151 lines
21 KiB
XML
151 lines
21 KiB
XML
<document id="29E3D0C11A220A37FB32802132DB3551" ID-DOI="10.5281/zenodo.190591" ID-GBIF-Dataset="d89826f0-deac-4609-b462-4080a4ad20ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="190591" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461115204573" checkinUser="plazi" docAuthor="Pinto-Da-Rocha, Ricardo & Hara, Marcos Ryotaro" docDate="2009" docId="039B87F0FF88EE79F2B6FC7F7B7FC623" docLanguage="en" docName="zt02241p046.pdf" docOrigin="Zootaxa 2241" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="minutus Roewer 1919" docType="treatment" docVersion="10" lastPageNumber="40" masterDocId="FFA2FF88FF8DEE7EF221FFE27A0CC750" masterDocTitle="New familial assignments for three species of Neotropical harvestmen based on cladistic analysis (Arachnida: Opiliones: Laniatores)" masterLastPageNumber="46" masterPageNumber="33" pageNumber="38" updateTime="1698595001921" updateUser="plazi">
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<mods:title id="9AE055066BEF3B7330D3BF673C8305EF">New familial assignments for three species of Neotropical harvestmen based on cladistic analysis (Arachnida: Opiliones: Laniatores)</mods:title>
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<mods:name id="1F932883482FAE917B7F19446323D0BC" type="personal">
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<mods:namePart id="2CED742CE33D17BFEAAD00940B984D7B">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:namePart id="E00157714E04FDF882585D91EEBCE8D3">Hara, Marcos Ryotaro</mods:namePart>
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<mods:typeOfResource id="06BB93405421B8C38CC59CEF119669A9">text</mods:typeOfResource>
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<mods:title id="C1EBA11C87BEB698E1435CC1E88A6419">Zootaxa</mods:title>
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<mods:part id="9256CE602858FD99E6C263E5674E879C">
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<mods:date id="703120D5D92B90B4F24CB8E1107F4E60">2009</mods:date>
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<mods:detail id="E0948A413F89AF1556D4843E64DEFBA2" type="volume">
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<mods:number id="D2C744922D27CE03518B9B160DB5AB5A">2241</mods:number>
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<mods:start id="BE768F0F238746F5400ABF5E82CF8727">33</mods:start>
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<mods:classification id="8FE0E7EEBA38F3FA8D8965D648A9AF24">journal article</mods:classification>
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<mods:identifier id="556D43E2AF5FE0DF13F80D543F2A3AEC" type="DOI">10.5281/zenodo.190591</mods:identifier>
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<mods:identifier id="2A2B815E8779CF4292ABCE08041F3CBF" type="GBIF-Dataset">d89826f0-deac-4609-b462-4080a4ad20ce</mods:identifier>
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<mods:identifier id="1870BD20D95AF983B34757AB45D9B6C0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3BA3BEAD2D20FC125C5D5D58F221BA11" type="Zenodo-Dep">190591</mods:identifier>
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<treatment id="039B87F0FF88EE79F2B6FC7F7B7FC623" ID-DOI="http://doi.org/10.5281/zenodo.6219188" ID-GBIF-Taxon="119624768" ID-Zenodo-Dep="6219188" LSID="urn:lsid:plazi:treatment:039B87F0FF88EE79F2B6FC7F7B7FC623" httpUri="http://treatment.plazi.org/id/039B87F0FF88EE79F2B6FC7F7B7FC623" lastPageId="7" lastPageNumber="40" pageId="5" pageNumber="38">
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<subSubSection id="C328656DFF88EE7BF2B6FC7F7B7DC48A" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph id="8B8D36E6FF88EE7BF2B6FC7F7941C4E7" blockId="5.[151,845,925,986]" box="[151,845,925,951]" pageId="5" pageNumber="38">
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<heading id="D0C5818AFF88EE7BF2B6FC7F7941C4E7" bold="true" box="[151,845,925,951]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<emphasis id="B946EAF4FF88EE7BF2B6FC7F7941C4E7" bold="true" box="[151,845,925,951]" pageId="5" pageNumber="38">
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<emphasis id="B946EAF4FF88EE7BF2B6FC7F7B05C4E7" bold="true" box="[151,265,925,951]" italics="true" pageId="5" pageNumber="38">Rivetinus</emphasis>
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<taxonomicName id="4C324D65FF88EE7BF331FC7F782FC4E7" authority="Roewer, 1919" authorityName="Roewer" authorityYear="1919" box="[272,547,925,951]" pageId="5" pageNumber="38" rank="species" species="minutus">
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<emphasis id="B946EAF4FF88EE7BF331FC7F7B7FC4E7" bold="true" box="[272,371,925,951]" italics="true" pageId="5" pageNumber="38">minutus</emphasis>
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<bibRefCitation id="EFA34B17FF88EE7BF35AFC7F782FC4E7" author="Roewer" box="[379,547,925,951]" pageId="5" pageNumber="46" refString="Roewer, C. F. (1919) Arachnida Opiliones. Mission du Service Geographique de l'armee pour la mesure d'un arc de meridien equatorial en Amerique du Sud, 10 (2): 121 - 141." type="journal article" year="1919">Roewer, 1919</bibRefCitation>
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</taxonomicName>
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, new family assignment
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D36E6FF88EE7BF2B6FC227B7DC48A" blockId="5.[151,845,925,986]" box="[151,369,960,986]" pageId="5" pageNumber="38">
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(
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<figureCitation id="13092A63FF88EE7BF2BEFC227AFDC48A" box="[159,241,960,986]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Figs. 2</figureCitation>
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and
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<figureCitation id="13092A63FF88EE7BF308FC227B35C48A" box="[297,313,960,986]" captionStart="FIGURE 4" captionStartId="9.[151,256,1819,1843]" captionTargetBox="[207,1393,339,1724]" captionTargetId="figure@9.[151,1436,296,1796]" captionTargetPageId="9" captionText="FIGURE 4. Distal part of penis: A – C: Globibunus rubrofemoratus Roewer, 1912, A, dorsal view; B, ventral view; C, right lateral view. D – E: Rivetinus minutus Roewer, 1919, D, dorsal view; E, right lateral view. F – G: Ramonus conifrons Roewer, 1956, F, dorsal view; G, right lateral view. A – C, at same scale; D and E at same scale; F and G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/190595/files/figure.png" pageId="5" pageNumber="38">4</figureCitation>
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D,E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C328656DFF88EE7BF2B6FBEF7BB8C3D4" pageId="5" pageNumber="38" type="reference_group">
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<paragraph id="8B8D36E6FF88EE7BF2B6FBEF7BB8C3D4" blockId="5.[151,1437,1037,1156]" pageId="5" pageNumber="38">
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<treatmentCitationGroup id="AB2211C8FF88EE7BF2B6FBEF7BB8C3D4" infered="true" pageId="5" pageNumber="38">
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<emphasis id="B946EAF4FF88EE7BF2B6FBEF7AF6C374" box="[151,250,1037,1060]" italics="true" pageId="5" pageNumber="38">Rivetinus</emphasis>
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<taxonomicName id="4C324D65FF88EE7BF321FBEF7816C374" authority="Roewer 1919:132" authorityName="Roewer" authorityPageNumber="132" authorityYear="1919" box="[256,538,1037,1060]" pageId="5" pageNumber="38" rank="species" species="minutus">
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<emphasis id="B946EAF4FF88EE7BF321FBEF7B59C374" box="[256,341,1037,1060]" italics="true" pageId="5" pageNumber="38">minutus</emphasis>
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<treatmentCitation id="0A9310F7FF88EE7BF37AFBEF7816C374" author="Roewer" box="[347,538,1037,1060]" page="132" pageId="5" pageNumber="38" year="1919">
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<bibRefCitation id="EFA34B17FF88EE7BF37AFBEF7BE1C374" author="Roewer" box="[347,493,1037,1060]" pageId="5" pageNumber="46" refString="Roewer, C. F. (1919) Arachnida Opiliones. Mission du Service Geographique de l'armee pour la mesure d'un arc de meridien equatorial en Amerique du Sud, 10 (2): 121 - 141." type="journal article" year="1919">Roewer 1919</bibRefCitation>
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:132
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</treatmentCitation>
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</taxonomicName>
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, pl. 13, Figs. 7 (dor hab), 7a–b (legI, IV), 1923:127 (rdesc), Fig. 133 (dor hab);
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<treatmentCitation id="0A9310F7FF88EE7BF744FBEF7B29C314" author="Kury" page="28" pageId="5" pageNumber="38" year="2003">
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<bibRefCitation id="EFA34B17FF88EE7BF744FBEF7B0CC314" author="Kury" pageId="5" pageNumber="46" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia vol. esp. monografico n o 1, 5 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
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: 28
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</treatmentCitation>
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(cat). (ma
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<typeStatus id="54898844FF88EE7BF3BBFBCF7BF5C314" box="[410,505,1069,1092]" pageId="5" pageNumber="38" type="paratype">paratype</typeStatus>
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;
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<collectingCountry id="F3257676FF88EE7BF022FBCF7852C314" box="[515,606,1069,1092]" name="Ecuador" pageId="5" pageNumber="38">Ecuador</collectingCountry>
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, [Carchí], El Pelado, collected at
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<quantity id="4CCA9B03FF88EE7BF1E2FBCF7E1BC314" box="[963,1047,1069,1092]" metricMagnitude="3" metricUnit="m" metricValue="4.15" pageId="5" pageNumber="38" unit="m" value="4150.0">4150 m</quantity>
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; without collector and date; SMF RI 311; examined). Note: Although not written in the label, according to the original description, the collector is Dr. Rivet, dated in
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<date id="FF8C1026FF88EE7BF34BFB8F7BBCC3D4" box="[362,432,1133,1156]" pageId="5" pageNumber="38" value="1903-01">i.1903</date>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328656DFF88EE79F2B6FB517B7FC623" lastPageId="7" lastPageNumber="40" pageId="5" pageNumber="38" type="diagnosis">
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<paragraph id="8B8D36E6FF88EE7BF2B6FB517B7CC212" blockId="5.[151,1437,1203,2003]" pageId="5" pageNumber="38">
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<emphasis id="B946EAF4FF88EE7BF2B6FB517B1CC39D" bold="true" box="[151,272,1203,1229]" pageId="5" pageNumber="38">Diagnosis</emphasis>
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: Zamorinae with unarmed ocularium, dorsal scutum mostly smooth, with mesotergal grooves I–IV backward, area I undivided, pedipalpal patella unarmed, coxae II–III without conspicuous armature, ventral plate of penis concave at the sides, with two distal roughly delimitated lobes, each one with acute and frontward corners.
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</paragraph>
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<paragraph id="8B8D36E6FF88EE7BF2E4FAAC7B2DC100" blockId="5.[151,1437,1203,2003]" pageId="5" pageNumber="38">
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<emphasis id="B946EAF4FF88EE7BF2E4FAAC7B78C238" bold="true" box="[197,372,1358,1384]" pageId="5" pageNumber="38">Redescription</emphasis>
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:
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<emphasis id="B946EAF4FF88EE7BF3A5FAAD7BCDC238" box="[388,449,1359,1384]" italics="true" pageId="5" pageNumber="38">Male</emphasis>
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(
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<typeStatus id="54898844FF88EE7BF3F0FAAC784CC238" box="[465,576,1358,1384]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
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): Dorsum (
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<figureCitation id="13092A63FF88EE7BF0E5FAAC7900C238" box="[708,780,1358,1384]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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A,C): Measurements: DSL 3.0; OMW 2.3; PL 1.4; PW 1.8; Leg I 5.2; II 7.5; III 5.7; IV 7.6. Dorsal scutum mostly smooth, with constriction near groove I and between grooves II–III, widening from area II to posterior margin; mesotergal area I slightly wider. Prosoma high behind ocularium. Ocularium, large, high, domed. Mesotergal area I entire; II–III with a pair of small tubercles; IV with a short row of four small tubercles; grooves I–IV backward. Posterior margin and free tergites with one row of larger tubercles (on posterior margin shorter, placed in the middle). Anal operculum tuberculate.
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</paragraph>
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<paragraph id="8B8D36E6FF88EE7BF2E7F9BF7BE5C1CE" blockId="5.[151,1437,1203,2003]" pageId="5" pageNumber="38">
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Venter (
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<figureCitation id="13092A63FF88EE7BF303F9BF7B65C127" box="[290,361,1629,1655]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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B): Coxae I–IV covered with tubercles, I with an anterior row of larger tubercles, two higher tubercles near coxapophysis.
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</paragraph>
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<paragraph id="8B8D36E6FF88EE7BF2E7F9487844C042" blockId="5.[151,1437,1203,2003]" pageId="5" pageNumber="38">
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Chelicera (
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<figureCitation id="13092A63FF88EE7BF366F9487B83C194" box="[327,399,1706,1732]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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A,C,D): Segment I with several small dorsal tubercles, dorso-prolaterally swollen, bulla well-marked; II swollen, with a frontal-median projection between base of fingers (
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<figureCitation id="13092A63FF88EE7BF687F9337EFEC1BB" box="[1190,1266,1745,1771]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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D). Fixed and movable finger with five small teeth.
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</paragraph>
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<paragraph id="8B8D36E6FF88EE7BF2E7F8FC7B67C0D6" blockId="5.[151,1437,1203,2003]" pageId="5" pageNumber="38">
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Pedipalpus (
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<figureCitation id="13092A63FF88EE7BF379F8FC7BACC068" box="[344,416,1822,1848]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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E,G): Coxa with large ventral tubercle. Trochanter with two ventral tubercles (median larger). Femur with five ventral tubercles (basal larger). Patella unarmed. Tibia mesal IIi, ectal iIi. Tarsus mesal Iii, ectal Iii.
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</paragraph>
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<paragraph id="8B8D36E6FF88EE78F2E7F87078A7C788" blockId="5.[151,1437,1203,2003]" lastBlockId="6.[151,1435,152,216]" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
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Legs (
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<figureCitation id="13092A63FF88EE7BF332F8707B51C0FC" box="[275,349,1938,1964]" captionStart="FIGURE 2" captionStartId="6.[151,255,1908,1932]" captionTargetBox="[234,1357,285,1871]" captionTargetId="figure@6.[202,1384,238,1910]" captionTargetPageId="6" captionText="FIGURE 2. Rivetinus minutus Roewer 1919. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II – III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur – tarsus, dorso-mesal view; H, right leg IV, prolateral view. A – C at same scale; E and G at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="5" pageNumber="38">Fig. 2</figureCitation>
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F,H): Small tuberculate. Coxae I–II each with two dorsal tubercles; III with one tubercle directed to another lateral of II; IV ending at area II, tuberculate. Trochanter I with three ventral larger tubercles; IV inflated. Femur IV slightly curved inward, with two ventral rows of tubercles increasing in size subapically. Tarsal formula: 6(3), 11 (4), 6, 6.
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</paragraph>
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<caption id="DF4D666EFF8BEE78F2B6F8967806C0A3" httpUri="https://zenodo.org/record/190593/files/figure.png" pageId="6" pageNumber="39" targetBox="[234,1357,285,1871]" targetPageId="6">
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<paragraph id="8B8D36E6FF8BEE78F2B6F8967806C0A3" blockId="6.[151,1436,1908,2035]" pageId="6" pageNumber="39">
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<emphasis id="B946EAF4FF8BEE78F2B6F8967B1FC0DC" bold="true" box="[151,275,1908,1932]" pageId="6" pageNumber="39">FIGURE 2</emphasis>
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.
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<emphasis id="B946EAF4FF8BEE78F303F8977B8AC0DC" box="[290,390,1909,1932]" italics="true" pageId="6" pageNumber="39">Rivetinus</emphasis>
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<taxonomicName id="4C324D65FF8BEE78F3ACF8977871C0DB" authority="Roewer 1919" authorityName="Roewer" authorityYear="1919" box="[397,637,1908,1932]" pageId="6" pageNumber="39" rank="species" species="minutus">
|
||
<emphasis id="B946EAF4FF8BEE78F3ACF8977BEEC0DC" box="[397,482,1909,1932]" italics="true" pageId="6" pageNumber="39">minutus</emphasis>
|
||
Roewer 1919
|
||
</taxonomicName>
|
||
. Male (paratype): A, habitus, dorsal view; B, idem ventral view; C, idem, left lateral view; D, left cheliceral segments II–III, frontal view; E, right pedipalp, ectal view; F, right femur IV, ventral view; G, right pedipalp, distal femur–tarsus, dorso-mesal view; H, right leg IV, prolateral view. A–C at same scale; E and G at same scale. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36E6FF8AEE79F2E7FF7A7B99C676" blockId="7.[151,1437,152,371]" pageId="7" pageNumber="40">
|
||
Penis (
|
||
<figureCitation id="13092A63FF8AEE79F33CFF7A7B67C7E2" box="[285,363,152,178]" captionStart="FIGURE 4" captionStartId="9.[151,256,1819,1843]" captionTargetBox="[207,1393,339,1724]" captionTargetId="figure@9.[151,1436,296,1796]" captionTargetPageId="9" captionText="FIGURE 4. Distal part of penis: A – C: Globibunus rubrofemoratus Roewer, 1912, A, dorsal view; B, ventral view; C, right lateral view. D – E: Rivetinus minutus Roewer, 1919, D, dorsal view; E, right lateral view. F – G: Ramonus conifrons Roewer, 1956, F, dorsal view; G, right lateral view. A – C, at same scale; D and E at same scale; F and G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/190595/files/figure.png" pageId="7" pageNumber="40">Fig. 4</figureCitation>
|
||
D,E): Truncus with three pairs of trifid long basal setae, two pairs of small, curved, unirramous setae on sides of stylus base and two unirramous pairs on ventral plate base. Ventral plate concave at the sides, with two distal roughly delimitated lobes, each one with acute and frontward corners. Stylus long, unique, without crest.
|
||
</paragraph>
|
||
<paragraph id="8B8D36E6FF8AEE79F2E7FED07B7FC623" blockId="7.[151,1437,152,371]" pageId="7" pageNumber="40">
|
||
Coloration: Specimen discolored. Dorsal scute, chelicera and pedipalp dark yellow, legs light yellow.
|
||
<emphasis id="B946EAF4FF8AEE79F2B6FEBB7AE3C622" box="[151,239,345,370]" italics="true" pageId="7" pageNumber="40">Female</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |