181 lines
23 KiB
XML
181 lines
23 KiB
XML
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<mods:title id="F28B4B98F5FE5B17F2DFBC71451ADBAD">On the Staphylinidae of the Greek island Samothraki (Insecta Coleoptera)</mods:title>
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<taxonomicName id="4C324D0EFFE7FFB5DACCFDE7FDD9FD87" ID-CoL="8Y6DC" authorityName="Assing" authorityYear="2019" box="[151,604,606,628]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="species" species="samothracicus" subGenus="Agnotyphlus">
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<emphasis id="B946EA9FFFE7FFB5DACCFDE7FEA7FD87" bold="true" box="[151,290,606,628]" italics="true" pageId="16" pageNumber="897">Metrotyphlus</emphasis>
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(
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<emphasis id="B946EA9FFFE7FFB5DB6AFDE7FE30FD87" bold="true" box="[305,437,606,628]" italics="true" pageId="16" pageNumber="897">Agnotyphlus</emphasis>
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)
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<emphasis id="B946EA9FFFE7FFB5DB9FFDE7FDD9FD87" bold="true" box="[452,604,606,628]" italics="true" pageId="16" pageNumber="897">samothracicus</emphasis>
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</taxonomicName>
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ASSING nov.sp. (
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<figureCitation id="13092A08FFE7FFB5D954FDE7FC04FD80" box="[783,897,606,628]" captionStart-0="Figs 9-18" captionStart-1="Figs 19-26" captionStartId-0="13.[151,191,1459,1481]" captionStartId-1="14.[151,191,1459,1481]" captionTargetBox-0="[156,1042,192,1449]" captionTargetBox-1="[159,1035,199,1435]" captionTargetId-0="figure@13.[152,1045,189,1454]" captionTargetId-1="figure@14.[152,1037,190,1444]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figs 9-18: Cyrtotyphlus samothracicus (9-14) and Metrotyphlus samothracicus (15-18): (9, 15) habitus; (10, 16) male sternite VIII; (11-14, 17-18) median lobe of aedeagus in lateral and in ventral view. Scale bars: 0.1 mm." captionText-1="Figs 19-26: Metrotyphlus samothracicus (19), Cephennium samothracicum (20-22), Scydmoraphes samothracicus (23-24), and Apimela procera (25-26): (19) hind leg; (20) habitus; (21-24) aedeagus in lateral and in ventral view; (25) male tergite VIII; (26) male sternite VIII. Scale bars: 20: 0.5 mm; 25-26: 0.2 mm; 19, 21-24: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3738349" figureDoi-1="http://doi.org/10.5281/zenodo.3738351" httpUri-0="https://zenodo.org/record/3738349/files/figure.png" httpUri-1="https://zenodo.org/record/3738351/files/figure.png" pageId="16" pageNumber="897">Figs 15-19</figureCitation>
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, 33)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D368DFFE7FFB5DACCFD3DFDD2FD34" blockId="16.[151,1058,641,711]" pageId="16" pageNumber="897">
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y p e m a t e r i a l:
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: "GR – Samothraki [27], Therma,
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,
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,
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,
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<emphasis id="B946EA9FFFE7FFB5DAABFD23FEC4FD5C" box="[240,321,666,687]" italics="true" pageId="16" pageNumber="897">Platanus</emphasis>
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forest
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, soil wash.,
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<collectingDate id="EFC8E9A5FFE7FFB5D84FFD23FDF9FD5C" box="[532,636,666,687]" pageId="16" pageNumber="897" value="2019-04-12">12.IV.2019</collectingDate>
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,
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<collectionCode id="ED23AE48FFE7FFB5D8CBFD23FD26FD5C" box="[656,675,666,687]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="897" type="Museum">V</collectionCode>
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.
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<collectingMunicipality id="6BE9ACF7FFE7FFB5D8EDFD23FD70FD43" box="[694,757,666,688]" pageId="16" pageNumber="897">Assing</collectingMunicipality>
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/
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<typeStatus id="5489882FFFE7FFB5D940FD23FCFCFD43" box="[795,889,666,688]" pageId="16" pageNumber="897" type="holotype">Holotypus</typeStatus>
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<specimenCount id="9D34FD04FFE7FFB5D9D2FD21FC1FFD5C" box="[905,922,664,687]" pageId="16" pageNumber="897" type="male">♂</specimenCount>
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<taxonomicName id="4C324D0EFFE7FFB5D9F2FD23FE9EFD35" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="species" species="samothracicus" status="sp. nov.">
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<emphasis id="B946EA9FFFE7FFB5D9F2FD23FE9EFD35" italics="true" pageId="16" pageNumber="897">Metrotyphlus samothracicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27557E4FFE7FFB5DB7AFD08FEC8FD35" box="[289,333,689,710]" pageId="16" pageNumber="897" rank="species">sp.n.</taxonomicNameLabel>
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det.
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<collectionCode id="ED23AE48FFE7FFB5DB22FD08FE09FD35" box="[377,396,689,710]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="897" type="Museum">V</collectionCode>
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.
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<collectingCounty id="62EC4E01FFE7FFB5DBCFFD08FE56FD34" box="[404,467,689,711]" pageId="16" pageNumber="897">Assing</collectingCounty>
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2019" (cAss).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286506FFE7FFB5DACCFD6CFC56FA64" pageId="16" pageNumber="897" type="description">
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<paragraph id="8B8D368DFFE7FFB5DACCFD6CFC56FD18" blockId="16.[151,979,725,747]" box="[151,979,725,747]" pageId="16" pageNumber="897">E t y m o l o g y: The specific epithet is an adjective derived from Samothrace.</paragraph>
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<paragraph id="8B8D368DFFE7FFB5DACCFD42FDBBFCB8" blockId="16.[151,1058,762,843]" pageId="16" pageNumber="897">
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D e s c r i p t i o n: Body length
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; length of forebody
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<quantity id="4CCA9B68FFE7FFB5D96DFD43FC0FFCE3" box="[822,906,762,784]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="897" unit="mm" value="0.5">0.5 mm</quantity>
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. Habitus as in
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<figureCitation id="13092A08FFE7FFB5DACCFCAEFF67FCDE" box="[151,226,791,813]" captionStart="Figs 9-18" captionStartId="13.[151,191,1459,1481]" captionTargetBox="[156,1042,192,1449]" captionTargetId="figure@13.[152,1045,189,1454]" captionTargetPageId="13" captionText="Figs 9-18: Cyrtotyphlus samothracicus (9-14) and Metrotyphlus samothracicus (15-18): (9, 15) habitus; (10, 16) male sternite VIII; (11-14, 17-18) median lobe of aedeagus in lateral and in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3738349" httpUri="https://zenodo.org/record/3738349/files/figure.png" pageId="16" pageNumber="897">Fig. 15</figureCitation>
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. Tarsal formula 2, 2, 2 (
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<figureCitation id="13092A08FFE7FFB5DB81FCAEFDA3FCDE" box="[474,550,791,813]" captionStart="Figs 19-26" captionStartId="14.[151,191,1459,1481]" captionTargetBox="[159,1035,199,1435]" captionTargetId="figure@14.[152,1037,190,1444]" captionTargetPageId="14" captionText="Figs 19-26: Metrotyphlus samothracicus (19), Cephennium samothracicum (20-22), Scydmoraphes samothracicus (23-24), and Apimela procera (25-26): (19) hind leg; (20) habitus; (21-24) aedeagus in lateral and in ventral view; (25) male tergite VIII; (26) male sternite VIII. Scale bars: 20: 0.5 mm; 25-26: 0.2 mm; 19, 21-24: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3738351" httpUri="https://zenodo.org/record/3738351/files/figure.png" pageId="16" pageNumber="897">Fig. 19</figureCitation>
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). Reliably distinguished from other species of the genus only by the male sexual characters.
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</paragraph>
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<paragraph id="8B8D368DFFE7FFB5DACCFCE1FE53FC3B" blockId="16.[151,1059,856,968]" pageId="16" pageNumber="897">
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♂: sternites III-VIII with shallow, but extensive median impressions; sternite VIII (
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||
<figureCitation id="13092A08FFE7FFB5D9A2FCE3FF34FC7E" captionStart="Figs 9-18" captionStartId="13.[151,191,1459,1481]" captionTargetBox="[156,1042,192,1449]" captionTargetId="figure@13.[152,1045,189,1454]" captionTargetPageId="13" captionText="Figs 9-18: Cyrtotyphlus samothracicus (9-14) and Metrotyphlus samothracicus (15-18): (9, 15) habitus; (10, 16) male sternite VIII; (11-14, 17-18) median lobe of aedeagus in lateral and in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3738349" httpUri="https://zenodo.org/record/3738349/files/figure.png" pageId="16" pageNumber="897">Fig. 16</figureCitation>
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||
) weakly oblong and nearly symmetric, posterior margin with a large median excision; aedeagus (
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<figureCitation id="13092A08FFE7FFB5DB58FC2CFEF6FC58" box="[259,371,917,939]" captionStart="Figs 9-18" captionStartId="13.[151,191,1459,1481]" captionTargetBox="[156,1042,192,1449]" captionTargetId="figure@13.[152,1045,189,1454]" captionTargetPageId="13" captionText="Figs 9-18: Cyrtotyphlus samothracicus (9-14) and Metrotyphlus samothracicus (15-18): (9, 15) habitus; (10, 16) male sternite VIII; (11-14, 17-18) median lobe of aedeagus in lateral and in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3738349" httpUri="https://zenodo.org/record/3738349/files/figure.png" pageId="16" pageNumber="897">Figs 17-18</figureCitation>
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)
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<quantity id="4CCA9B68FFE7FFB5DBD8FC2DFE58FC59" box="[387,477,916,938]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="897" unit="mm" value="0.15">0.15 mm</quantity>
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long and, except internal structures, symmetric; internal structures of distinctive shapes.
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</paragraph>
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<paragraph id="8B8D368DFFE7FFB5DACCFC6EFEFCFB48" blockId="16.[151,1058,983,1211]" pageId="16" pageNumber="897">
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C o m p a r a t i v e n o t e s:
|
||
<taxonomicName id="4C324D0EFFE7FFB5D858FC6EFD0FFC1F" authorityName="Coiffait" authorityYear="1959" box="[515,650,983,1004]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="genus">
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<emphasis id="B946EA9FFFE7FFB5D858FC6EFD0FFC1F" box="[515,650,983,1004]" italics="true" pageId="16" pageNumber="897">Metrotyphlus</emphasis>
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||
</taxonomicName>
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||
COIFFAIT, 1959 previously included twelve species in two subgenera, four in the nominal subgenus and eight in the subgenus
|
||
<taxonomicName id="4C324D0EFFE7FFB5DACCFBABFE93FBD4" box="[151,278,1042,1063]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="subGenus" subGenus="Agnotyphlus">
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||
<emphasis id="B946EA9FFFE7FFB5DACCFBABFE93FBD4" box="[151,278,1042,1063]" italics="true" pageId="16" pageNumber="897">Agnotyphlus</emphasis>
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</taxonomicName>
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||
<bibRefCitation id="EFA34B7CFFE7FFB5DB72FBABFE51FBD4" author="COIFFAIT H" box="[297,468,1042,1064]" pageId="16" pageNumber="897" pagination="1 - 651" refId="ref14361" refString="COIFFAIT H. (1972): Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, sous-familles: Xantholininae et Leptotyphlinae. - Supplement a la Nouvelle Revue d'Entomologie 2 (2): 1 - 651." type="journal article" year="1972">COIFFAIT, 1972</bibRefCitation>
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||
(SCHÜLKE & SMETANA 2015). Those assigned to
|
||
<taxonomicName id="4C324D0EFFE7FFB5DACCFB96FE93FBB7" box="[151,278,1071,1092]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="subGenus" subGenus="Agnotyphlus">
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||
<emphasis id="B946EA9FFFE7FFB5DACCFB96FE93FBB7" box="[151,278,1071,1092]" italics="true" pageId="16" pageNumber="897">Agnotyphlus</emphasis>
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||
</taxonomicName>
|
||
had been recorded from the Alpi Apuane and across the southern Alps southeastwards to
|
||
<collectingRegion id="49F6F86FFFE7FFB5DB07FBF4FE15FB90" box="[348,400,1101,1123]" country="Croatia" name="Istarska" pageId="16" pageNumber="897">Istria</collectingRegion>
|
||
(
|
||
<collectingCountry id="F325761DFFE7FFB5DBF9FBF4FE76FB90" box="[418,499,1101,1123]" name="Croatia" pageId="16" pageNumber="897">Croatia</collectingCountry>
|
||
). Thus, the record of
|
||
<taxonomicName id="4C324D0EFFE7FFB5D8BBFBF4FC1EFB91" authorityName="Assing" authorityYear="2019" box="[736,923,1101,1122]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="species" species="samothracicus">
|
||
<emphasis id="B946EA9FFFE7FFB5D8BBFBF4FC1EFB91" box="[736,923,1101,1122]" italics="true" pageId="16" pageNumber="897">M. samothracicus</emphasis>
|
||
</taxonomicName>
|
||
expands the known range of both the genus and the subgenus southeastwards by a distance of more than
|
||
<quantity id="4CCA9B68FFE7FFB5DA91FB3EFEAFFB6E" box="[202,298,1159,1181]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="897" unit="km" value="1000.0">1,000 km</quantity>
|
||
and suggests that at least
|
||
<taxonomicName id="4C324D0EFFE7FFB5D86DFB3EFD30FB6F" box="[566,693,1159,1180]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="subGenus" subGenus="Agnotyphlus">
|
||
<emphasis id="B946EA9FFFE7FFB5D86DFB3EFD30FB6F" box="[566,693,1159,1180]" italics="true" pageId="16" pageNumber="897">Agnotyphlus</emphasis>
|
||
</taxonomicName>
|
||
is far more widespread and diverse than presently known.
|
||
</paragraph>
|
||
<paragraph id="8B8D368DFFE7FFB5DACCFB73FB81FAE8" blockId="16.[151,1058,1226,1307]" pageId="16" pageNumber="897">
|
||
The new species differs from other
|
||
<taxonomicName id="4C324D0EFFE7FFB5D853FB73FD02FB2C" box="[520,647,1226,1247]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="subGenus" subGenus="Agnotyphlus">
|
||
<emphasis id="B946EA9FFFE7FFB5D853FB73FD02FB2C" box="[520,647,1226,1247]" italics="true" pageId="16" pageNumber="897">Agnotyphlus</emphasis>
|
||
</taxonomicName>
|
||
species by the distinctive male primary and secondary sexual characters, particularly the internal structures of the aedeagus. For illustrations of the previously described species see
|
||
<bibRefCitation id="EFA34B7CFFE7FFB5D8FEFABCFCC9FAE8" author="COIFFAIT H" box="[677,844,1285,1307]" pageId="16" pageNumber="897" pagination="1 - 651" refId="ref14361" refString="COIFFAIT H. (1972): Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, sous-familles: Xantholininae et Leptotyphlinae. - Supplement a la Nouvelle Revue d'Entomologie 2 (2): 1 - 651." type="journal article" year="1972">COIFFAIT (1972)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFA34B7CFFE7FFB5D926FABCFB84FAE8" author="PACE R" box="[893,1025,1285,1307]" pageId="16" pageNumber="897" pagination="1 - 328" refId="ref14571" refString="PACE R. (1997): Coleoptera Staphylinidae Leptotyphlinae. Fauna d'Italia Vol. 34. - Edizioni Calderini, Bologna: i-viii + 1 - 328." type="book chapter" year="1997">PACE (1997)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8D368DFFE7FFB5DACCFA92FC56FA64" blockId="16.[151,1058,1322,1432]" pageId="16" pageNumber="897">
|
||
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y:
|
||
<taxonomicName id="4C324D0EFFE7FFB5D8A1FA93FBA7FACC" authorityName="Assing" authorityYear="2019" box="[762,1058,1322,1343]" class="Insecta" family="Staphylinidae" genus="Metrotyphlus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="897" phylum="Arthropoda" rank="species" species="samothracicus">
|
||
<emphasis id="B946EA9FFFE7FFB5D8A1FA93FBA7FACC" box="[762,1058,1322,1343]" italics="true" pageId="16" pageNumber="897">Metrotyphlus samothracicus</emphasis>
|
||
</taxonomicName>
|
||
is most likely endemic to Samothraki. Despite extensive soil washing in numerous localities, only a single male was found. It was collected in the north of the island in a moist
|
||
<emphasis id="B946EA9FFFE7FFB5DA8DFA3BFEB5FA64" box="[214,304,1410,1431]" italics="true" pageId="16" pageNumber="897">Platanus</emphasis>
|
||
forest at an altitude of
|
||
<specimenCount id="9D34FD04FFE7FFB5D840FA3BFDE7FA6B" box="[539,610,1410,1432]" pageId="16" pageNumber="897" type="generic">
|
||
<quantity id="4CCA9B68FFE7FFB5D840FA3BFDD8FA6B" box="[539,605,1410,1432]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="897" unit="m" value="110.0">110 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The locality is illustrated in
|
||
<figureCitation id="13092A08FFE7FFB5D9DFFA3BFC4AFA64" box="[900,975,1410,1432]" captionStart="Figs 33-34" captionStartId="17.[151,191,1459,1481]" captionTargetBox="[153,1038,191,1445]" captionTargetId="figure@17.[153,1038,191,1445]" captionTargetPageId="17" captionText="Figs 33-34: Type localities of Metrotyphlus samothracicus (sample number 20) (above) and of Tychobythinus assingi (sample number 14) (below). In the latter locality, Cyrtotyphlus samothracicus was found, too." figureDoi="http://doi.org/10.5281/zenodo.3738355" httpUri="https://zenodo.org/record/3738355/files/figure.png" pageId="16" pageNumber="897">Fig. 33</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |