654 lines
113 KiB
XML
654 lines
113 KiB
XML
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<mods:title id="11A51AD5B99CC4FD4C7FAB82587CF28E">On the systematic placement of Montezumellidae Ossó & Domínguez, 2013 (Decapoda: Brachyura: Eubrachyura)</mods:title>
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<mods:namePart id="58F4F524A9655FCED678BBDE66B5D277">Ossó, Àlex</mods:namePart>
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<mods:namePart id="1DCE0F9B1F9420C0A829ED96D78CE413">Domínguez, José Luis</mods:namePart>
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<mods:title id="B68C9CB56B1F27192213A0C68FF87C97">Zootaxa</mods:title>
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<mods:date id="3B2AD2CA8B279A0C5B05405956D8EBC0">2019</mods:date>
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<subSubSection id="C32B6503FFF67050FF49FC47FD0BCE8F" box="[151,690,981,1008]" pageId="2" pageNumber="177" type="nomenclature">
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<paragraph id="8B8E3688FFF67050FF49FC47FD0BCE8F" blockId="2.[151,690,981,1008]" box="[151,690,981,1008]" pageId="2" pageNumber="177">
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<heading id="D0C681E4FFF67050FF49FC47FD0BCE8F" bold="true" box="[151,690,981,1008]" fontSize="11" level="1" pageId="2" pageNumber="177" reason="1">
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<taxonomicName id="4C314D0BFFF67050FF49FC47FD0BCE8F" ID-CoL="8ee8962f-7f88-44c7-bcc8-4dc97de0b26a" authority="Osso & Dominguez, 2013" authorityName="Osso & Dominguez" authorityYear="2013" box="[151,690,981,1008]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="177" phylum="Arthropoda" rank="family">
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<emphasis id="B945EA9AFFF67050FF49FC47FD0BCE8F" bold="true" box="[151,690,981,1008]" pageId="2" pageNumber="177">
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Montezumellidae
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<bibRefCitation id="EFA04B79FFF67050FEA9FC47FD0BCE8F" author="Osso, A. & Dominguez, J. L." box="[375,690,981,1008]" pageId="2" pageNumber="177" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez, 2013</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6503FFF67051FF49FB8FFC18CCEB" lastPageId="3" lastPageNumber="178" pageId="2" pageNumber="177" type="diagnosis">
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<paragraph id="8B8E3688FFF67050FF49FB8FFC91C8A3" blockId="2.[151,1437,1053,2016]" pageId="2" pageNumber="177">
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<emphasis id="B945EA9AFFF67050FF49FB8FFE2FC958" bold="true" box="[151,406,1053,1080]" pageId="2" pageNumber="177">Diagnosis (emended).</emphasis>
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Carapace from small/medium size (
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<taxonomicName id="4C314D0BFFF67050FCF0FB8FFC74C957" authorityName="Rathbun" authorityYear="1930" box="[814,973,1053,1079]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">
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<emphasis id="B945EA9AFFF67050FCF0FB8FFC74C957" box="[814,973,1053,1079]" italics="true" pageId="2" pageNumber="177">Montezumella</emphasis>
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</taxonomicName>
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) to large size (
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<taxonomicName id="4C314D0BFFF67050FBABFB8FFB51C957" box="[1141,1256,1053,1079]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">
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<emphasis id="B945EA9AFFF67050FBABFB8FFB51C957" box="[1141,1256,1053,1079]" italics="true" pageId="2" pageNumber="177">Moianella</emphasis>
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</taxonomicName>
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), from elongate to ovate subhexagonal shape, from slightly longer than wide to as wide as long, almost flattened; surface rugose, with coarse granules, and rows of granules forming short ridges of squamous aspect in the posterior half. Dorsal regions slightly swollen, well defined by shallow grooves, usually smooth. Hepatic region slightly swollen. Protogastric lobes swollen, anterior half of lobes both subdivided longitudinally by a median groove forming convergent ridges, that extend to the four frontal lobes or teeth; mesogastric region large, narrowing distally; metagastric region inflated; urogastric region narrow. Metagastric and cardiac regions well delimited transversally, and bounded laterally by a groove connecting the gastro-hepatic and cervical groove to the branchiocardiac groove. Cardiac region large. Intestinal region flat. Epibranchial lobes inflated. Mesobranchial and metabranchial regions slightly inflated. Front wide, bilobed, each lobe with two teeth or spines, axially divided by a notch. Orbits oblique, oval, complete; supra-orbital margin with three teeth separated by two fissures; serrated orbital corner continuous with the infraorbital margin which bears a prominent infra-orbital tooth.
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</paragraph>
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<paragraph id="8B8E3688FFF67050FF19FA5FFC51CB33" blockId="2.[151,1437,1053,2016]" pageId="2" pageNumber="177">Anterolateral margin short, straight to slightly convex, with four teeth or spines (excluding the outer orbital tooth) forward directed, subequal or decreasing in size. Posterolateral margin straight to convex, unarmed, stepped, edge decorated with tubercles or ridges; marked re-entrant of fifth pereiopod; branchiostegite not completely folded ventrally. Posterior margin from straight to slightly concave, short, rimmed.</paragraph>
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<paragraph id="8B8E3688FFF67050FF19F9CFFBD1CABF" blockId="2.[151,1437,1053,2016]" pageId="2" pageNumber="177">Antennular fossa deep, transverse, subtrapezoidal; basal article folded transversely. Basal article of antenna 2+3 subrectangular, elongate, directed obliquely between infraorbital tooth and antennular basal article. Proepistome thick, bluntly triangular. Ischium of third maxilliped with convex inner margin; merus subtrapezoidal. Male thoracic sternum narrow, elongate, metamerised. Suture 1/2 not recognizable, 3/4 only present laterally; sutures 2/3 and 4/5 to 7/8 complete; episternites well developed, delineated by complete sulci. Sternites 1 and 2 subtriangular; sternite 3 short, gynglyme (socket) receiving the coxo-sternal condyle of the mxp3 wide; sternite 4 elongate; sternites 5, 6, 7 subtrapezoidal. Median line present along the sternite 7 and extending until sternite 4. Press-button on sternite 5 close to suture 5/6. Sterno-pleonal cavity deep, narrow, V-shaped, reaching the half of sternite 4. Male pleon narrow, with all free somites and telson; somites 1, 2 not folded ventrally and in dorsal position; somites rectangular, somite 6 elongate. Female thoracic sternum larger than in males. Vulvae small, located inside deep sterno-pleonal cavity, on thoracic sternite 6 close to suture 5/6, completely covered by wide pleonal somite 6.</paragraph>
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<paragraph id="8B8E3688FFF77051FF19FF0BFC18CCEB" blockId="3.[151,1437,153,647]" pageId="3" pageNumber="178">Chelipeds strong, slightly heterochelous in males and females. Merus robust, not fused with basis-ischium, subtriangular in section, inner surface concave, outer surface granulated and ridged. Carpus strong, massive, densely granulated, upper margin with two forward directed spines on inner and outer angles. Propodus strong, upper margin strongly tuberculated or spiny, outer margin convex, with two marked transverse spiny keels, the second one reaching the tip of pollex; dactylus curved, as long as propodus length, proximal third of upper margin spiny; both, pollex and dactylus almost homodontous, with central tooth of cutting edge slightly more developed. Ambulatory legs long, robust, sub-oval in section; dactylus long, acute lanceolate.</paragraph>
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</subSubSection>
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<subSubSection id="C32B6503FFF77054FF19FE07FCCACABF" lastPageId="6" lastPageNumber="181" pageId="3" pageNumber="178" type="discussion">
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<paragraph id="8B8E3688FFF77051FF19FE07FCD8CC97" blockId="3.[151,1437,153,647]" pageId="3" pageNumber="178">
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<emphasis id="B945EA9AFFF77051FF19FE07FE82CCCF" bold="true" box="[199,315,405,431]" pageId="3" pageNumber="178">Remarks.</emphasis>
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The new specimen of
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<taxonomicName id="4C314D0BFFF77051FD9BFE07FC71CCCF" authority="Via, 1959" authorityName="Via" authorityYear="1959" box="[581,968,404,431]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="amenosi">
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<emphasis id="B945EA9AFFF77051FD9BFE07FCF2CCCF" box="[581,843,405,431]" italics="true" pageId="3" pageNumber="178">Montezumella amenosi</emphasis>
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<bibRefCitation id="EFA04B79FFF77051FC8AFE07FC71CCCF" author="Via, L." box="[852,968,404,431]" pageId="3" pageNumber="178" pagination="331 - 402" refId="ref9906" refString="Via, L. (1959) Decapodos fosiles del Eoceno espanol. Boletin del Instituto Geologico y Minero de Espana, 70, 331 - 402." type="journal article" year="1959">Vía, 1959</bibRefCitation>
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</taxonomicName>
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, which have well-preserved cephalic appendages, and a female of
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<taxonomicName id="4C314D0BFFF77051FE17FE2BFD0ECCB3" authorityName="Osso & Dominguez" authorityYear="2013" box="[457,695,441,467]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="cervantesi">
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<emphasis id="B945EA9AFFF77051FE17FE2BFD0ECCB3" box="[457,695,441,467]" italics="true" pageId="3" pageNumber="178">Moianella cervantesi</emphasis>
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</taxonomicName>
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with well-preserved thoracic and pleonal elements, allow and, at the same time, require a review of the family
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<taxonomicName id="4C314D0BFFF77051FD4BFE4FFCE4CC97" authorityName="Osso & Dominguez" authorityYear="2013" box="[661,861,477,503]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B8E3688FFF77051FF19FD93FC97CFE7" blockId="3.[151,1437,153,647]" pageId="3" pageNumber="178">
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Although
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<taxonomicName id="4C314D0BFFF77051FEE7FD93FE44CF7B" authorityName="Osso & Dominguez" authorityYear="2013" box="[313,509,513,539]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
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shares similar thoracic sternal pattern with Cancroidea (
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<emphasis id="B945EA9AFFF77051FBA1FD90FB04CF7B" box="[1151,1213,514,539]" italics="true" pageId="3" pageNumber="178">sensu</emphasis>
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Guinot
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<emphasis id="B945EA9AFFF77051FAC5FD90FAF7CF7B" box="[1307,1358,513,539]" italics="true" pageId="3" pageNumber="178">et al</emphasis>
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. 2008;
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<bibRefCitation id="EFA04B79FFF77051FF49FDB7FE66CF5F" author="Schubart, C. D. & Reuschel, S." box="[151,479,549,575]" pageId="3" pageNumber="178" pagination="533 - 549" refId="ref9281" refString="Schubart, C. D. & Reuschel, S. (2009) A proposal for a new classification of Portunoidea and Cancroidea (Brachyura: Heterotremata) based on two independent molecular phylogenies. In: Martin, J. W., Crandall, K. A. & Felder, D. L. (Eds.), " Decapod Crustacean Phylogenetics ". Crustacean Issues, 18, pp. 533 - 549. https: // doi. org / 10.1201 / 9781420092592 - c 27" type="journal article" year="2009">Schubart and Reuschel 2009</bibRefCitation>
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), it cannot be retained in this superfamily in view of the clear differences, now confirmed by the data provided by the new specimens of
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<taxonomicName id="4C314D0BFFF77051FD22FDDBFC22CF03" authorityName="Rathbun" authorityYear="1930" box="[764,923,585,611]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="genus">
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<emphasis id="B945EA9AFFF77051FD22FDDBFC22CF03" box="[764,923,585,611]" italics="true" pageId="3" pageNumber="178">Montezumella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C314D0BFFF77051FC0DFDDBFBFFCF03" box="[979,1094,585,611]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="genus">
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<emphasis id="B945EA9AFFF77051FC0DFDDBFBFFCF03" box="[979,1094,585,611]" italics="true" pageId="3" pageNumber="178">Moianella</emphasis>
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</taxonomicName>
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. Hence, below we summarize the main differences after comparison with extant material.
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</paragraph>
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<caption id="DF4E6600FFF77051FF49F975FDF7CAB8" httpUri="https://zenodo.org/record/3253512/files/figure.png" pageId="3" pageNumber="178" startId="3.[151,250,1767,1792]" targetBox="[151,1436,669,1748]" targetPageId="3">
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<paragraph id="8B8E3688FFF77051FF49F975FDF7CAB8" blockId="3.[151,1437,1767,2008]" pageId="3" pageNumber="178">
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<emphasis id="B945EA9AFFF77051FF49F975FEB2CA60" bold="true" box="[151,267,1767,1792]" pageId="3" pageNumber="178">FIGURE 1</emphasis>
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. A:
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<taxonomicName id="4C314D0BFFF77051FEEDF97AFD3DCA60" authority="Via, 1959" authorityName="Via" authorityYear="1959" box="[307,644,1767,1792]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="amenosi">
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<emphasis id="B945EA9AFFF77051FEEDF97AFDA2CA60" box="[307,539,1768,1792]" italics="true" pageId="3" pageNumber="178">Montezumella amenosi</emphasis>
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<bibRefCitation id="EFA04B79FFF77051FDFEF97AFD3DCA60" author="Via, L." box="[544,644,1767,1792]" pageId="3" pageNumber="178" pagination="331 - 402" refId="ref9906" refString="Via, L. (1959) Decapodos fosiles del Eoceno espanol. Boletin del Instituto Geologico y Minero de Espana, 70, 331 - 402." type="journal article" year="1959">Vía, 1959</bibRefCitation>
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</taxonomicName>
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, holotype MGSB 16313, dorsal view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). B:
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<taxonomicName id="4C314D0BFFF77051FE15F89EFC00CA44" authority="Osso & Dominguez, 2012" authorityName="Osso & Dominguez" authorityYear="2012" box="[459,953,1804,1828]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="cervantesi">
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<emphasis id="B945EA9AFFF77051FE15F89EFD1ACA44" box="[459,675,1804,1828]" italics="true" pageId="3" pageNumber="178">Moianella cervantesi</emphasis>
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Ossó & Domínguez, 2012
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</taxonomicName>
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, holotype MGB 59624 dorsal view, Priabonian (late Eocene), Moià, Moianès (Catalonia). C–D:
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<taxonomicName id="4C314D0BFFF77051FDA1F8A2FCDECA28" authorityName="Via" authorityYear="1959" box="[639,871,1840,1864]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="amenosi">
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<emphasis id="B945EA9AFFF77051FDA1F8A2FCDECA28" box="[639,871,1840,1864]" italics="true" pageId="3" pageNumber="178">Montezumella amenosi</emphasis>
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</taxonomicName>
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, MGB 88019, Collbàs Fm., Bartonian (Middle Eocene), La Pobla de Claramunt, Anoia (Catalonia); C: ventral view; D; frontal view. Abbreviations, a: pleonal somite; aa: basal antennular article; ab: basal antennal article 2+3; ca: carpus; cmxp3: coxa of third maxilliped; en: endostome; ep: epistome; ex exopod of mxp3; ey: eyestalk; is: ischium of mxp3; ma: mandibles; me: merus of mxp3; mxp3: third maxilliped; pr: proepistome; ua: urinary article of antenna. Scale bar = 1cm.
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</paragraph>
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</caption>
|
||
<caption id="DF4E6600FFF07056FF49F948FB20CA8F" httpUri="https://zenodo.org/record/3253514/files/figure.png" pageId="4" pageNumber="179" startId="4.[151,250,1754,1779]" targetBox="[181,1405,166,1734]" targetPageId="4">
|
||
<paragraph id="8B8E3688FFF07056FF49F948FB20CA8F" blockId="4.[151,1437,1754,2031]" pageId="4" pageNumber="179">
|
||
<emphasis id="B945EA9AFFF07056FF49F948FEB6CB93" bold="true" box="[151,271,1754,1779]" pageId="4" pageNumber="179">FIGURE 2</emphasis>
|
||
. A:
|
||
<taxonomicName id="4C314D0BFFF07056FEE0F948FE77CB92" authorityName="Rathbun" authorityYear="1930" box="[318,462,1754,1778]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF07056FEE0F948FE77CB92" box="[318,462,1754,1778]" italics="true" pageId="4" pageNumber="179">Montezumella</emphasis>
|
||
</taxonomicName>
|
||
sp., MGB 59626, male ventral view, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). B:
|
||
<taxonomicName id="4C314D0BFFF07056FF68F96CFD19CA77" authority="Osso & Dominguez, 2013" authorityName="Osso & Dominguez" authorityYear="2013" box="[182,672,1790,1815]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="cervantesi">
|
||
<emphasis id="B945EA9AFFF07056FF68F96CFE34CA76" box="[182,397,1790,1814]" italics="true" pageId="4" pageNumber="179">Moianella cervantesi</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF07056FE4AF96CFD19CA77" author="Osso, A. & Dominguez, J. L." box="[404,672,1790,1815]" pageId="4" pageNumber="179" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, MGB 59624, dorsal view, Priabonian (late Eocene), Moià, Bages (Catalonia). C:
|
||
<taxonomicName id="4C314D0BFFF07056FE88F8B0FD0BCA5B" authority="Via, 1959" authorityName="Via" authorityYear="1959" box="[342,690,1826,1851]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="amenosi">
|
||
<emphasis id="B945EA9AFFF07056FE88F8B0FDFBCA5A" box="[342,578,1826,1850]" italics="true" pageId="4" pageNumber="179">Montezumella amenosi</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF07056FD94F8B0FD0BCA5B" author="Via, L." box="[586,690,1826,1851]" pageId="4" pageNumber="179" pagination="331 - 402" refId="ref9906" refString="Via, L. (1959) Decapodos fosiles del Eoceno espanol. Boletin del Instituto Geologico y Minero de Espana, 70, 331 - 402." type="journal article" year="1959">Vía, 1959</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, MGSB 16313, ventral view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). D:
|
||
<taxonomicName id="4C314D0BFFF07056FDFAF8D4FD0DCA3E" authorityName="Rathbun" authorityYear="1930" box="[548,692,1862,1886]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF07056FDFAF8D4FD0DCA3E" box="[548,692,1862,1886]" italics="true" pageId="4" pageNumber="179">Montezumella</emphasis>
|
||
</taxonomicName>
|
||
sp., MGB 59627, female ventral view showing the vulvae, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). E:
|
||
<taxonomicName id="4C314D0BFFF07056FDBCF8F8FD4BCAE2" authorityName="Rathbun" authorityYear="1930" box="[610,754,1898,1922]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF07056FDBCF8F8FD4BCAE2" box="[610,754,1898,1922]" italics="true" pageId="4" pageNumber="179">Montezumella</emphasis>
|
||
</taxonomicName>
|
||
sp., MGB 59629, female ventral view, with pleon, Bartonian (Middle Eocene), Huesca, Aragón (Spain). F, G:
|
||
<taxonomicName id="4C314D0BFFF07056FD87F81CFBFDCAC7" authority="Osso & Dominguez, 2013" authorityName="Osso & Dominguez" authorityYear="2013" box="[601,1092,1934,1959]" class="Malacostraca" family="Montezumellidae" genus="Moianella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="cervantesi">
|
||
<emphasis id="B945EA9AFFF07056FD87F81CFC89CAC6" box="[601,816,1934,1958]" italics="true" pageId="4" pageNumber="179">Moianella cervantesi</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF07056FCE9F81CFBFDCAC7" author="Osso, A. & Dominguez, J. L." box="[823,1092,1934,1959]" pageId="4" pageNumber="179" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, MGB 87858, female, Priabonian (Late Eocene), Moià, Moianès (Catalonia); F: ventral view, with pleon. G: close up of the press button.Abbreviations, a: pleonal somites; ml: median line; pb: press button; st: thoracic sternites; t: telson; v: vulvae. Scale bar = 1cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E6600FFF17057FF49FA79FC18CBD8" httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="5" pageNumber="180" startId="5.[151,250,1515,1540]" targetBox="[151,1436,181,1496]" targetPageId="5">
|
||
<paragraph id="8B8E3688FFF17057FF49FA79FC18CBD8" blockId="5.[151,1436,1515,1720]" pageId="5" pageNumber="180">
|
||
<emphasis id="B945EA9AFFF17057FF49FA79FEB7CB64" bold="true" box="[151,270,1515,1540]" pageId="5" pageNumber="180">FIGURE 3</emphasis>
|
||
. Extant species of Cancroidea. A, D, H
|
||
<taxonomicName id="4C314D0BFFF17057FD76FA7EFC4ACB64" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[680,1011,1515,1540]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="pagurus">
|
||
<emphasis id="B945EA9AFFF17057FD76FA7EFCF4CB64" box="[680,845,1516,1540]" italics="true" pageId="5" pageNumber="180">Cancer pagurus</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF17057FC8AFA79FC4ACB64" author="Linnaeus, C." box="[852,1011,1515,1540]" pageId="5" pageNumber="180" refId="ref7859" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Vol. 1. 10 th Edition. Impensis Direct. Laurentii Salvii, Holmiae, iii + 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F
|
||
<taxonomicName id="4C314D0BFFF17057FEF9F982FD26CB48" authority="(Olivi, 1792)" authorityName="Olivi" authorityYear="1792" baseAuthorityName="Olivi" baseAuthorityYear="1792" box="[295,671,1551,1576]" class="Malacostraca" family="Atelecyclidae" genus="Atelecyclus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="rotundatus">
|
||
<emphasis id="B945EA9AFFF17057FEF9F982FDA9CB48" box="[295,528,1552,1576]" italics="true" pageId="5" pageNumber="180">Atelecyclus rotundatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF17057FDC1F99DFD21CB48" author="Olivi, G." box="[543,664,1551,1576]" pageId="5" pageNumber="180" refId="ref8395" refString="Olivi, G. (1792) Zoologia adriatica: ossia catalogo ragionato degli animali del Golfo e delle Lagune di Venezia: preceduto da una dissertazione sull storia fisica e naturale del Golfo: e accompagnato da memorie, ed osservazioni di fisica storia naturale ed economia. G. Remondini e fl., Bassano, xxxii + 334 pp. https: // doi. org / 10.5962 / bhl. title. 60887" type="book" year="1792">Olivi, 1792</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G
|
||
<taxonomicName id="4C314D0BFFF17057FB21F982FE74CB2C" authority="(Herbst, 1783)" authorityName="Herbst" authorityYear="1783" baseAuthorityName="Herbst" baseAuthorityYear="1783" class="Malacostraca" family="Atelecyclidae" genus="Atelecyclus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="undecimdentatus">
|
||
<emphasis id="B945EA9AFFF17057FB21F982FE90CB2C" italics="true" pageId="5" pageNumber="180">Atelecyclus undecimdentatus</emphasis>
|
||
(Herbst, 1783)
|
||
</taxonomicName>
|
||
, male, MNHN-IU-2017-9091 (= MNHN-B30431). I
|
||
<taxonomicName id="4C314D0BFFF17057FC21F9A6FB72CB2C" baseAuthorityName="Herbst" baseAuthorityYear="1783" box="[1023,1227,1588,1612]" class="Malacostraca" family="Atelecyclidae" genus="Atelecyclus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="undecimdentatus">
|
||
<emphasis id="B945EA9AFFF17057FC21F9A6FB72CB2C" box="[1023,1227,1588,1612]" italics="true" pageId="5" pageNumber="180">A. undecimdentatus</emphasis>
|
||
</taxonomicName>
|
||
female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E3688FFF17054FF19F974FC90CD9B" blockId="5.[151,1436,1766,2045]" lastBlockId="6.[151,1437,153,2015]" lastPageId="6" lastPageNumber="181" pageId="5" pageNumber="180">
|
||
Members of
|
||
<taxonomicName id="4C314D0BFFF17057FE84F974FDA7CA60" authorityName="Osso & Dominguez" authorityYear="2013" box="[346,542,1766,1792]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
possess: a dorsal carapace slightly longer than wide, ornate with strong granules, tubercles and ridges, with regions well marked (
|
||
<emphasis id="B945EA9AFFF17057FD31F899FCBFCA44" box="[751,774,1803,1828]" italics="true" pageId="5" pageNumber="180">vs</emphasis>
|
||
. carapace wider than long in
|
||
<taxonomicName id="4C314D0BFFF17057FB8FF898FB7FCA44" box="[1105,1222,1802,1828]" class="Malacostraca" family="Cancridae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Cancridae</taxonomicName>
|
||
; ovate, and usually smooth in both families,
|
||
<taxonomicName id="4C314D0BFFF17057FE6FF8BCFD9ACA28" box="[433,547,1838,1864]" class="Malacostraca" family="Cancridae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Cancridae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C314D0BFFF17057FD87F8BCFD41CA29" authorityName="Ortmann" authorityYear="1893" box="[601,760,1838,1865]" class="Malacostraca" family="Atelecyclidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Atelecyclidae</taxonomicName>
|
||
); only four anterolateral spines or teeth in
|
||
<taxonomicName id="4C314D0BFFF17057FB06F8BCFA25CA28" authorityName="Osso & Dominguez" authorityYear="2013" box="[1240,1436,1838,1864]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF17057FF7FF8C1FF01CA0C" box="[161,184,1875,1900]" italics="true" pageId="5" pageNumber="180">vs</emphasis>
|
||
. higher number of anterolateral teeth in Cancroidea); bilobed front, broader in
|
||
<taxonomicName id="4C314D0BFFF17057FBE8F8C0FB43CA0C" authorityName="Osso & Dominguez" authorityYear="2013" box="[1078,1274,1874,1900]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF17057FAD5F8C1FA9BCA0C" box="[1291,1314,1875,1900]" italics="true" pageId="5" pageNumber="180">vs</emphasis>
|
||
. front narrow and uneven in Cancroidea); relatively broad and separated orbits in
|
||
<taxonomicName id="4C314D0BFFF17057FC0CF8E4FB2FCAF0" authorityName="Osso & Dominguez" authorityYear="2013" box="[978,1174,1910,1936]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="180" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF17057FB79F8E5FB07CAF0" box="[1191,1214,1911,1936]" italics="true" pageId="5" pageNumber="180">vs</emphasis>
|
||
. small and close in Cancroidea) (
|
||
<figureCitation id="130A2A0DFFF17057FEF1F808FE11CAD4" box="[303,424,1946,1972]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="5" pageNumber="180">Fig. 3A, C</figureCitation>
|
||
; also see.
|
||
<bibRefCitation id="EFA04B79FFF17057FDC6F808FD7ECAD4" author="Rathbun, M. J." box="[536,711,1946,1972]" pageId="5" pageNumber="180" pagination="1 - 609" refId="ref8852" refString="Rathbun, M. J. (1930 b) The Cancroid Crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae, and Xanthidae. United States National Museum Bulletin, 152, 1 - 609. https: // doi. org / 10.5479 / si. 03629236.152. i" type="journal article" year="1930">Rathbun 1930b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF17057FD0FF808FCD3CAD4" author="Nations, J. D." box="[721,874,1945,1972]" pageId="5" pageNumber="180" pagination="1 - 104" refId="ref8204" refString="Nations, J. D. (1975) The genus Cancer (Crustacea: Brachyura): Systematics, Biogeography and Fossil Record. Natural History Museum of Los Angeles County Science Bulletin, 23, 1 - 104." type="journal article" year="1975">Nations 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF17057FCABF808FB7ECAD4" author="Schweitzer, C. C. & Feldmann, R. M." box="[885,1223,1946,1972]" pageId="5" pageNumber="180" pagination="636 - 653" refId="ref9451" refString="Schweitzer, C. C. & Feldmann, R. M. (2000) Re-evaluation of the Cancridae Latreille, 1802 (Decapoda: Brachyura) including three new genera and three new species. Journal of Paleontology, 74 (4), 636 - 653. https: // doi. org / 10.1017 / S 0022336000032765" type="journal article" year="2000">Schweitzer & Feldmann 2000</bibRefCitation>
|
||
); antennular fossae transversely subtrapezoidal, in which the antennules are folded transversely (
|
||
<emphasis id="B945EA9AFFF17057FC28F82DFBB4CAB8" box="[1014,1037,1983,2008]" italics="true" pageId="5" pageNumber="180">vs</emphasis>
|
||
. antennular fossae subquadrate and antennules folding longitudinally or slightly obliquely in Cancroidea); antennal basal article 2+3 narrower than basal (urinary) article (
|
||
<emphasis id="B945EA9AFFF27054FE44FF08FE08CDD3" box="[410,433,154,179]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. much larger antennal basal article in Cancroidea); thick proepistome (
|
||
<emphasis id="B945EA9AFFF27054FB0FFF08FB51CDD3" box="[1233,1256,154,179]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. very narrow in Cancroidea) (
|
||
<figureCitation id="130A2A0DFFF27054FEF1FF2FFE1FCDB7" box="[303,422,189,215]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">Fig. 3D, E</figureCitation>
|
||
; also see
|
||
<bibRefCitation id="EFA04B79FFF27054FDD0FF2FFD4ACDB7" author="Rathbun, M. J." box="[526,755,189,215]" pageId="6" pageNumber="181" pagination="1 - 609" refId="ref8852" refString="Rathbun, M. J. (1930 b) The Cancroid Crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae, and Xanthidae. United States National Museum Bulletin, 152, 1 - 609. https: // doi. org / 10.5479 / si. 03629236.152. i" type="journal article" year="1930">Rathbun 1930b: 176</bibRefCitation>
|
||
, pl. 87–88;
|
||
<bibRefCitation id="EFA04B79FFF27054FCABFF2FFBD2CDB7" author="Via Boada, L & Martinell, J. & Domenech, R." box="[885,1131,189,215]" pageId="6" pageNumber="181" pagination="245 - 254" refId="ref9934" refString="Via Boada, L, Martinell, J. & Domenech, R. (1982) Presencia de Cancer sismondai, Meyer 1842 (Crustacea, Decapoda) en el Plioceno espanol. Boletin de la Real Sociedad Espanola de Historia natural, Geologia, 80, 245 - 254." type="journal article" year="1982">
|
||
Vía Boada
|
||
<emphasis id="B945EA9AFFF27054FC2CFF2CFB9BCDB7" box="[1010,1058,189,215]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 1982
|
||
</bibRefCitation>
|
||
: fig. 2d;
|
||
<bibRefCitation id="EFA04B79FFF27054FB12FF2FFF6BCD9B" author="Salva, E. W. & Feldmann, R. M." pageId="6" pageNumber="181" pagination="9 - 62" refId="ref9086" refString="Salva, E. W. & Feldmann, R. M. (2001) Reevaluation of the family Atelecyclidae (Decapoda: Brachyura). Kirtlandia, 52, 9 - 62." type="journal article" year="2001">Salva & Feldmann 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FF00FF73FE5ACD9B" author="Spiridonov, V. A. & Neretina, T. V. & Schepetov, D." box="[222,483,225,251]" pageId="6" pageNumber="181" pagination="404 - 429" refId="ref9575" refString="Spiridonov, V. A., Neretina, T. V. & Schepetov, D. (2014) Morphological characterization and molecular phylogeny of Portunoidea Rafinesque, 1815 (Crustacea Brachyura): Implications for understanding evolution of swimming capacity and revision of the family-level classification. Zoologischer Anzeiger, 253, 404 - 429. https: // doi. org / 10.1016 / j. jcz. 2014.03.003" type="journal article" year="2014">
|
||
Spiridonov
|
||
<emphasis id="B945EA9AFFF27054FEBFFF70FE2ACD9B" box="[353,403,225,251]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
., 2014
|
||
</bibRefCitation>
|
||
: fig. 2B;
|
||
<bibRefCitation id="EFA04B79FFF27054FD95FF73FCA4CD9B" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." box="[587,797,224,251]" pageId="6" pageNumber="181" pagination="1049 - 1130" refId="ref6600" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Systematics and classification of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C), (I), Decapoda: Brachyura (Part 2), 1049 - 1130. https: // doi. org / 10.1163 / 9789004190832 _ 021" type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B945EA9AFFF27054FD4BFF70FD7FCD9B" box="[661,710,225,251]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 2015a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF27054FF19FE97FBE1CF03" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="181">
|
||
Which respect to thoracic sternal and pleonal features, differences are present as well. In
|
||
<taxonomicName id="4C314D0BFFF27054FB76FE97FAD6CC7F" authorityName="Osso & Dominguez" authorityYear="2013" box="[1192,1391,261,287]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
, the suture 2/3 is complete (
|
||
<emphasis id="B945EA9AFFF27054FE41FEB8FE0FCC23" box="[415,438,298,323]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. absent in
|
||
<taxonomicName id="4C314D0BFFF27054FDEEFEBBFD1BCC23" box="[560,674,297,323]" class="Malacostraca" family="Cancridae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Cancridae</taxonomicName>
|
||
[
|
||
<figureCitation id="130A2A0DFFF27054FD6EFEBBFCB2CC23" box="[688,779,296,323]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">Fig. 3H</figureCitation>
|
||
], and well-marked but only laterally in
|
||
<taxonomicName id="4C314D0BFFF27054FB1BFEBBFAE6CC23" authorityName="Ortmann" authorityYear="1893" box="[1221,1375,297,323]" class="Malacostraca" family="Atelecyclidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Atelecyclidae</taxonomicName>
|
||
[
|
||
<figureCitation id="130A2A0DFFF27054FAB0FEBBFF07CC07" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">Fig. 3G</figureCitation>
|
||
]); suture 3/4 is only marked laterally in
|
||
<taxonomicName id="4C314D0BFFF27054FD4EFEDFFCEDCC07" authorityName="Osso & Dominguez" authorityYear="2013" box="[656,852,333,359]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF27054FCB8FEDCFCC4CC07" box="[870,893,334,359]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. almost complete in
|
||
<taxonomicName id="4C314D0BFFF27054FBB1FEDFFAB7CC08" authorityName="Ortmann" authorityYear="1893" box="[1135,1294,333,360]" class="Malacostraca" family="Atelecyclidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Atelecyclidae</taxonomicName>
|
||
); the sternopleonal cavity is deep and reaches only half of sternite
|
||
<quantity id="4CC99B6DFFF27054FCCCFEE3FC86CCEB" box="[786,831,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="181" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C314D0BFFF27054FC96FEE3FBB5CCEB" authorityName="Osso & Dominguez" authorityYear="2013" box="[840,1036,369,395]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF27054FBC0FEE0FB8CCCEB" box="[1054,1077,370,395]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. sterno-pleonal cavity reaching sternite
|
||
<quantity id="4CC99B6DFFF27054FF2EFE07FEA3CCCF" box="[240,282,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="181" unit="in" value="3.0">3 in</quantity>
|
||
Cancroidea) (
|
||
<bibRefCitation id="EFA04B79FFF27054FE69FE07FDF0CCCF" author="Guinot, D." box="[439,585,405,431]" pageId="6" pageNumber="181" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot 1979</bibRefCitation>
|
||
: pl. 9, figs. 4–7;
|
||
<bibRefCitation id="EFA04B79FFF27054FCDFFE07FC47CCCF" author="Tavares, M. & Cleva, R." box="[769,1022,405,431]" pageId="6" pageNumber="181" pagination="97 - 157" refId="ref9801" refString="Tavares, M. & Cleva, R. (2010) Trichopeltariidae (Crustacea, Decapoda, Brachyura), a new family and superfamily of eubrachyuran crabs with description of one new genus and five new species. Papeis Avulsos Zoologia, 50 (9), 97 - 157. https: // doi. org / 10.1590 / S 0031 - 10492010000900001" type="journal article" year="2010">Tavares & Cleva 2010</bibRefCitation>
|
||
: fig. 5E; Schram & Ng. 2012: fig. 3); vulvae are small in females of
|
||
<taxonomicName id="4C314D0BFFF27054FE33FE2BFD08CCB3" authorityName="Osso & Dominguez" authorityYear="2013" box="[493,689,441,467]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF27054FD1FFE28FD61CCB3" box="[705,728,442,467]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. larger in Cancroidea [
|
||
<figureCitation id="130A2A0DFFF27054FC06FE2BFB9ECCB3" box="[984,1063,440,467]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">
|
||
Fig.
|
||
<date id="FF8F1048FFF27054FBD3FE2AFB9ECCB3" box="[1037,1063,440,467]" pageId="6" pageNumber="181">3I</date>
|
||
</figureCitation>
|
||
]); pleonal somites 3, 4, 5 are free in males of
|
||
<taxonomicName id="4C314D0BFFF27054FEC1FE4FFE5ACC97" authorityName="Osso & Dominguez" authorityYear="2013" box="[287,483,477,503]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF27054FE2AFE4CFDB2CC97" box="[500,523,478,503]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. fused in Cancroidea [
|
||
<figureCitation id="130A2A0DFFF27054FCD3FE4FFC36CC97" box="[781,911,476,503]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">Fig. 3G, H</figureCitation>
|
||
]); the pleonal somites 1, 2 are not folded ventrally, thus exposed in dorsal position, in
|
||
<taxonomicName id="4C314D0BFFF27054FDBCFD93FC9FCF7B" authorityName="Osso & Dominguez" authorityYear="2013" box="[610,806,513,539]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
(
|
||
<emphasis id="B945EA9AFFF27054FCE8FD90FCF4CF7B" box="[822,845,514,539]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. ventrally folded in
|
||
<taxonomicName id="4C314D0BFFF27054FBEEFD93FB1BCF7B" box="[1072,1186,513,539]" class="Malacostraca" family="Cancridae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Cancridae</taxonomicName>
|
||
[e.g.:
|
||
<bibRefCitation id="EFA04B79FFF27054FB37FD93FA21CF7B" author="Rathbun, M. J." box="[1257,1432,513,539]" pageId="6" pageNumber="181" pagination="1 - 609" refId="ref8852" refString="Rathbun, M. J. (1930 b) The Cancroid Crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae, and Xanthidae. United States National Museum Bulletin, 152, 1 - 609. https: // doi. org / 10.5479 / si. 03629236.152. i" type="journal article" year="1930">Rathbun 1930b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FF49FDB7FE73CF5F" author="Bonfiglio, L. & Donadeo, G." box="[151,458,548,575]" pageId="6" pageNumber="181" pagination="255 - 296" refId="ref6396" refString="Bonfiglio, L. & Donadeo, G. (1982) Cancer sismondai Meyer nel Pliocene di Torre Dell'Orso (Puglia). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 123, 255 - 296." type="journal article" year="1982">Bonfiglio & Donadeo 1982</bibRefCitation>
|
||
] and in
|
||
<taxonomicName id="4C314D0BFFF27054FDC0FDB7FD01CF5F" authorityName="Ortmann" authorityYear="1893" box="[542,696,549,575]" class="Malacostraca" family="Atelecyclidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Atelecyclidae</taxonomicName>
|
||
[
|
||
<bibRefCitation id="EFA04B79FFF27054FD1AFDB7FC7BCF5F" author="Tavares, M. & Cleva, R." box="[708,962,549,575]" pageId="6" pageNumber="181" pagination="97 - 157" refId="ref9801" refString="Tavares, M. & Cleva, R. (2010) Trichopeltariidae (Crustacea, Decapoda, Brachyura), a new family and superfamily of eubrachyuran crabs with description of one new genus and five new species. Papeis Avulsos Zoologia, 50 (9), 97 - 157. https: // doi. org / 10.1590 / S 0031 - 10492010000900001" type="journal article" year="2010">Tavares & Cleva 2010</bibRefCitation>
|
||
, fig. 3A;
|
||
<figureCitation id="130A2A0DFFF27054FBF9FDB7FB25CF5F" box="[1063,1180,548,575]" captionStart="FIGURE 3" captionStartId="5.[151,250,1515,1540]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure@5.[151,1436,181,1496]" captionTargetPageId="5" captionText="FIGURE 3. Extant species of Cancroidea. A, D, H Cancer pagurus Linnaeus, 1758, male MNHN-IU-2017–9094 (= MNHN- B6388). B, F Atelecyclus rotundatus (Olivi, 1792), male MNHN-IU-2017–9085 (= MNHN-B5424). C, E, G Atelecyclus undecimdentatus (Herbst, 1783), male, MNHN-IU-2017-9091 (= MNHN-B30431). I A. undecimdentatus female MNHN-IU- 2017–9092 (= MNHN-B5432). Abbreviations. a: pleonal somites; aa: basal antennular article; ab: basal antennal article 2+3; a: pleonal somites; ep: epistome; ex: exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml: median line; pb: press button; pr: proepistome; st: thoracic sternites; t: telson; v: vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253516/files/figure.png" pageId="6" pageNumber="181">Fig. 3B, C</figureCitation>
|
||
]) (e.g.
|
||
<bibRefCitation id="EFA04B79FFF27054FB34FDB7FA21CF5F" author="Rathbun, M. J." box="[1258,1432,549,575]" pageId="6" pageNumber="181" pagination="1 - 609" refId="ref8852" refString="Rathbun, M. J. (1930 b) The Cancroid Crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae, and Xanthidae. United States National Museum Bulletin, 152, 1 - 609. https: // doi. org / 10.5479 / si. 03629236.152. i" type="journal article" year="1930">Rathbun 1930b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FF49FDDBFE9ECF03" author="Guinot, D." box="[151,295,585,611]" pageId="6" pageNumber="181" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FEEDFDDBFD3FCF03" author="Schweitzer, C. C. & Feldmann, R. M." box="[307,646,585,611]" pageId="6" pageNumber="181" pagination="636 - 653" refId="ref9451" refString="Schweitzer, C. C. & Feldmann, R. M. (2000) Re-evaluation of the Cancridae Latreille, 1802 (Decapoda: Brachyura) including three new genera and three new species. Journal of Paleontology, 74 (4), 636 - 653. https: // doi. org / 10.1017 / S 0022336000032765" type="journal article" year="2000">Schweitzer & Feldmann 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FD4FFDDBFCD6CF03" author="Schram, F. R. & Ng, P. K. L." box="[657,879,585,611]" pageId="6" pageNumber="181" pagination="665 - 672" refId="ref9233" refString="Schram, F. R. & Ng, P. K. L. (2012) What is Cancer? Journal of Crustacean Biology, 32 (4), 665 - 672. https: // doi. org / 10.1163 / 193724012 X 640650" type="journal article" year="2012">Schram & Ng 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FCA4FDDBFBF5CF03" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." box="[890,1100,584,611]" pageId="6" pageNumber="181" pagination="1049 - 1130" refId="ref6600" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Systematics and classification of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C), (I), Decapoda: Brachyura (Part 2), 1049 - 1130. https: // doi. org / 10.1163 / 9789004190832 _ 021" type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B945EA9AFFF27054FC1AFDD8FC4CCF03" box="[964,1013,585,611]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 2015a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF27054FF19FDFFFDECCFCB" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="181">
|
||
The morphological differences cited here, therefore are deemed enough to remove
|
||
<taxonomicName id="4C314D0BFFF27054FB81FDFFFA9ACFE7" authorityName="Osso & Dominguez" authorityYear="2013" box="[1119,1315,621,647]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
from Cancroidea and raise it to superfamily rank.
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF27054FF19FD27FB81CEE4" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="181">
|
||
The differences between Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FD65FD27FCF2CFAF" bold="true" box="[699,843,692,719]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
. and apparently related taxa, are summarized herein, making emphasis on the new characters now available, such as the cephalic appendages. Formerly, in introducing
|
||
<taxonomicName id="4C314D0BFFF27054FF49FD6FFEE2CE77" authorityName="Osso & Dominguez" authorityYear="2013" box="[151,347,765,791]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
as a new family,
|
||
<bibRefCitation id="EFA04B79FFF27054FDC1FD6FFCEBCE77" author="Osso, A. & Dominguez, J. L." box="[543,850,765,791]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez (2013)</bibRefCitation>
|
||
, largely discussed the traits shared with closest relatives and listed its clearly distinguishable characters. Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FC1BFCB3FBECCE5B" bold="true" box="[965,1109,800,827]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
. shares with the families and genera grouped by
|
||
<bibRefCitation id="EFA04B79FFF27054FEB5FCD7FDECCE3F" author="Guinot, D." box="[363,597,837,863]" pageId="6" pageNumber="181" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot (1979: 81–86</bibRefCitation>
|
||
, table 1), higher level characters such as the thoracic sternum with complete sutures 4/5 to 7/8 (see
|
||
<bibRefCitation id="EFA04B79FFF27054FE4AFCFBFC9CCEE3" author="Osso, A. & Dominguez, J. L." box="[404,805,873,899]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez 2013: 289–293</bibRefCitation>
|
||
, and references therein).
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF27054FF19FC1FFA89CA67" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="181">
|
||
An array of differences amongst
|
||
<taxonomicName id="4C314D0BFFF27054FDE5FC1FFD67CEC7" authorityName="Ortmann" authorityYear="1893" box="[571,734,909,935]" class="Malacostraca" family="Cheiragonidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Cheiragonidae</taxonomicName>
|
||
(Cheiragonoidea
|
||
<bibRefCitation id="EFA04B79FFF27054FC77FC1FFBEACEC7" author="Ortmann, A. E." box="[937,1107,909,935]" pageId="6" pageNumber="181" pagination="411 - 495" refId="ref8472" refString="Ortmann, A. E. (1893) Die Decapoden-Krebse des Strassburger Museum. VII. Theil. Abtheilung: Brachyura (Brachyura genuina Boas) II. Unterabtheilung: Cancroidea, 2, Section: Cancrinea, I. Gruppe: Cyclometopa. Zoologischen Jahrbuchern, Abtheilung fur Systematic, Geographie und Biologie der Thiere, 7, 411 - 495. https: // doi. org / 10.5962 / bhl. part. 24064" type="journal article" year="1893">Ortmann, 1893</bibRefCitation>
|
||
[
|
||
<emphasis id="B945EA9AFFF27054FBBAFC1CFB1BCEC7" box="[1124,1186,910,935]" italics="true" pageId="6" pageNumber="181">sensu</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF27054FB74FC1FFAE9CEC7" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1194,1360,908,935]" pageId="6" pageNumber="181" pagination="1 - 286" refId="ref8316" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286. [updates at http: // rmbr. nus. edu. sg / research / cotw / supplement 17. php]" type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="B945EA9AFFF27054FB0AFC1CFABFCEC7" box="[1236,1286,909,935]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
]) (
|
||
<figureCitation id="130A2A0DFFF27054FAB1FC1FFF1ECEAB" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">Fig. 4</figureCitation>
|
||
),
|
||
<taxonomicName id="4C314D0BFFF27054FF63FC23FEE5CEAB" authorityName="Rathbun" authorityYear="1930" box="[189,348,945,971]" class="Malacostraca" family="Cheiragonidae" genus="Montezumella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF27054FF63FC23FEE5CEAB" box="[189,348,945,971]" italics="true" pageId="6" pageNumber="181">Montezumella</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C314D0BFFF27054FE45FC23FDDBCEAB" authorityName="Osso & Dominguez" authorityYear="2013" box="[411,610,945,971]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
, was already listed by
|
||
<bibRefCitation id="EFA04B79FFF27054FCBDFC23FAB2CEAB" author="Osso, A. & Dominguez, J. L." box="[867,1291,945,971]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez (2013: 289–292)</bibRefCitation>
|
||
. In any case, the most distinctive character that excludes members of Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FC33FC47FBC4CE8F" bold="true" box="[1005,1149,980,1007]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
. from Cheiragonoidea, is that the female of Cheiragonoidea possess exposed vulvae located laterally on the sternite 6 (
|
||
<figureCitation id="130A2A0DFFF27054FB7FFC6BFAA3C973" box="[1185,1306,1017,1043]" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">Fig. 4A, B</figureCitation>
|
||
; also
|
||
<bibRefCitation id="EFA04B79FFF27054FA87FC6BFE82C957" author="Benedict, J. E." pageId="6" pageNumber="181" pagination="223 - 230" refId="ref6350" refString="Benedict, J. E. (1892) Corystoid crabs of the genera Telmessus and Erimacrus. Proceedings of the United States National Museum, 15, 223 - 230. https: // doi. org / 10.5479 / si. 00963801.15 - 900.223" type="journal article" year="1892">Benedict 1892: 223</bibRefCitation>
|
||
, pl. 26, figs. 3, 6;
|
||
<bibRefCitation id="EFA04B79FFF27054FDDEFB8FFD7FC957" author="Guinot, D." box="[512,710,1053,1079]" pageId="6" pageNumber="181" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot 1979: 178</bibRefCitation>
|
||
, pl. 25, figs. 1–3; 1993: 1232;
|
||
<bibRefCitation id="EFA04B79FFF27054FBC7FB8EFB14C957" author="Stevcic, Z." box="[1049,1197,1052,1079]" pageId="6" pageNumber="181" pagination="1 - 14" refId="ref9648" refString="Stevcic, Z. (1988) The status of the family Cheiragonidae Ortmann, 1893. Oebalia, 14, New Series, 1 - 14. https: // doi. org / 10.1111 / j. 1752 - 0606.1988. tb 01645. x" type="book chapter" year="1988">Števčić 1988</bibRefCitation>
|
||
; Schweitzer & Salva, 2000: 285;
|
||
<bibRefCitation id="EFA04B79FFF27054FEC9FBD3FD19C93B" author="Karasawa, H. & Schweitzer, C. E." box="[279,672,1089,1115]" pageId="6" pageNumber="181" pagination="143 - 154" refId="ref7524" refString="Karasawa, H. & Schweitzer, C. E. (2004) Revision of the genus Glyphithyreus Reuss, 1859 (Crustacea, Decapoda, Brachyura: Xanthoidea) and recognition of a new genus: Paleontological Research, 8 (3), 143 - 154. https: // doi. org / 10.2517 / prpsj. 8.143" type="journal article" year="2004">Karasawa & Schweitzer 2004: 144</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FD75FBD3FC1DC93B" author="Guinot, D. & Tavares, M. & Castro, P." box="[683,932,1089,1115]" pageId="6" pageNumber="181" pagination="1 - 414" refId="ref7280" refString="Guinot, D., Tavares, M. & Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
|
||
Guinot
|
||
<emphasis id="B945EA9AFFF27054FCDEFBD0FC88C93B" box="[768,817,1089,1115]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 2013: 28
|
||
</bibRefCitation>
|
||
, fig. 3). By contrast, in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FA58FBD3FEB0C91F" bold="true" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
., the vulvae are located inside the sterno-pleonal cavity and are completely covered by the pleon, as usual in Eubrachyura (
|
||
<figureCitation id="130A2A0DFFF27054FE8FFB1BFE5CC9C3" box="[337,485,1161,1187]" captionStart="FIGURE 2" captionStartId="4.[151,250,1754,1779]" captionTargetBox="[181,1405,166,1734]" captionTargetId="figure@4.[181,1405,166,1734]" captionTargetPageId="4" captionText="FIGURE 2. A: Montezumella sp., MGB 59626, male ventral view, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). B: Moianella cervantesi Ossó & Domínguez, 2013, holotype, MGB 59624, dorsal view, Priabonian (late Eocene), Moià, Bages (Catalonia). C: Montezumella amenosi Vía, 1959, holotype, MGSB 16313, ventral view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). D: Montezumella sp., MGB 59627, female ventral view showing the vulvae, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). E: Montezumella sp., MGB 59629, female ventral view, with pleon, Bartonian (Middle Eocene), Huesca, Aragón (Spain). F, G: Moianella cervantesi Ossó & Domínguez, 2013, MGB 87858, female, Priabonian (Late Eocene), Moià, Moianès (Catalonia); F: ventral view, with pleon. G: close up of the press button.Abbreviations, a: pleonal somites; ml: median line; pb: press button; st: thoracic sternites; t: telson; v: vulvae. Scale bar = 1cm." httpUri="https://zenodo.org/record/3253514/files/figure.png" pageId="6" pageNumber="181">Fig. 2D, E, F</figureCitation>
|
||
; also
|
||
<bibRefCitation id="EFA04B79FFF27054FDFDFB1BFCFBC9C3" author="Osso, A. & Dominguez, J. L." box="[547,834,1161,1187]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez 2013</bibRefCitation>
|
||
: fig. 5 b, b’, c–e). In Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FADEFB1BFA36C9C3" bold="true" box="[1280,1423,1160,1187]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
., the somites 1, 2 are in dorsal position, whereas in
|
||
<taxonomicName id="4C314D0BFFF27054FD14FB3FFCD4C9A7" authorityName="Ortmann" authorityYear="1893" box="[714,877,1197,1223]" class="Malacostraca" family="Cheiragonidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Cheiragonidae</taxonomicName>
|
||
the pleon appears only partially exposed dorsally, with only the first somite appearing in dorsal view (at least the somite 1 plus the entire somite 2 or only a part), thus the pleon is also incompletely folded, but to a lesser extent (see
|
||
<bibRefCitation id="EFA04B79FFF27054FCB4FB67FBA8C86F" author="Benedict, J. E." box="[874,1041,1269,1295]" pageId="6" pageNumber="181" pagination="223 - 230" refId="ref6350" refString="Benedict, J. E. (1892) Corystoid crabs of the genera Telmessus and Erimacrus. Proceedings of the United States National Museum, 15, 223 - 230. https: // doi. org / 10.5479 / si. 00963801.15 - 900.223" type="journal article" year="1892">Benedict 1892</bibRefCitation>
|
||
: pls. 25–27). Furthermore, the new available specimens show the antennal basal article narrower than the preceding urinary article, and antennae 2+ 3 clearly divergent (
|
||
<figureCitation id="130A2A0DFFF27054FEBBFAAFFE58C837" box="[357,481,1341,1367]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[151,1436,669,1748]" captionTargetId="figure@3.[151,1436,669,1748]" captionTargetPageId="3" captionText="FIGURE 1. A: Montezumella amenosi Vía, 1959, holotype MGSB 16313, dorsal view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). B: Moianella cervantesi Ossó & Domínguez, 2012, holotype MGB 59624 dorsal view, Priabonian (late Eocene), Moià, Moianès (Catalonia). C–D: Montezumella amenosi, MGB 88019, Collbàs Fm., Bartonian (Middle Eocene), La Pobla de Claramunt, Anoia (Catalonia); C: ventral view; D; frontal view. Abbreviations, a: pleonal somite; aa: basal antennular article; ab: basal antennal article 2+3; ca: carpus; cmxp3: coxa of third maxilliped; en: endostome; ep: epistome; ex exopod of mxp3; ey: eyestalk; is: ischium of mxp3; ma: mandibles; me: merus of mxp3; mxp3: third maxilliped; pr: proepistome; ua: urinary article of antenna. Scale bar = 1cm." httpUri="https://zenodo.org/record/3253512/files/figure.png" pageId="6" pageNumber="181">Fig. 1C, D</figureCitation>
|
||
), in contrast to species of Cheiragonoidea where antennae 2+3 are stout and convergent, respectively (
|
||
<figureCitation id="130A2A0DFFF27054FEB3FAF3FE7EC81B" box="[365,455,1377,1403]" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">Fig. 4C</figureCitation>
|
||
). In Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FD0AFAF3FCDDC81B" bold="true" box="[724,868,1376,1403]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
., the exopod of the third maxilliped is longer than the merus, whereas in Cheiragonoidea it is not as long as the merus (
|
||
<figureCitation id="130A2A0DFFF27054FC4FFA17FBACC8FF" box="[913,1045,1413,1439]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[151,1436,669,1748]" captionTargetId="figure@3.[151,1436,669,1748]" captionTargetPageId="3" captionText="FIGURE 1. A: Montezumella amenosi Vía, 1959, holotype MGSB 16313, dorsal view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). B: Moianella cervantesi Ossó & Domínguez, 2012, holotype MGB 59624 dorsal view, Priabonian (late Eocene), Moià, Moianès (Catalonia). C–D: Montezumella amenosi, MGB 88019, Collbàs Fm., Bartonian (Middle Eocene), La Pobla de Claramunt, Anoia (Catalonia); C: ventral view; D; frontal view. Abbreviations, a: pleonal somite; aa: basal antennular article; ab: basal antennal article 2+3; ca: carpus; cmxp3: coxa of third maxilliped; en: endostome; ep: epistome; ex exopod of mxp3; ey: eyestalk; is: ischium of mxp3; ma: mandibles; me: merus of mxp3; mxp3: third maxilliped; pr: proepistome; ua: urinary article of antenna. Scale bar = 1cm." httpUri="https://zenodo.org/record/3253512/files/figure.png" pageId="6" pageNumber="181">Figs. 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="130A2A0DFFF27054FBFEFA17FBFDC8FF" box="[1056,1092,1413,1439]" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">4C</figureCitation>
|
||
); additionally, the proepistome is larger than in Cheiragonoidea (
|
||
<figureCitation id="130A2A0DFFF27054FDD2FA3BFDEDC8A3" box="[524,596,1449,1475]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[151,1436,669,1748]" captionTargetId="figure@3.[151,1436,669,1748]" captionTargetPageId="3" captionText="FIGURE 1. A: Montezumella amenosi Vía, 1959, holotype MGSB 16313, dorsal view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). B: Moianella cervantesi Ossó & Domínguez, 2012, holotype MGB 59624 dorsal view, Priabonian (late Eocene), Moià, Moianès (Catalonia). C–D: Montezumella amenosi, MGB 88019, Collbàs Fm., Bartonian (Middle Eocene), La Pobla de Claramunt, Anoia (Catalonia); C: ventral view; D; frontal view. Abbreviations, a: pleonal somite; aa: basal antennular article; ab: basal antennal article 2+3; ca: carpus; cmxp3: coxa of third maxilliped; en: endostome; ep: epistome; ex exopod of mxp3; ey: eyestalk; is: ischium of mxp3; ma: mandibles; me: merus of mxp3; mxp3: third maxilliped; pr: proepistome; ua: urinary article of antenna. Scale bar = 1cm." httpUri="https://zenodo.org/record/3253512/files/figure.png" pageId="6" pageNumber="181">Figs. 1</figureCitation>
|
||
C–D, 4C; cf.
|
||
<bibRefCitation id="EFA04B79FFF27054FD30FA3BFC2DC8A3" author="Benedict, J. E." box="[750,916,1449,1475]" pageId="6" pageNumber="181" pagination="223 - 230" refId="ref6350" refString="Benedict, J. E. (1892) Corystoid crabs of the genera Telmessus and Erimacrus. Proceedings of the United States National Museum, 15, 223 - 230. https: // doi. org / 10.5479 / si. 00963801.15 - 900.223" type="journal article" year="1892">Benedict 1892</bibRefCitation>
|
||
, pl. 26, figs. 2, 5;
|
||
<bibRefCitation id="EFA04B79FFF27054FB82FA3BFA91C8A3" author="Kamio, M. & Matsunaga, S. & Fusetani, N." box="[1116,1320,1449,1475]" pageId="6" pageNumber="181" pagination="731 - 733" refId="ref7466" refString="Kamio, M., Matsunaga, S. & Fusetani, N. (2000) Studies on Sex Pheromones of the Helmet Crab, Telmessus cheiragonus 1. An Assay Based on Precopulatory Mate-Guarding. Zoological Science, 17, 731 - 733. https: // doi. org / 10.2108 / zsj. 17.731" type="journal article" year="2000">
|
||
Kamio
|
||
<emphasis id="B945EA9AFFF27054FB71FA38FB59C8A3" box="[1199,1248,1449,1475]" italics="true" pageId="6" pageNumber="181">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
, fig. 1). In adult females of
|
||
<taxonomicName id="4C314D0BFFF27054FE8BFA5CFD51C887" authority="(Brandt, 1848)" authorityName="Brandt" authorityYear="1848" baseAuthorityName="Brandt" baseAuthorityYear="1848" box="[341,744,1485,1511]" class="Malacostraca" family="Cheiragonidae" genus="Erimacrus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="isenbecki">
|
||
<emphasis id="B945EA9AFFF27054FE8BFA5CFD83C887" box="[341,570,1485,1511]" italics="true" pageId="6" pageNumber="181">Erimacrus isenbecki</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF27054FD94FA5FFD59C887" author="Brandt, J. F." box="[586,736,1485,1511]" pageId="6" pageNumber="181" pagination="1 - 179" refId="ref6502" refString="Brandt, J. F. (1848) Vorlaufige Bemerkungen uber eine neue eigenthumliche, der Fauna Russlands angehorige Gattung der Krabben. Bulletin physico-mathematique de l'Academie de Saint-Petersbourg, 7, 1 - 179." type="journal article" year="1848">Brandt, 1848</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C314D0BFFF27054FCFEFA5FFB64C887" authority="(Tilesius, 1812)" authorityName="Tilesius" authorityYear="1812" baseAuthorityName="Tilesius" baseAuthorityYear="1812" box="[800,1245,1485,1511]" class="Malacostraca" family="Cheiragonidae" genus="Telmessus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="cheiragonus">
|
||
<emphasis id="B945EA9AFFF27054FCFEFA5FFB9BC887" box="[800,1058,1485,1511]" italics="true" pageId="6" pageNumber="181">Telmessus cheiragonus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF27054FBECFA5FFB6FC887" author="Tilesius, G." box="[1074,1238,1485,1511]" pageId="6" pageNumber="181" pagination="311 - 405" refId="ref9867" refString="Tilesius, G. (1812) De Cancris camtschaticis, onoscis, entomostracis et cancellis marinis microscopicis nictilucentibus. Memoires de l'Academie Imperale des sciences de Saint Petersburg, 5 (5), 311 - 405." type="journal article" year="1812">Tilesius, 1812</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the press button is present but vestigial in mature females (
|
||
<emphasis id="B945EA9AFFF27054FDADFA60FD33CB6B" box="[627,650,1522,1547]" italics="true" pageId="6" pageNumber="181">vs</emphasis>
|
||
. present and well-developed in adult females of Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FA58FA63FEB0CB4F" bold="true" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
. (
|
||
<figureCitation id="130A2A0DFFF27054FEC1F987FE0DCB4F" box="[287,436,1557,1583]" captionStart="FIGURE 2" captionStartId="4.[151,250,1754,1779]" captionTargetBox="[181,1405,166,1734]" captionTargetId="figure@4.[181,1405,166,1734]" captionTargetPageId="4" captionText="FIGURE 2. A: Montezumella sp., MGB 59626, male ventral view, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). B: Moianella cervantesi Ossó & Domínguez, 2013, holotype, MGB 59624, dorsal view, Priabonian (late Eocene), Moià, Bages (Catalonia). C: Montezumella amenosi Vía, 1959, holotype, MGSB 16313, ventral view, Collbàs Fm., Bartonian (Middle Eocene), Carme, Anoia (Catalonia). D: Montezumella sp., MGB 59627, female ventral view showing the vulvae, Lutetian (Middle Eocene), Vic zone, Osona (Catalonia). E: Montezumella sp., MGB 59629, female ventral view, with pleon, Bartonian (Middle Eocene), Huesca, Aragón (Spain). F, G: Moianella cervantesi Ossó & Domínguez, 2013, MGB 87858, female, Priabonian (Late Eocene), Moià, Moianès (Catalonia); F: ventral view, with pleon. G: close up of the press button.Abbreviations, a: pleonal somites; ml: median line; pb: press button; st: thoracic sternites; t: telson; v: vulvae. Scale bar = 1cm." httpUri="https://zenodo.org/record/3253514/files/figure.png" pageId="6" pageNumber="181">Fig. 2E, F, G</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFA04B79FFF27054FE1EF987FD36CB4F" author="Guinot, D." box="[448,655,1557,1583]" pageId="6" pageNumber="181" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot, 1979: 137</bibRefCitation>
|
||
, pl. 25, figs 1–3;
|
||
<bibRefCitation id="EFA04B79FFF27054FC8DF987FBCCCB4F" author="Guinot D. & Bouchard J. M." box="[851,1141,1557,1583]" pageId="6" pageNumber="181" pagination="613 - 694" refId="ref7053" refString="Guinot D. & Bouchard J. M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20, 613 - 694." type="journal article" year="1998">Guinot & Bouchard 1998</bibRefCitation>
|
||
). Reexamination of extant material in the
|
||
<collectingRegion id="49F5F86AFFF27054FE9CF9ABFE21CB33" box="[322,408,1593,1619]" country="Ireland" name="Monaghan" pageId="6" pageNumber="181">MNHN</collectingRegion>
|
||
collections, revealed that females of
|
||
<taxonomicName id="4C314D0BFFF27054FCE3F9A8FB9ACB33" baseAuthorityName="Brandt" baseAuthorityYear="1848" box="[829,1059,1593,1619]" class="Malacostraca" family="Cheiragonidae" genus="Erimacrus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="isenbecki">
|
||
<emphasis id="B945EA9AFFF27054FCE3F9A8FB9ACB33" box="[829,1059,1593,1619]" italics="true" pageId="6" pageNumber="181">Erimacrus isenbecki</emphasis>
|
||
</taxonomicName>
|
||
present a small press button near the suture 5/6, not covered by the pleon, and thus exposed (
|
||
<figureCitation id="130A2A0DFFF27054FC92F9CFFC10CB17" box="[844,937,1629,1655]" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">Fig. 4D</figureCitation>
|
||
). Likewise, females of
|
||
<taxonomicName id="4C314D0BFFF27054FB69F9CFFEEBCBFB" authority="(Stimpson 1858)" authorityName="Stimpson" authorityYear="1858" baseAuthorityName="Stimpson" baseAuthorityYear="1858" class="Malacostraca" family="Cheiragonidae" genus="Telmessus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="acutidens">
|
||
<emphasis id="B945EA9AFFF27054FB69F9CFFA25CB17" box="[1207,1436,1629,1655]" italics="true" pageId="6" pageNumber="181">Telmessus acutidens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF27054FF41F913FEF3CBFB" author="Stimpson, W." box="[159,330,1664,1691]" pageId="6" pageNumber="181" pagination="31 - 41" refId="ref9727" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars IV. Crustacea Cancroidea et Corystoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 31 - 41. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C314D0BFFF27054FE59F910FD92CBFB" baseAuthorityName="Tilesius" baseAuthorityYear="1812" box="[391,555,1665,1691]" class="Malacostraca" family="Cheiragonidae" genus="Telmessus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="cheiragonus">
|
||
<emphasis id="B945EA9AFFF27054FE59F910FD92CBFB" box="[391,555,1665,1691]" italics="true" pageId="6" pageNumber="181">T. cheiragonus</emphasis>
|
||
</taxonomicName>
|
||
possess a small vestigial press button (
|
||
<figureCitation id="130A2A0DFFF27054FC03F913FBEACBFB" box="[989,1107,1665,1691]" captionStart="FIGURE 4" captionStartId="8.[151,250,1854,1879]" captionTargetBox="[213,1375,166,1834]" captionTargetId="figure@8.[213,1375,166,1834]" captionTargetPageId="8" captionText="FIGURE 4. Extant species of Cheiragonoidea. A, D Erimacrus isenbecki (Brandt, 1848), female, MNHN-IU-2017–9089 (= MNHN-B5404). B Telmessus acutidens (Stimpson, 1858), female, MNHN-IU-2017–9086 (= MNHN-B5408). C, E Telmessus cheiragonus (Tilesius, 1812); C, male, MNHN-IU-2017-9095 (= MNHN-B5410); E, female, MNHN-IU-2017–9097 (= MNHN-B5445). Abbreviations. a: basal antennular article; ab: basal antennal article 2+3; ex, exopod of mxp3; is: ischium of mxp3; me: merus of mxp3; ml= median line; pb= press button; pl= pleopods; v= vulvae. ©MNHN-À.Ossó." httpUri="https://zenodo.org/record/3253518/files/figure.png" pageId="6" pageNumber="181">Fig. 4B, E</figureCitation>
|
||
). However, they are not functional due to the presence of pleopods and the lack of the socket on the corresponding part of the pleon (somite 6), as is often the case in mature female brachyurans. As previously stated by
|
||
<bibRefCitation id="EFA04B79FFF27054FC62F95BFAA6CB83" author="Osso, A. & Dominguez, J. L." box="[956,1311,1737,1763]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez (2013: 289</bibRefCitation>
|
||
, 292, 296), according to the available fossil record, we do not recognize any fossil representative of Cheiragonoidea.
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF27054FF19F883FCCACABF" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="181">
|
||
An extensive and detailed comparison between
|
||
<taxonomicName id="4C314D0BFFF27054FD00F883FC1BCA4B" authorityName="Osso & Dominguez" authorityYear="2013" box="[734,930,1809,1835]" class="Malacostraca" family="Montezumellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="181" phylum="Arthropoda" rank="family">Montezumellidae</taxonomicName>
|
||
and other taxa besides Cancroidea and Cheiragonoidea, but with which it also shares similar thoracic structures, e.g., a narrow and completely metamerised sternum and similar dorsal features, was carried out by
|
||
<bibRefCitation id="EFA04B79FFF27054FCD4F8CBFBC9CA13" author="Osso, A. & Dominguez, J. L." box="[778,1136,1881,1907]" pageId="6" pageNumber="181" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez (2013: 292</bibRefCitation>
|
||
, 293). These comparisons, which support a clear distinction between these taxa, are also valid for Montezumelloidea
|
||
<emphasis id="B945EA9AFFF27054FB5AF8EFFAADCAF7" bold="true" box="[1156,1300,1916,1943]" pageId="6" pageNumber="181">n. superfam</emphasis>
|
||
. and are not be repeated herein. Nevertheless, a comparison taking into account the sterno-abdominal features and cephalic appendages, preserved in the new specimens, is summarized below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6503FFF37055FF19FF0BFACECEE3" pageId="7" pageNumber="182" type="reference_group">
|
||
<paragraph id="8B8E3688FFF37055FF19FF0BFACECEE3" blockId="7.[151,1437,152,899]" pageId="7" pageNumber="182">
|
||
In
|
||
<taxonomicName id="4C314D0BFFF37055FF39FF0BFD12CDD3" ID-CoL="YSVB" authority="(Pennant, 1777)" authorityName="Pennant" authorityYear="1777" baseAuthorityName="Pennant" baseAuthorityYear="1777" box="[231,683,153,179]" class="Malacostraca" family="Corystidae" genus="Corystes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="cassivelaunus">
|
||
<emphasis id="B945EA9AFFF37055FF39FF0BFE57CDD3" box="[231,494,153,179]" italics="true" pageId="7" pageNumber="182">Corystes cassivelaunus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF37055FE20FF0BFD1ACDD3" author="Pennant, T." box="[510,675,153,179]" pageId="7" pageNumber="182" refId="ref8710" refString="Pennant, T. (1777) British Zoology. 4. Crustacea, Mollusca, Testacea. Benjamin White, London, XCIII pp." type="book" year="1777">Pennant, 1777</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(Fig. 5A, B)
|
||
<typeStatus id="548A882AFFF37055FC97FF0BFCC0CDD3" box="[841,889,153,179]" pageId="7" pageNumber="182">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C314D0BFFF37055FC22FF0BFB53CDD3" ID-CoL="7NTM6" authority="Bosc, 1802" authorityName="Bosc" authorityYear="1802" box="[1020,1258,153,179]" class="Malacostraca" family="Corystidae" genus="Corystes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF37055FC22FF0BFBE7CDD3" box="[1020,1118,153,179]" italics="true" pageId="7" pageNumber="182">Corystes</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF37055FBB9FF0BFB53CDD3" author="Bosc, L. A. G." box="[1127,1258,153,179]" pageId="7" pageNumber="182" refId="ref6443" refString="Bosc, L. A. G. (1802) Histoire naturelle des Crustaces, contenant leur description et leurs moeurs, avec figures dessinees d'apres nature. Vol. 1 & 2. De Guilleminet, Paris, 258 & 296 pp. https: // doi. org / 10.5962 / bhl. title. 39831" type="book" year="1802">Bosc, 1802</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="548A882AFFF37055FAFCFF0BFAEBCDD3" box="[1314,1362,153,179]" pageId="7" pageNumber="182">type</typeStatus>
|
||
genus of Corystoidea
|
||
<bibRefCitation id="EFA04B79FFF37055FE96FF2FFDBFCDB7" author="Samouelle, G." box="[328,518,189,215]" pageId="7" pageNumber="182" refId="ref9121" refString="Samouelle, G. (1819) The entomologist's useful compendium, or an introduction to the British insects, etc. T. Boys, London, 496 pp." type="book" year="1819">Samouelle, 1819</bibRefCitation>
|
||
(
|
||
<emphasis id="B945EA9AFFF37055FDC6FF2CFDEFCDB7" box="[536,598,190,215]" italics="true" pageId="7" pageNumber="182">sensu</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF37055FD81FF2FFCB4CDB7" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[607,781,189,215]" pageId="7" pageNumber="182" pagination="1 - 286" refId="ref8316" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286. [updates at http: // rmbr. nus. edu. sg / research / cotw / supplement 17. php]" type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="B945EA9AFFF37055FD55FF2CFD07CDB7" box="[651,702,189,215]" italics="true" pageId="7" pageNumber="182">et al</emphasis>
|
||
., 2008
|
||
</bibRefCitation>
|
||
[see
|
||
<bibRefCitation id="EFA04B79FFF37055FC92FF2FFBD9CDB7" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." box="[844,1120,188,215]" pageId="7" pageNumber="182" pagination="1049 - 1130" refId="ref6729" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 b) Phylogeny of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology-The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C), (I), Decapoda: Brachyura (Part 2), 1049 - 1130." type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B945EA9AFFF37055FC49FF2CFC72CDB7" box="[919,971,189,215]" italics="true" pageId="7" pageNumber="182">et al</emphasis>
|
||
. 2015b: 945
|
||
</bibRefCitation>
|
||
]), the antennules are folded longitudinally (
|
||
<emphasis id="B945EA9AFFF37055FE98FF70FEE4CD9B" box="[326,349,226,251]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. transversely in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FD29FF73FC31CD9B" bold="true" box="[759,904,224,251]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.), and a pleonal-locking mechanism is lacking (
|
||
<emphasis id="B945EA9AFFF37055FF7FFE94FF01CC7F" box="[161,184,262,287]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. present in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FDC3FE97FD16CC7F" bold="true" box="[541,687,260,287]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.); the first pleonal somites is visible dorsally, as in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FF3DFEBBFECACC23" bold="true" box="[227,371,296,323]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
. (also
|
||
<bibRefCitation id="EFA04B79FFF37055FE60FEBBFD3ECC23" author="Guinot, D." box="[446,647,297,323]" pageId="7" pageNumber="182" pagination="1 - 354" refId="ref7013" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum National d'Histoire Naturelle, A, Zoologie, 112, 1 - 354." type="journal article" year="1979">Guinot 1979: 132</bibRefCitation>
|
||
; Ng.
|
||
<emphasis id="B945EA9AFFF37055FD1DFEB8FD4CCC23" box="[707,757,297,323]" italics="true" pageId="7" pageNumber="182">et al</emphasis>
|
||
. 2008: fig. 39;
|
||
<bibRefCitation id="EFA04B79FFF37055FC7CFEBBFB03CC23" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." box="[930,1210,296,323]" pageId="7" pageNumber="182" pagination="1049 - 1130" refId="ref6600" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Systematics and classification of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C), (I), Decapoda: Brachyura (Part 2), 1049 - 1130. https: // doi. org / 10.1163 / 9789004190832 _ 021" type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B945EA9AFFF37055FC32FEB8FBA4CC23" box="[1004,1053,297,323]" italics="true" pageId="7" pageNumber="182">et al</emphasis>
|
||
. 2015a: 1075
|
||
</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C314D0BFFF37055FB32FEBBFE40CC07" ID-CoL="58GK7" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" class="Malacostraca" family="Trichopeltariidae" genus="Trichopeltarion" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="nobile">
|
||
<emphasis id="B945EA9AFFF37055FB32FEBBFF64CC07" italics="true" pageId="7" pageNumber="182">Trichopeltarion nobile</emphasis>
|
||
A.
|
||
<bibRefCitation id="EFA04B79FFF37055FEDBFEDFFE40CC07" author="Milne-Edwards, A." box="[261,505,333,359]" pageId="7" pageNumber="182" pagination="1 - 68" refId="ref7926" refString="Milne-Edwards, A. (1880) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the United States Coast Survey Steamer " Blake, " Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII. Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology at Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">Milne-Edwards, 1880</bibRefCitation>
|
||
</taxonomicName>
|
||
(Fig. 5C),
|
||
<typeStatus id="548A882AFFF37055FDAAFEDFFD1DCC07" box="[628,676,333,359]" pageId="7" pageNumber="182">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C314D0BFFF37055FCFFFEDFFB49CC07" ID-CoL="7Q54Q" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[801,1264,333,359]" class="Malacostraca" family="Trichopeltariidae" genus="Trichopeltarion" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF37055FCFFFEDFFC68CC07" box="[801,977,333,359]" italics="true" pageId="7" pageNumber="182">Trichopeltarion</emphasis>
|
||
A.
|
||
<bibRefCitation id="EFA04B79FFF37055FC27FEDFFB49CC07" author="Milne-Edwards, A." box="[1017,1264,333,359]" pageId="7" pageNumber="182" pagination="1 - 68" refId="ref7926" refString="Milne-Edwards, A. (1880) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the United States Coast Survey Steamer " Blake, " Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII. Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology at Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">Milne-Edwards, 1880</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="548A882AFFF37055FAFAFEDFFAEDCC07" box="[1316,1364,333,359]" pageId="7" pageNumber="182">type</typeStatus>
|
||
genus of Trichopeltarioidea
|
||
<bibRefCitation id="EFA04B79FFF37055FE56FEE3FD32CCEB" author="Tavares, M. & Cleva, R." box="[392,651,369,395]" pageId="7" pageNumber="182" pagination="97 - 157" refId="ref9801" refString="Tavares, M. & Cleva, R. (2010) Trichopeltariidae (Crustacea, Decapoda, Brachyura), a new family and superfamily of eubrachyuran crabs with description of one new genus and five new species. Papeis Avulsos Zoologia, 50 (9), 97 - 157. https: // doi. org / 10.1590 / S 0031 - 10492010000900001" type="journal article" year="2010">Tavares & Cleva, 2010</bibRefCitation>
|
||
, the antennulae are folded longitudinally (
|
||
<emphasis id="B945EA9AFFF37055FBBFFEE0FBC1CCEB" box="[1121,1144,370,395]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. transversely in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FED3FE07FE25CCCF" bold="true" box="[269,412,404,431]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.) the basal article of antennae 2+3 rises convergently to the axis (
|
||
<emphasis id="B945EA9AFFF37055FBA3FE04FB2DCCCF" box="[1149,1172,406,431]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. divergently to the axis in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FE53FE2BFDA5CCB3" bold="true" box="[397,540,440,467]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.). Additionally, the limit between the endostome and epistome is poorly defined (
|
||
<emphasis id="B945EA9AFFF37055FF7FFE4CFF01CC97" box="[161,184,478,503]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. well defined in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FD90FE4FFD64CC97" bold="true" box="[590,733,476,503]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.). The first pleonal somite is visible dorsally, as in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FF29FD93FE3ECF7B" bold="true" box="[247,391,512,539]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
. (see
|
||
<bibRefCitation id="EFA04B79FFF37055FE17FD93FD70CF7B" author="Tavares, M. & Cleva, R." box="[457,713,513,539]" pageId="7" pageNumber="182" pagination="97 - 157" refId="ref9801" refString="Tavares, M. & Cleva, R. (2010) Trichopeltariidae (Crustacea, Decapoda, Brachyura), a new family and superfamily of eubrachyuran crabs with description of one new genus and five new species. Papeis Avulsos Zoologia, 50 (9), 97 - 157. https: // doi. org / 10.1590 / S 0031 - 10492010000900001" type="journal article" year="2010">Tavares & Cleva 2010</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C314D0BFFF37055FD24FD90FBC1CF7B" ID-CoL="6NMMN" authority="(Krauss, 1843)" authorityName="Krauss" authorityYear="1843" baseAuthorityName="Krauss" baseAuthorityYear="1843" box="[762,1144,513,539]" class="Malacostraca" family="Xanthidae" genus="Kraussia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="rugulosa">
|
||
<emphasis id="B945EA9AFFF37055FD24FD90FC72CF7B" box="[762,971,513,539]" italics="true" pageId="7" pageNumber="182">Kraussia rugulosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF37055FC07FD93FBC9CF7B" author="Krauss, F." box="[985,1136,513,539]" pageId="7" pageNumber="182" refId="ref7725" refString="Krauss, F. (1843) Die Sudafrikanischen Crustaceen. Eine Zusammenstellung aller bekannten Malacostraca, Bemerkungen uber deren Lebensweise und geographische Verbreitung, nebst Beschrwibung und Abbildung mehrer neuen Arten. Schweizerbartsche, Stuttgart, 68 pp. https: // doi. org / 10.5962 / bhl. title. 4825" type="book" year="1843">Krauss, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(Fig. 5D, F),
|
||
<typeStatus id="548A882AFFF37055FACAFD93FAFDCF7B" box="[1300,1348,513,539]" pageId="7" pageNumber="182">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C314D0BFFF37055FF69FDB4FE13CF5F" ID-CoL="7P7X2" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[183,426,548,575]" class="Malacostraca" family="Xanthidae" genus="Kraussia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF37055FF69FDB4FEA5CF5F" box="[183,284,550,575]" italics="true" pageId="7" pageNumber="182">Kraussia</emphasis>
|
||
<bibRefCitation id="EFA04B79FFF37055FEFBFDB7FE13CF5F" author="Dana, J. D." box="[293,426,548,575]" pageId="7" pageNumber="182" pagination="1 - 1618" refId="ref6539" refString="Dana, J. D. (1852) Crustacea. Parts I and II. U. S. Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N. C., Sherman, Philadelphia, 13, 1 - 1618." type="journal article" year="1852">Dana, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="548A882AFFF37055FE3FFDB7FDA8CF5F" box="[481,529,549,575]" pageId="7" pageNumber="182">type</typeStatus>
|
||
genus of
|
||
<taxonomicName id="4C314D0BFFF37055FD5DFDB7FCC3CF5F" ID-CoL="847GB" authority="Ng, 1993" authorityName="Ng" authorityYear="1993" box="[643,890,549,575]" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="subFamily" subFamily="Kraussiinae">
|
||
Kraussiinae
|
||
<bibRefCitation id="EFA04B79FFF37055FCCEFDB7FCC3CF5F" author="Ng, P. K. L." box="[784,890,549,575]" pageId="7" pageNumber="182" pagination="133 - 157" refId="ref8245" refString="Ng, P. K. L. (1993) Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with description of two new species from Singapore and the Philippines. The Raffles Bulletin of Zoology, 41 (1), 133 - 157." type="journal article" year="1993">Ng, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C314D0BFFF37055FC54FDB7FB00CF5F" ID-CoL="7NLKC" authority="MacLeay, 1838" authorityName="MacLeay" authorityYear="1838" box="[906,1209,549,575]" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="family">
|
||
Xanthidae
|
||
<bibRefCitation id="EFA04B79FFF37055FBD9FDB7FB00CF5F" author="MacLeay, W. S." box="[1031,1209,549,575]" pageId="7" pageNumber="182" pagination="53 - 71" refId="ref8093" refString="MacLeay, W. S. (1838) On the brachyurous decapod Crustacea brought from the Cape by Dr. Smith. In: Smith, A. (Ed.), Illustrations of the Annulosa of South Africa; being a portion of the objects of natural history chiefly collected during an expedition into the interior of South Africa, under the direction of Dr. Andrew Smith, in the years 1834, 1835 and 1836; fitted out by " The Cape of Good Hope Association for Exploring Central Africa ". Smith, Elder, and Co., London, 53 - 71." type="book chapter" year="1838">MacLeay, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
), the antennules are folded obliquely (
|
||
<emphasis id="B945EA9AFFF37055FE83FDD8FECDCF03" box="[349,372,586,611]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. transversely in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FCDCFDDBFC29CF03" bold="true" box="[770,912,584,611]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.); the antennae 2+3 rise longitudinally directed (
|
||
<emphasis id="B945EA9AFFF37055FF7FFDFCFF01CFE7" box="[161,184,622,647]" italics="true" pageId="7" pageNumber="182">vs</emphasis>
|
||
. obliquely divergent in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FD7DFDFFFC8DCFE7" bold="true" box="[675,820,620,647]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.); the pleonal somites 1, 2 appear not folded ventrally and are in dorsal position, as in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FD0FFD03FCD9CFCB" bold="true" box="[721,864,656,683]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
. (Figs.5D, F; also
|
||
<bibRefCitation id="EFA04B79FFF37055FBEEFD03FB2DCFCB" author="Ng, P. K. L." box="[1072,1172,657,683]" pageId="7" pageNumber="182" pagination="133 - 157" refId="ref8245" refString="Ng, P. K. L. (1993) Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with description of two new species from Singapore and the Philippines. The Raffles Bulletin of Zoology, 41 (1), 133 - 157." type="journal article" year="1993">Ng 1993</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C314D0BFFF37055FB1BFD03FECCCFB0" ID-CoL="4HVQ4" authority="(Poeppig, 1836)" authorityName="Poeppig" authorityYear="1836" baseAuthorityName="Poeppig" baseAuthorityYear="1836" class="Malacostraca" family="Pilumnoididae" genus="Pilumnoides" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="perlatus">
|
||
<emphasis id="B945EA9AFFF37055FB1BFD03FF01CFAF" italics="true" pageId="7" pageNumber="182">Pilumnoides perlatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA04B79FFF37055FF19FD27FED5CFB0" author="Poeppig, E." box="[199,364,693,720]" pageId="7" pageNumber="182" pagination="133 - 145" refId="ref8736" refString="Poeppig, E. (1836) Crustacea Chilensia nova aut minus nota descripsit. Archiv fur Naturgeschichte, 2 (1), 133 - 145." type="journal article" year="1836">Poeppig, 1836</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(Fig. 5G, I),
|
||
<typeStatus id="548A882AFFF37055FDD5FD27FD82CFAF" box="[523,571,693,719]" pageId="7" pageNumber="182">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C314D0BFFF37055FD65FD27FA8ECFAF" ID-CoL="7PS44" authority="Lucas in H. Milne Edwards & Lucas, 1844" authorityName="Lucas in H. Milne Edwards & Lucas" authorityYear="1844" box="[699,1335,693,719]" class="Malacostraca" family="Pilumnoididae" genus="Pilumnoides" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EA9AFFF37055FD65FD27FCFECFAF" box="[699,839,693,719]" italics="true" pageId="7" pageNumber="182">Pilumnoides</emphasis>
|
||
Lucas
|
||
<emphasis id="B945EA9AFFF37055FC47FD24FC16CFAF" box="[921,943,694,719]" italics="true" pageId="7" pageNumber="182">in</emphasis>
|
||
H. Milne Edwards & Lucas, 1844
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="548A882AFFF37055FAB2FD27FA25CFAF" box="[1388,1436,693,719]" pageId="7" pageNumber="182">type</typeStatus>
|
||
genus of
|
||
<taxonomicName id="4C314D0BFFF37055FEDEFD4BFD4ACF93" ID-CoL="7NKH2" authority="Guinot & Macpherson, 1987" authorityName="Guinot & Macpherson" authorityYear="1987" box="[256,755,729,755]" class="Malacostraca" family="Pilumnoididae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="family">
|
||
Pilumnoididae
|
||
<bibRefCitation id="EFA04B79FFF37055FE75FD4BFD4ACF93" author="Guinot, D. & Macpherson, E." box="[427,755,729,755]" pageId="7" pageNumber="182" pagination="211 - 247" refId="ref7174" refString="Guinot, D. & Macpherson, E. (1987) Revision du genre Pilumnoides Lucas, 1844, avec description de quatre especes nouvelles et creation de Pilumnoidinae subfam. nov. (Crustacea Decapoda Brachyura). Bulletin du Museum National d'Histoire Naturelle, 4 (9 A / 1), 211 - 247." type="journal article" year="1987">Guinot & Macpherson, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
, the antennules are folded nearly transversely (Fig. 5H; also
|
||
<bibRefCitation id="EFA04B79FFF37055FF49FD6FFE01CE78" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." box="[151,440,764,792]" pageId="7" pageNumber="182" pagination="1049 - 1130" refId="ref6600" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Systematics and classification of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C), (I), Decapoda: Brachyura (Part 2), 1049 - 1130. https: // doi. org / 10.1163 / 9789004190832 _ 021" type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B945EA9AFFF37055FF3DFD6CFEAFCE77" box="[227,278,765,791]" italics="true" pageId="7" pageNumber="182">et al</emphasis>
|
||
. 2015a: 1106
|
||
</bibRefCitation>
|
||
;), as in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FD32FD6FFCC7CE77" bold="true" box="[748,894,764,791]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
.; however, in Montezumelloidea
|
||
<emphasis id="B945EA9AFFF37055FB23FD6FFA36CE77" bold="true" box="[1277,1423,764,791]" pageId="7" pageNumber="182">n. superfam</emphasis>
|
||
., the somites 1, 2 are not folded ventrally and in dorsal position, whereas in
|
||
<taxonomicName id="4C314D0BFFF37055FC0CFCB0FBF0CE5B" ID-CoL="4HVQ4" baseAuthorityName="Poeppig" baseAuthorityYear="1836" box="[978,1097,801,827]" class="Malacostraca" family="Pilumnoididae" genus="Pilumnoides" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="perlatus">
|
||
<emphasis id="B945EA9AFFF37055FC0CFCB0FBF0CE5B" box="[978,1097,801,827]" italics="true" pageId="7" pageNumber="182">P. perlatus</emphasis>
|
||
</taxonomicName>
|
||
they are folded ventrally (Fig. 5G; also
|
||
<bibRefCitation id="EFA04B79FFF37055FF22FCD7FD82CE3F" author="Guinot, D. & Macpherson, E." box="[252,571,837,863]" pageId="7" pageNumber="182" pagination="211 - 247" refId="ref7174" refString="Guinot, D. & Macpherson, E. (1987) Revision du genre Pilumnoides Lucas, 1844, avec description de quatre especes nouvelles et creation de Pilumnoidinae subfam. nov. (Crustacea Decapoda Brachyura). Bulletin du Museum National d'Histoire Naturelle, 4 (9 A / 1), 211 - 247." type="journal article" year="1987">Guinot & Macpherson 1987</bibRefCitation>
|
||
: pl. 1, fig. A). In any case, the differences highlighted by
|
||
<bibRefCitation id="EFA04B79FFF37055FB60FCD7FEADCEE3" author="Osso, A. & Dominguez, J. L." pageId="7" pageNumber="182" pagination="285 - 298" refId="ref8623" refString="Osso, A. & Dominguez, J. L. (2013) A proposal for a new family: Montezumellidae (Crustacea, Decapoda, Brachyura) and description of new genus and species Moianella cervantesi from the Priabonian (late Eocene) of Catalonia (NE of Iberian Peninsula). Boletin de la Sociedad Geologica Mexicana, 65 (2), 285 - 298. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 10" type="journal article" year="2013">Ossó & Domínguez (2013: 292</bibRefCitation>
|
||
, 293) precludes any close relationship among both taxa, and likewise with all the taxa listed therein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |