="Equatorial Guinea" latitude="1.15" longLatPrecision="1308" longitude="9.55" municipality="Between Cogo" name="Rio Muni" pageId="6" pageNumber="537" stateProvince="Kongo-Central">Rio Muni</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFAB0FB89BE9CFBDA" box="[1350,1432,1108,1135]" degrees="1" direction="north" minutes="09" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="1.15">1°09′N</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFF61FBA5BBE2FB26" box="[151,230,1144,1171]" degrees="9" direction="east" minutes="33" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="9.55">9°33′E</geoCoordinate>
, secondary forest, 
<date id="FFA5105AFF93FF9DFE31FBAAB95AFB24" box="[455,606,1143,1169]" pageId="6" pageNumber="537" value="1989-07-20">
<collectingDate id="EFE1E9B2FF93FF9DFE31FBAAB95AFB24" box="[455,606,1143,1169]" pageId="6" pageNumber="537" value="1989-07-20">20 Jul. 1989</collectingDate>
</date>
, 
<collectorName id="26EE534CFF93FF9DFD98FBAAB832FB24" box="[622,822,1143,1169]" pageId="6" pageNumber="537">M. Alderweireldt</collectorName>
, 
<specimenCount id="9D1DFD13FF93FF9DFCB0FBAAB86EFB24" box="[838,874,1143,1169]" pageId="6" pageNumber="537" type="female">4♀</specimenCount>
(
<specimenCode id="DBBD9EE1FF93FF9DFC8BFBAABF36FB24" box="[893,1074,1143,1169]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 170101</specimenCode>
)
</materialsCitation>
.— 
<materialsCitation id="3B733CC7FF93FF9DFBADFBAABFF3FB00" collectingDate="1985-10" collectingDateMax="1985-11-31" collectingDateMin="1985-10" collectionCode="MRAC" collectorName="A. Pauly" country="Gabon" latitude="0.38333333" location="Ntoum" longLatPrecision="1308" longitude="9.783334" pageId="6" pageNumber="537" specimenCode="MRAC 172791" specimenCount="1" stateProvince="Estuaire">
<collectingCountry id="F30C760AFF93FF9DFBADFBAABFC0FB24" box="[1115,1220,1143,1169]" name="Gabon" pageId="6" pageNumber="537">GABON</collectingCountry>
. 
<emphasis id="B96FEA88FF93FF9DFB22FBAABE3FFB25" box="[1236,1339,1143,1168]" italics="true" pageId="6" pageNumber="537">
<collectingRegion id="49DFF878FF93FF9DFB22FBAABE31FB25" box="[1236,1333,1143,1168]" country="Gabon" name="Estuaire" pageId="6" pageNumber="537">Estuaire</collectingRegion>
:
</emphasis>
<location id="8EC46041FF93FF9DFAB1FBAABE9CFB24" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFAB1FBAABE9CFB24" box="[1351,1432,1143,1169]" country="Gabon" latitude="0.38333333" longLatPrecision="1308" longitude="9.783334" name="Ntoum" pageId="6" pageNumber="537" stateProvince="Estuaire">Ntoum</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFF61FB41BBEDFB02" box="[151,233,1180,1207]" degrees="0" direction="north" minutes="23" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="0.38333333">0°23′N</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFF00FB41BA41FB02" box="[246,325,1180,1207]" degrees="9" direction="east" minutes="47" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="9.783334">9°47′E</geoCoordinate>
, piège, bac d’eau, forêt, 
<date id="FFA5105AFF93FF9DFDA8FB46B811FB00" box="[606,789,1178,1205]" pageId="6" pageNumber="537" value="1985-10" valueMax="1985-11-31" valueMin="1985-10-00">
<collectingDate id="EFE1E9B2FF93FF9DFDA8FB46B811FB00" box="[606,789,1178,1205]" pageId="6" pageNumber="537" value="1985-10" valueMax="1985-11-31" valueMin="1985-10-00">Oct.–Nov. 1985</collectingDate>
</date>
, 
<collectorName id="26EE534CFF93FF9DFCE9FB46B880FB00" box="[799,900,1179,1205]" pageId="6" pageNumber="537">A. Pauly</collectorName>
, 2♁ (
<specimenCode id="DBBD9EE1FF93FF9DFC35FB46BF70FB00" box="[963,1140,1179,1205]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 172791</specimenCode>
&amp; 172836)
</materialsCitation>
.— 
<materialsCitation id="3B733CC7FF93FF9DFAEEFB46BFEFFB6C" collectingDate="1979-04" collectionCode="MRAC" collectorName="M. E. Baddeley" country="Namibia" latitude="-18.066668" location="Andara" longLatPrecision="1276" longitude="21.483334" pageId="6" pageNumber="537" specimenCode="MRAC 170589" specimenCount="1" specimenCount-female="1" stateProvince="Kavango East">
<collectingCountry id="F30C760AFF93FF9DFAEEFB46BE9DFB00" box="[1304,1433,1179,1205]" name="Namibia" pageId="6" pageNumber="537">NAMIBIA</collectingCountry>
. 
<emphasis id="B96FEA88FF93FF9DFF61FB62BA45FB6D" box="[151,321,1215,1240]" italics="true" pageId="6" pageNumber="537">
<collectingRegion id="49DFF878FF93FF9DFF61FB62BA3FFB6D" box="[151,315,1215,1240]" country="Namibia" name="Kavango East" pageId="6" pageNumber="537">Kavango East</collectingRegion>
:
</emphasis>
<location id="8EC46041FF93FF9DFEBCFB62BA9BFB6C" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFEBCFB62BA9BFB6C" box="[330,415,1215,1241]" country="Namibia" latitude="-18.066668" longLatPrecision="1276" longitude="21.483334" name="Andara" pageId="6" pageNumber="537" stateProvince="Kavango East">Andara</location>
, 
<geoCoordinate id="EE2F505DFF93FF9DFE5BFB1DB90CFB6E" box="[429,520,1216,1243]" degrees="18" direction="south" minutes="04" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-18.066668">18°04′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFDE1FB1DB970FB6E" box="[535,628,1216,1243]" degrees="21" direction="east" minutes="29" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="21.483334">21°29′E</geoCoordinate>
, 
<date id="FFA5105AFF93FF9DFD77FB62B9F3FB6C" box="[641,759,1215,1241]" pageId="6" pageNumber="537" value="1979-04">
<collectingDate id="EFE1E9B2FF93FF9DFD77FB62B9F3FB6C" box="[641,759,1215,1241]" pageId="6" pageNumber="537" value="1979-04">Apr. 1979</collectingDate>
</date>
, 
<collectorName id="26EE534CFF93FF9DFCF3FB62B8B8FB6C" box="[773,956,1215,1241]" pageId="6" pageNumber="537">M. E. Baddeley</collectorName>
, 2♁ 
<specimenCount id="9D1DFD13FF93FF9DFC01FB62BF1EFB6C" box="[1015,1050,1215,1241]" pageId="6" pageNumber="537" type="female">1♀</specimenCount>
(
<specimenCode id="DBBD9EE1FF93FF9DFBDDFB62BFE7FB6C" box="[1067,1251,1214,1241]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 170589</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B733CC7FF93FF9DFB0FFB62B97BFB48" collectingDate="1979-05" collectingDateMax="1979-06-31" collectingDateMin="1979-05" collectionCode="MRAC" collectorName="Baddeley" country="Namibia" pageId="6" pageNumber="537" specimenCode="MRAC 170638" specimenCount="1" stateProvince="Kavango East">
do., 
<date id="FFA5105AFF93FF9DFADBFB62BBD6FB48" pageId="6" pageNumber="537" value="1979-05" valueMax="1979-06-31" valueMin="1979-05-00">
<collectingDate id="EFE1E9B2FF93FF9DFADBFB62BBD6FB48" pageId="6" pageNumber="537" value="1979-05" valueMax="1979-06-31" valueMin="1979-05-00">May–Jun. 1979</collectingDate>
</date>
, M. E. 
<collectorName id="26EE534CFF93FF9DFED6FB3EBA8FFB48" box="[288,395,1251,1277]" pageId="6" pageNumber="537">Baddeley</collectorName>
, 1♁ (
<specimenCode id="DBBD9EE1FF93FF9DFE33FB3EB97CFB48" box="[453,632,1251,1277]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 170638</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B733CC7FF93FF9DFD7CFB3EB920FA94" collectingDate="1979-10" collectingDateMax="1979-11-31" collectingDateMin="1979-10" collectionCode="MRAC" collectorName="Baddeley" country="Namibia" pageId="6" pageNumber="537" specimenCode="MRAC 170640" specimenCount="2" specimenCount-female="2" stateProvince="Kavango East">
do., 
<date id="FFA5105AFF93FF9DFD4FFB3EB869FB48" box="[697,877,1251,1277]" pageId="6" pageNumber="537" value="1979-10" valueMax="1979-11-31" valueMin="1979-10-00">
<collectingDate id="EFE1E9B2FF93FF9DFD4FFB3EB869FB48" box="[697,877,1251,1277]" pageId="6" pageNumber="537" value="1979-10" valueMax="1979-11-31" valueMin="1979-10-00">Oct.–Nov. 1979</collectingDate>
</date>
, M. E. 
<collectorName id="26EE534CFF93FF9DFC4CFB3EBF21FB48" box="[954,1061,1251,1277]" pageId="6" pageNumber="537">Baddeley</collectorName>
, 
<specimenCount id="9D1DFD13FF93FF9DFBD9FB3EBF56FB48" box="[1071,1106,1251,1277]" pageId="6" pageNumber="537" type="female">2♀</specimenCount>
(
<specimenCode id="DBBD9EE1FF93FF9DFBA9FB3EBE0BFB48" box="[1119,1295,1251,1277]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="537" type="Museum">MRAC 170640</specimenCode>
) (previously identified as 
<taxonomicName id="4C1B4D19FF93FF9DFED8FADAB91DFA94" authorityName="Simon" authorityYear="1898" box="[302,537,1287,1313]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96FEA88FF93FF9DFED8FADAB91DFA94" box="[302,537,1287,1313]" italics="true" pageId="6" pageNumber="537">Pardosa manubriata</emphasis>
</taxonomicName>
)
</materialsCitation>
. 
<materialsCitation id="3B733CC7FF93FF9DFDC5FADABA06FAF0" collectionCode="SMF" country="Namibia" latitude="-21.983334" location="Okahandja" longLatPrecision="1262" longitude="16.916666" pageId="6" pageNumber="537" specimenCode="SMF 9910988" specimenCount="1" stateProvince="Otjozondjupa">
<emphasis id="B96FEA88FF93FF9DFDC5FADAB9D1FA94" box="[563,725,1287,1313]" italics="true" pageId="6" pageNumber="537">
<collectingRegion id="49DFF878FF93FF9DFDC5FADAB9CBFA94" box="[563,719,1287,1313]" country="Namibia" name="Otjozondjupa" pageId="6" pageNumber="537">Otjozondjupa</collectingRegion>
:
</emphasis>
<location id="8EC46041FF93FF9DFD29FADAB85FFA94" LSID="urn:lsid:plazi:treatment:03B2878CFF93FF93FF61FDB7BF57FC58:8EC46041FF93FF9DFD29FADAB85FFA94" box="[735,859,1287,1313]" country="Namibia" latitude="-21.983334" longLatPrecision="1262" longitude="16.916666" name="Okahandja" pageId="6" pageNumber="537" stateProvince="Otjozondjupa">Okahandja</location>
, [
<geoCoordinate id="EE2F505DFF93FF9DFC87FAD5B8C9FA96" box="[881,973,1288,1315]" degrees="21" direction="south" minutes="59" orientation="latitude" pageId="6" pageNumber="537" precision="925" value="-21.983334">21°59′S</geoCoordinate>
, 
<geoCoordinate id="EE2F505DFF93FF9DFC2BFADBBF39FA94" box="[989,1085,1286,1313]" degrees="16" direction="east" minutes="55" orientation="longitude" pageId="6" pageNumber="537" precision="925" value="16.916666">
16°55 
<emphasis id="B96FEA88FF93FF9DFBD6FADABF2CFA94" bold="true" box="[1056,1064,1287,1313]" italics="true" pageId="6" pageNumber="537">′</emphasis>
E
</geoCoordinate>
], 1♁, ex RII/10988/648 (
<specimenCode id="DBBD9EE1FF93FF9DFA95FADABBFFFAF0" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="537">SMF 9910988</specimenCode>
)
</materialsCitation>
(together with 
<specimenCount id="9D1DFD13FF93FF9DFE58FAF6BAD5FAF0" box="[430,465,1323,1349]" pageId="6" pageNumber="537" type="female">1♀</specimenCount>
of 
<taxonomicName id="4C1B4D19FF93FF9DFE01FAF6B998FAF0" authorityName="Simon" authorityYear="1898" box="[503,668,1323,1349]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96FEA88FF93FF9DFE01FAF6B998FAF0" box="[503,668,1323,1349]" italics="true" pageId="6" pageNumber="537">W. manubriata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016511FF93FF9DFF31FA92B956F9A8" pageId="6" pageNumber="537" type="discussion">
<paragraph id="8BA4369AFF93FF9DFF31FA92B956F9A8" blockId="6.[151,1437,782,2033]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31FA92BA34FADC" bold="true" box="[199,304,1359,1385]" pageId="6" pageNumber="537">Remark.</emphasis>
<bibRefCitation id="EF8A4B6BFF93FF9DFECCFA92BAFDFADC" author="Roewer, C. F." box="[314,505,1358,1385]" pageId="6" pageNumber="537" pagination="1 - 518" refId="ref11081" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II (Lycosidae). Exploration du Parc National de l'Upemba, Mission G. F. De Witte, 55, 1 - 518." type="journal article" year="1959">Roewer’s (1959)</bibRefCitation>
description of 
<taxonomicName id="4C1B4D19FF93FF9DFD5DFA92B84EFADC" authorityName="Roewer" authorityYear="1959" box="[683,842,1359,1385]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="mabweana">
<emphasis id="B96FEA88FF93FF9DFD5DFA92B84EFADC" box="[683,842,1359,1385]" italics="true" pageId="6" pageNumber="537">W. mabweana</emphasis>
</taxonomicName>
was based on 
<specimenCount id="9D1DFD13FF93FF9DFC0FFA92BF19FADC" box="[1017,1053,1359,1385]" pageId="6" pageNumber="537" type="female">7♀</specimenCount>
named ‘Typus’ from 
<collectingCountry id="F30C760AFF93FF9DFAECFA92BE9CFADC" box="[1306,1432,1359,1385]" name="Democratic Republic of the Congo" pageId="6" pageNumber="537">DR Congo</collectingCountry>
, Upemba National Park, “Mabwe, riv. est du Lac Upemba, 
<quantity id="4CE39B7FFF93FF9DFCCEFAAFB887FA38" box="[824,899,1394,1421]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="6" pageNumber="537" unit="m" value="585.0">585 m</quantity>
, 
<date id="FFA5105AFF93FF9DFC67FAAEBF45FA38" box="[913,1089,1395,1421]" pageId="6" pageNumber="537" value="1947-08-01" valueMax="1947-08-12" valueMin="1947-08-01">1–12.VIII.1947</date>
” and 
<specimenCount id="9D1DFD13FF93FF9DFB70FAAFBFADFA38" box="[1158,1193,1394,1421]" pageId="6" pageNumber="537" type="female">5♀</specimenCount>
named ’Paratypoide’ from Upemba National Park, “Buye-Bala, affl[uent] g[auche] Muye et s[ous]-affl[uent] dr[oit] Lufira, 
<quantity id="4CE39B7FFF93FF9DFACBFA4BBE93FA04" box="[1341,1431,1430,1457]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="537" unit="m" value="1750.0">1750 m</quantity>
, 
<date id="FFA5105AFF93FF9DFF61FA66BA18FA60" box="[151,284,1467,1493]" pageId="6" pageNumber="537" value="1948-09-27">27.IX.1948</date>
”. As no 
<typeStatus id="54A08838FF93FF9DFE8CFA66BAD8FA60" box="[378,476,1467,1493]" pageId="6" pageNumber="537" type="holotype">holotype</typeStatus>
was formally selected, all original materials are regarded as 
<typeStatus id="54A08838FF93FF9DFB7EFA66BFEEFA60" box="[1160,1258,1467,1493]" pageId="6" pageNumber="537" type="syntype">syntypes</typeStatus>
and a 
<typeStatus id="54A08838FF93FF9DFAC0FA66BE99FA60" box="[1334,1437,1467,1493]" pageId="6" pageNumber="537" type="lectotype">lectotype</typeStatus>
(previously labelled ‘holotype’) is here designated. Remaining females of the material listed in 
<bibRefCitation id="EF8A4B6BFF93FF9DFB4FFA02BE65FA4C" author="Roewer, C. F." box="[1209,1377,1502,1529]" pageId="6" pageNumber="537" pagination="1 - 518" refId="ref11081" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II (Lycosidae). Exploration du Parc National de l'Upemba, Mission G. F. De Witte, 55, 1 - 518." type="journal article" year="1959">Roewer (1959)</bibRefCitation>
have not been located and are presumed lost.
</paragraph>
</subSubSection>
<subSubSection id="C3016511FF93FF9DFF31F9FBBF34F964" pageId="6" pageNumber="537" type="diagnosis">
<paragraph id="8BA4369AFF93FF9DFF31F9FBBF34F964" blockId="6.[151,1437,782,2033]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31F9FBBA46F9F5" bold="true" box="[199,322,1574,1600]" pageId="6" pageNumber="537">Diagnosis.</emphasis>
The male differs from other 
<taxonomicName id="4C1B4D19FF93FF9DFD64F9FAB85CF9F4" box="[658,856,1575,1601]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFD64F9FAB9FAF9F4" box="[658,766,1575,1601]" italics="true" pageId="6" pageNumber="537">Wadicosa</emphasis>
species
</taxonomicName>
except 
<taxonomicName id="4C1B4D19FF93FF9DFC45F9FABF41F9F4" authorityName="Kronestedt" authorityYear="2023" baseAuthorityName="Roewer" baseAuthorityYear="1959" box="[947,1093,1575,1601]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="enucleata">
<emphasis id="B96FEA88FF93FF9DFC45F9FABF41F9F4" box="[947,1093,1575,1601]" italics="true" pageId="6" pageNumber="537">W. enucleata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1B4D19FF93FF9DFB89F9FABE23F9F4" authorityName="Simon" authorityYear="1898" box="[1151,1319,1575,1601]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96FEA88FF93FF9DFB89F9FABE23F9F4" box="[1151,1319,1575,1601]" italics="true" pageId="6" pageNumber="537">W. manubriata</emphasis>
</taxonomicName>
by a long and looped embolus. It differs from 
<taxonomicName id="4C1B4D19FF93FF9DFDDBF996B9CDF9D0" authorityName="Roewer" authorityYear="1959" box="[557,713,1611,1637]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="mabweana">
<emphasis id="B96FEA88FF93FF9DFDDBF996B9CDF9D0" box="[557,713,1611,1637]" italics="true" pageId="6" pageNumber="537">W. mabweana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1B4D19FF93FF9DFCF6F996B8A2F9D0" authorityName="Simon" authorityYear="1898" box="[768,934,1611,1637]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96FEA88FF93FF9DFCF6F996B8A2F9D0" box="[768,934,1611,1637]" italics="true" pageId="6" pageNumber="537">W. manubriata</emphasis>
</taxonomicName>
by a weakly developed posterior retrolateral tegular process in the bulbus (
<figureCitation id="13202A1FFF93FF9DFE06F9B2B94EF93C" box="[496,586,1647,1673]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="6" pageNumber="537">Fig. 4E</figureCitation>
: 
<emphasis id="B96FEA88FF93FF9DFDA0F9B2B997F93C" box="[598,659,1647,1673]" italics="true" pageId="6" pageNumber="537">p.rl.p</emphasis>
, 4H: 
<emphasis id="B96FEA88FF93FF9DFD20F9B2B81DF93D" box="[726,793,1647,1672]" italics="true" pageId="6" pageNumber="537">arrow</emphasis>
). The female is similar to 
<taxonomicName id="4C1B4D19FF93FF9DFBBAF9B2BFDAF93C" authorityName="Kronestedt" authorityYear="2023" baseAuthorityName="Roewer" baseAuthorityYear="1959" box="[1100,1246,1647,1673]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="enucleata">
<emphasis id="B96FEA88FF93FF9DFBBAF9B2BFDAF93C" box="[1100,1246,1647,1673]" italics="true" pageId="6" pageNumber="537">W. enucleata</emphasis>
</taxonomicName>
in configuration of the epigyne but differs by the cavity in 
<taxonomicName id="4C1B4D19FF93FF9DFD83F94EB815F918" authorityName="Roewer" authorityYear="1959" box="[629,785,1683,1709]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="mabweana">
<emphasis id="B96FEA88FF93FF9DFD83F94EB815F918" box="[629,785,1683,1709]" italics="true" pageId="6" pageNumber="537">W. mabweana</emphasis>
</taxonomicName>
often being less wide and the foveolae less distinct when observed under light microscope (
<emphasis id="B96FEA88FF93FF9DFDE0F96AB936F964" box="[534,562,1719,1745]" italics="true" pageId="6" pageNumber="537">cf.</emphasis>
<figureCitation id="13202A1FFF93FF9DFDCEF96AB9FCF964" box="[568,760,1718,1745]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[163,1427,1037,1869]" captionTargetId="figure-375@8.[151,1436,1026,1874]" captionTargetPageId="8" captionText="FIGURE 5. Epigyne, ventral view. A. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, C. W. mabweana (Roewer), from DR Congo: Mikembo. D. W. manubriata (Simon), from Zimbabwe: Umguza River. E. W. sordidecolorata (Strand), from Cameroon: Koum. Scale bar: 0.2 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.7523279" httpUri="https://zenodo.org/record/7523279/files/figure.png" pageId="6" pageNumber="537">Figs 5B–C and A</figureCitation>
as well as 6C–D and A–B).
</paragraph>
</subSubSection>
<subSubSection id="C3016511FF93FF93FF31F907BE8EFC7C" lastPageId="8" lastPageNumber="539" pageId="6" pageNumber="537" type="description">
<paragraph id="8BA4369AFF93FF9DFF31F907BFD5F940" blockId="6.[151,1437,782,2033]" box="[199,1233,1754,1781]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31F907BA9FF940" bold="true" box="[199,411,1754,1781]" pageId="6" pageNumber="537">
Description. 
<emphasis id="B96FEA88FF93FF9DFE96F907BA9FF940" bold="true" box="[352,411,1754,1781]" italics="true" pageId="6" pageNumber="537">Male</emphasis>
</emphasis>
(
<collectingCountry id="F30C760AFF93FF9DFE5FF906B92CF940" box="[425,552,1755,1781]" name="Democratic Republic of the Congo" pageId="6" pageNumber="537">DR Congo</collectingCountry>
: Mikembo). Total length 5.1. Carapace 2.80 long, 2.15 wide.
</paragraph>
<paragraph id="8BA4369AFF93FF9DFF31F922B918F81C" blockId="6.[151,1437,782,2033]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31F922BA6BF8AC" box="[199,367,1791,1817]" italics="true" pageId="6" pageNumber="537">Cephalothorax</emphasis>
(
<figureCitation id="13202A1FFF93FF9DFE76F922BAF8F8AC" box="[384,508,1791,1817]" captionStart="FIGURE 2" captionStartId="4.[151,250,1888,1912]" captionTargetBox="[193,1395,179,1862]" captionTargetId="figure-18@4.[192,1395,179,1862]" captionTargetPageId="4" captionText="FIGURE 2. Habitus, dorsal view.A–C. Wadicosa enucleata (Roewer).A ♁ from Tanzania: Mkomazi, B ♁ from Nigeria: Ilorin. C ♀ from Tanzania: Kyela. D–F. W. mabweana (Roewer). D ♁ and F ♀ from DR Congo: Mikembo, E ♁ from Namibia:Andara. G–I. W. manubriata (Simon). G ♁ from Namibia: Brukkaros Crater. H ♁ from South Africa: Dendron. I ♀ from Zimbabwe: Umguza River. Scale bar: 2 mm (A–I)." figureDoi="http://doi.org/10.5281/zenodo.7523273" httpUri="https://zenodo.org/record/7523273/files/figure.png" pageId="6" pageNumber="537">Fig. 2D–E</figureCitation>
). Carapace brown to blackish with oval yellowish to brownish median field, edge somewhat jagged, in thoracic part. No lateral bands. Sides of carapace with short black pubescence. Median field with recumbent longer white pubescence, often more or less missing due to wear. Recumbent white hairs in between eyes forwards to AME. Clypeus yellowish to dark brown. Chelicerae yellowish to dark brown with sooty longitudinal markings. Sternum greyish brown.
</paragraph>
<paragraph id="8BA4369AFF93FF9DFF31F86EBFAFF844" blockId="6.[151,1437,782,2033]" pageId="6" pageNumber="537">
<emphasis id="B96FEA88FF93FF9DFF31F86EBA00F879" box="[199,260,1971,1996]" italics="true" pageId="6" pageNumber="537">Eyes.</emphasis>
Width of row I 50 (slightly procurved as seen from front), row II 64, row III 80, row II–III 63. Diameter of AME 13, ALE 10, PME 24, PLE 20. Distance between AME 8, between AME and ALE 2.
</paragraph>
<caption id="DF646612FF92FF9CFF61F931BBD2F8C4" ID-DOI="http://doi.org/10.5281/zenodo.7523277" ID-Zenodo-Dep="7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="7" pageNumber="538" startId="7.[151,250,1772,1796]" targetBox="[151,1436,179,1747]" targetPageId="7">
<paragraph id="8BA4369AFF92FF9CFF61F931BBD2F8C4" blockId="7.[151,1437,1772,1905]" pageId="7" pageNumber="538">
<emphasis id="B96FEA88FF92FF9CFF61F931BA10F8B0" bold="true" box="[151,276,1772,1797]" pageId="7" pageNumber="538">FIGURE 4.</emphasis>
Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. 
<taxonomicName id="4C1B4D19FF92FF9CFACDF930BA5BF89C" authority="(Roewer)" authorityName="Kronestedt" authorityYear="2023" baseAuthorityName="Roewer" baseAuthorityYear="1959" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="enucleata">
<emphasis id="B96FEA88FF92FF9CFACDF930BBFDF89C" italics="true" pageId="7" pageNumber="538">Wadicosa enucleata</emphasis>
(Roewer)
</taxonomicName>
from Tanzania: Kyela. B, E, H. 
<taxonomicName id="4C1B4D19FF92FF9CFD52F8CCB893F89C" authority="(Roewer)" baseAuthorityName="Roewer" box="[676,919,1809,1833]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="mabweana">
<emphasis id="B96FEA88FF92FF9CFD52F8CCB836F89C" box="[676,818,1809,1833]" italics="true" pageId="7" pageNumber="538">W. mabweana</emphasis>
(Roewer)
</taxonomicName>
from DR Congo: Mikembo. C, F, I. 
<taxonomicName id="4C1B4D19FF92FF9CFAF1F8CCBBEEF8F8" authority="(Simon)" baseAuthorityName="Simon" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96FEA88FF92FF9CFAF1F8CCBE98F89C" box="[1287,1436,1809,1833]" italics="true" pageId="7" pageNumber="538">W. manubriata</emphasis>
(Simon)
</taxonomicName>
from Namibia: Windhoek. 
<emphasis id="B96FEA88FF92FF9CFDF1F8E8B955F8F9" box="[519,593,1845,1868]" italics="true" pageId="7" pageNumber="538">Arrows</emphasis>
point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I).
</paragraph>
</caption>
<paragraph id="8BA4369AFF92FF9CFF31F842BE99F80C" blockId="7.[151,1437,1951,2013]" box="[199,1437,1951,1977]" pageId="7" pageNumber="538">
<emphasis id="B96FEA88FF92FF9CFF31F842BA35F80C" box="[199,305,1951,1977]" italics="true" pageId="7" pageNumber="538">Abdomen</emphasis>
(
<figureCitation id="13202A1FFF92FF9CFEB1F842BACCF80C" box="[327,456,1951,1977]" captionStart="FIGURE 2" captionStartId="4.[151,250,1888,1912]" captionTargetBox="[193,1395,179,1862]" captionTargetId="figure-18@4.[192,1395,179,1862]" captionTargetPageId="4" captionText="FIGURE 2. Habitus, dorsal view.A–C. Wadicosa enucleata (Roewer).A ♁ from Tanzania: Mkomazi, B ♁ from Nigeria: Ilorin. C ♀ from Tanzania: Kyela. D–F. W. mabweana (Roewer). D ♁ and F ♀ from DR Congo: Mikembo, E ♁ from Namibia:Andara. G–I. W. manubriata (Simon). G ♁ from Namibia: Brukkaros Crater. H ♁ from South Africa: Dendron. I ♀ from Zimbabwe: Umguza River. Scale bar: 2 mm (A–I)." figureDoi="http://doi.org/10.5281/zenodo.7523273" httpUri="https://zenodo.org/record/7523273/files/figure.png" pageId="7" pageNumber="538">Fig. 2D–E</figureCitation>
). Dorsum anteriorly with greyish brown lanceolate stripe bordered by dark spots.
</paragraph>
<paragraph id="8BA4369AFF92FF93FF61F81EB892FF4C" blockId="7.[151,1437,1951,2013]" lastBlockId="8.[151,1437,151,1005]" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538">
Lanceolate stripe with a large whitish to yellowish elongated patch on each side (
<figureCitation id="13202A1FFF92FF9CFBD6F81EBF7DF868" box="[1056,1145,1987,2013]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1751]" captionTargetBox="[159,1419,416,1689]" captionTargetId="figure-117@3.[159,1429,416,1703]" captionTargetPageId="3" captionText="FIGURE 1. A–C. Wadicosa manubriata (Simon), male. A. Right palp in retrolateral view, from Simon (1898: fig. 356). B–C. Holotype, right palp in retrolateral view (B), and left palp in antero-ventral view (C). Photographs by C. Hervé, MNHN. D–E. W. sordidecolorata (Strand), epigynes, from Strand (1908): D pl. 8, fig. 16 (“Lycosa sordide-colorata”) and E pl. 8, fig.9 (“Lycosa darolii”). F. W. enucleata (Roewer), epigyne, from Roewer (1959, fig. 62b). G–I. W. manubriata (Simon), from Roewer 1959, fig. 63a–c), G. Right palp in retrolateral view. H. Left palp in ventral view. I. Epigyne. (G and H obviously drawn from microslide SMF 13616 (RII 4914/252) in which the embolus in ventral view is distorted similarly to that shown in H.) K. W. mabweana (Roewer), epigyne, from Roewer (1959, fig. 54a). L–M. W. sordidecolorata (Strand), from Roewer (1959, fig. 30a–c). L. Right male palp. M. “Lamella characteristica”= tegular apophysis. N. Epigyne. O. Pardosa darolii (Strand). from Roewer (1959, fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.7523271" httpUri="https://zenodo.org/record/7523271/files/figure.png" pageId="7" pageNumber="538">Fig. 1D</figureCitation>
). From back of lanceolate stripe and rearwards dark chevrons and transverse bars on whitish to yellowish ground. Whitish to yellowish colour due to underlying guanocytes. Venter whitish yellow with recumbent white pubescence and scattered short dark hairs. Sides light brown to blackish brown, dotted with lighter spots.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FEDEBFE0FEF4" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FEDEBBF8FEA9" box="[199,252,259,284]" italics="true" pageId="8" pageNumber="539">Legs</emphasis>
(Table I). Yellowish to greyish brown with darker annulation. In dark specimens, most of Fe appear dark and other segments more or less darkened. Coxae light yellowish, more or less suffused with grey.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FE96BF11FDAC" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FE96BBF8FED0" box="[199,252,331,357]" italics="true" pageId="8" pageNumber="539">Palp</emphasis>
(3B, E, 4B, E, H). Pt 0.50, Ti 0.45, Cy 1.30. Fe mostly dark grey, other segments greyish brown to yellowish, partly suffused with dark grey. Cy brownish, more or less suffused with grey, distally lighter Tegular apophysis in ventral view (
<figureCitation id="13202A1FFF9DFF93FE3AFE4EB92EFE18" box="[460,554,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[163,1419,185,1757]" captionTargetId="figure-15@5.[151,1436,179,1772]" captionTargetPageId="5" captionText="FIGURE 3. Right male palp, ventral view (A–C) and retrolateral view (D–F). A, D. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E. W. mabweana (Roewer) from DR Congo: Mikembo. C, F. W. manubriata (Simon) from Namibia: Brukkaros Crater. Scale bar: 0.5 mm (A–F)." figureDoi="http://doi.org/10.5281/zenodo.7523275" httpUri="https://zenodo.org/record/7523275/files/figure.png" pageId="8" pageNumber="539">Figs 3B</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FDCEFE4EB95EFE18" box="[568,602,403,429]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="8" pageNumber="539">4B</figureCitation>
) longer than wide, widest distally, with hook-like process in lower distal “corner”. Anterior retrolateral tegular process in retrolateral view (
<figureCitation id="13202A1FFF9DFF93FC68FE6AB8F8FE64" box="[926,1020,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[163,1419,185,1757]" captionTargetId="figure-15@5.[151,1436,179,1772]" captionTargetPageId="5" captionText="FIGURE 3. Right male palp, ventral view (A–C) and retrolateral view (D–F). A, D. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E. W. mabweana (Roewer) from DR Congo: Mikembo. C, F. W. manubriata (Simon) from Namibia: Brukkaros Crater. Scale bar: 0.5 mm (A–F)." figureDoi="http://doi.org/10.5281/zenodo.7523275" httpUri="https://zenodo.org/record/7523275/files/figure.png" pageId="8" pageNumber="539">Figs 3E</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FBFDFE6ABF55FE64" box="[1035,1105,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="8" pageNumber="539">4E, H</figureCitation>
) comparatively long, widest proximally, obliquely cut distally. Proximal retrolateral tegular process resembling a low crest (
<figureCitation id="13202A1FFF9DFF93FB4FFE06BE10FE40" box="[1209,1300,475,501]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="8" pageNumber="539">Fig. 4H</figureCitation>
). Conductor not discernible. Embolus (
<figureCitation id="13202A1FFF9DFF93FE4BFE22B93CFDAC" box="[445,568,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[163,1419,185,1757]" captionTargetId="figure-15@5.[151,1436,179,1772]" captionTargetPageId="5" captionText="FIGURE 3. Right male palp, ventral view (A–C) and retrolateral view (D–F). A, D. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E. W. mabweana (Roewer) from DR Congo: Mikembo. C, F. W. manubriata (Simon) from Namibia: Brukkaros Crater. Scale bar: 0.5 mm (A–F)." figureDoi="http://doi.org/10.5281/zenodo.7523275" httpUri="https://zenodo.org/record/7523275/files/figure.png" pageId="8" pageNumber="539">Figs 3B, E</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FDB2FE22B980FDAC" box="[580,644,511,537]" captionStart="FIGURE 4" captionStartId="7.[151,250,1772,1796]" captionTargetBox="[151,1436,179,1747]" captionTargetId="figure-15@7.[151,1436,179,1747]" captionTargetPageId="7" captionText="FIGURE 4. Bulbus of left male palp in ventral (A–C) and retrolateral (D–F, detail enlarged in G–I), view. A, D, G. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon) from Namibia: Windhoek. Arrows point at posterior retrolateral tegular process. Scale bars: 300 µm (A–F), 200 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523277" httpUri="https://zenodo.org/record/7523277/files/figure.png" pageId="8" pageNumber="539">4B, E</figureCitation>
) long, looped, wide, tapering at tip.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FDFFBF51FD88" blockId="8.[151,1437,151,1005]" box="[199,1109,546,573]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FDFFBA1AFD88" bold="true" box="[199,286,546,573]" italics="true" pageId="8" pageNumber="539">Female</emphasis>
(
<collectingCountry id="F30C760AFF9DFF93FEDBFDFEBAAFFD88" box="[301,427,547,573]" name="Democratic Republic of the Congo" pageId="8" pageNumber="539">DR Congo</collectingCountry>
: Mikembo). Total length 7.6. Carapace 3.40 long, 2.60 wide.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FD9AB932FD30" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FD9AB909FDD4" box="[199,525,583,609]" italics="true" pageId="8" pageNumber="539">Cephalothorax and abdomen</emphasis>
(
<figureCitation id="13202A1FFF9DFF93FDEAFD9AB975FDD4" box="[540,625,583,609]" captionStart="FIGURE 2" captionStartId="4.[151,250,1888,1912]" captionTargetBox="[193,1395,179,1862]" captionTargetId="figure-18@4.[192,1395,179,1862]" captionTargetPageId="4" captionText="FIGURE 2. Habitus, dorsal view.A–C. Wadicosa enucleata (Roewer).A ♁ from Tanzania: Mkomazi, B ♁ from Nigeria: Ilorin. C ♀ from Tanzania: Kyela. D–F. W. mabweana (Roewer). D ♁ and F ♀ from DR Congo: Mikembo, E ♁ from Namibia:Andara. G–I. W. manubriata (Simon). G ♁ from Namibia: Brukkaros Crater. H ♁ from South Africa: Dendron. I ♀ from Zimbabwe: Umguza River. Scale bar: 2 mm (A–I)." figureDoi="http://doi.org/10.5281/zenodo.7523273" httpUri="https://zenodo.org/record/7523273/files/figure.png" pageId="8" pageNumber="539">Fig. 2F</figureCitation>
). Similar to male. Lateral bands on carapace may be present as indistinct yellowish patches with whitish hairs.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FD52BFAEFD78" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FD52BA00FD1D" box="[199,260,655,680]" italics="true" pageId="8" pageNumber="539">Eyes.</emphasis>
Width of row I 58 (slightly procurved as seen from front), row II 74, row III 93, row II–III 74. Diameter of AME 14, ALE 11, PME 27, PLE 24. Distance between AME 9, between AME and ALE 2.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FD0AB8E8FCA0" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FD0ABBF8FD45" box="[199,252,727,752]" italics="true" pageId="8" pageNumber="539">Legs</emphasis>
(
<tableCitation id="C6990321FF9DFF93FEFBFD0ABA66FD44" box="[269,354,727,753]" captionStart="TABLE 1" captionStartId="14.[152,244,151,175]" captionText="TABLE 1. Leg measurements (in mm) of Wadicosa enucleata, W. mabweana, W. manubriata and W. sordidecolorata." pageId="8" pageNumber="539">Table 1</tableCitation>
). Yellowish with greyish annulation. Darker specimens with darkened legs: Fe may be mostly black with annulation of whitish hairs. Coxae yellowish, suffused with grey.
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FCC2B9D9FC10" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FCC2BA81FC8C" box="[199,389,799,825]" italics="true" pageId="8" pageNumber="539">Female genitalia</emphasis>
(
<figureCitation id="13202A1FFF9DFF93FE64FCC2B909FC8C" box="[402,525,798,825]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[163,1427,1037,1869]" captionTargetId="figure-375@8.[151,1436,1026,1874]" captionTargetPageId="8" captionText="FIGURE 5. Epigyne, ventral view. A. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, C. W. mabweana (Roewer), from DR Congo: Mikembo. D. W. manubriata (Simon), from Zimbabwe: Umguza River. E. W. sordidecolorata (Strand), from Cameroon: Koum. Scale bar: 0.2 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.7523279" httpUri="https://zenodo.org/record/7523279/files/figure.png" pageId="8" pageNumber="539">Figs 5B–C</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FDE1FCC2B958FC8C" box="[535,604,799,825]" captionStart="FIGURE 6" captionStartId="9.[151,250,878,902]" captionTargetBox="[151,1436,179,852]" captionTargetId="figure-15@9.[151,1436,179,852]" captionTargetPageId="9" captionText="FIGURE 6. Epigyne, ventral view.A–B. Wadicosa enucleata (Roewer), A holotype, B from Zimbabwe: Batoka Gorge. C–D. W. mabweana (Roewer), C lectotype, D from Namibia:Andara. E–F. W. manubriata (Simon), E from Namibia: Windhoek, F from Namibia: Brukkaros Crater. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7523281" httpUri="https://zenodo.org/record/7523281/files/figure.png" pageId="8" pageNumber="539">6C–D</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FD90FCC2B9CDFC8C" box="[614,713,799,825]" captionStart="FIGURE 7" captionStartId="10.[151,250,1840,1864]" captionTargetBox="[151,1435,846,1814]" captionTargetId="figure-325@10.[151,1435,846,1814]" captionTargetPageId="10" captionText="FIGURE 7. Epigyne in ventral (A, B, C) and female genitalia in dorsal (D–F, details in G–I) view.A, D, G. Wadicosa enucleata (Roewer) from Zimbabwe: Batoka Gorge. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon), C from Namibia: Windhoek, F from South Africa: Paarl (F). Scale bars: 300 µm (A–F), 50 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523285" httpUri="https://zenodo.org/record/7523285/files/figure.png" pageId="8" pageNumber="539">7B, E, H</figureCitation>
). Epigyne with median cavity wider than long, anteriorly covered by cuticular extension forming a transverse edge posteriorly (
<figureCitation id="13202A1FFF9DFF93FCA8FC9EB8DAFCE8" box="[862,990,834,861]" captionStart="FIGURE 5" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[163,1427,1037,1869]" captionTargetId="figure-375@8.[151,1436,1026,1874]" captionTargetPageId="8" captionText="FIGURE 5. Epigyne, ventral view. A. Wadicosa enucleata (Roewer) from Tanzania: Kyela. B, C. W. mabweana (Roewer), from DR Congo: Mikembo. D. W. manubriata (Simon), from Zimbabwe: Umguza River. E. W. sordidecolorata (Strand), from Cameroon: Koum. Scale bar: 0.2 mm (A–E)." figureDoi="http://doi.org/10.5281/zenodo.7523279" httpUri="https://zenodo.org/record/7523279/files/figure.png" pageId="8" pageNumber="539">Figs 5B–C</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FC1BFC9EBF36FCE8" box="[1005,1074,835,861]" captionStart="FIGURE 6" captionStartId="9.[151,250,878,902]" captionTargetBox="[151,1436,179,852]" captionTargetId="figure-15@9.[151,1436,179,852]" captionTargetPageId="9" captionText="FIGURE 6. Epigyne, ventral view.A–B. Wadicosa enucleata (Roewer), A holotype, B from Zimbabwe: Batoka Gorge. C–D. W. mabweana (Roewer), C lectotype, D from Namibia:Andara. E–F. W. manubriata (Simon), E from Namibia: Windhoek, F from Namibia: Brukkaros Crater. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7523281" httpUri="https://zenodo.org/record/7523281/files/figure.png" pageId="8" pageNumber="539">6C–D</figureCitation>
, 
<figureCitation id="13202A1FFF9DFF93FBB6FC9EBF61FCE8" box="[1088,1125,835,861]" captionStart="FIGURE 7" captionStartId="10.[151,250,1840,1864]" captionTargetBox="[151,1435,846,1814]" captionTargetId="figure-325@10.[151,1435,846,1814]" captionTargetPageId="10" captionText="FIGURE 7. Epigyne in ventral (A, B, C) and female genitalia in dorsal (D–F, details in G–I) view.A, D, G. Wadicosa enucleata (Roewer) from Zimbabwe: Batoka Gorge. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon), C from Namibia: Windhoek, F from South Africa: Paarl (F). Scale bars: 300 µm (A–F), 50 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523285" httpUri="https://zenodo.org/record/7523285/files/figure.png" pageId="8" pageNumber="539">7B</figureCitation>
). Anterior to the latter two foveolae, not distinctly separated when seen under light microscope (
<figureCitation id="13202A1FFF9DFF93FC4CFCBABF3DFC34" box="[954,1081,871,897]" captionStart="FIGURE 6" captionStartId="9.[151,250,878,902]" captionTargetBox="[151,1436,179,852]" captionTargetId="figure-15@9.[151,1436,179,852]" captionTargetPageId="9" captionText="FIGURE 6. Epigyne, ventral view.A–B. Wadicosa enucleata (Roewer), A holotype, B from Zimbabwe: Batoka Gorge. C–D. W. mabweana (Roewer), C lectotype, D from Namibia:Andara. E–F. W. manubriata (Simon), E from Namibia: Windhoek, F from Namibia: Brukkaros Crater. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7523281" httpUri="https://zenodo.org/record/7523281/files/figure.png" pageId="8" pageNumber="539">Fig. 6C–D</figureCitation>
). Accessory structures close to midline, partly surrounding spermathecae (
<figureCitation id="13202A1FFF9DFF93FD8FFC56B9D4FC10" box="[633,720,907,933]" captionStart="FIGURE 7" captionStartId="10.[151,250,1840,1864]" captionTargetBox="[151,1435,846,1814]" captionTargetId="figure-325@10.[151,1435,846,1814]" captionTargetPageId="10" captionText="FIGURE 7. Epigyne in ventral (A, B, C) and female genitalia in dorsal (D–F, details in G–I) view.A, D, G. Wadicosa enucleata (Roewer) from Zimbabwe: Batoka Gorge. B, E, H. W. mabweana (Roewer) from DR Congo: Mikembo. C, F, I. W. manubriata (Simon), C from Namibia: Windhoek, F from South Africa: Paarl (F). Scale bars: 300 µm (A–F), 50 µm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.7523285" httpUri="https://zenodo.org/record/7523285/files/figure.png" pageId="8" pageNumber="539">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BA4369AFF9DFF93FF31FC73BE8EFC7C" blockId="8.[151,1437,151,1005]" box="[199,1418,942,969]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FC73BA76FC7D" bold="true" box="[199,370,942,968]" pageId="8" pageNumber="539">Size variation.</emphasis>
Carapace lengths of material measured: males 2.30–3.15 (N=15), females 2.70–3.60 (N=15).
</paragraph>
</subSubSection>
<subSubSection id="C3016511FF9DFF93FF31FC0FBF57FC58" box="[199,1107,978,1005]" pageId="8" pageNumber="539" type="distribution">
<paragraph id="8BA4369AFF9DFF93FF31FC0FBF57FC58" blockId="8.[151,1437,151,1005]" box="[199,1107,978,1005]" pageId="8" pageNumber="539">
<emphasis id="B96FEA88FF9DFF93FF31FC0FBA5DFC59" bold="true" box="[199,345,978,1004]" pageId="8" pageNumber="539">Distribution</emphasis>
(
<figureCitation id="13202A1FFF9DFF93FE9FFC0EBAB8FC58" box="[361,444,979,1005]" captionStart="FIGURE 10" captionStartId="15.[151,250,1891,1915]" captionTargetBox="[151,1436,479,1865]" captionTargetId="figure-76@15.[151,1436,479,1865]" captionTargetPageId="15" captionText="FIGURE 10. Collection localities for Wadicosa enucleata (Roewer), W. mabweana (Roewer), W. manubriata (Simon) and W. sordidecolorata (Strand)." figureDoi="http://doi.org/10.5281/zenodo.7523291" httpUri="https://zenodo.org/record/7523291/files/figure.png" pageId="8" pageNumber="539">Fig. 10</figureCitation>
). 
<collectingCountry id="F30C760AFF9DFF93FE26FC0EB921FC58" box="[464,549,979,1005]" name="Angola" pageId="8" pageNumber="539">Angola</collectingCountry>
, 
<collectingCountry id="F30C760AFF9DFF93FDC7FC0EB9A8FC58" box="[561,684,979,1005]" name="Democratic Republic of the Congo" pageId="8" pageNumber="539">DR Congo</collectingCountry>
, 
<collectingCountry id="F30C760AFF9DFF93FD41FC0EB883FC58" box="[695,903,979,1005]" name="Equatorial Guinea" pageId="8" pageNumber="539">Equatorial Guinea</collectingCountry>
, 
<collectingCountry id="F30C760AFF9DFF93FC64FC0EB8DBFC58" box="[914,991,979,1005]" name="Gabon" pageId="8" pageNumber="539">Gabon</collectingCountry>
, 
<collectingCountry id="F30C760AFF9DFF93FC1DFC0EBF4BFC58" box="[1003,1103,979,1005]" name="Namibia" pageId="8" pageNumber="539">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>