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<document id="8ADADF0B6DFBA09A624B3F5F298914FA" ID-CLB-Dataset="21861" ID-DOI="10.5852/ejt.2016.243" ID-GBIF-Dataset="3aa325f2-3f96-4380-8145-ee552b0bd9c9" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850482" ID-ZooBank="F54A0E2E-1399-4A45-8B7A-0027A8713B33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590445888376" checkinUser="valdenar" docAuthor="Zhang, Lulu, Yang, Yuxia &amp; Okushima, Yûichi" docDate="2016" docId="039B87F51E6DFFC5725719AA8759477B" docLanguage="en" docName="ejt-2016-243.pdf.imf" docOrigin="European Journal of Taxonomy 243" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Fissocantharis nantouensis" docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="FFA2FF8D1E63FFCA70461A5886424576" masterDocTitle="A contribution to the knowledge of the genera Fissocantharis Pic, 1921 and Micropodabrus Pic, 1920 from Taiwan (Coleoptera, Cantharidae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="15" updateTime="1698841283492" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="58C466A95C22135B249533F211C377AF">A contribution to the knowledge of the genera Fissocantharis Pic, 1921 and Micropodabrus Pic, 1920 from Taiwan (Coleoptera, Cantharidae)</mods:title>
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<mods:namePart id="3C57947F4B2973039F619CA1F3D90112">Zhang, Lulu</mods:namePart>
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<mods:affiliation id="B8D8181EC856CF886683A704A9FDE851">The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding 071002, Hebei Province, China. &amp; Email: 2940331171 @ qq. com &amp; urn: lsid: zoobank. org: author: 2 D 7 E 44 B 4 - 1472 - 43 C 0 - A 669 - B 86433 EEA 6 DE</mods:affiliation>
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<mods:namePart id="0022DD9F7DC5D4BB4440E6F5D7FD17F3">Yang, Yuxia</mods:namePart>
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<mods:affiliation id="F0C3636E3510369CF9990831BF0CF247">The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding 071002, Hebei Province, China. &amp; yxyang @ hbu. edu. cn, &amp; urn: lsid: zoobank. org: author: CA 819 BAF-F 411 - 4917 - 9205 - 4 F 1 C 36 AC 4 DC 9 &amp; Co-corresponding authors:</mods:affiliation>
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<mods:namePart id="C574D41BC37B566E7E0B96601B896976">Okushima, Yûichi</mods:namePart>
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<mods:affiliation id="13B481C53D673021737867EC881696C3">Kurashiki Museum of Natural History, Chûô 2 - 6 - 1, Kurashiki, Okayama, 710 - 0046 Japan. &amp; musoku @ city. kurashiki. okayama. jp &amp; urn: lsid: zoobank. org: author: BB 590861 - E 5 AE- 438 B-AC 0 E-A 8724 F 8 FAAB 5 &amp; Co-corresponding authors:</mods:affiliation>
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<treatment id="039B87F51E6DFFC5725719AA8759477B" ID-DOI="http://doi.org/10.5281/zenodo.3854559" ID-GBIF-Taxon="164283178" ID-Zenodo-Dep="3854559" LSID="urn:lsid:plazi:treatment:039B87F51E6DFFC5725719AA8759477B" httpUri="http://treatment.plazi.org/id/039B87F51E6DFFC5725719AA8759477B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C32865681E6DFFC4725719AA8260417B" box="[529,1058,1010,1037]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8B8D36E31E6DFFC4725719AA8260417B" blockId="14.[529,1058,1010,1074]" box="[529,1058,1010,1037]" pageId="14" pageNumber="15">
<heading id="D0C5818F1E6DFFC4725719AA8260417B" box="[529,1058,1010,1037]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C324D601E6DFFC4725719AA8260417B" authority="(Wittmer, 1995)" baseAuthorityName="Wittmer" baseAuthorityYear="1995" box="[529,1058,1010,1037]" class="Insecta" family="Cantharidae" genus="Fissocantharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B946EAF11E6DFFC4725719AA851D417A" bold="true" box="[529,863,1010,1037]" italics="true" pageId="14" pageNumber="15">Fissocantharis nantouensis</emphasis>
(
<bibRefCitation id="EFA34B121E6DFFC4732819AA8258417B" author="Wittmer W." box="[878,1050,1010,1037]" pageId="14" pageNumber="15" pagination="109 - 169" refId="ref11421" refString="Wittmer W. 1995. Neue Cantharidae (Col.) aus dem indo-malaiischen und palaearktischen Faunengebiet mit Mutationen. Entomologica Basiliensia 18: 109 - 169." type="journal article" year="1995">Wittmer, 1995</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C32865681E6DFFC472E31E4085D24144" box="[677,912,1048,1074]" pageId="14" pageNumber="15" type="description">
<paragraph id="8B8D36E31E6DFFC472E31E4085D24144" blockId="14.[529,1058,1010,1074]" box="[677,912,1048,1074]" pageId="14" pageNumber="15">
<figureCitation id="13092A661E6DFFC472E31E4084AE4144" box="[677,748,1048,1074]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[357,1250,284,1869]" captionTargetId="figure@13.[224,1363,274,1886]" captionTargetPageId="13" captionText="Fig. 6. Aedeagus (A, D, G: ventral view; B, E, H: dorsal view; C, F, I: lateral view). AC. Fissocantharis lishanensis (Wittmer, 1982). DF. F. nantouensis (Wittmer, 1995). GI. F. nigriceps Y.Yang &amp; Okushima sp. nov. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3850504" httpUri="https://zenodo.org/record/3850504/files/figure.png" pageId="14" pageNumber="15">Figs 6</figureCitation>
DF, 8B, 11A
</paragraph>
</subSubSection>
<subSubSection id="C32865681E6DFFC470FB1E03855241CC" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8B8D36E31E6DFFC470FB1E0385264100" blockId="14.[189,868,1115,1142]" box="[189,868,1115,1142]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2211CD1E6DFFC470FB1E0385264100" box="[189,868,1115,1142]" pageId="14" pageNumber="15">
<treatmentCitation id="0A9310F21E6DFFC470FB1E0384B04100" author="Wittmer W." box="[189,754,1115,1142]" page="113" pageId="14" pageNumber="15" year="1995">
<taxonomicName id="4C324D601E6DFFC470FB1E0384B04100" authority="Wittmer, 1995: 113" authorityName="Wittmer" authorityPageNumber="113" authorityYear="1995" box="[189,754,1115,1142]" class="Insecta" family="Cantharidae" genus="Micropodabrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B946EAF11E6DFFC470FB1E0384444103" box="[189,518,1115,1141]" italics="true" pageId="14" pageNumber="15">Micropodabrus nantouensis</emphasis>
<bibRefCitation id="EFA34B121E6DFFC4724B1E0384B04100" author="Wittmer W." box="[525,754,1115,1142]" pageId="14" pageNumber="15" pagination="109 - 169" refId="ref11421" refString="Wittmer W. 1995. Neue Cantharidae (Col.) aus dem indo-malaiischen und palaearktischen Faunengebiet mit Mutationen. Entomologica Basiliensia 18: 109 - 169." type="journal article" year="1995">Wittmer, 1995: 113</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 3, 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1EC7855241CC" blockId="14.[189,784,1183,1210]" box="[189,784,1183,1210]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2211CD1E6DFFC470FB1EC7855241CC" box="[189,784,1183,1210]" pageId="14" pageNumber="15">
<taxonomicName id="4C324D601E6DFFC470FB1EC787BD41CF" baseAuthorityName="Wittmer" baseAuthorityYear="1995" box="[189,511,1183,1209]" class="Insecta" family="Cantharidae" genus="Fissocantharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B946EAF11E6DFFC470FB1EC787BD41CF" box="[189,511,1183,1209]" italics="true" pageId="14" pageNumber="15">Fissocantharis nantouensis</emphasis>
</taxonomicName>
<treatmentCitation id="0A9310F21E6DFFC4725D1EF8854E41CC" author="Yang Y. X. &amp; Okushima Y. &amp; Yang X. K." box="[539,780,1183,1210]" page="52" pageId="14" pageNumber="15" year="2012">
<bibRefCitation id="EFA34B121E6DFFC4725D1EF8854E41CC" author="Yang Y. X. &amp; Okushima Y. &amp; Yang X. K." box="[539,780,1183,1210]" pageId="14" pageNumber="15" pagination="46 - 53" refId="ref11525" refString="Yang Y. X., Okushima Y. &amp; Yang X. K. 2012. Synonym, new species and checklist of the genus Fissocantharis Pic from Taiwan (Coleoptera, Cantharidae). Zootaxa 3262: 46 - 53." type="journal article" year="2012">
Yang
<emphasis id="B946EAF11E6DFFC472191EF884D841CF" box="[607,666,1183,1209]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2012: 52
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32865681E6DFFC470FB1EBB8771429D" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8B8D36E31E6DFFC470FB1EBB87E8418B" blockId="14.[189,426,1251,1277]" box="[189,426,1251,1277]" pageId="14" pageNumber="15">
<heading id="D0C5818F1E6DFFC470FB1EBB87E8418B" bold="true" box="[189,426,1251,1277]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B946EAF11E6DFFC470FB1EBB87E8418B" bold="true" box="[189,426,1251,1277]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1F4C821E4025" blockId="14.[189,1399,1300,1364]" pageId="14" pageNumber="15">
<collectingCountry id="F32576731E6DFFC470FB1F4C87714058" box="[189,307,1300,1326]" name="Taiwan" pageId="14" pageNumber="15">TAIWAN</collectingCountry>
:
<collectingRegion id="49F6F8011E6DFFC471791F4C87A54058" box="[319,487,1300,1326]" country="Taiwan" name="Nantou" pageId="14" pageNumber="15">Nantou Hsien</collectingRegion>
:
<specimenCount id="9D34FD6A1E6DFFC471B51F4C84624058" box="[499,544,1300,1326]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
, Meixi,
<date id="FF8C10231E6DFFC472C01F4C85614058" box="[646,803,1300,1326]" pageId="14" pageNumber="15" value="1991-04-10">10 Apr. 1991</date>
, leg.
<collectionCode id="ED23AE261E6DFFC473221F4C85354058" box="[868,887,1300,1326]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="15">T</collectionCode>
. Kishimoto; 1 Ƌ, Mt. Howanshan,
<date id="FF8C10231E6DFFC4755C1F4C86B84025" pageId="14" pageNumber="15" value="1991-03-17">17 Mar. 1991</date>
, leg. C.-C. Lo; 1 Ƌ, Sungkang,
<date id="FF8C10231E6DFFC472281F61854B4025" box="[622,777,1337,1363]" pageId="14" pageNumber="15" value="1977-04-26">26 Apr. 1977</date>
, leg. S. Saito (all in
<collectionCode id="ED23AE261E6DFFC473B11F6182124025" box="[1015,1104,1337,1363]" pageId="14" pageNumber="15">KURA</collectionCode>
).
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1F25845340E1" blockId="14.[189,529,1405,1431]" box="[189,529,1405,1431]" pageId="14" pageNumber="15">
<heading id="D0C5818F1E6DFFC470FB1F25845340E1" bold="true" box="[189,529,1405,1431]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B946EAF11E6DFFC470FB1F25845340E1" bold="true" box="[189,529,1405,1431]" pageId="14" pageNumber="15">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1FF6849D434E" blockId="14.[189,1398,1453,1592]" pageId="14" pageNumber="15">
AEDEAGUS (
<figureCitation id="13092A661E6DFFC4710C1FF587CD40BE" box="[330,399,1453,1480]" captionStart="Fig" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[357,1250,284,1869]" captionTargetId="figure@13.[224,1363,274,1886]" captionTargetPageId="13" captionText="Fig. 6. Aedeagus (A, D, G: ventral view; B, E, H: dorsal view; C, F, I: lateral view). AC. Fissocantharis lishanensis (Wittmer, 1982). DF. F. nantouensis (Wittmer, 1995). GI. F. nigriceps Y.Yang &amp; Okushima sp. nov. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3850504" httpUri="https://zenodo.org/record/3850504/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
DF). Ventral process of each paramere wide, slightly narrowed apically, moderately hooked at apex; conjoint dorsal plate of parameres moderately developed, about half length of ventral processes and as wide as half of the distance between outer margins of ventral processes, distinctly narrowed apically, with straight apical margin.
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1C3A848043B0" blockId="14.[189,1398,1633,1735]" pageId="14" pageNumber="15">
ABDOMINAL STERNITE VIII Oŀ ŀEMALE (
<figureCitation id="13092A661E6DFFC472CE1C3984B7430A" box="[648,757,1633,1660]" captionStart="Fig" captionStartId="23.[189,232,1930,1956]" captionTargetBox="[195,1391,637,1875]" captionTargetId="figure@23.[189,1399,627,1888]" captionTargetPageId="23" captionText="Fig. 11. Abdominal sternite VIII of female, ventral view. A. Fissocantharis nantouensis (Wittmer, 1995). B. F. piluchiensis (Wittmer, 1995). C. F. pingtungensis (Wittmer, 1982). D. F. ssulingensis (Wittmer, 1995). E. F. taipeiana (Wittmer, 1982). F. F. uenoi (Wittmer, 1982). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3850522" httpUri="https://zenodo.org/record/3850522/files/figure.png" pageId="14" pageNumber="15">Fig. 11A</figureCitation>
). Distinctly protuberant in middle of posterior margin, the middle protuberance moderately and triangularly emarginate at apex, forming a pair of median lobes, which are widely triangular at apices.
</paragraph>
<paragraph id="8B8D36E31E6DFFC470FB1CA88771429D" blockId="14.[189,1399,1776,2027]" pageId="14" pageNumber="15">
<collectionCode id="ED23AE261E6DFFC470FB1CA887CB427C" box="[189,393,1776,1802]" pageId="14" pageNumber="15">INTERNAL ORGAN</collectionCode>
Oŀ REPRODUCTIVE SYSTEM (
<figureCitation id="13092A661E6DFFC4728D1CA88565427C" box="[715,807,1776,1802]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[242,1368,295,1839]" captionTargetId="figure@18.[213,1374,285,1888]" captionTargetPageId="18" captionText="Fig. 8. Female genitalia, lateral view. A. Fissocantharis lishanensis (Wittmer, 1982). B. F. nantouensis (Wittmer, 1995). C. F. piluchiensis (Wittmer, 1995). D. F. pingtungensis (Wittmer, 1982). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3850510" httpUri="https://zenodo.org/record/3850510/files/figure.png" pageId="14" pageNumber="15">Fig. 8B</figureCitation>
). Vagina stout, abruptly thinned and extended into a slightly long duct at ventroapical portion; diverticulum and spermathecal duct arising from the end of the duct of vagina; diverticulum moderately long, spiral and distinctly thinned apically; spermathecal duct distinctly shorter than diverticulum and thinner than basal portion of diverticulum; spermatheca composed of a spiral tube which is slightly longer than diverticulum, provided with a thin accessory gland which is slightly shorter than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
</paragraph>
</subSubSection>
<subSubSection id="C32865681E6CFFC570FB1B53876C44EA" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8B8D36E31E6CFFC570FB1B53876D4453" blockId="15.[189,303,267,293]" box="[189,303,267,293]" pageId="15" pageNumber="16">
<heading id="D0C5818F1E6CFFC570FB1B53876D4453" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B946EAF11E6CFFC570FB1B53876D4453" bold="true" box="[189,303,267,293]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36E31E6CFFC570FB1B63876C44EA" blockId="15.[189,1399,314,412]" pageId="15" pageNumber="16">
In the original publication (
<bibRefCitation id="EFA34B121E6CFFC572451B6284EF4423" author="Wittmer W." box="[515,685,314,341]" pageId="15" pageNumber="16" pagination="109 - 169" refId="ref11421" refString="Wittmer W. 1995. Neue Cantharidae (Col.) aus dem indo-malaiischen und palaearktischen Faunengebiet mit Mutationen. Entomologica Basiliensia 18: 109 - 169." type="journal article" year="1995">Wittmer 1995</bibRefCitation>
), this species was only illustrated with male antennae. Here the genitalia of both sexes and abdominal sternite VIII of female are illustrated and described for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C32865681E6CFFC570FB1B9C8759477B" box="[189,342,452,525]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8B8D36E31E6CFFC570FB1B9C871444A8" blockId="15.[189,342,452,478]" box="[189,342,452,478]" pageId="15" pageNumber="16">
<heading id="D0C5818F1E6CFFC570FB1B9C871444A8" bold="true" box="[189,342,452,478]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B946EAF11E6CFFC570FB1B9C871444A8" bold="true" box="[189,342,452,478]" pageId="15" pageNumber="16">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36E31E6CFFC570FB1BAB8759477B" blockId="15.[189,283,499,525]" box="[189,283,499,525]" pageId="15" pageNumber="16">
<collectingCountry id="F32576731E6CFFC570FB1BAB8755477B" box="[189,279,499,525]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
.
</paragraph>
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