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<document id="046C71BE07F920F84A3E59AC22FBC6AF" ID-CLB-Dataset="298832" ID-DOI="10.11646/zootaxa.5471.2.8" ID-GBIF-Dataset="f86db238-ec9b-4c73-923d-c67b5d5524a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190027" ID-ZooBank="E02B0AE1-E256-4620-90A0-4917D6A5608D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718893184829" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="039B87876828FF92BEDD0775FDB6FC1F" docLanguage="en" docName="zootaxa.5471.2.8.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pomphopsilla bifurcata Jałoszyński 2024, sp. n." docType="treatment" docVersion="6" lastPageNumber="290" masterDocId="FFA2FFFF682AFF91BE4A022BFFFCFF9E" masterDocTitle="Three new species of Pomphopsilla Jałoszyński in Togo (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="292" masterPageNumber="287" pageNumber="289" updateTime="1719234991119" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B2435C09464E83657D520C672A6BD278">Three new species of Pomphopsilla Jałoszyński in Togo (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:date id="3C37CE72A9BB63C614A976D4F82A7159">2024</mods:date>
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<treatment id="039B87876828FF92BEDD0775FDB6FC1F" ID-DOI="http://doi.org/10.5281/zenodo.12190745" ID-GBIF-Taxon="233743413" ID-Zenodo-Dep="12190745" LSID="urn:lsid:plazi:treatment:039B87876828FF92BEDD0775FDB6FC1F" httpUri="http://treatment.plazi.org/id/039B87876828FF92BEDD0775FDB6FC1F" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289">
<subSubSection id="C328651A6828FF93BEDD0775FE01FAE6" box="[151,509,1374,1401]" pageId="2" pageNumber="289" type="nomenclature">
<paragraph id="8B8D36916828FF93BEDD0775FE01FAE6" blockId="2.[151,509,1374,1437]" box="[151,509,1374,1401]" pageId="2" pageNumber="289">
<heading id="D0C581FD6828FF93BEDD0775FE01FAE6" bold="true" box="[151,509,1374,1401]" fontSize="11" level="1" pageId="2" pageNumber="289" reason="1">
<emphasis id="B946EA836828FF93BEDD0775FE01FAE6" bold="true" box="[151,509,1374,1401]" pageId="2" pageNumber="289">
<taxonomicName id="4C324D126828FF93BEDD0775FE48FAE7" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[151,436,1374,1401]" class="Insecta" family="Staphylinidae" genus="Pomphopsilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="bifurcata" status="sp. nov.">
<emphasis id="B946EA836828FF93BEDD0775FE48FAE7" bold="true" box="[151,436,1374,1401]" italics="true" pageId="2" pageNumber="289">Pomphopsilla bifurcata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557F86828FF93BFF10775FE01FAE6" box="[443,509,1374,1400]" pageId="2" pageNumber="289" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C328651A6828FF93BEDD07A9FEDBFA02" box="[151,295,1410,1437]" pageId="2" pageNumber="289" type="description">
<paragraph id="8B8D36916828FF93BEDD07A9FEDBFA02" blockId="2.[151,509,1374,1437]" box="[151,295,1410,1437]" pageId="2" pageNumber="289">
(
<figureCitation id="13092A146828FF93BED507A9FF14FA02" box="[159,232,1410,1436]" captionStart="FIGURES 13" captionStartId="1.[151,264,1977,2001]" captionTargetBox="[157,1430,1132,1948]" captionTargetId="figure-189@1.[151,1436,1126,1954]" captionTargetPageId="1" captionText="FIGURES 13. Dorsal habitus of holotype males. Pomphopsilla spinosa sp. n. (1); P. bifurcata sp. n. (2); P. klotoensis sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.12190029" httpUri="https://zenodo.org/record/12190029/files/figure.png" pageId="2" pageNumber="289">Figs 2</figureCitation>
,
<figureCitation id="13092A146828FF93BEBE07A9FEF8FA03" box="[244,260,1410,1437]" captionStart="FIGURES 46" captionStartId="3.[151,264,1648,1672]" captionTargetBox="[155,1432,925,1618]" captionTargetId="figure-272@3.[151,1436,921,1625]" captionTargetPageId="3" captionText="FIGURES 46. Aedeagus in ventral view. Pomphopsilla spinosa sp. n. (4); P. bifurcata sp. n. (5); P. klotoensis sp. n. (6)." figureDoi="http://doi.org/10.5281/zenodo.12190031" httpUri="https://zenodo.org/record/12190031/files/figure.png" pageId="2" pageNumber="289">5</figureCitation>
,
<figureCitation id="13092A146828FF93BF5A07A9FEE3FA02" box="[272,287,1410,1436]" captionStart="FIGURE 7" captionStartId="4.[151,250,1654,1678]" captionTargetBox="[157,1435,525,1623]" captionTargetId="figure-176@4.[151,1435,525,1631]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of nominal Pomphopsilla species, with aedeagus of each species in ventral view (P. felicitas is known from a single female, and P. crenata is known from a single male with a strongly distorted aedeagus)." figureDoi="http://doi.org/10.5281/zenodo.12190035" httpUri="https://zenodo.org/record/12190035/files/figure.png" pageId="2" pageNumber="289">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C328651A6828FF93BEDD07E1FF02F9EA" pageId="2" pageNumber="289" type="materials_examined">
<paragraph id="8B8D36916828FF93BEDD07E1FF02F9EA" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="289">
<emphasis id="B946EA836828FF93BEDD07E1FEB4FA7A" bold="true" box="[151,328,1482,1508]" pageId="2" pageNumber="289">Type material.</emphasis>
<materialsCitation id="3B5A3CCC6828FF93BF1B07E1FD5CF9B2" ID-GBIF-Occurrence="4899212305" collectionCode="PALIME, MHNG" collectorName="S. Vit" country="Togo" location="Foret de Klouto" pageId="2" pageNumber="289" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B946EA836828FF93BF1B07E1FE3BFA7A" bold="true" box="[337,455,1482,1508]" pageId="2" pageNumber="289">
<typeStatus id="548988336828FF93BF1B07E1FE3EFA7A" box="[337,450,1482,1508]" pageId="2" pageNumber="289" type="holotype">Holotype</typeStatus>
:
</emphasis>
(
<emphasis id="B946EA836828FF93BF9007E1FDBCFA7A" bold="true" box="[474,576,1482,1508]" pageId="2" pageNumber="289">
<collectingCountry id="F32576016828FF93BF9007E1FDCFFA7A" box="[474,563,1482,1508]" name="Togo" pageId="2" pageNumber="289">TOGO</collectingCountry>
):
</emphasis>
<specimenCount id="9D34FD186828FF93BC0007E0FD9DFA7B" box="[586,609,1483,1509]" count="1" pageId="2" pageNumber="289" type="male"></specimenCount>
, three labels: “TOGO
<collectionCode id="ED23AE546828FF93BD2D07E1FC31FA7A" box="[871,973,1482,1508]" pageId="2" pageNumber="289">PALIME</collectionCode>
/
<location id="8EED604A6828FF93BDA207E1FB63FA7A" LSID="urn:lsid:plazi:treatment:039B87876828FF92BEDD0775FDB6FC1F:8EED604A6828FF93BDA207E1FB63FA7A" box="[1000,1183,1482,1508]" country="Togo" name="Foret de Klouto" pageId="2" pageNumber="289">Forêt de Klouto</location>
/ 20-24-IV-74
<collectorName id="26C753476828FF93BB0407E1FA6FFA7A" box="[1358,1427,1482,1508]" pageId="2" pageNumber="289">S. Vit</collectorName>
” [white, printed], “
<collectorName id="26C753476828FF93BF1507C5FE22F996" box="[351,478,1518,1544]" pageId="2" pageNumber="289">Coll. S. Vit</collectorName>
/
<collectionCode id="ED23AE546828FF93BFBE07C5FDB6F996" box="[500,586,1518,1544]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="289" type="Museum">MHNG</collectionCode>
- 2021” [white, printed], “
<taxonomicName id="4C324D126828FF93BD3F07C4FBBFF997" authorityName="Jaloszynski" authorityYear="2011" box="[885,1091,1519,1545]" class="Insecta" family="Staphylinidae" genus="Pomphopsilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="289" phylum="Arthropoda" rank="genus">
<emphasis id="B946EA836828FF93BD3F07C4FBBFF997" box="[885,1091,1519,1545]" italics="true" pageId="2" pageNumber="289">POMPHOPSILLA</emphasis>
</taxonomicName>
/
<taxonomicName id="4C324D126828FF93BA1007C5FB3CF996" authorityName="Jałoszyński" authorityYear="2024" box="[1114,1216,1518,1544]" class="Insecta" family="Staphylinidae" genus="Pomphopsilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis id="B946EA836828FF93BA1007C5FB3CF996" box="[1114,1216,1518,1544]" italics="true" pageId="2" pageNumber="289">bifurcata</emphasis>
</taxonomicName>
m. /
<collectorName id="26C753476828FF93BAB307C4FA64F996" box="[1273,1432,1518,1545]" pageId="2" pageNumber="289">P. Jałoszyński</collectorName>
, '24 / HOLOTYPUS” [red, printed] (
<collectionCode id="ED23AE546828FF93BC710439FD6BF9B2" box="[571,663,1554,1580]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="289" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5A3CCC6828FF93BCE70439FF06F9EA" ID-GBIF-Occurrence="4899212303" collectionCode="MHNG" collectorName="S. Vit" country="Togo" location="Foret de Klouto" pageId="2" pageNumber="289" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B946EA836828FF93BCE70439FCD9F9B2" bold="true" box="[685,805,1554,1580]" pageId="2" pageNumber="289">
<typeStatus id="548988336828FF93BCE70439FCD9F9B2" box="[685,805,1554,1580]" pageId="2" pageNumber="289" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 exx):
<specimenCount id="9D34FD186828FF93BDDF0439FC25F9B3" box="[917,985,1554,1581]" count="2" pageId="2" pageNumber="289" type="male">2 ♂♂</specimenCount>
, same data as for holotype except that “Klouto” is crossed out and replaced with handwritten “Kpimé”; paratypes with yellow “PARATYPUS” labels (cPJ,
<collectionCode id="ED23AE546828FF93BEDD0471FF0EF9EA" box="[151,242,1626,1652]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="289" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651A6828FF93BE8D0455FEDAF89A" pageId="2" pageNumber="289" type="diagnosis">
<paragraph id="8B8D36916828FF93BE8D0455FEDAF89A" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="289">
<emphasis id="B946EA836828FF93BE8D0455FEBEF906" bold="true" box="[199,322,1662,1688]" pageId="2" pageNumber="289">Diagnosis.</emphasis>
Body (
<figureCitation id="13092A146828FF93BFD10455FE1DF906" box="[411,481,1662,1688]" captionStart="FIGURES 13" captionStartId="1.[151,264,1977,2001]" captionTargetBox="[157,1430,1132,1948]" captionTargetId="figure-189@1.[151,1436,1126,1954]" captionTargetPageId="1" captionText="FIGURES 13. Dorsal habitus of holotype males. Pomphopsilla spinosa sp. n. (1); P. bifurcata sp. n. (2); P. klotoensis sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.12190029" httpUri="https://zenodo.org/record/12190029/files/figure.png" pageId="2" pageNumber="289">Fig. 2</figureCitation>
) light brown; pronotum and elytra with inconspicuously fine punctures; pronotum with five antebasal pits: two pairs of circular and small lateral pits and slightly transverse median pit; aedeagus in ventral view (
<figureCitation id="13092A146828FF93BF1E04EDFE64F97F" box="[340,408,1734,1761]" captionStart="FIGURES 46" captionStartId="3.[151,264,1648,1672]" captionTargetBox="[155,1432,925,1618]" captionTargetId="figure-272@3.[151,1436,921,1625]" captionTargetPageId="3" captionText="FIGURES 46. Aedeagus in ventral view. Pomphopsilla spinosa sp. n. (4); P. bifurcata sp. n. (5); P. klotoensis sp. n. (6)." figureDoi="http://doi.org/10.5281/zenodo.12190031" httpUri="https://zenodo.org/record/12190031/files/figure.png" pageId="2" pageNumber="289">Fig. 5</figureCitation>
) with deeply bifurcate apex forming lateral lobes and with weakly sclerotized and simple endophallus.
</paragraph>
</subSubSection>
<subSubSection id="C328651A6828FF92BE8D0525FE62FD6F" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289" type="description">
<paragraph id="8B8D36916828FF93BE8D0525FAB8F8B7" blockId="2.[151,1437,1482,2013]" box="[199,1348,1806,1833]" pageId="2" pageNumber="289">
<emphasis id="B946EA836828FF93BE8D0525FEA5F8B6" bold="true" box="[199,345,1806,1832]" pageId="2" pageNumber="289">Description.</emphasis>
Body of male (
<figureCitation id="13092A146828FF93BC4D0525FDB7F8B6" box="[519,587,1806,1832]" captionStart="FIGURES 13" captionStartId="1.[151,264,1977,2001]" captionTargetBox="[157,1430,1132,1948]" captionTargetId="figure-189@1.[151,1436,1126,1954]" captionTargetPageId="1" captionText="FIGURES 13. Dorsal habitus of holotype males. Pomphopsilla spinosa sp. n. (1); P. bifurcata sp. n. (2); P. klotoensis sp. n. (3)." figureDoi="http://doi.org/10.5281/zenodo.12190029" httpUri="https://zenodo.org/record/12190029/files/figure.png" pageId="2" pageNumber="289">Fig. 2</figureCitation>
) strongly elongate, weakly convex, light brown. BL
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.
</paragraph>
<paragraph id="8B8D36916828FF92BE8D0519FB4DFF4B" blockId="2.[151,1437,1482,2013]" lastBlockId="3.[151,1437,150,897]" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289">
Head broadest at large, strongly convex and coarsely faceted eyes, HL
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, HW
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; vertex strongly transverse, weakly convex, anteriorly confluent with subtriangular and similarly weakly convex frons; supraantennal tubercles small but distinct; punctures on frons and vertex fine and inconspicuous; setae very short and sparse, recumbent; frontal glands present, each situated behind supraantennal tubercle and near mesal margin of eye. Antenna moderately long, with abruptly broadened club composed of two antennomeres, AnL
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, antennomeres 1 and 2 each elongate, 3 indistinctly transverse, 4 as long as broad, 59 each indistinctly elongate, 10 indistinctly transverse, 11 slightly broader and slightly longer than 10, about as long as broad.
</paragraph>
<paragraph id="8B8D36916829FF92BE8D02F4FE9FFE4F" blockId="3.[151,1437,150,897]" pageId="3" pageNumber="290">
Pronotum subrectangular with broadly rounded anterior corners, broadest near anterior fourth; PL
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, PW
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.Anterior margin nearly straight; anterior corners in strictly dorsal view not visible; lateral margins rounded in anterior third and nearly straight in posterior half, indistinctly microserrate; posterior corners obtuse-angled; posterior margin with narrow and rounded emargination in front of mesoscutellum, sides of posterior margin barely noticeably concave; pronotal base with five pits: inner and outer lateral pairs of circular and similarly small pits, and with slightly transverse distinct median pit. Punctures on pronotal disc inconspicuously fine; setae short, moderately dense, recumbent.
</paragraph>
<paragraph id="8B8D36916829FF92BE8D03F0FAECFD87" blockId="3.[151,1437,150,897]" pageId="3" pageNumber="290">
Elytra oval, broadest slightly anterior to middle; EL
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, EW
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, EI 1.331.42; humeral calli small but distinctly elevated, slightly elongate. Punctures and setae similar as those on pronotum.
</paragraph>
<paragraph id="8B8D36916829FF92BE8D0008FDFEFDA3" blockId="3.[151,1437,150,897]" box="[199,514,547,573]" pageId="3" pageNumber="290">Hind wings well developed.</paragraph>
<paragraph id="8B8D36916829FF92BE8D006CFD24FDFF" blockId="3.[151,1437,150,897]" box="[199,728,583,609]" pageId="3" pageNumber="290">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="8B8D36916829FF92BE8D0040FC24FD53" blockId="3.[151,1437,150,897]" pageId="3" pageNumber="290">
Aedeagus (
<figureCitation id="13092A146829FF92BF0D0040FE77FD1B" box="[327,395,618,645]" captionStart="FIGURES 46" captionStartId="3.[151,264,1648,1672]" captionTargetBox="[155,1432,925,1618]" captionTargetId="figure-272@3.[151,1436,921,1625]" captionTargetPageId="3" captionText="FIGURES 46. Aedeagus in ventral view. Pomphopsilla spinosa sp. n. (4); P. bifurcata sp. n. (5); P. klotoensis sp. n. (6)." figureDoi="http://doi.org/10.5281/zenodo.12190031" httpUri="https://zenodo.org/record/12190031/files/figure.png" pageId="3" pageNumber="290">Fig. 5</figureCitation>
) stout, AeL
<quantity id="4CCA9B746829FF92BC510040FD78FD1B" box="[539,644,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="290" unit="mm" value="0.1">0.10 mm</quantity>
; median lobe in ventral view drop-shaped with deeply bifurcate apex forming a fishtail-like structure; endophallus weakly sclerotized, with median tubular structure; parameres slender, their apices not reaching apex of aedeagus, each with one long apical seta.
</paragraph>
<paragraph id="8B8D36916829FF92BE8D00FCFE62FD6F" blockId="3.[151,1437,150,897]" box="[199,414,727,753]" pageId="3" pageNumber="290">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C328651A6829FF92BE8D00D1FD92FC8B" box="[199,622,762,789]" pageId="3" pageNumber="290" type="distribution">
<paragraph id="8B8D36916829FF92BE8D00D1FD92FC8B" blockId="3.[151,1437,150,897]" box="[199,622,762,789]" pageId="3" pageNumber="290">
<emphasis id="B946EA836829FF92BE8D00D1FE9DFC8A" bold="true" box="[199,353,762,788]" pageId="3" pageNumber="290">Distribution.</emphasis>
Southern
<collectingCountry id="F32576016829FF92BF9900D0FDF0FC8B" box="[467,524,763,789]" name="Togo" pageId="3" pageNumber="290">Togo</collectingCountry>
(
<figureCitation id="13092A146829FF92BC5100D0FD9CFC8B" box="[539,608,763,789]" captionStart="FIGURE 7" captionStartId="4.[151,250,1654,1678]" captionTargetBox="[157,1435,525,1623]" captionTargetId="figure-176@4.[151,1435,525,1631]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of nominal Pomphopsilla species, with aedeagus of each species in ventral view (P. felicitas is known from a single female, and P. crenata is known from a single male with a strongly distorted aedeagus)." figureDoi="http://doi.org/10.5281/zenodo.12190035" httpUri="https://zenodo.org/record/12190035/files/figure.png" pageId="3" pageNumber="290">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328651A6829FF92BE8D0134FAF9FCA7" box="[199,1285,799,825]" pageId="3" pageNumber="290" type="etymology">
<paragraph id="8B8D36916829FF92BE8D0134FAF9FCA7" blockId="3.[151,1437,150,897]" box="[199,1285,799,825]" pageId="3" pageNumber="290">
<emphasis id="B946EA836829FF92BE8D0134FEB2FCA7" bold="true" box="[199,334,799,825]" pageId="3" pageNumber="290">Etymology.</emphasis>
The specific adjective
<taxonomicName id="4C324D126829FF92BC1B0134FD4BFCA7" authorityName="Jałoszyński" authorityYear="2024" box="[593,695,799,825]" class="Insecta" family="Staphylinidae" genus="Pomphopsilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis id="B946EA836829FF92BC1B0134FD4BFCA7" box="[593,695,799,825]" italics="true" pageId="3" pageNumber="290">bifurcata</emphasis>
</taxonomicName>
refers to the unusually shaped apex of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C328651A6829FF92BE8D0168FDB6FC1F" pageId="3" pageNumber="290" type="discussion">
<paragraph id="8B8D36916829FF92BE8D0168FDB6FC1F" blockId="3.[151,1437,150,897]" pageId="3" pageNumber="290">
<emphasis id="B946EA836829FF92BE8D0168FEC7FCC3" bold="true" box="[199,315,835,861]" pageId="3" pageNumber="290">Remarks.</emphasis>
This species is externally unremarkable, but the bifurcate aedeagal apex is a unique feature and allows for unambiguous identification.
</paragraph>
</subSubSection>
</treatment>
</document>