439 lines
50 KiB
XML
439 lines
50 KiB
XML
<document id="2BCBBA20BAD900B57AF0A92F7D2DBD40" ID-DOI="10.5281/zenodo.183986" ID-GBIF-Dataset="4ced8216-c041-4541-81aa-55e2f672a565" ID-ISSN="1175-5326" ID-Zenodo-Dep="183986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459937723978" checkinUser="plazi" docAuthor="Schaaf-Da, Jayna A. & Ebert, David A." docDate="2008" docId="0398879EFFF5FFA9FF49B5BEFD64FD77" docLanguage="en" docName="zt01872p028.pdf" docOrigin="Zootaxa 1872" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cephaloscyllium pardelotum Schaaf-Da & Ebert, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="FFA1FFE6FFF7FFAFFFDEB04BFF84FF95" masterDocTitle="A revision of the western North Pacific swellsharks, genus Cephaloscyllium Gill 1862 (Chondrichthyes: Carcharhiniformes: Scyliorhinidae), including descriptions of two new species" masterLastPageNumber="28" masterPageNumber="1" pageNumber="3" updateTime="1698230248003" updateUser="plazi">
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<mods:title id="2D9859823E534FC23F3BCCB64EBA495D">A revision of the western North Pacific swellsharks, genus Cephaloscyllium Gill 1862 (Chondrichthyes: Carcharhiniformes: Scyliorhinidae), including descriptions of two new species</mods:title>
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<mods:name id="6096A0454CA1E7D6433D268CFB89EC07" type="personal">
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<mods:roleTerm id="B75A346A863CE3EC8888E5CC74B66149">Author</mods:roleTerm>
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<mods:namePart id="701F7FEA1DBD29D6288156D740FEA452">Schaaf-Da, Jayna A.</mods:namePart>
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<mods:name id="67E8B56005142EA6EAD45934AC7B9B07" type="personal">
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<mods:roleTerm id="3CCF31FC2291B7AB356ADAE49F9E4473">Author</mods:roleTerm>
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<mods:namePart id="FC7B44696E3D7F156569A62F85A88721">Ebert, David A.</mods:namePart>
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<mods:typeOfResource id="02030748551EB33B66D38AFBD21CD5F1">text</mods:typeOfResource>
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<mods:title id="8BA91B38A9860D0AF22147E584FADD55">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A0491D4E296C701785073DD3E7B660BF">
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<mods:date id="EFB991D41254B535B6E75299D987EBFE">2008</mods:date>
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<mods:detail id="3493AE8D969B1F06C873E757FD88739F" type="volume">
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<mods:number id="5FC5DD1C2E35F0784FBB9829848C7925">1872</mods:number>
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<mods:classification id="265F03C09ABC9700F22631A986CD3D3D">journal article</mods:classification>
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<mods:identifier id="F12258CC805424F5950F084CB5AF0254" type="DOI">10.5281/zenodo.183986</mods:identifier>
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<mods:identifier id="6D8CFBB5B6EDD6950CDCF4140AEDEB11" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0FDC7700E54929C03325FE05B7934F7A" type="Zenodo-Dep">183986</mods:identifier>
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<treatment id="0398879EFFF5FFA9FF49B5BEFD64FD77" ID-DOI="http://doi.org/10.5281/zenodo.5613443" ID-GBIF-Taxon="119375611" ID-Zenodo-Dep="5613443" LSID="urn:lsid:plazi:treatment:0398879EFFF5FFA9FF49B5BEFD64FD77" httpUri="http://treatment.plazi.org/id/0398879EFFF5FFA9FF49B5BEFD64FD77" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
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<subSubSection id="C32B6503FFF5FFADFF49B5BEFE31F9A7" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B8E3688FFF5FFADFF49B5BEFDD3F99A" blockId="2.[151,599,1525,1586]" box="[151,599,1525,1551]" pageId="2" pageNumber="3">
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<heading id="D0C681E4FFF5FFADFF49B5BEFDD3F99A" bold="true" box="[151,599,1525,1551]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="B945EA9AFFF5FFADFF49B5BEFDD3F99A" bold="true" box="[151,599,1525,1551]" pageId="2" pageNumber="3">
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<taxonomicName id="4C314D0BFFF5FFADFF49B5BEFE76F99A" ID-CoL="5XJXJ" box="[151,498,1525,1551]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
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<emphasis id="B945EA9AFFF5FFADFF49B5BEFE76F99A" bold="true" box="[151,498,1525,1551]" italics="true" pageId="2" pageNumber="3">Cephaloscyllium pardelotum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27657E1FFF5FFADFE24B5BEFDD3F99A" box="[506,599,1525,1551]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8E3688FFF5FFADFF49B653FE31F9A7" blockId="2.[151,599,1525,1586]" box="[151,437,1560,1586]" pageId="2" pageNumber="3">
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(
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<figureCitation id="130A2A0DFFF5FFADFF41B653FF75F9A7" box="[159,241,1560,1586]" captionStart="FIGURE 1" captionStartId="3.[151,255,1912,1936]" captionTargetBox="[151,1435,924,1888]" captionTargetId="figure@3.[151,1436,924,1888]" captionTargetPageId="3" captionText="FIGURE 1. Cephaloscyllium pardelotum sp. nov. holotype (CAS 224876); a. lateral view; b. dorsal view. Photos by D. A. Ebert" httpUri="https://zenodo.org/record/183987/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
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,
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<figureCitation id="130A2A0DFFF5FFADFF21B653FE8AF9A7" box="[255,270,1560,1586]" captionStart="FIGURE 5" captionStartId="9.[151,255,1897,1921]" captionTargetBox="[195,1374,202,1863]" captionTargetId="figure@9.[181,1405,194,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lateral views of the WNP Cephaloscyllium, all about 220 mm TL; a. Cephaloscyllium pardelotum sp. nov.; b. Cephaloscyllium maculatum sp. nov.; c. Cephaloscyllium fasciatum; d. Cephaloscyllium umbratile; e. Cephaloscyllium sarawakensis; f. Cephaloscyllium circulopullum. Illustration by J. A. S. - Da Silva." httpUri="https://zenodo.org/record/183992/files/figure.png" pageId="2" pageNumber="3">5</figureCitation>
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a, 7a;
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<tableCitation id="C6B30333FFF5FFADFE8AB653FE2DF9A7" box="[340,425,1560,1586]" captionStart="TABLE 1" captionStartId="6.[151,239,946,970]" captionTargetBox="[159,1427,1141,2015]" captionTargetPageId="6" captionText="TABLE 1. Morphometric and meristic data for the holotype (HT) specimens for Cephaloscyllium pardelotum sp. nov. (CAS 224876) and Cephaloscyllium maculatum sp. nov. (CAS 224877). TL is the actual measurement in millimeters; all other measurements are expressed as percentage of TL. VERT—total vertebral count, mono—monospondylous centra count, predip—precaudal diplospondylous centra count, caudaldip—caudal diplospondylous centra count, U tot—tooth count of upper jaw, L tot—tooth count of lower jaw. Abbreviations are listed in Appendix 1." httpUri="http://table.plazi.org/id/DF4E6600FFF1FFA9FF49B3F9FBF4FBC1" pageId="2" pageNumber="3" tableUuid="DF4E6600FFF1FFA9FF49B3F9FBF4FBC1">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6503FFF5FFADFF49B62CFB5DF93F" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8B8E3688FFF5FFADFF49B62CFB5DF93F" blockId="2.[151,1436,1639,2026]" pageId="2" pageNumber="3">
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<emphasis id="B945EA9AFFF5FFADFF49B62CFE51F914" bold="true" box="[151,469,1639,1665]" pageId="2" pageNumber="3">
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<typeStatus id="548A882AFFF5FFADFF49B62CFF50F914" box="[151,212,1639,1665]" pageId="2" pageNumber="3">Type</typeStatus>
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Series and Locality.
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</emphasis>
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<typeStatus id="548A882AFFF5FFADFE00B623FDCAF917" box="[478,590,1640,1666]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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,
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<collectionCode id="ED20AE4DFFF5FFADFD85B623FD10F917" LSID="urn:lsid:biocol.org:col:34700" box="[603,660,1640,1666]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="2" pageNumber="3">CAS</collectionCode>
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224876, female,
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<quantity id="4CC99B6DFFF5FFADFCBBB623FC4CF917" box="[869,968,1640,1666]" metricMagnitude="-1" metricUnit="m" metricValue="2.02" pageId="2" pageNumber="3" unit="mm" value="202.0">202 mm</quantity>
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TL, collected from Tungkang,
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<collectingCountry id="F3267618FFF5FFADFAE0B623FA13F917" box="[1342,1431,1640,1666]" name="Taiwan" pageId="2" pageNumber="3">Taiwan</collectingCountry>
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,
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<geoCoordinate id="EE05504FFFF5FFADFF49B6DBFE95F93F" box="[151,273,1680,1706]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="22.46">22°27.6'N</geoCoordinate>
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,
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<geoCoordinate id="EE05504FFFF5FFADFEC2B6DBFE25F93F" box="[284,417,1680,1706]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="120.44">120°26.4'E</geoCoordinate>
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, collected by David A. Ebert on
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<date id="FF8F1048FFF5FFADFCFFB6DBFC4CF93F" box="[801,968,1680,1706]" pageId="2" pageNumber="3" value="1988-04-21">21 April 1988</date>
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, in excellent condition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6503FFF5FFADFF18B6FCFB6BF80F" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8B8E3688FFF5FFADFF18B6FCFB6BF80F" blockId="2.[151,1436,1639,2026]" pageId="2" pageNumber="3">
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<emphasis id="B945EA9AFFF5FFADFF18B6FCFEC2F944" bold="true" box="[198,326,1719,1745]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Body comparatively slender, head short, pre-first dorsal length 48% TL. Mouth wide, its width
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<quantity id="4CC99B6DFFF5FFADFF49B6ABFF6CF96F" box="[151,232,1760,1786]" metricMagnitude="-2" metricUnit="m" metricValue="1.3462" pageId="2" pageNumber="3" unit="in" value="0.53">0.53 in</quantity>
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head length. Snout very short, moderately flattened, broadly rounded; anterior nasal flaps not overlapping mouth posteriorly, narrowly lobate, inner margin with a deep notch. First dorsal fin relatively small; considerably larger than second dorsal fin; originates behind half of pelvic-fin base. Pectoral fins relatively large and broad; anal fin relatively large. Color pattern elaborate; speckled dorsal saddles; three lateral clusters of spots arranged into rosettes between paired fins; dark lateral blotch between paired fins absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B6503FFF5FFA9FF18B7ECFE64FE37" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
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<paragraph id="8B8E3688FFF5FFACFF18B7ECFD03FD2F" blockId="2.[151,1436,1639,2026]" lastBlockId="3.[151,1437,152,858]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<emphasis id="B945EA9AFFF5FFADFF18B7ECFEDAF854" bold="true" box="[198,350,1959,1985]" pageId="2" pageNumber="3">Description.</emphasis>
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Proportional dimensions in percentage of TL are shown in
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<tableCitation id="C6B30333FFF5FFADFBEEB7E3FB0FF857" box="[1072,1163,1960,1986]" captionStart="TABLE 1" captionStartId="6.[151,239,946,970]" captionTargetBox="[159,1427,1141,2015]" captionTargetPageId="6" captionText="TABLE 1. Morphometric and meristic data for the holotype (HT) specimens for Cephaloscyllium pardelotum sp. nov. (CAS 224876) and Cephaloscyllium maculatum sp. nov. (CAS 224877). TL is the actual measurement in millimeters; all other measurements are expressed as percentage of TL. VERT—total vertebral count, mono—monospondylous centra count, predip—precaudal diplospondylous centra count, caudaldip—caudal diplospondylous centra count, U tot—tooth count of upper jaw, L tot—tooth count of lower jaw. Abbreviations are listed in Appendix 1." httpUri="http://table.plazi.org/id/DF4E6600FFF1FFA9FF49B3F9FBF4FBC1" pageId="2" pageNumber="3" tableUuid="DF4E6600FFF1FFA9FF49B3F9FBF4FBC1">Table 1</tableCitation>
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. Head short, its length 0.27 times in precaudal length; pre-pectoral length 18.3% TL. Snout moderately flattened, broadly rounded, and short, preorbital length 0.30 times head length; 0.72 times interorbital width. Eye small, its horizontal eye length
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<quantity id="4CC99B6DFFF4FFACFF37B08BFEB8FF4F" box="[233,316,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="4.826" pageId="3" pageNumber="4" unit="in" value="0.19">0.19 in</quantity>
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head length; cat-like. Spiracle small, its diameter 0.27 times horizontal eye length. Gill openings very small, vertical length of first gill opening
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<quantity id="4CC99B6DFFF4FFACFD2CB0A3FCC0FE97" box="[754,836,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="4" unit="in" value="0.09">0.09 in</quantity>
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head length. Mouth relatively small, its width 0.53 times head length; labial furrows absent. Nostrils large; internarial distance
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<quantity id="4CC99B6DFFF4FFACFBEAB15BFB0EFEBF" box="[1076,1162,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" pageId="3" pageNumber="4" unit="in" value="0.24">0.24 in</quantity>
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mouth width; anterior nasal flaps elongate, lobed, with deep notch on inner margin, not overlapping mouth posteriorly. First dorsalfin small; first dorsal-fin height
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<quantity id="4CC99B6DFFF4FFACFDD5B12BFDD8FEEF" box="[523,604,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="4" unit="in" value="0.4">0.40 in</quantity>
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length; origin about opposite or behind half of pelvic-fin base; pre-first dorsal length 48.0% TL. Second dorsal fin small, only slightly smaller than first; second dorsal-fin height
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<quantity id="4CC99B6DFFF4FFACFAB7B1C3FF2AFE5F" metricMagnitude="-3" metricUnit="m" metricValue="9.144" pageId="3" pageNumber="4" unit="in" value="0.36">0.36 in</quantity>
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length; pre-second dorsal length 0.84 times precaudal length; second dorsal-fin height
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<quantity id="4CC99B6DFFF4FFACFB66B1FBFA8FFE5F" box="[1208,1291,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="1.8034" pageId="3" pageNumber="4" unit="in" value="0.71">0.71 in</quantity>
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first dorsalfin height. Pectoral fins relatively large and broad; anterior margin 0.63 times head length; base
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<quantity id="4CC99B6DFFF4FFACFAD9B193FADEFE67" box="[1287,1370,472,498]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="3" pageNumber="4" unit="in" value="0.72">0.72 in</quantity>
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anterior margin. Pelvic fin small; its length 0.14 times precaudal length; base
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<quantity id="4CC99B6DFFF4FFACFBD5B24BFBDAFD8F" box="[1035,1118,512,538]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="3" pageNumber="4" unit="in" value="0.66">0.66 in</quantity>
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length. Anal fin relatively large; its height 0.05 times precaudal length and its length 0.14 times precaudal length; larger than either dorsal fin; second dorsal-fin height
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<quantity id="4CC99B6DFFF4FFACFDC6B21BFDEFFDFF" box="[536,619,592,618]" metricMagnitude="-2" metricUnit="m" metricValue="1.8034" pageId="3" pageNumber="4" unit="in" value="0.71">0.71 in</quantity>
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||
anal-fin height; first dorsal-fin height about equal to anal-fin height. Caudal fin broad, asymmetrical, with subterminal notch; dorsal-lobe length 0.37 times precaudal length; ventral-lobe length
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||
<quantity id="4CC99B6DFFF4FFACFE8CB2EBFE27FD2F" box="[338,419,672,698]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" pageId="3" pageNumber="4" unit="in" value="0.35">0.35 in</quantity>
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dorsal-lobe length.
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</paragraph>
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<paragraph id="8B8E3688FFF4FFACFF1BB283FBCAFCA7" blockId="3.[151,1437,152,858]" pageId="3" pageNumber="4">Body firm, thick-skinned, with well calcified dermal denticles; no ridge of enlarged denticles along dorsal caudal-fin margin. Teeth very small and numerous; similar in both jaws; three cusps in females. Total vertebrae 120, monospondylous 44, precaudal 74, and pre-caudal diplospondylous 30.</paragraph>
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<paragraph id="8B8E3688FFF4FFACFF18B374FD7FFCCF" blockId="3.[151,1437,152,858]" box="[198,763,831,858]" pageId="3" pageNumber="4">
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<emphasis id="B945EA9AFFF4FFACFF18B374FD84FCCC" bold="true" box="[198,512,831,857]" pageId="3" pageNumber="4">Size and sexual maturity.</emphasis>
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Unknown at present.
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</paragraph>
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<caption id="DF4E6600FFF4FFACFF49B733FE8FF826" httpUri="https://zenodo.org/record/183987/files/figure.png" pageId="3" pageNumber="4" targetBox="[151,1435,924,1888]" targetPageId="3">
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||
<paragraph id="8B8E3688FFF4FFACFF49B733FE8FF826" blockId="3.[151,1436,1912,1971]" pageId="3" pageNumber="4">
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||
<emphasis id="B945EA9AFFF4FFACFF49B733FE9FF805" bold="true" box="[151,283,1912,1936]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
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<taxonomicName id="4C314D0BFFF4FFACFEFBB732FDDFF805" box="[293,603,1913,1936]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
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<emphasis id="B945EA9AFFF4FFACFEFBB732FDDFF805" box="[293,603,1913,1936]" italics="true" pageId="3" pageNumber="4">Cephaloscyllium pardelotum</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A27657E1FFF4FFACFDBAB733FD38F805" box="[612,700,1912,1936]" pageId="3" pageNumber="4" rank="species">
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||
<emphasis id="B945EA9AFFF4FFACFDBAB733FD38F805" bold="true" box="[612,700,1912,1936]" pageId="3" pageNumber="4">sp. nov.</emphasis>
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</taxonomicNameLabel>
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||
holotype (CAS 224876); a. lateral view; b. dorsal view. Photos by D.A. Ebert
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||
</paragraph>
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</caption>
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||
<caption id="DF4E6600FFF3FFABFF49B444FF6FFBDC" httpUri="https://zenodo.org/record/183988/files/figure.png" pageId="4" pageNumber="5" targetBox="[155,1430,199,1008]" targetPageId="4">
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||
<paragraph id="8B8E3688FFF3FFABFF49B444FF6FFBDC" blockId="4.[151,1436,1039,1097]" pageId="4" pageNumber="5">
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<emphasis id="B945EA9AFFF3FFABFF49B444FE9EFBB2" bold="true" box="[151,282,1039,1063]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
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Scatter diagram of proportional dimensions of anal-fin posterior margin length for select
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<taxonomicName id="4C314D0BFFF3FFABFB37B444FA18FBB3" box="[1257,1436,1039,1062]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B945EA9AFFF3FFABFB37B444FA18FBB3" box="[1257,1436,1039,1062]" italics="true" pageId="4" pageNumber="5">Cephaloscyllium</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E3688FFF3FFABFF18B437FC17FA43" blockId="4.[151,1437,1148,2017]" pageId="4" pageNumber="5">
|
||
<emphasis id="B945EA9AFFF3FFABFF18B437FED1FB03" bold="true" box="[198,341,1148,1174]" pageId="4" pageNumber="5">Coloration.</emphasis>
|
||
Background color golden to light brown dorsally; pale ventrally; small brown specks cover dorsal and lateral surfaces; dark lateral blotch between paired fins absent. Seven dorsal H-shaped saddles clearly defined; first in line with the gills, second above pectoral-fin insertion, third just before pelvic-fin origin, then one at each dorsal-fin base, one at caudal-fin origin, and dorsal-caudal lobe; saddles open, outline brown and variegated, not dark and solid. Rosette pattern present, consisting of a circle of 5–6 small brown speckles; three rosettes present laterally between the pectoral and pelvic fins; rosettes on head include one behind each orbit, one between orbits, one over each gill series, and a few on snout; each paired fin has one rosette placed dorsally, larger rosettes on larger fins; paired rosettes are present laterally above pelvic-fin origin, between dorsal fins, caudal peduncle, and lower caudal lobe.
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF3FFA9FF18B5AFFE64FE37" blockId="4.[151,1437,1148,2017]" lastBlockId="6.[151,1437,152,738]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
|
||
<emphasis id="B945EA9AFFF3FFABFF18B5AFFDD4FA6B" bold="true" box="[198,592,1508,1534]" pageId="4" pageNumber="5">Comparison with other species.</emphasis>
|
||
Morphometric data are presented in
|
||
<tableCitation id="C6B30333FFF3FFABFBDBB5AFFBD9FA6B" box="[1029,1117,1508,1534]" captionStart="TABLE 1" captionStartId="6.[151,239,946,970]" captionTargetBox="[159,1427,1141,2015]" captionTargetPageId="6" captionText="TABLE 1. Morphometric and meristic data for the holotype (HT) specimens for Cephaloscyllium pardelotum sp. nov. (CAS 224876) and Cephaloscyllium maculatum sp. nov. (CAS 224877). TL is the actual measurement in millimeters; all other measurements are expressed as percentage of TL. VERT—total vertebral count, mono—monospondylous centra count, predip—precaudal diplospondylous centra count, caudaldip—caudal diplospondylous centra count, U tot—tooth count of upper jaw, L tot—tooth count of lower jaw. Abbreviations are listed in Appendix 1." httpUri="http://table.plazi.org/id/DF4E6600FFF1FFA9FF49B3F9FBF4FBC1" pageId="4" pageNumber="5" tableUuid="DF4E6600FFF1FFA9FF49B3F9FBF4FBC1">Table 1</tableCitation>
|
||
.
|
||
<taxonomicName id="4C314D0BFFF3FFABFBB5B5AEFF47F9B3" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFBB5B5AEFF47F9B3" italics="true" pageId="4" pageNumber="5">Cephaloscyllium pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFF13B647FEA9F9B3" bold="true" box="[205,301,1548,1574]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFF13B647FEA9F9B3" box="[205,301,1548,1574]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C314D0BFFF3FFABFE0EB646FD1CF9B3" box="[464,664,1548,1574]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="circulopullum">
|
||
<emphasis id="B945EA9AFFF3FFABFE0EB646FE67F9B3" box="[464,483,1549,1574]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFE2AB646FD1CF9B3" box="[500,664,1549,1574]" italics="true" pageId="4" pageNumber="5">circulopullum</emphasis>
|
||
</taxonomicName>
|
||
in lacking a dark lateral blotch between the pectoral and pelvic fins, and possessing a lobed anterior nasal flap.
|
||
<taxonomicName id="4C314D0BFFF3FFABFD0BB67EFBADF9DB" box="[725,1065,1589,1614]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFD0BB67EFBADF9DB" box="[725,1065,1589,1614]" italics="true" pageId="4" pageNumber="5">Cephaloscyllium pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFBECB67FFB15F9DB" bold="true" box="[1074,1169,1588,1614]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFBECB67FFB15F9DB" box="[1074,1169,1588,1614]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a remarkably distinct color pattern from
|
||
<taxonomicName id="4C314D0BFFF3FFABFE6AB616FDC9F9E3" box="[436,589,1628,1654]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="fasciatum">
|
||
<emphasis id="B945EA9AFFF3FFABFE6AB616FE43F9E3" box="[436,455,1629,1654]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFE09B616FDC9F9E3" box="[471,589,1629,1654]" italics="true" pageId="4" pageNumber="5">fasciatum</emphasis>
|
||
</taxonomicName>
|
||
. Although the two species have open dorsal saddles,
|
||
<taxonomicName id="4C314D0BFFF3FFABFB19B616FAF4F9E3" box="[1223,1392,1628,1654]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFB19B616FB5EF9E3" box="[1223,1242,1629,1654]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFB34B616FAF4F9E3" box="[1258,1392,1629,1654]" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFAA7B617FF4EF90B" bold="true" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFAA7B617FF4EF90B" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possesses saddle borders that are not dark and solid as in
|
||
<taxonomicName id="4C314D0BFFF3FFABFCA8B6CEFB8FF90B" box="[886,1035,1668,1694]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="fasciatum">
|
||
<emphasis id="B945EA9AFFF3FFABFCA8B6CEFC0DF90B" box="[886,905,1669,1694]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFC47B6CEFB8FF90B" box="[921,1035,1669,1694]" italics="true" pageId="4" pageNumber="5">fasciatum</emphasis>
|
||
</taxonomicName>
|
||
. In contrast, the saddle borders of
|
||
<taxonomicName id="4C314D0BFFF3FFABFF49B6E6FEBBF953" box="[151,319,1708,1734]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFF49B6E6FF2EF953" box="[151,170,1709,1734]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFF67B6E6FEBBF953" box="[185,319,1709,1734]" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFE99B6E7FE21F953" bold="true" box="[327,421,1708,1734]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFE99B6E7FE21F953" box="[327,421,1708,1734]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are variegated and patchy and form an H-shape. Also, this new species has distinctive rosettes and no reticulated lines. The anal fin is slightly larger in the new species. Anal-fin posterior margin is 0.07 times in precaudal length for
|
||
<taxonomicName id="4C314D0BFFF3FFABFDF2B6B6FD50F883" box="[556,724,1788,1814]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFDF2B6B6FDBBF883" box="[556,575,1789,1814]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFD90B6B6FD50F883" box="[590,724,1789,1814]" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFD02B6B7FCBEF883" bold="true" box="[732,826,1788,1814]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFD02B6B7FCBEF883" box="[732,826,1788,1814]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but 0.03–0.04 times in precaudal length for
|
||
<taxonomicName id="4C314D0BFFF3FFABFA93B6B6FF63F8AB" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="fasciatum">
|
||
<emphasis id="B945EA9AFFF3FFABFA93B6B6FAE4F883" box="[1357,1376,1789,1814]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFAAEB6B6FF63F8AB" italics="true" pageId="4" pageNumber="5">fasciatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130A2A0DFFF3FFABFF29B76FFEC4F8AB" box="[247,320,1828,1854]" captionStart="FIGURE 2" captionStartId="4.[151,255,1039,1063]" captionTargetBox="[155,1430,199,1008]" captionTargetId="figure@4.[151,1436,194,1014]" captionTargetPageId="4" captionText="FIGURE 2. Scatter diagram of proportional dimensions of anal-fin posterior margin length for select Cephaloscyllium species." httpUri="https://zenodo.org/record/183988/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
).
|
||
<taxonomicName id="4C314D0BFFF3FFABFE89B76EFD2EF8AB" box="[343,682,1829,1854]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFE89B76EFD2EF8AB" box="[343,682,1829,1854]" italics="true" pageId="4" pageNumber="5">Cephaloscyllium pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFD6CB76FFC95F8AB" bold="true" box="[690,785,1828,1854]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFD6CB76FFC95F8AB" box="[690,785,1828,1854]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C314D0BFFF3FFABFC70B76EFBEAF8AB" box="[942,1134,1828,1854]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF3FFABFC70B76EFC45F8AB" box="[942,961,1829,1854]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFC0FB76EFBEAF8AB" box="[977,1134,1829,1854]" italics="true" pageId="4" pageNumber="5">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
in color pattern, and size and position of the dorsal fins. The new species does not display the distinctive polka-dots of juvenile
|
||
<taxonomicName id="4C314D0BFFF3FFABFA5FB706FEB1F805" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF3FFABFA5FB706FA10F8F3" box="[1409,1428,1869,1894]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFF49B73CFEB1F805" box="[151,309,1911,1936]" italics="true" pageId="4" pageNumber="5">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
, nor the dark lateral blotch of
|
||
<taxonomicName id="4C314D0BFFF3FFABFD78B73CFCEDF805" box="[678,873,1910,1936]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF3FFABFD78B73CFD3DF805" box="[678,697,1911,1936]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFD15B73CFCEDF805" box="[715,873,1911,1936]" italics="true" pageId="4" pageNumber="5">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
. Overall, both dorsal fins of
|
||
<taxonomicName id="4C314D0BFFF3FFABFB18B73CFAF4F805" box="[1222,1392,1910,1936]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFB18B73CFB5DF805" box="[1222,1241,1911,1936]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFB34B73CFAF4F805" box="[1258,1392,1911,1936]" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFAA7B73DFF4EF82C" bold="true" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFAA7B73DFF4EF82C" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are smaller than in other WNP species of swellshark. The first dorsal-fin base is relatively short in
|
||
<taxonomicName id="4C314D0BFFF3FFABFA90B7D4FF71F875" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFA90B7D4FAE5F82D" box="[1358,1377,1951,1976]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFAB7B7D4FF71F875" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFF22B78CFEDEF874" bold="true" box="[252,346,1991,2017]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFF22B78CFEDEF874" box="[252,346,1991,2017]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130A2A0DFFF3FFABFEB4B78CFE35F874" box="[362,433,1991,2017]" captionStart="FIGURE 3" captionStartId="5.[151,255,1012,1036]" captionTargetBox="[155,1430,192,996]" captionTargetId="figure@5.[151,1436,189,996]" captionTargetPageId="5" captionText="FIGURE 3. Scatter diagram of proportional dimensions of first dorsal-fin height for select Cephaloscyllium species." httpUri="https://zenodo.org/record/183989/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
|
||
). This length is 0.08 times in precaudal length in
|
||
<taxonomicName id="4C314D0BFFF3FFABFC28B78CFB1AF875" box="[1014,1182,1991,2017]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF3FFABFC28B78CFB8DF875" box="[1014,1033,1991,2016]" italics="true" pageId="4" pageNumber="5">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF3FFABFBC6B78CFB1AF875" box="[1048,1182,1991,2016]" italics="true" pageId="4" pageNumber="5">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF3FFABFB78B78CFA87F874" bold="true" box="[1190,1283,1991,2017]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="A27657E1FFF3FFABFB78B78CFA87F874" box="[1190,1283,1991,2017]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, whereas
|
||
<taxonomicName id="4C314D0BFFF1FFA9FF49B0D3FED3FF24" box="[151,343,152,178]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF1FFA9FF49B0D3FF2EFF24" box="[151,170,152,177]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FF67B0D3FED3FF24" box="[185,343,152,177]" italics="true" pageId="6" pageNumber="7">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
has a dorsal-fin base between
|
||
<quantity id="4CC99B6DFFF1FFA9FD65B0D3FCCAFF27" box="[699,846,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="6" pageNumber="7" unit="in" value="0.105" valueMax="0.12" valueMin="0.09">0.09–0.12 in</quantity>
|
||
precaudal length. The first dorsal-fin height is relatively short (
|
||
<figureCitation id="130A2A0DFFF1FFA9FEE2B08BFE00FF4F" box="[316,388,192,218]" captionStart="FIGURE 4" captionStartId="5.[151,255,1958,1982]" captionTargetBox="[163,1427,1102,1939]" captionTargetId="figure@5.[151,1436,1090,1941]" captionTargetPageId="5" captionText="FIGURE 4. Scatter diagram of proportional dimensions of first dorsal-fin base length for select Cephaloscyllium species." httpUri="https://zenodo.org/record/183990/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
|
||
) in
|
||
<taxonomicName id="4C314D0BFFF1FFA9FE68B08BFDE4FF4C" box="[438,608,192,218]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FE68B08BFE4DFF4C" box="[438,457,192,217]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FE04B08BFDE4FF4C" box="[474,608,192,217]" italics="true" pageId="6" pageNumber="7">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FDB7B0F4FD4CFF4C" bold="true" box="[617,712,191,217]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FDB7B0F4FD4CFF4C" box="[617,712,191,217]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, at
|
||
<quantity id="4CC99B6DFFF1FFA9FD29B08BFCCEFF4F" box="[759,842,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="in" value="0.05">0.05 in</quantity>
|
||
precaudal length. This measurement is 0.06–0.07 times in precaudal length in
|
||
<taxonomicName id="4C314D0BFFF1FFA9FE39B0A3FD2CFE94" box="[487,680,232,258]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF1FFA9FE39B0A3FE7EFE94" box="[487,506,232,257]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FDD4B0A3FD2CFE94" box="[522,680,232,257]" italics="true" pageId="6" pageNumber="7">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C314D0BFFF1FFA9FD09B0A3FC04FE94" box="[727,896,232,258]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FD09B0A3FD6EFE94" box="[727,746,232,257]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FD24B0A3FC04FE94" box="[762,896,232,257]" italics="true" pageId="6" pageNumber="7">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FC56B0ACFC6AFE94" bold="true" box="[904,1006,231,257]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FC56B0ACFC6CFE94" box="[904,1000,231,257]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
the first dorsal fin originates behind the anterior half of the pelvic-fin base. In
|
||
<taxonomicName id="4C314D0BFFF1FFA9FDA2B15BFCB8FEBC" box="[636,828,272,298]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF1FFA9FDA2B15BFD0BFEBC" box="[636,655,272,297]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FD40B15BFCB8FEBC" box="[670,828,272,297]" italics="true" pageId="6" pageNumber="7">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
, however, the first dorsal fin originates closer to the pelvic-fin origin.
|
||
<taxonomicName id="4C314D0BFFF1FFA9FEB7B173FD39FEC4" box="[361,701,312,337]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FEB7B173FD39FEC4" box="[361,701,312,337]" italics="true" pageId="6" pageNumber="7">Cephaloscyllium pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FD18B17CFCA1FEC4" bold="true" box="[710,805,311,337]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FD18B17CFCA1FEC4" box="[710,805,311,337]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from adult
|
||
<taxonomicName id="4C314D0BFFF1FFA9FB5AB173FA9FFEC4" box="[1156,1307,312,338]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="umbratile">
|
||
<emphasis id="B945EA9AFFF1FFA9FB5AB173FB13FEC4" box="[1156,1175,312,337]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FB77B173FA9FFEC4" box="[1193,1307,312,337]" italics="true" pageId="6" pageNumber="7">umbratile</emphasis>
|
||
</taxonomicName>
|
||
by overall color pattern, and morphology of the anterior nasal flap.
|
||
<taxonomicName id="4C314D0BFFF1FFA9FCE9B12BFBF2FEEC" box="[823,1142,352,377]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="umbratile">
|
||
<emphasis id="B945EA9AFFF1FFA9FCE9B12BFBF2FEEC" box="[823,1142,352,377]" italics="true" pageId="6" pageNumber="7">Cephaloscyllium umbratile</emphasis>
|
||
</taxonomicName>
|
||
does not have open saddles, nor are they H-shaped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4E6600FFF2FFAAFF49B3BFFAF7FB9E" box="[151,1395,1012,1036]" httpUri="https://zenodo.org/record/183989/files/figure.png" pageId="5" pageNumber="6" targetBox="[155,1430,192,996]" targetPageId="5">
|
||
<paragraph id="8B8E3688FFF2FFAAFF49B3BFFAF7FB9E" blockId="5.[151,1395,1012,1036]" box="[151,1395,1012,1036]" pageId="5" pageNumber="6">
|
||
<emphasis id="B945EA9AFFF2FFAAFF49B3BFFE9DFB99" bold="true" box="[151,281,1012,1036]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
Scatter diagram of proportional dimensions of first dorsal-fin height for select
|
||
<taxonomicName id="4C314D0BFFF2FFAAFBBBB3BEFA9CFB99" box="[1125,1304,1013,1036]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B945EA9AFFF2FFAAFBBBB3BEFA9CFB99" box="[1125,1304,1013,1036]" italics="true" pageId="5" pageNumber="6">Cephaloscyllium</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E6600FFF2FFAAFF49B7EDFF4CF875" httpUri="https://zenodo.org/record/183990/files/figure.png" pageId="5" pageNumber="6" targetBox="[163,1427,1102,1939]" targetPageId="5">
|
||
<paragraph id="8B8E3688FFF2FFAAFF49B7EDFF4CF875" blockId="5.[151,1436,1958,2016]" pageId="5" pageNumber="6">
|
||
<emphasis id="B945EA9AFFF2FFAAFF49B7EDFE9FF82B" bold="true" box="[151,283,1958,1982]" pageId="5" pageNumber="6">FIGURE 4.</emphasis>
|
||
Scatter diagram of proportional dimensions of first dorsal-fin base length for select
|
||
<taxonomicName id="4C314D0BFFF2FFAAFB6AB7EDFAE3F828" box="[1204,1383,1958,1981]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B945EA9AFFF2FFAAFB6AB7EDFAE3F828" box="[1204,1383,1958,1981]" italics="true" pageId="5" pageNumber="6">Cephaloscyllium</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32B6503FFF1FFA9FF18B1E4FBAAFDD7" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8B8E3688FFF1FFA9FF18B1E4FBAAFDD7" blockId="6.[151,1437,152,738]" pageId="6" pageNumber="7">
|
||
<emphasis id="B945EA9AFFF1FFA9FF18B1E4FEC4FE5C" bold="true" box="[198,320,431,457]" pageId="6" pageNumber="7">Remarks.</emphasis>
|
||
If young similar-sized specimens of
|
||
<taxonomicName id="4C314D0BFFF1FFA9FD2EB1FBFC1CFE5C" box="[752,920,432,458]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FD2EB1FBFC87FE5C" box="[752,771,432,457]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FCCCB1FBFC1CFE5C" box="[786,920,432,457]" italics="true" pageId="6" pageNumber="7">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FC41B1E4FC78FE5C" bold="true" box="[927,1020,431,457]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FC41B1E4FC78FE5C" box="[927,1020,431,457]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C314D0BFFF1FFA9FBD5B1FBFB55FE5C" box="[1035,1233,432,458]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="circulopullum">
|
||
<emphasis id="B945EA9AFFF1FFA9FBD5B1FBFB9AFE5C" box="[1035,1054,432,457]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FBF3B1FBFB55FE5C" box="[1069,1233,432,457]" italics="true" pageId="6" pageNumber="7">circulopullum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C314D0BFFF1FFA9FB01B1FBFAF7FE5C" box="[1247,1395,432,458]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="fasciatum">
|
||
<emphasis id="B945EA9AFFF1FFA9FB01B1FBFB76FE5C" box="[1247,1266,432,457]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FADFB1FBFAF7FE5C" box="[1281,1395,432,457]" italics="true" pageId="6" pageNumber="7">fasciatum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C314D0BFFF1FFA9FA5FB1FBFEB1FE64" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="sarawakensis">
|
||
<emphasis id="B945EA9AFFF1FFA9FA5FB1FBFA10FE5C" box="[1409,1428,432,457]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FF49B193FEB1FE64" box="[151,309,472,497]" italics="true" pageId="6" pageNumber="7">sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C314D0BFFF1FFA9FEAAB193FD83FE64" box="[372,519,472,498]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="umbratile">
|
||
<emphasis id="B945EA9AFFF1FFA9FEAAB193FE03FE64" box="[372,391,472,497]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FE4BB193FD83FE64" box="[405,519,472,497]" italics="true" pageId="6" pageNumber="7">umbratile</emphasis>
|
||
</taxonomicName>
|
||
are examined side-by-side, it is fairly easy to tell that they are all separate species (
|
||
<figureCitation id="130A2A0DFFF1FFA9FF09B24BFEA5FD8F" box="[215,289,512,538]" captionStart="FIGURE 5" captionStartId="9.[151,255,1897,1921]" captionTargetBox="[195,1374,202,1863]" captionTargetId="figure@9.[181,1405,194,1870]" captionTargetPageId="9" captionText="FIGURE 5. Lateral views of the WNP Cephaloscyllium, all about 220 mm TL; a. Cephaloscyllium pardelotum sp. nov.; b. Cephaloscyllium maculatum sp. nov.; c. Cephaloscyllium fasciatum; d. Cephaloscyllium umbratile; e. Cephaloscyllium sarawakensis; f. Cephaloscyllium circulopullum. Illustration by J. A. S. - Da Silva." httpUri="https://zenodo.org/record/183992/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
|
||
). Although the color pattern of adult
|
||
<taxonomicName id="4C314D0BFFF1FFA9FD36B24BFC17FD8C" box="[744,915,512,538]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FD36B24BFD7FFD8C" box="[744,763,512,537]" italics="true" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<emphasis id="B945EA9AFFF1FFA9FCD3B24BFC17FD8C" box="[781,915,512,537]" italics="true" pageId="6" pageNumber="7">pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FC43B1B4FC79FD8C" bold="true" box="[925,1021,511,537]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FC43B1B4FC79FD8C" box="[925,1021,511,537]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is presently unknown, its juvenile markings are quite distinct from any other
|
||
<taxonomicName id="4C314D0BFFF1FFA9FD55B263FCD4FDD4" box="[651,848,552,577]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B945EA9AFFF1FFA9FD55B263FCD4FDD4" box="[651,848,552,577]" italics="true" pageId="6" pageNumber="7">Cephaloscyllium</emphasis>
|
||
</taxonomicName>
|
||
within this region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6503FFF1FFA9FF18B204FA1FFDFF" box="[198,1435,591,618]" pageId="6" pageNumber="7" type="distribution">
|
||
<paragraph id="8B8E3688FFF1FFA9FF18B204FA1FFDFF" blockId="6.[151,1437,152,738]" box="[198,1435,591,618]" pageId="6" pageNumber="7">
|
||
<emphasis id="B945EA9AFFF1FFA9FF18B204FEE3FDFC" bold="true" box="[198,359,591,617]" pageId="6" pageNumber="7">Distribution.</emphasis>
|
||
<taxonomicName id="4C314D0BFFF1FFA9FEB0B21BFD44FDFC" box="[366,704,592,617]" class="Elasmobranchii" family="Scyliorhinidae" genus="Cephaloscyllium" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pardelotum" status="sp. nov.">
|
||
<emphasis id="B945EA9AFFF1FFA9FEB0B21BFD44FDFC" box="[366,704,592,617]" italics="true" pageId="6" pageNumber="7">Cephaloscyllium pardelotum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B945EA9AFFF1FFA9FD19B204FCA1FDFC" bold="true" box="[711,805,591,617]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="A27657E1FFF1FFA9FD19B204FCA1FDFC" box="[711,805,591,617]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from off the coast of southern
|
||
<collectingCountry id="F3267618FFF1FFA9FAE0B21BFA1CFDFF" box="[1342,1432,592,618]" name="Taiwan" pageId="6" pageNumber="7">Taiwan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6503FFF1FFA9FF18B23CFD64FD77" pageId="6" pageNumber="7" type="etymology">
|
||
<paragraph id="8B8E3688FFF1FFA9FF18B23CFDC0FD2F" blockId="6.[151,1437,152,738]" pageId="6" pageNumber="7">
|
||
<emphasis id="B945EA9AFFF1FFA9FF18B23CFED7FD04" bold="true" box="[198,339,631,657]" pageId="6" pageNumber="7">Etymology.</emphasis>
|
||
The species is named after the Latin
|
||
<emphasis id="B945EA9AFFF1FFA9FCC3B233FCE9FD04" box="[797,877,632,657]" italics="true" pageId="6" pageNumber="7">pardus</emphasis>
|
||
for leopard, referring to the rosette-type spots along the dorsal and lateral surfaces.
|
||
</paragraph>
|
||
<paragraph id="8B8E3688FFF1FFA9FF18B28CFD64FD77" blockId="6.[151,1437,152,738]" box="[198,736,711,738]" pageId="6" pageNumber="7">
|
||
<emphasis id="B945EA9AFFF1FFA9FF18B28CFE09FD74" bold="true" box="[198,397,711,737]" pageId="6" pageNumber="7">Common name.</emphasis>
|
||
Leopard-spotted swellshark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |