434 lines
56 KiB
XML
434 lines
56 KiB
XML
<document id="0822BBBA2CCA2B7E69B85561B71219D8" ID-CLB-Dataset="21210" ID-DOI="10.1080/00222933.2016.1193649" ID-GBIF-Dataset="2a2f38ca-c6d9-4512-b977-e157be51e4ca" ID-ISSN="1464-5262" ID-Zenodo-Dep="3994511" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598016783362" checkinUser="carolina" docAuthor="Sun, Jing, Jiang, Jibao, Hu, Feng & Qiu, Jiangping" docDate="2016" docId="039B87C9CA6CFFA228E4FD7EFE9D39E6" docLanguage="en" docName="JNATHIST.50.39-40.2499-2513.pdf" docOrigin="Journal of Natural History 50" docSource="http://dx.doi.org/10.1080/00222933.2016.1193649" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Amynthas leshanensis Sun and Qiu 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="2503" masterDocId="FFA2FFB1CA6DFFA62963FFB7FFF43A25" masterDocTitle="Four new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Mount Emei, Sichuan Province, China" masterLastPageNumber="2513" masterPageNumber="2499" pageNumber="2500" updateTime="1698845083862" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="39493AE9BF08ADE97694C9CF55AB6193">Four new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Mount Emei, Sichuan Province, China</mods:title>
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<mods:name id="33359A5701947A6FFC9E87837F9292C8" type="personal">
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<mods:role id="D0853BE8FB2213B8D6B3532D683E2E0C">
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<mods:roleTerm id="68B71E5B71F608523544DE3A8D44DD93">Author</mods:roleTerm>
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<mods:namePart id="E884367F25F0F19240035DB5DC4ECCED">Sun, Jing</mods:namePart>
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<mods:affiliation id="B6F916B9BFDDA581222FEBEF09477269">College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China; & School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China;</mods:affiliation>
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<mods:name id="3094EFC24F4A36991A309451A3051663" type="personal">
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<mods:roleTerm id="089B6E2C92992A73CBAC6C912CEF7308">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4FBA8A6C61B22BCF3842AE95DE18A2AC">Jiang, Jibao</mods:namePart>
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<mods:affiliation id="DDC2FEE3D4A7006992D447BCC5C201B4">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China; & Key Laboratory of Urban Agriculture (South), Ministry of Agriculture, Shanghai, China</mods:affiliation>
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</mods:name>
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<mods:name id="034F037F3669536E8C91ADCF4816DE38" type="personal">
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<mods:roleTerm id="B7CF5A6FF6BA504B1CE04EC4B1C376F1">Author</mods:roleTerm>
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<mods:namePart id="51568F0840AC8932DB909B2AD9207118">Hu, Feng</mods:namePart>
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<mods:affiliation id="A339A2B2C80F67184BF238C69EB41B8F">College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China;</mods:affiliation>
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</mods:role>
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<mods:namePart id="21F00B3749E56EF23D569A9CB9C96E77">Qiu, Jiangping</mods:namePart>
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<mods:affiliation id="A7C81C48AD142DC13E404724EB1340CE">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China; & Key Laboratory of Urban Agriculture (South), Ministry of Agriculture, Shanghai, China</mods:affiliation>
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<mods:title id="68760CEC10C234F19124B4B4C4C73290">Journal of Natural History</mods:title>
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<mods:part id="81D0C6A1D6FB2D8B08890D3E9C665C0C">
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<mods:date id="6EDBC9CAE092BAE3534BE686C2CEB347">2016</mods:date>
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<mods:number id="B9BD360EBB31F504287764BC617D8AE4">2016-07-31</mods:number>
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<mods:number id="AD9B41BB8363FDA7D8EFAEFAF0C35630">50</mods:number>
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<treatment id="039B87C9CA6CFFA228E4FD7EFE9D39E6" ID-DOI="http://doi.org/10.5281/zenodo.4328446" ID-GBIF-Taxon="167288267" ID-Zenodo-Dep="4328446" LSID="urn:lsid:plazi:treatment:039B87C9CA6CFFA228E4FD7EFE9D39E6" httpUri="http://treatment.plazi.org/id/039B87C9CA6CFFA228E4FD7EFE9D39E6" lastPageId="4" lastPageNumber="2503" pageId="1" pageNumber="2500">
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<subSubSection id="C3286554CA6CFFA728E4FD7EFC6938C7" box="[391,925,713,738]" pageId="1" pageNumber="2500" type="nomenclature">
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<paragraph id="8B8D36DFCA6CFFA728E4FD7EFC6938C7" blockId="1.[391,925,713,773]" box="[391,925,713,738]" pageId="1" pageNumber="2500">
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<heading id="D0C581B3CA6CFFA728E4FD7EFC6938C7" box="[391,925,713,738]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2500" reason="2">
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<taxonomicName id="4C324D5CCA6CFFA728E4FD7EFCC238C7" authority="Sun and Qiu" authorityName="Sun and Qiu" authorityYear="2016" box="[391,822,713,738]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="2500" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
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<emphasis id="B946EACDCA6CFFA728E4FD7EFD6C38C7" bold="true" box="[391,664,713,738]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">Amynthas leshanensis</emphasis>
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Sun and Qiu
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</taxonomicName>
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<taxonomicNameLabel id="A27557B6CA6CFFA72A23FD7EFC6938C7" box="[832,925,713,738]" pageId="1" pageNumber="2500" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286554CA6CFFA72B58FD5BFD1C3920" box="[571,744,748,773]" pageId="1" pageNumber="2500" type="description">
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<paragraph id="8B8D36DFCA6CFFA72B58FD5BFD1C3920" blockId="1.[391,925,713,773]" box="[571,744,748,773]" pageId="1" pageNumber="2500">
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(
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<figureCitation id="13092A5ACA6CFFA72B20FD5BFD543920" box="[579,672,748,773]" captionStart="Figure 1" captionStartId="2.[160,225,1146,1169]" captionTargetBox="[163,1153,142,1115]" captionTargetId="figure@2.[163,1153,142,1115]" captionTargetPageId="2" captionText="Figure 1. (a) Amynthas leshanensis Sun and Qiu sp. nov., holotype, spermathecal pores; (b) A. leshanensis sp. nov., male pore region; (c) A. leshanensis sp. nov., intestinal caecum; (d) A. leshanensis sp. nov., spermathecae. (e) Amynthas gilvus Sun and Qiu sp. nov., holotype, male pore region; (f) A. gilvus sp. nov., intestinal caecum; (g) A. gilvus sp. nov., spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="1" pageNumber="2500">Figure 1</figureCitation>
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(a–d))
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286554CA6CFFA729C3FC8BFCF63E27" pageId="1" pageNumber="2500" type="materials_examined">
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<paragraph id="8B8D36DFCA6CFFA729C3FC8BFEB13970" blockId="1.[160,1156,828,1026]" box="[160,325,828,853]" pageId="1" pageNumber="2500">
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<heading id="D0C581B3CA6CFFA729C3FC8BFEB13970" box="[160,325,828,853]" fontSize="11" level="3" pageId="1" pageNumber="2500" reason="5">
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<emphasis id="B946EACDCA6CFFA729C3FC8BFEB13970" bold="true" box="[160,325,828,853]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D36DFCA6CFFA729C3FCE9FCF63E27" blockId="1.[160,1156,828,1026]" pageId="1" pageNumber="2500">
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<materialsCitation id="3B5A3C82CA6CFFA729C3FCE9FCA23998" ID-GBIF-Occurrence="2846790306" collectingDate="2011-07-27" collectorName="J. Sun & J. B. Jiang & X. D. Lei & H. W. Feng" country="China" elevation="1300" latitude="29.591389" location="Mt. Emei" longLatPrecision="20" longitude="103.28667" municipality="One" pageId="1" pageNumber="2500" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
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<emphasis id="B946EACDCA6CFFA729C3FCE9FEE53952" bold="true" box="[160,273,862,887]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">
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<typeStatus id="5489887DCA6CFFA729C3FCE9FEFF3952" box="[160,267,862,887]" pageId="1" pageNumber="2500" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingMunicipality id="6BE9ACA5CA6CFFA72842FCE8FEA43952" box="[289,336,863,887]" pageId="1" pageNumber="2500">One</collectingMunicipality>
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clitellate (C-SC201102-02A) specimen:
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<collectingCountry id="F325764FCA6CFFA72A65FCE8FCBD3952" box="[774,841,863,887]" name="China" pageId="1" pageNumber="2500">China</collectingCountry>
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,
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<collectingRegion id="49F6F83DCA6CFFA72A30FCE8FBE23952" box="[851,1046,863,887]" country="China" name="Sichuan" pageId="1" pageNumber="2500">Sichuan Province</collectingRegion>
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||
,
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<location id="8EED6004CA6CFFA72D42FCE8FB773952" LSID="urn:lsid:plazi:treatment:039B87C9CA6CFFA228E4FD7EFE9D39E6:8EED6004CA6CFFA72D42FCE8FB773952" box="[1057,1155,863,887]" country="China" latitude="29.591389" longLatPrecision="20" longitude="103.28667" municipality="One" name="Mt. Emei" pageId="1" pageNumber="2500" stateProvince="Sichuan">Mt. Emei</location>
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(
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<geoCoordinate id="EE065018CA6CFFA729C5FC35FED139BF" box="[166,293,898,922]" degrees="29" direction="north" minutes="35" orientation="latitude" pageId="1" pageNumber="2500" precision="15" seconds="29" value="29.591389">29°35′29″N</geoCoordinate>
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,
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<geoCoordinate id="EE065018CA6CFFA72852FC35FE4D39BF" box="[305,441,898,922]" degrees="103" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="2500" precision="15" seconds="12" value="103.28667">103°17′12″E</geoCoordinate>
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),
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<quantity id="4CCA9B3ACA6CFFA728A8FC35FDEB39BF" box="[459,543,898,922]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="2500" unit="m" value="1300.0">
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<elevation id="001FD1ECCA6CFFA728A8FC35FDEB39BF" box="[459,543,898,922]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="2500" unit="m" value="1300.0">1300 m</elevation>
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</quantity>
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elevation, woods and bamboo, in dark cinnamon soil,
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<date id="FF8C101FCA6CFFA729C3FC12FEC13998" box="[160,309,933,957]" pageId="1" pageNumber="2500" value="2011-07-27">
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<collectingDate id="EFC8E9F7CA6CFFA729C3FC12FEC13998" box="[160,309,933,957]" pageId="1" pageNumber="2500" value="2011-07-27">27 July 2011</collectingDate>
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</date>
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,
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<collectorName id="26C75309CA6CFFA72820FC12FE7E3998" box="[323,394,933,957]" pageId="1" pageNumber="2500">J. Sun</collectorName>
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,
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<collectorName id="26C75309CA6CFFA728FBFC12FDF73998" box="[408,515,933,957]" pageId="1" pageNumber="2500">J.B. Jiang</collectorName>
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,
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<collectorName id="26C75309CA6CFFA72B72FC12FD9C3998" box="[529,616,933,957]" pageId="1" pageNumber="2500">X.D. Lei</collectorName>
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and
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<collectorName id="26C75309CA6CFFA72BCBFC12FCEA3998" box="[680,798,933,957]" pageId="1" pageNumber="2500">H.W. Feng</collectorName>
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coll.
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</materialsCitation>
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<typeStatus id="5489887DCA6CFFA72A03FC12FC243998" box="[864,976,933,957]" pageId="1" pageNumber="2500" type="paratype">Paratypes</typeStatus>
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: A total of two specimens, as follows: One clitellate (C-SC201102-02B): Same data as for holotype.
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<materialsCitation id="3B5A3C82CA6CFFA72D36FC70FD0A3E27" ID-GBIF-Occurrence="2846790317" collectingDate="2011-07-27" collectorName="J. Sun & J. B. Jiang & X. D. Lei & H. W. Feng" country="China" elevation="1300" latitude="29.591389" location="Mt. Emei" longLatPrecision="20" longitude="103.28667" municipality="One" pageId="1" pageNumber="2500" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">One clitellate (C-SC201102-14): Same data as for holotype</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286554CA6CFFA729C3FB86FC183E48" box="[160,1004,1073,1133]" pageId="1" pageNumber="2500" type="etymology">
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||
<paragraph id="8B8D36DFCA6CFFA729C3FB86FEE93E6F" blockId="1.[160,1004,1073,1133]" box="[160,285,1073,1098]" pageId="1" pageNumber="2500">
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||
<heading id="D0C581B3CA6CFFA729C3FB86FEE93E6F" box="[160,285,1073,1098]" fontSize="11" level="3" pageId="1" pageNumber="2500" reason="5">
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<emphasis id="B946EACDCA6CFFA729C3FB86FEE93E6F" bold="true" box="[160,285,1073,1098]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B8D36DFCA6CFFA729C3FBE2FC183E48" blockId="1.[160,1004,1073,1133]" box="[160,1004,1108,1133]" pageId="1" pageNumber="2500">
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This species was named after the collection location – Leshan City,
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<collectingCountry id="F325764FCA6CFFA72AC6FBE2FC1C3E48" box="[933,1000,1109,1133]" name="China" pageId="1" pageNumber="2500">China</collectingCountry>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3286554CA6CFFA729C3FB2BFC4E3F47" pageId="1" pageNumber="2500" type="diagnosis">
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||
<paragraph id="8B8D36DFCA6CFFA729C3FB2BFEE03E90" blockId="1.[160,1155,1180,1378]" box="[160,276,1180,1205]" pageId="1" pageNumber="2500">
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||
<heading id="D0C581B3CA6CFFA729C3FB2BFEE03E90" box="[160,276,1180,1205]" fontSize="11" level="3" pageId="1" pageNumber="2500" reason="5">
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||
<emphasis id="B946EACDCA6CFFA729C3FB2BFEE03E90" bold="true" box="[160,276,1180,1205]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">Diagnosis</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B8D36DFCA6CFFA729C3FB08FC4E3F47" blockId="1.[160,1155,1180,1378]" pageId="1" pageNumber="2500">Three pairs of spermathecal pores in 6/7–8/9, 0.4 of circumference ventrally apart; each male pore on an elliptical glandular flat-top pad, surrounded by a large irregular shaped pad which extends from 17/18–½ XIX; intestinal caeca complex with three long and broad finger-shaped sacs; spermathecal diverticulum shorter than main spermathecal axis by 0.42, distal 0.5 dilated into elongated ovoid seminal chamber.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3286554CA6CFFA529C3FA26FDE23824" lastPageId="3" lastPageNumber="2502" pageId="1" pageNumber="2500" type="description">
|
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<paragraph id="8B8D36DFCA6CFFA729C3FA26FED23F8F" blockId="1.[160,1156,1425,1727]" box="[160,294,1425,1450]" pageId="1" pageNumber="2500">
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||
<heading id="D0C581B3CA6CFFA729C3FA26FED23F8F" box="[160,294,1425,1450]" fontSize="11" level="3" pageId="1" pageNumber="2500" reason="5">
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<emphasis id="B946EACDCA6CFFA729C3FA26FED23F8F" bold="true" box="[160,294,1425,1450]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">Description</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B8D36DFCA6CFFA429C3FA03FED53C1C" blockId="1.[160,1156,1425,1727]" lastBlockId="2.[160,1156,1326,1593]" lastPageId="2" lastPageNumber="2501" pageId="1" pageNumber="2500">
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<emphasis id="B946EACDCA6CFFA729C3FA03FE7A3FE8" bold="true" box="[160,398,1460,1485]" inLineHeading="true" italics="true" pageId="1" pageNumber="2500" reason="3">External characters.</emphasis>
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||
Preserved specimens dark grey on dorsum, lacking pigment on ventrum. Mid-dorsal line pigmented. Dimensions
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||
<quantity id="4CCA9B3ACA6CFFA72BB2FA60FCBD3FCA" box="[721,841,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" metricValueMax="9.3" metricValueMin="8.8" pageId="1" pageNumber="2500" unit="mm" value="90.5" valueMax="93.0" valueMin="88.0">88–93 mm</quantity>
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||
by 3.0–
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|
||
at clitellum, segments number 95–120. Body cylindrical in cross section, and gradually tapered towards head and tail. Secondary annulations absent. Prostomium ½ epilobous. First dorsal pore in 11/12. Clitellum annular in XIV–XVI, pale, smooth, swollen, no visible setae externally, and dorsal pore usually absent on clitellum. Setae uniformly distributed, dorsal breaks more distinct than ventral, 22–24 at III, 30–35 at V, 28–36 at VIII, 36–42 at XX, 38–44 at XXV; 10 between male pores; 11–14/VII, 11–14/VIII between spermathecal pores, setal formula AA = 1–1.2AB, ZZ = 2–2.2ZY. Spermathecal pores three pairs in 6/7–8/9, intersegmental, about 0.4 of circumference ventrally apart, with tumid epidermis around each spermathecal pore (
|
||
<figureCitation id="13092A5ACA6FFFA42B8BFAC3FCB63FA9" box="[744,834,1396,1420]" captionStart="Figure 1" captionStartId="2.[160,225,1146,1169]" captionTargetBox="[163,1153,142,1115]" captionTargetId="figure@2.[163,1153,142,1115]" captionTargetPageId="2" captionText="Figure 1. (a) Amynthas leshanensis Sun and Qiu sp. nov., holotype, spermathecal pores; (b) A. leshanensis sp. nov., male pore region; (c) A. leshanensis sp. nov., intestinal caecum; (d) A. leshanensis sp. nov., spermathecae. (e) Amynthas gilvus Sun and Qiu sp. nov., holotype, male pore region; (f) A. gilvus sp. nov., intestinal caecum; (g) A. gilvus sp. nov., spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="2" pageNumber="2501">Figure 1</figureCitation>
|
||
(a)). Genital papillae absent around spermathecal region. Male pores paired in XVIII, about 0.33 of circumference ventrally apart, each on a small elliptical glandular flat-top pad, swollen, and surrounded by a larger irregular-shaped pad which extends from 17/18–½ XIX. No genital papillae in this region (
|
||
<figureCitation id="13092A5ACA6FFFA42848FA48FE703C32" box="[299,388,1535,1559]" captionStart="Figure 1" captionStartId="2.[160,225,1146,1169]" captionTargetBox="[163,1153,142,1115]" captionTargetId="figure@2.[163,1153,142,1115]" captionTargetPageId="2" captionText="Figure 1. (a) Amynthas leshanensis Sun and Qiu sp. nov., holotype, spermathecal pores; (b) A. leshanensis sp. nov., male pore region; (c) A. leshanensis sp. nov., intestinal caecum; (d) A. leshanensis sp. nov., spermathecae. (e) Amynthas gilvus Sun and Qiu sp. nov., holotype, male pore region; (f) A. gilvus sp. nov., intestinal caecum; (g) A. gilvus sp. nov., spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="2" pageNumber="2501">Figure 1</figureCitation>
|
||
(b)). Female pore single, med-ventral in XIV, elliptical, situated in a depression.
|
||
</paragraph>
|
||
<caption id="DF4D6657CA6FFFA429C3FBCDFCAB3ECC" ID-DOI="http://doi.org/10.5281/zenodo.3994513" ID-Zenodo-Dep="3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="2" pageNumber="2501" startId="2.[160,225,1146,1169]" targetBox="[163,1153,142,1115]" targetPageId="2">
|
||
<paragraph id="8B8D36DFCA6FFFA429C3FBCDFCAB3ECC" blockId="2.[160,1156,1146,1257]" pageId="2" pageNumber="2501">
|
||
<emphasis id="B946EACDCA6FFFA429C3FBCDFEF43EB4" bold="true" box="[160,256,1146,1169]" pageId="2" pageNumber="2501">Figure 1.</emphasis>
|
||
(a)
|
||
<taxonomicName id="4C324D5CCA6FFFA4284DFBCDFD6C3EB4" authority="Sun and Qiu" authorityName="Sun and Qiu" authorityYear="2016" box="[302,664,1146,1169]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA4284DFBCDFDF23EB4" box="[302,518,1146,1169]" italics="true" pageId="2" pageNumber="2501">Amynthas leshanensis</emphasis>
|
||
Sun and Qiu
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA42BC7FBCDFD0E3EB4" box="[676,762,1146,1169]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, spermathecal pores; (b)
|
||
<taxonomicName id="4C324D5CCA6FFFA42D12FBCDFEF83E8B" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA42D12FBCDFEF83E8B" italics="true" pageId="2" pageNumber="2501">A. leshanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA42870FB20FE963E8B" box="[275,354,1175,1198]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male pore region; (c)
|
||
<taxonomicName id="4C324D5CCA6FFFA42B23FB20FD313E8B" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" box="[576,709,1175,1198]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA42B23FB20FD313E8B" box="[576,709,1175,1198]" italics="true" pageId="2" pageNumber="2501">A. leshanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA42BAFFB20FCEF3E8B" box="[716,795,1175,1198]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, intestinal caecum; (d)
|
||
<taxonomicName id="4C324D5CCA6FFFA42A9DFB20FB773E8B" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" box="[1022,1155,1175,1198]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA42A9DFB20FB773E8B" box="[1022,1155,1175,1198]" italics="true" pageId="2" pageNumber="2501">A. leshanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA429C3FB02FF053EE9" box="[160,241,1205,1228]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, spermathecae. (e)
|
||
<taxonomicName id="4C324D5CCA6FFFA428D0FB02FD213EE9" authority="Sun and Qiu" authorityName="Sun and Qiu" authorityYear="2016" box="[435,725,1205,1228]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="gilvus" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA428D0FB02FDA53EE9" box="[435,593,1205,1228]" italics="true" pageId="2" pageNumber="2501">Amynthas gilvus</emphasis>
|
||
Sun and Qiu
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA42BBFFB02FCD93EE9" box="[732,813,1205,1228]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, male pore region; (f)
|
||
<taxonomicName id="4C324D5CCA6FFFA42D12FB02FF233ECC" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="gilvus" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA42D12FB02FF233ECC" italics="true" pageId="2" pageNumber="2501">A. gilvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA429BCFB65FEC43ECC" box="[223,304,1234,1257]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, intestinal caecum; (g)
|
||
<taxonomicName id="4C324D5CCA6FFFA42B7AFB65FD9F3ECC" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" box="[537,619,1234,1257]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="2501" phylum="Annelida" rank="species" species="gilvus" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6FFFA42B7AFB65FD9F3ECC" box="[537,619,1234,1257]" italics="true" pageId="2" pageNumber="2501">A. gilvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6FFFA42B10FB65FD313ECC" box="[627,709,1234,1257]" pageId="2" pageNumber="2501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, spermathecae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36DFCA6FFFA429C3F9CCFB773C92" blockId="2.[160,1155,1659,1719]" pageId="2" pageNumber="2501">
|
||
<emphasis id="B946EACDCA6FFFA429C3F9CCFE643CB1" bold="true" box="[160,400,1659,1684]" inLineHeading="true" italics="true" pageId="2" pageNumber="2501" reason="3">Internal characters.</emphasis>
|
||
Septa 5/6–7/8 and 10/11–12/13 slightly thickened, 8/9–9/10 absent. Dorsal blood vessel single, continuous onto pharynx; esophageal hearts four
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA6EFFA529C3FF39FDE23824" blockId="3.[160,1156,142,513]" pageId="3" pageNumber="2502">
|
||
pairs in X–XIII. Gizzard barrel-like, in VIII–X; intestine beginning to swell in XVI; intestinal caeca complex, with three long and broad finger-shaped sacs, originating in XXVII and extending forward to XXIV (
|
||
<figureCitation id="13092A5ACA6EFFA528BFFF63FDC73AC9" box="[476,563,212,236]" captionStart="Figure 1" captionStartId="2.[160,225,1146,1169]" captionTargetBox="[163,1153,142,1115]" captionTargetId="figure@2.[163,1153,142,1115]" captionTargetPageId="2" captionText="Figure 1. (a) Amynthas leshanensis Sun and Qiu sp. nov., holotype, spermathecal pores; (b) A. leshanensis sp. nov., male pore region; (c) A. leshanensis sp. nov., intestinal caecum; (d) A. leshanensis sp. nov., spermathecae. (e) Amynthas gilvus Sun and Qiu sp. nov., holotype, male pore region; (f) A. gilvus sp. nov., intestinal caecum; (g) A. gilvus sp. nov., spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="3" pageNumber="2502">Figure 1</figureCitation>
|
||
(c)). Male sexual system is holandric. Testis sacs two pairs, small, ventral in X and XI, connected with membrane on ventrum. Seminal vesicles paired in XI and XII, developed, broadly connected with each other on ventrum; prostate glands developed, coarsely lobate, in ½ XVI–XXI, ducts U-curved, distal end stouter. No visible accessory glands around base of each duct. Female organs: Ovaries in XIII. Spermathecae paired in VII–IX, about
|
||
<quantity id="4CCA9B3ACA6EFFA52B03FE36FD4D3BBC" box="[608,697,385,409]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="2502" unit="mm" value="2.6">2.6 mm</quantity>
|
||
long; ampulla heart-shaped, with very short duct, about
|
||
<quantity id="4CCA9B3ACA6EFFA52816FE13FE243B99" box="[373,464,420,444]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="2502" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; diverticulum shorter than main spermathecal axis by 0.42, slender, distal 0.5 dilated into elongated ovoid seminal chamber; no nephridia on spermathecal ducts (
|
||
<figureCitation id="13092A5ACA6EFFA528EEFE5EFE123824" box="[397,486,489,513]" captionStart="Figure 1" captionStartId="2.[160,225,1146,1169]" captionTargetBox="[163,1153,142,1115]" captionTargetId="figure@2.[163,1153,142,1115]" captionTargetPageId="2" captionText="Figure 1. (a) Amynthas leshanensis Sun and Qiu sp. nov., holotype, spermathecal pores; (b) A. leshanensis sp. nov., male pore region; (c) A. leshanensis sp. nov., intestinal caecum; (d) A. leshanensis sp. nov., spermathecae. (e) Amynthas gilvus Sun and Qiu sp. nov., holotype, male pore region; (f) A. gilvus sp. nov., intestinal caecum; (g) A. gilvus sp. nov., spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3994513" httpUri="https://zenodo.org/record/3994513/files/figure.png" pageId="3" pageNumber="2502">Figure 1</figureCitation>
|
||
(d)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3286554CA6EFFA229C3FD99FE9D39E6" lastPageId="4" lastPageNumber="2503" pageId="3" pageNumber="2502" type="discussion">
|
||
<paragraph id="8B8D36DFCA6EFFA529C3FD99FEF23862" blockId="3.[160,1156,558,1727]" box="[160,262,558,583]" pageId="3" pageNumber="2502">
|
||
<heading id="D0C581B3CA6EFFA529C3FD99FEF23862" box="[160,262,558,583]" fontSize="11" level="3" pageId="3" pageNumber="2502" reason="5">
|
||
<emphasis id="B946EACDCA6EFFA529C3FD99FEF23862" bold="true" box="[160,262,558,583]" inLineHeading="true" italics="true" pageId="3" pageNumber="2502" reason="3">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA6EFFA529C3FDE6FDAD391F" blockId="3.[160,1156,558,1727]" pageId="3" pageNumber="2502">
|
||
<taxonomicName id="4C324D5CCA6EFFA529C3FDE6FE67384F" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" box="[160,403,593,618]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA6EFFA529C3FDE6FE67384F" box="[160,403,593,618]" italics="true" pageId="3" pageNumber="2502">Amynthas leshanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557B6CA6EFFA528FEFDE5FE0C384F" box="[413,504,594,618]" pageId="3" pageNumber="2502" rank="species">sp. nov.</taxonomicNameLabel>
|
||
keys to the
|
||
<taxonomicName id="4C324D5CCA6EFFA52BECFDE6FCAD384F" baseAuthorityName="Horst" baseAuthorityYear="1883" box="[655,857,593,618]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="sieboldi">
|
||
<emphasis id="B946EACDCA6EFFA52BECFDE6FCAD384F" box="[655,857,593,618]" italics="true" pageId="3" pageNumber="2502">Amynthas sieboldi</emphasis>
|
||
</taxonomicName>
|
||
group according to
|
||
<bibRefCitation id="EFA34B2ECA6EFFA52D2CFDE5FE9038A8" author="Sims RW & Easton EG" pageId="3" pageNumber="2502" pagination="169 - 268" refId="ref8133" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton, 1972</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFA34B2ECA6EFFA52811FDC2FD8438A8" author="Sims RW & Easton EG" box="[370,624,629,653]" pageId="3" pageNumber="2502" pagination="169 - 268" refId="ref8133" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton 1972</bibRefCitation>
|
||
). Members of this group can be distinguished from those of the other species of the genus
|
||
<taxonomicName id="4C324D5CCA6EFFA52BCCFD21FCE8388A" box="[687,796,662,687]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="genus">
|
||
<emphasis id="B946EACDCA6EFFA52BCCFD21FCE8388A" box="[687,796,662,687]" italics="true" pageId="3" pageNumber="2502">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
by three pairs of spermathecal pores intersegmental in 6/7–8/9 and a holandric male sexual system. In many principal respects the present new species is relatively similar to
|
||
<taxonomicName id="4C324D5CCA6EFFA52A7EFD6BFF2B3932" authority="(Chen, 1930)" baseAuthorityName="Chen" baseAuthorityYear="1930" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA52A7EFD6BFBC638D0" box="[797,1074,732,757]" italics="true" pageId="3" pageNumber="2502">Amynthas obscuritoporus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA34B2ECA6EFFA52D21FD6BFF233932" author="Chen Y" pageId="3" pageNumber="2502" pagination="11 - 37" refId="ref7817" refString="Chen Y. 1930. On some new earthworms from Nanking, China. Science Reports. Vol. 1. Nanjing: National Central University, College of Science, Nanking; p. 11 - 37." type="book chapter" year="1930">Chen, 1930</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D5CCA6EFFA52874FD49FD5D3932" authority="editus (Chen, 1946)" authorityName="editus (Chen" authorityYear="1946" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[279,681,766,791]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA6EFFA52874FD49FDE03932" box="[279,532,766,791]" italics="true" pageId="3" pageNumber="2502">Amynthas editus editus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA34B2ECA6EFFA52B43FD48FD543932" author="Chen Y" box="[544,672,767,791]" pageId="3" pageNumber="2502" pagination="83 - 141" refId="ref7925" refString="Chen Y. 1946. On the terrestrial Oligochaeta from Szechuan, = 3 \ * ROMAN III. West China Border Res Soc. 16: 83 - 141." type="journal article" year="1946">Chen, 1946</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Moreover, they share the same collection location – Mt. Emei (
|
||
<bibRefCitation id="EFA34B2ECA6EFFA528E8FC95FDF3391F" author="Chen Y" box="[395,519,802,826]" pageId="3" pageNumber="2502" pagination="11 - 37" refId="ref7817" refString="Chen Y. 1930. On some new earthworms from Nanking, China. Science Reports. Vol. 1. Nanjing: National Central University, College of Science, Nanking; p. 11 - 37." type="book chapter" year="1930">Chen 1930</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA34B2ECA6EFFA52B76FC95FDB8391F" author="Chen Y" box="[533,588,802,826]" pageId="3" pageNumber="2502" pagination="83 - 141" refId="ref7925" refString="Chen Y. 1946. On the terrestrial Oligochaeta from Szechuan, = 3 \ * ROMAN III. West China Border Res Soc. 16: 83 - 141." type="journal article" year="1946">1946</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA6EFFA529A3FCF3FDE13FCA" blockId="3.[160,1156,558,1727]" pageId="3" pageNumber="2502">
|
||
The species
|
||
<taxonomicName id="4C324D5CCA6EFFA5282BFCF3FDF03978" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[328,516,836,861]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA5282BFCF3FDF03978" box="[328,516,836,861]" italics="true" pageId="3" pageNumber="2502">
|
||
A.
|
||
<materialsCitation id="3B5A3C82CA6EFFA52807FCF3FDF03978" ID-GBIF-Occurrence="2846790321" box="[356,516,836,861]" collectorName="Suzhou City & Nanjing City" country="China" county="Mt" location="Wuxi City" municipality="Yixing City" pageId="3" pageNumber="2502" specimenCount="1" stateProvince="Jiangsu">obscuritoporus</materialsCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, which was initially collected in
|
||
<collectorName id="26C75309CA6EFFA52A0EFCF3FC073979" box="[877,1011,836,860]" pageId="3" pageNumber="2502">Suzhou City</collectorName>
|
||
and
|
||
<collectorName id="26C75309CA6EFFA52D48FCF3FF3A395A" pageId="3" pageNumber="2502">Nanjing City</collectorName>
|
||
,
|
||
<collectingRegion id="49F6F83DCA6EFFA529B9FCD0FE6F395A" box="[218,411,871,895]" country="China" name="Jiangsu" pageId="3" pageNumber="2502">Jiangsu Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F325764FCA6EFFA528CBFCD0FE1F395A" box="[424,491,871,895]" name="China" pageId="3" pageNumber="2502">China</collectingCountry>
|
||
, and subsequently gathered in
|
||
<location id="8EED6004CA6EFFA52A3AFCD0FC32395A" LSID="urn:lsid:plazi:treatment:039B87C9CA6CFFA228E4FD7EFE9D39E6:8EED6004CA6EFFA52A3AFCD0FC32395A" box="[857,966,871,895]" country="China" county="Mt" municipality="Yixing City" name="Wuxi City" pageId="3" pageNumber="2502" stateProvince="Jiangsu">Wuxi City</location>
|
||
and
|
||
<collectingMunicipality id="6BE9ACA5CA6EFFA52D67FCD0FB74395A" box="[1028,1152,871,895]" pageId="3" pageNumber="2502">Yixing City</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49F6F83DCA6EFFA529C3FC3DFEAA3987" box="[160,350,906,930]" country="China" name="Jiangsu" pageId="3" pageNumber="2502">Jiangsu Province</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F6F83DCA6EFFA5280BFC3DFDC03987" box="[360,564,906,930]" country="China" name="Zhejiang" pageId="3" pageNumber="2502">Zhejiang Province</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F6F83DCA6EFFA52B5CFC3DFD1D3987" box="[575,745,906,930]" country="China" name="Anhui" pageId="3" pageNumber="2502">Anhui Province</collectingRegion>
|
||
and
|
||
<collectingCounty id="62EC4E53CA6EFFA52A41FC3DFCB63987" box="[802,834,906,930]" pageId="3" pageNumber="2502">Mt</collectingCounty>
|
||
. Emei,
|
||
<collectingRegion id="49F6F83DCA6EFFA52AF3FC3DFBA73987" box="[912,1107,906,930]" country="China" name="Sichuan" pageId="3" pageNumber="2502">Sichuan Province</collectingRegion>
|
||
, has several similar characteristics to the new species, as follows: (1) a dark colour on the dorsum and lacking pigment on the ventrum; (2) spermathecal pores three pairs in 6/7– 8/9; (3) spermathecal ampulla heart-shaped with the duct shorter than the ampulla; (4) spermathecal diverticulum is usually about half the length of the main spermathecal axis, and the seminal chamber was described as elongated ovoid or bulb-like; (5) the left and right lobe of testis sacs connected ventro-medially in the form of a transverse band. However, the two are chiefly differentiated from each other as follows: (1) the clitellum of the new species present in XIV–XVI, smooth and swollen, but not present in all cases for
|
||
<taxonomicName id="4C324D5CCA6EFFA529A4FB76FE703EFF" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[199,388,1217,1242]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA529A4FB76FE703EFF" box="[199,388,1217,1242]" italics="true" pageId="3" pageNumber="2502">A. obscuritoporus</emphasis>
|
||
</taxonomicName>
|
||
; (2) even though both have the spermathecal pores in 6/7–8/9, the pores of
|
||
<taxonomicName id="4C324D5CCA6EFFA52860FB53FE343ED8" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[259,448,1252,1277]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA52860FB53FE343ED8" box="[259,448,1252,1277]" italics="true" pageId="3" pageNumber="2502">A. obscuritoporus</emphasis>
|
||
</taxonomicName>
|
||
are too small to recognise, comparing with the new species in which the spermathecal pores are surrounded by a relatively large swollen area; (3) the male pores of the new species are surrounded by a larger irregular-shaped pad, while the male pore region of
|
||
<taxonomicName id="4C324D5CCA6EFFA528D2FAFBFD993F40" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[433,621,1356,1381]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA528D2FAFBFD993F40" box="[433,621,1356,1381]" italics="true" pageId="3" pageNumber="2502">A. obscuritoporus</emphasis>
|
||
</taxonomicName>
|
||
is an obscure whitish patch and appears vague under the skin, or looks like a small crescent-shaped groove in some cases; (4) the intestinal caeca of the new species is complex, while it is simple for
|
||
<taxonomicName id="4C324D5CCA6EFFA52AFEFA26FBAD3F8F" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[925,1113,1425,1450]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA52AFEFA26FBAD3F8F" box="[925,1113,1425,1450]" italics="true" pageId="3" pageNumber="2502">A. obscuritoporus</emphasis>
|
||
</taxonomicName>
|
||
; (5) the prostate glands of the new species are developed, by comparison, but are small or rudimentary in
|
||
<taxonomicName id="4C324D5CCA6EFFA52832FA61FDFA3FCA" baseAuthorityName="Chen" baseAuthorityYear="1930" box="[337,526,1494,1519]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="species" species="obscuritoporus">
|
||
<emphasis id="B946EACDCA6EFFA52832FA61FDFA3FCA" box="[337,526,1494,1519]" italics="true" pageId="3" pageNumber="2502">A. obscuritoporus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA6EFFA229A3FA4DFD073B99" blockId="3.[160,1156,558,1727]" lastBlockId="4.[160,1156,142,963]" lastPageId="4" lastPageNumber="2503" pageId="3" pageNumber="2502">
|
||
Another species,
|
||
<taxonomicName id="4C324D5CCA6EFFA528EAFA4EFDC33C37" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[393,567,1529,1554]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA6EFFA528EAFA4EFDC33C37" box="[393,567,1529,1554]" italics="true" pageId="3" pageNumber="2502">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
, also collected in Mt. Emei,
|
||
<collectingRegion id="49F6F83DCA6EFFA52AEBFA4DFC173C37" box="[904,995,1530,1554]" country="China" name="Sichuan" pageId="3" pageNumber="2502">Sichuan</collectingRegion>
|
||
, is somewhat similar to the new species in that they have a similar density of setae, the spermathecal pores are all located in 6/7–8/9, the male pore papillae occupy more than one segment, the spermathecal ampulla is heart-shaped, the seminal chamber is elongated, and the prostate glands are developed with a U-curved duct. However, they differ in the following main characteristics: (1) the pigment of
|
||
<taxonomicName id="4C324D5CCA6EFFA52B87F911FC7A3C9A" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[740,910,1702,1727]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="2502" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA6EFFA52B87F911FC7A3C9A" box="[740,910,1702,1727]" italics="true" pageId="3" pageNumber="2502">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
is pale both dorsally and ventrally, but just ventrally for the new species; (2) the first two pairs of spermathecal pores of
|
||
<taxonomicName id="4C324D5CCA69FFA22848FF07FE253AEC" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[299,465,176,201]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA69FFA22848FF07FE253AEC" box="[299,465,176,201]" italics="true" pageId="4" pageNumber="2503">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
are extremely close – one seta interval, compared to the last pair – while the distance is equal in the new species; (3) there is a large lens-shaped papilla placed medio-ventrally in the male pore region of
|
||
<taxonomicName id="4C324D5CCA69FFA22A4FFF42FC253B2B" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[812,977,245,270]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA69FFA22A4FFF42FC253B2B" box="[812,977,245,270]" italics="true" pageId="4" pageNumber="2503">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
but no papillae for the new species; (4) the intestinal caeca is simple for
|
||
<taxonomicName id="4C324D5CCA69FFA22A42FEAFFC333B14" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[801,967,280,305]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA69FFA22A42FEAFFC333B14" box="[801,967,280,305]" italics="true" pageId="4" pageNumber="2503">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
but complex for the new species; (5) the spermathecal diverticulum is very long and the ectal portion of duct coiled in
|
||
<taxonomicName id="4C324D5CCA69FFA22827FEEAFE1D3B53" authority="editus" authorityName="editus (Chen" authorityYear="1946" box="[324,489,349,374]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA69FFA22827FEEAFE1D3B53" box="[324,489,349,374]" italics="true" pageId="4" pageNumber="2503">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
, compared to a shorter and more slender duct in the new species; (6) the first pair of seminal vesicles in XI are enclosed in testis sacs for
|
||
<taxonomicName id="4C324D5CCA69FFA22D46FE37FF153B99" authority="editus" authorityName="editus (Chen" authorityYear="1946" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="subSpecies" species="editus" subSpecies="editus">
|
||
<emphasis id="B946EACDCA69FFA22D46FE37FF153B99" italics="true" pageId="4" pageNumber="2503">A. editus editus</emphasis>
|
||
</taxonomicName>
|
||
but are clearly separated for the new species.
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA69FFA229A3FE71FF0C391C" blockId="4.[160,1156,142,963]" pageId="4" pageNumber="2503">
|
||
Two other recently reported
|
||
<taxonomicName id="4C324D5CCA69FFA22B75FE72FD143BFB" baseAuthorityName="Horst" baseAuthorityYear="1883" box="[534,736,453,478]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="sieboldi">
|
||
<emphasis id="B946EACDCA69FFA22B75FE72FD143BFB" box="[534,736,453,478]" italics="true" pageId="4" pageNumber="2503">Amynthas sieboldi</emphasis>
|
||
</taxonomicName>
|
||
-group species –
|
||
<taxonomicName id="4C324D5CCA69FFA22ACDFE72FE443824" authority="Bantaowong and Panha" authorityName="Bantaowong and Panha" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="arenulus">
|
||
<emphasis id="B946EACDCA69FFA22ACDFE72FB773BFB" box="[942,1155,453,478]" italics="true" pageId="4" pageNumber="2503">Amynthas arenulus</emphasis>
|
||
Bantaowong and Panha
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D5CCA69FFA2288DFE5FFC063824" authority="Bantaowong and Panha" authorityName="Bantaowong and Panha" box="[494,1010,488,513]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="longicaeca">
|
||
<emphasis id="B946EACDCA69FFA2288DFE5FFD2D3824" box="[494,729,488,513]" italics="true" pageId="4" pageNumber="2503">Amynthas longicaeca</emphasis>
|
||
Bantaowong and Panha
|
||
</taxonomicName>
|
||
– are somewhat similar to the new species according to the big pad on the male pore region at first glance, and the distance between male pores and spermathecal pores. However, they have many differences. For example: (1) comparing with
|
||
<taxonomicName id="4C324D5CCA69FFA22A51FDE7FC59384C" box="[818,941,592,617]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="arenulus">
|
||
<emphasis id="B946EACDCA69FFA22A51FDE7FC59384C" box="[818,941,592,617]" italics="true" pageId="4" pageNumber="2503">A. arenulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D5CCA69FFA22A88FDE7FB8A384C" box="[1003,1150,592,617]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="longicaeca">
|
||
<emphasis id="B946EACDCA69FFA22A88FDE7FB8A384C" box="[1003,1150,592,617]" italics="true" pageId="4" pageNumber="2503">A. longicaeca</emphasis>
|
||
</taxonomicName>
|
||
, the new species has a smaller body size; (2) the spermathecal pores have tumid epidermis for the new species, but not for
|
||
<taxonomicName id="4C324D5CCA69FFA22B1DFD22FD03388B" box="[638,759,661,686]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="arenulus">
|
||
<emphasis id="B946EACDCA69FFA22B1DFD22FD03388B" box="[638,759,661,686]" italics="true" pageId="4" pageNumber="2503">A. arenulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D5CCA69FFA22A4CFD22FC34388B" box="[815,960,661,686]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="longicaeca">
|
||
<emphasis id="B946EACDCA69FFA22A4CFD22FC34388B" box="[815,960,661,686]" italics="true" pageId="4" pageNumber="2503">A. longicaeca</emphasis>
|
||
</taxonomicName>
|
||
; (3) the intestinal caeca of the new species are complex; however, they are simple for
|
||
<taxonomicName id="4C324D5CCA69FFA22AD5FD0FFBC538F4" box="[950,1073,696,721]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="arenulus">
|
||
<emphasis id="B946EACDCA69FFA22AD5FD0FFBC538F4" box="[950,1073,696,721]" italics="true" pageId="4" pageNumber="2503">A. arenulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D5CCA69FFA22D0DFD0FFEE138D6" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="longicaeca">
|
||
<emphasis id="B946EACDCA69FFA22D0DFD0FFEE138D6" italics="true" pageId="4" pageNumber="2503">A. longicaeca</emphasis>
|
||
</taxonomicName>
|
||
; (4) the duct of spermathecae is stout for
|
||
<taxonomicName id="4C324D5CCA69FFA22B95FD6DFC8438D6" box="[758,880,730,755]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="arenulus">
|
||
<emphasis id="B946EACDCA69FFA22B95FD6DFC8438D6" box="[758,880,730,755]" italics="true" pageId="4" pageNumber="2503">A. arenulus</emphasis>
|
||
</taxonomicName>
|
||
, and the diverticulum is very small with ovate knob for
|
||
<taxonomicName id="4C324D5CCA69FFA2289EFD4AFD7A3933" box="[509,654,765,790]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="longicaeca">
|
||
<emphasis id="B946EACDCA69FFA2289EFD4AFD7A3933" box="[509,654,765,790]" italics="true" pageId="4" pageNumber="2503">A. longicaeca</emphasis>
|
||
</taxonomicName>
|
||
, which are both very different from the new species.
|
||
</paragraph>
|
||
<paragraph id="8B8D36DFCA69FFA229A3FCF4FE9D39E6" blockId="4.[160,1156,142,963]" pageId="4" pageNumber="2503">
|
||
<materialsCitation id="3B5A3C82CA69FFA229A3FCF4FB8B397E" ID-GBIF-Occurrence="2846790307" box="[192,1151,834,859]" collectionCode="COI" location="The" pageId="4" pageNumber="2503" specimenCount="1" typeStatus="holotype">
|
||
<location id="8EED6004CA69FFA229A3FCF4FF1D397E" LSID="urn:lsid:plazi:treatment:039B87C9CA6CFFA228E4FD7EFE9D39E6:8EED6004CA69FFA229A3FCF4FF1D397E" box="[192,233,835,859]" name="The" pageId="4" pageNumber="2503">The</location>
|
||
partial cytochrome c oxidase subunit I (
|
||
<collectionCode id="ED23AE1ACA69FFA22BB3FCF4FD08397E" box="[720,764,835,859]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="4" pageNumber="2503" type="Herbarium">COI</collectionCode>
|
||
) sequence of the
|
||
<typeStatus id="5489887DCA69FFA22AB9FCF4FBCA397E" box="[986,1086,835,859]" pageId="4" pageNumber="2503" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C324D5CCA69FFA22D0DFCF5FB8B397E" authority="Sun & Jiang & Hu & Qiu, 2016" authorityName="Sun & Jiang & Hu & Qiu" authorityYear="2016" box="[1134,1151,834,859]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="2503" phylum="Annelida" rank="species" species="leshanensis" status="sp. nov.">
|
||
<emphasis id="B946EACDCA69FFA22D0DFCF5FB8B397E" box="[1134,1151,834,859]" italics="true" pageId="4" pageNumber="2503">A</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
<emphasis id="B946EACDCA69FFA22DE3FCF5FEEF395B" italics="true" pageId="4" pageNumber="2503">. leshanensis</emphasis>
|
||
<taxonomicNameLabel id="A27557B6CA69FFA22848FCD1FE7F395B" box="[299,395,870,894]" pageId="4" pageNumber="2503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<materialsCitation id="3B5A3C82CA69FFA228F9FCD1FE9139E6" ID-GBIF-Occurrence="2846790313" collectingDate="02-11-20" collectionCode="NCBI" location="National Center of Biotechnology Information" pageId="4" pageNumber="2503" specimenCount="1" typeStatus="holotype">
|
||
has been deposited in the
|
||
<location id="8EED6004CA69FFA22B91FCD1FED03984" LSID="urn:lsid:plazi:treatment:039B87C9CA6CFFA228E4FD7EFE9D39E6:8EED6004CA69FFA22B91FCD1FED03984" name="National Center of Biotechnology Information" pageId="4" pageNumber="2503">National Center of Biotechnology Information</location>
|
||
(
|
||
<collectionCode id="ED23AE1ACA69FFA22829FC3EFE703984" box="[330,388,905,929]" pageId="4" pageNumber="2503">NCBI</collectionCode>
|
||
) GenBank with accession number
|
||
<accessionNumber id="9461AB3CCA69FFA22AE1FC3EFC043984" box="[898,1008,905,929]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF205474" pageId="4" pageNumber="2503">KF205474</accessionNumber>
|
||
(specimen C-SC
|
||
<date id="FF8C101FCA69FFA229BFFC1CFEDB39E6" box="[220,303,939,963]" pageId="4" pageNumber="2503" value="02-11-20">
|
||
<collectingDate id="EFC8E9F7CA69FFA229BFFC1CFEDB39E6" box="[220,303,939,963]" pageId="4" pageNumber="2503" value="02-11-20">201102</collectingDate>
|
||
</date>
|
||
-02A)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |