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<emphasis id="B954EAA4FFD04B33B388A4FE1162FA8C" bold="true" box="[926,1144,1275,1299]" italics="true" pageId="42" pageNumber="267">Quedius hispanicus</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B468A4F91023FA8C" author="BERNHAUER M." box="[1150,1337,1276,1299]" pageId="42" pageNumber="267" pagination="338 - 341" refId="ref80314" refString="BERNHAUER M. 1898: Neue Staphyliniden aus Europa. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 48: 338 - 341." type="journal article" year="1898">Bernhauer, 1898</bibRefCitation>
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<paragraph id="8B9F36B6FFD04B33B3FCA51F11F4FAB2" blockId="42.[828,1437,1306,1600]" box="[1002,1262,1306,1325]" pageId="42" pageNumber="267">
(
<figureCitation id="131B2A33FFD04B33B3E6A51F113CFAB2" box="[1008,1062,1306,1325]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="42" pageNumber="267">Figs 1</figureCitation>
, 
<figureCitation id="131B2A33FFD04B33B427A51F1157FAB2" box="[1073,1101,1306,1325]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="42" pageNumber="267">2D</figureCitation>
, 
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<figureCitation id="131B2A33FFD04B33B47DA51F1180FAB2" box="[1131,1178,1306,1325]" captionStart="Fig" captionStartId="19.[151,182,2028,2047]" captionTargetBox="[203,1360,198,1958]" captionTargetId="figure-13@19.[183,1405,186,1971]" captionTargetPageId="19" captionText="Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915." figureDoi="http://doi.org/10.5281/zenodo.7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="42" pageNumber="267">9A,B</figureCitation>
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<figureCitation id="131B2A33FFD04B33B4B5A51F11D2FAB2" box="[1187,1224,1306,1325]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="42" pageNumber="267">14C</figureCitation>
, 
<figureCitation id="131B2A33FFD04B33B4C7A51F11FDFAB2" box="[1233,1255,1306,1325]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="42" pageNumber="267">19</figureCitation>
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<emphasis id="B954EAA4FFD04B33B32AA53816F8FACF" box="[828,994,1341,1360]" italics="true" pageId="42" pageNumber="267">Quedius hispanicus</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B3F1A5381187FACF" author="BERNHAUER M." box="[999,1181,1341,1360]" pageId="42" pageNumber="267" pagination="338 - 341" refId="ref80314" refString="BERNHAUER M. 1898: Neue Staphyliniden aus Europa. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 48: 338 - 341." type="journal article" year="1898">Bernhauer, 1898: 341</bibRefCitation>
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locality: Cartagena und Cadix in 
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<emphasis id="B954EAA4FFD04B33B3EFA552115DFAF5" bold="true" box="[1017,1095,1367,1386]" pageId="42" pageNumber="267">stat. rev.</emphasis>
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<emphasis id="B954EAA4FFD04B33B32AA57716A5FA1A" box="[828,959,1394,1413]" italics="true" pageId="42" pageNumber="267">Quedius cobosi</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B3D5A5771178FA1A" author="COIFFAIT H." box="[963,1122,1394,1413]" pageId="42" pageNumber="267" pagination="293 - 296" refId="ref81616" refString="COIFFAIT H. 1964: Deux nouveaux Quedius s. str. de la region mediterraneenne occidentale. Revue Francaise d'Entomologie 31: 293 - 296." type="journal article" year="1964">Coiffait, 1964: 293</bibRefCitation>
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locality: 
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: Province de Almeria, Seron], 
<emphasis id="B954EAA4FFD04B33B3F8A5881127FA3F" bold="true" box="[1006,1085,1421,1440]" pageId="42" pageNumber="267">syn. nov.</emphasis>
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<emphasis id="B954EAA4FFD04B33B32AA5A216FDFA25" box="[828,999,1447,1466]" italics="true" pageId="42" pageNumber="267">Quedius hammianus</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B3FCA5A21176FA25" author="SHARP D." box="[1002,1132,1447,1466]" pageId="42" pageNumber="267" pagination="57" refId="ref91034" refString="SHARP D. 1911: A new British Quedius. Entomologist's Monthly Magazine 47: 57." type="journal article" year="1911">Sharp, 1911:57</bibRefCitation>
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locality:Anglia:Deal, Strood, Lymington, and Hayling Island], 
<emphasis id="B954EAA4FFD04B33B460A5C711DBFA4A" bold="true" box="[1142,1217,1474,1493]" pageId="42" pageNumber="267">syn. rev.</emphasis>
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<treatmentCitationGroup id="AB301198FFD04B33B32AA5D816BFF995" pageId="42" pageNumber="267">
<treatmentCitation id="0A8110A7FFD04B33B32AA5D8116BFA6F" author="BLUMML E. K." box="[828,1137,1501,1520]" page="216" pageId="42" pageNumber="267" year="1898">
<taxonomicName id="4C204D35FFD04B33B32AA5D8116BFA6F" ID-CoL="4R3VB" authority="Blumml, 1898: 216" authorityName="Blumml" authorityPageNumber="216" authorityYear="1898" box="[828,1137,1501,1520]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="rufulus">
<emphasis id="B954EAA4FFD04B33B32AA5D816DEFA6F" box="[828,964,1501,1520]" italics="true" pageId="42" pageNumber="267">Quedius rufulus</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B3DCA5D8116BFA6F" author="BLUMML E. K." box="[970,1137,1501,1520]" pageId="42" pageNumber="267" pagination="216" refId="ref80473" refString="BLUMML E. K. 1898: Eine neue Kafer-Variation. Illustrierte Zeitschrift fur Entomologie 3: 216." type="journal article" year="1898">Blümml, 1898: 216</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="549B8814FFD04B33B46DA5D811B2FA6F" box="[1147,1192,1501,1520]" pageId="42" pageNumber="267">Type</typeStatus>
locality: 
<collectingCountry id="F3377626FFD04B33B4EEA5D81025FA6F" box="[1272,1343,1501,1520]" name="Spain" pageId="42" pageNumber="267">Spanien</collectingCountry>
, 
<collectingCountry id="F3377626FFD04B33B55EA5D8108FFA6F" box="[1352,1429,1501,1520]" name="Grenada" pageId="42" pageNumber="267">Granada</collectingCountry>
], 
<emphasis id="B954EAA4FFD04B33B34CA5F216BFF995" bold="true" box="[858,933,1527,1546]" pageId="42" pageNumber="267">syn. rev.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9F36B6FFD04B33B32AA6171161F9DF" blockId="42.[828,1437,1306,1600]" pageId="42" pageNumber="267">
<taxonomicName id="4C204D35FFD04B33B32AA6171137F9BA" ID-CoL="4R3VL" authority="Last, 1952" authorityName="Last" authorityYear="1952" box="[828,1069,1554,1573]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B954EAA4FFD04B33B32AA61716C9F9BA" box="[828,979,1554,1573]" italics="true" pageId="42" pageNumber="267">Quedius secundus</emphasis>
<bibRefCitation id="EFB14B47FFD04B33B3C0A6171137F9BA" author="LAST H." box="[982,1069,1554,1573]" pageId="42" pageNumber="267" pagination="148 - 150" refId="ref86758" refString="LAST H. 1952: Taxonomic notes on Quedius molochinus Grav. and Q. pallipes Lucas (Col., Staphylinidae). Entomologist's Monthly Magazine 88: 148 - 150." type="journal article" year="1952">Last, 1952</bibRefCitation>
</taxonomicName>
[
<typeStatus id="549B8814FFD04B33B420A6171178F9BA" box="[1078,1122,1554,1573]" pageId="42" pageNumber="267">Type</typeStatus>
locality:Studland, 
<collectingRegion id="49E4F854FFD04B33B517A6171026F9BA" box="[1281,1340,1554,1573]" country="United Kingdom" name="Dorset" pageId="42" pageNumber="267">Dorset</collectingRegion>
; Mudeford, 
<collectingRegion id="49E4F854FFD04B33B34CA62816A0F9DF" box="[858,954,1581,1600]" country="United Kingdom" name="Hampshire" pageId="42" pageNumber="267">Hampshire</collectingRegion>
; Chichester], 
<emphasis id="B954EAA4FFD04B33B426A6281161F9DF" bold="true" box="[1072,1147,1581,1600]" pageId="42" pageNumber="267">syn. rev.</emphasis>
</paragraph>
<paragraph id="8B9F36B6FFD04B33B32AA6531128F78B" blockId="42.[828,1438,1622,2068]" pageId="42" pageNumber="267">
<emphasis id="B954EAA4FFD04B33B32AA65316BEF9F6" bold="true" box="[828,932,1622,1641]" pageId="42" pageNumber="267">References.</emphasis>
Fൺඎඏൾඅ (1865): 16 (synonym of 
<taxonomicName id="4C204D35FFD04B33B4D1A653105FF9F6" ID-CoL="4R2YX" box="[1223,1349,1622,1641]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFD04B33B4D1A653105FF9F6" box="[1223,1349,1622,1641]" italics="true" pageId="42" pageNumber="267">Q. molochinus</emphasis>
</taxonomicName>
); Kඋൺൺඍඓ (1867): 414 (synonym of 
<taxonomicName id="4C204D35FFD04B33B435A67411B9F91B" ID-CoL="4R2YX" box="[1059,1187,1649,1668]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFD04B33B435A67411B9F91B" box="[1059,1187,1649,1668]" italics="true" pageId="42" pageNumber="267">Q. molochinus</emphasis>
</taxonomicName>
); GൺඇGඅൻൺඎൾඋ (1895): 403 (synonym of 
<taxonomicName id="4C204D35FFD04B33B3A3A689112CF900" ID-CoL="4R2YX" box="[949,1078,1676,1695]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFD04B33B3A3A689112CF900" box="[949,1078,1676,1695]" italics="true" pageId="42" pageNumber="267">Q. molochinus</emphasis>
</taxonomicName>
); BൾඋඇHൺඎൾඋ (1900): 46 (distribution); Pඈඋඍൺ (1907): 129 (synonym of 
<taxonomicName id="4C204D35FFD04B33B44FA6A311CCF926" ID-CoL="4R2YX" box="[1113,1238,1702,1721]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFD04B33B44FA6A311CCF926" box="[1113,1238,1702,1721]" italics="true" pageId="42" pageNumber="267">Q. molochinus</emphasis>
</taxonomicName>
); Gඋංൽൾඅඅං (1924): 81 (subspecies of 
<taxonomicName id="4C204D35FFD04B33B3ABA6C41121F94B" ID-CoL="4R2YX" box="[957,1083,1729,1748]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFD04B33B3ABA6C41121F94B" box="[957,1083,1729,1748]" italics="true" pageId="42" pageNumber="267">Q. molochinus</emphasis>
</taxonomicName>
; characters); SർHൾൾඋඉൾඅඍඓ (1933): 1457 (synonym of 
<taxonomicName id="4C204D35FFD04B33B3A6A6D91139F970" ID-CoL="4RKPS" authorityName="Stephens" authorityYear="1832" box="[944,1059,1756,1775]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B954EAA4FFD04B33B3A6A6D91139F970" box="[944,1059,1756,1775]" italics="true" pageId="42" pageNumber="267">Q. picipennis</emphasis>
</taxonomicName>
); MඣඊඎංGඇඈඇ (1937): 3 (as 
<taxonomicName id="4C204D35FFD04B33B508A6D91665F896" ID-CoL="4R2YZ" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="subSpecies" species="molochinus" subSpecies="pallipes">
<emphasis id="B954EAA4FFD04B33B508A6D91665F896" italics="true" pageId="42" pageNumber="267">Q. molochinus pallipes</emphasis>
</taxonomicName>
); KඈඋGൾ (1960b):69 (synonym of 
<taxonomicName id="4C204D35FFD04B33B4B5A6F311E5F896" ID-CoL="4R2YZ" authorityName="Lucas" authorityYear="1846" box="[1187,1279,1782,1801]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD04B33B4B5A6F311E5F896" box="[1187,1279,1782,1801]" italics="true" pageId="42" pageNumber="267">Q. pallipes</emphasis>
</taxonomicName>
); Cඈංൿൿൺංඍ (1961): 56 (characters and distribution); (1978):196 (characters); Hൺඇඌൾඇ (1962): 319 (characters and distribution); Sආൾඍൺඇൺ (1962a): 134 (characters); (1963): 39; (1970): 61 (distribution); Lൺඌඍ (1963): 45 (synonym of 
<taxonomicName id="4C204D35FFD04B33B591A74316B1F8EB" ID-CoL="4R2Z4" authorityName="Fairmaire" authorityYear="1861" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD04B33B591A74316B1F8EB" italics="true" pageId="42" pageNumber="267">Q. simplicifrons</emphasis>
</taxonomicName>
); LඈHඌൾ (1964): 211 (characters and distribution); Hඈඋං- ඈඇ (1965): 276 (distribution); Bඈඋൽඈඇං (1976a): 100 (synonym of 
<taxonomicName id="4C204D35FFD04B33B591A779169AF836" ID-CoL="4R2YZ" authorityName="Lucas" authorityYear="1846" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD04B33B591A779169AF836" italics="true" pageId="42" pageNumber="267">Q. pallipes</emphasis>
</taxonomicName>
); Pඈඉൾ (1977): 31 (distribution); LඎർHඍ (1987): 109 (distribution); LඈHඌൾ &amp; LඎർHඍ (1989): 172 (distribution); Iඌඋൺൾඅඌඈඇ (1990): 3 (distribution); BඈඋGൾඌ (1990): Table II (distribution); WൾඅർH (1993): 229 (ovariole number and ovary structure); Oඎඍൾඋൾඅඈ et al. (1995): 80 (distribution; as 
<taxonomicName id="4C204D35FFD04B33B3DEA804113CF78B" ID-CoL="4R2YZ" authorityName="Lucas" authorityYear="1846" box="[968,1062,2049,2068]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="267" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD04B33B3DEA804113CF78B" box="[968,1062,2049,2068]" italics="true" pageId="42" pageNumber="267">Q. pallipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD14B32B081A0B811E6FE63" pageId="43" pageNumber="268" type="materials_examined">
<paragraph id="8B9F36B6FFD14B32B081A0B81751FE99" blockId="43.[150,761,184,2075]" pageId="43" pageNumber="268">
<emphasis id="B954EAA4FFD14B32B081A0B8149FFF4F" bold="true" box="[151,389,189,208]" pageId="43" pageNumber="268">Type material examined.</emphasis>
<taxonomicName id="4C204D35FFD14B32B19BA0B81702FF4F" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[397,536,189,208]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD14B32B19BA0B81702FF4F" box="[397,536,189,208]" italics="true" pageId="43" pageNumber="268">Quedius cobosi</emphasis>
</taxonomicName>
: Pൺඋൺඍඒඉൾ: J (
<collectionCode id="ED31AE73FFD14B32B2B1A0B817F4FF4F" box="[679,750,189,208]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="43" pageNumber="268">MNHN</collectionCode>
), labelled: “ 
<typeStatus id="549B8814FFD14B32B0FAA0DD1442FF74" box="[236,344,216,235]" pageId="43" pageNumber="268" type="paratype">PARATYPE</typeStatus>
[red label] / 
<taxonomicName id="4C204D35FFD14B32B1D5A0DD17E2FF74" authority="Coiff. H. COIFFAIT" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[451,760,216,235]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="cobosi">Quedius cobosi Coiff. H. COIFFAIT</taxonomicName>
det. 1964 / Laujar Almeria - 
<collectingCountry id="F3377626FFD14B32B192A0F614D9FE99" box="[388,451,243,262]" name="Spain" pageId="43" pageNumber="268">España</collectingCountry>
A. Cobos coll.”
</paragraph>
<paragraph id="8B9F36B6FFD14B32B081A10811E6FE63" blockId="43.[150,761,184,2075]" lastBlockId="43.[828,1437,189,508]" pageId="43" pageNumber="268">
<emphasis id="B954EAA4FFD14B32B081A108172BFEBF" bold="true" box="[151,561,269,288]" pageId="43" pageNumber="268">
Additional material examined. 
<collectingCountry id="F3377626FFD14B32B1AFA1081737FEBF" box="[441,557,269,288]" name="Denmark" pageId="43" pageNumber="268">DENMARK</collectingCountry>
:
</emphasis>
Lakolk, [55.14, 8.49], debris on beach, 
<date id="FF9E1076FFD14B32B130A12D1464FEA4" box="[294,382,296,315]" pageId="43" pageNumber="268" value="2013-11-01">1.XI.2013</date>
, leg. J. Pedersen &amp; A. Brunke (1 J 
<collectionCode id="ED31AE73FFD14B32B2BFA12D17F4FEA4" box="[681,750,296,315]" pageId="43" pageNumber="268">NHMD</collectionCode>
); Skallingen, [55.50, 8.25], 
<date id="FF9E1076FFD14B32B163A14614CBFEC9" box="[373,465,323,342]" pageId="43" pageNumber="268" value="1959-10-17">17.X.1959</date>
, leg. F. Bangsholt (2 JJ 
<collectionCode id="ED31AE73FFD14B32B2BCA14617F5FEC9" box="[682,751,323,342]" pageId="43" pageNumber="268">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B081A1581417FEEF" bold="true" box="[151,269,349,368]" pageId="43" pageNumber="268">
<collectingCountry id="F3377626FFD14B32B081A1581413FEEF" box="[151,265,349,368]" name="Germany" pageId="43" pageNumber="268">GERMANY</collectingCountry>
:
</emphasis>
Archsum, [Sylt], [54.87, 8.39], 
<date id="FF9E1076FFD14B32B230A1581776FEEF" box="[550,620,349,368]" pageId="43" pageNumber="268" value="1901-09">IX.1901</date>
(1 
<collectionCode id="ED31AE73FFD14B32B29DA15817A4FEEF" box="[651,702,349,368]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Insel Borkum, [53.58, 6.66], 
<date id="FF9E1076FFD14B32B148A17D14D5FE14" box="[350,463,376,395]" pageId="43" pageNumber="268" value="1937-08-12">12.VIII.1937</date>
, leg. Struse (
<specimenCount id="9D26FD3FFFD14B32B22DA17D1772FE14" box="[571,616,370,395]" pageId="43" pageNumber="268" type="female">2 ♀♀</specimenCount>
4 JJ 
<collectionCode id="ED31AE73FFD14B32B2B6A17D17C0FE14" box="[672,730,376,395]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="268">NMW</collectionCode>
); N Dagebüll, Nordfriesland, 
<geoCoordinate id="EE145071FFD14B32B17BA19614AEFE39" box="[365,436,403,422]" degrees="54.7361" direction="north" orientation="latitude" pageId="43" pageNumber="268" precision="5" value="54.7361">54.7361</geoCoordinate>
, 
<geoCoordinate id="EE145071FFD14B32B1AAA19614E3FE39" box="[444,505,403,422]" degrees="8.7094" direction="east" orientation="longitude" pageId="43" pageNumber="268" precision="5" value="8.7094">8.7094</geoCoordinate>
, salt pasture among stones and clay on dam, 
<date id="FF9E1076FFD14B32B11AA1A81464FE5F" box="[268,382,429,448]" pageId="43" pageNumber="268" value="2017-08-25">25.VIII.2017</date>
, leg. Wrase &amp; Laser (1 J cSch); Memmert, Juist, [53.64, 6.88], 
<date id="FF9E1076FFD14B32B15AA1CD14A4FE44" box="[332,446,456,475]" pageId="43" pageNumber="268" value="1917-08-13">13.VIII.1917</date>
, leg. Alfrum (1 
<collectionCode id="ED31AE73FFD14B32B245A1CD179DFE44" box="[595,647,456,475]" pageId="43" pageNumber="268">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B281A1CD17E2FE44" bold="true" box="[663,760,456,475]" pageId="43" pageNumber="268">
<collectingCountry id="F3377626FFD14B32B281A1CD17EEFE44" box="[663,756,456,475]" name="France" pageId="43" pageNumber="268">FRANCE</collectingCountry>
:
</emphasis>
Bordeuax, St. Louis, [44.94, -0.52], 
<date id="FF9E1076FFD14B32B1D0A1E61704FE69" box="[454,542,483,502]" pageId="43" pageNumber="268" value="1964-02-22">22.II.1964</date>
leg. C. Jeanne (1 JcBor); Camargue, Etang du Fangassier, [43.42, 4.63], 
<date id="FF9E1076FFD14B32B209A1F8176CFD8F" box="[543,630,509,528]" pageId="43" pageNumber="268" value="1994-05-20">20.V.1994</date>
salt marsh, leg. Schülke &amp; Grünberg (1 J cSch); Sainte Marie la Mer, [42.72, 3.03], 
<date id="FF9E1076FFD14B32B081A23615EDFDD9" box="[151,247,563,582]" pageId="43" pageNumber="268" value="1955-04-28">28.IV.1955</date>
, leg. J.T. Skovgaard (
<specimenCount id="9D26FD3FFFD14B32B1B8A23614D0FDDA" box="[430,458,556,582]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B1C6A236170FFDD9" box="[464,533,563,582]" pageId="43" pageNumber="268">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B234A23617BBFDD9" bold="true" box="[546,673,563,582]" pageId="43" pageNumber="268">
<collectingCountry id="F3377626FFD14B32B234A2361784FDD9" box="[546,670,563,582]" name="Portugal" pageId="43" pageNumber="268">PORTUGAL</collectingCountry>
:
</emphasis>
<collectingRegion id="49E4F854FFD14B32B2B3A23617EEFDD9" box="[677,756,563,582]" country="Portugal" name="Lisboa" pageId="43" pageNumber="268">Lissabon</collectingRegion>
, [38.76, -8.87], leg. Von Heyden (1 J 
<specimenCount id="9D26FD3FFFD14B32B1C0A248171EFDFF" box="[470,516,583,608]" pageId="43" pageNumber="268" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B21CA2481727FDFF" box="[522,573,589,608]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Lusitania, 
<collectingRegion id="49E4F854FFD14B32B2B3A24817EEFDFF" box="[677,756,589,608]" country="Portugal" name="Lisboa" pageId="43" pageNumber="268">Lissabon</collectingRegion>
, [38.76, -8.87], 
<date id="FF9E1076FFD14B32B105A26D146BFDE4" box="[275,369,616,635]" pageId="43" pageNumber="268" value="1910-04" valueMax="1910-05-31" valueMin="1910-04-00">IV.-V.1910</date>
, leg.A. Schartzmayr (1 J 
<specimenCount id="9D26FD3FFFD14B32B25FA26D177CFDE4" box="[585,614,610,635]" pageId="43" pageNumber="268" type="female">1♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B27DA26D1784FDE4" box="[619,670,616,635]" pageId="43" pageNumber="268">SDEI</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B2BCA26D17E2FDE4" bold="true" box="[682,760,616,635]" pageId="43" pageNumber="268">Aඓඈඋൾඌ:</emphasis>
Faial.Baia da Areia das Fontes 
<quantity id="4CD89B53FFD14B32B188A28614DEFD09" box="[414,452,643,662]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="268" unit="km" value="1.0">1km</quantity>
N of Praia do Norte, [38.61, -28.74], under stones, grassy ground, near rocky shore, 
<date id="FF9E1076FFD14B32B20AA2981768FD2F" box="[540,626,669,688]" pageId="43" pageNumber="268" value="1957-04-02">2.IV.1957</date>
, leg.Brinck and Dahl (
<specimenCount id="9D26FD3FFFD14B32B0D8A2BD141AFD54" box="[206,256,690,715]" pageId="43" pageNumber="268" type="female">5 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B110A2BD145CFD54" box="[262,326,696,715]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Faial, Caldeira, [38.58, -28.71], 
<date id="FF9E1076FFD14B32B278A2BD17D5FD54" box="[622,719,696,715]" pageId="43" pageNumber="268" value="1957-04-10">10.IV.1957</date>
. leg. Brinck and Dahl (
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<collectionCode id="ED31AE73FFD14B32B14AA2D61486FD79" box="[348,412,723,742]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Faial, Horta, 
<collectingRegion id="49E4F854FFD14B32B202A2D61758FD79" box="[532,578,723,742]" country="Portugal" name="Porto" pageId="43" pageNumber="268">Porto</collectingRegion>
Pim., [38.52, -28.62], sandy ground, 
<date id="FF9E1076FFD14B32B104A2E8147FFC9F" box="[274,357,749,768]" pageId="43" pageNumber="268" value="1957-04-07">7.IV.1957</date>
leg.Brinck and Dahl (
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<collectionCode id="ED31AE73FFD14B32B255A2E81799FC9F" box="[579,643,749,768]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Faial. Nasce Agua 
<quantity id="4CD89B53FFD14B32B0DCA30D15E9FC84" box="[202,243,776,795]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="268" unit="km" value="3.0">3 km</quantity>
S of Cedros, [38.60, -28.68], ravine, under stones, 
<date id="FF9E1076FFD14B32B289A30D17EEFC84" box="[671,756,776,795]" pageId="43" pageNumber="268" value="1957-04-05">5.IV.1957</date>
, leg. Brinck and Dahl (1 J 
<collectionCode id="ED31AE73FFD14B32B16BA32614A7FCA9" box="[381,445,803,822]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Faial, Praia do Almoxarife, [38.55, -28.61], sandy grassy ground, under stones, 
<date id="FF9E1076FFD14B32B21AA3381775FCCF" box="[524,623,829,848]" pageId="43" pageNumber="268" value="1957-03-31">31.III.1957</date>
. leg. Brinck and Dahl (
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<collectionCode id="ED31AE73FFD14B32B0F8A35D1434FCF4" box="[238,302,856,875]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); São Miguel,Atalhada, [37.74, -25.58], natural forest, 
<date id="FF9E1076FFD14B32B081A37615EAFC19" box="[151,240,883,902]" pageId="43" pageNumber="268" value="1999-09-04">4.IX.1999</date>
, leg. H. Enghoff (
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<collectionCode id="ED31AE73FFD14B32B1C6A376170CFC19" box="[464,534,883,902]" pageId="43" pageNumber="268">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B230A376179BFC19" bold="true" box="[550,641,883,902]" pageId="43" pageNumber="268">Mൺൽൾංඋൺ:</emphasis>
Campanario, Fontes, [32.69, -17.01], 
<quantity id="4CD89B53FFD14B32B17DA38814C1FC3F" box="[363,475,909,928]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="43" pageNumber="268" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200-1400m</quantity>
, 
<date id="FF9E1076FFD14B32B1F3A3881753FC3F" box="[485,585,909,928]" pageId="43" pageNumber="268" value="2014-09-16">16.IX.2014</date>
, leg. Kleeberg (1 J cSch); Funchal, [32.67, -16.91], 
<quantity id="4CD89B53FFD14B32B1B1A3AD14EAFC24" box="[423,496,936,955]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="43" pageNumber="268" unit="m" value="250.0" valueMax="500.0" valueMin="0.0">0-500 m</quantity>
, 
<date id="FF9E1076FFD14B32B1E1A3AD172DFC24" box="[503,567,936,955]" pageId="43" pageNumber="268" value="1966-02">II.1966</date>
, leg. Palm (2 JJ 
<specimenCount id="9D26FD3FFFD14B32B2DCA3AD17ECFC24" box="[714,758,930,955]" pageId="43" pageNumber="268" type="female">3♀♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B081A3C615CDFC49" box="[151,215,963,982]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Pico do Castelo, [33.08, -16.33], 17.19. 
<date id="FF9E1076FFD14B32B23DA3C61770FC49" box="[555,618,963,982]" pageId="43" pageNumber="268" value="1977-05">V.1977</date>
, leg.Palm (2 JJ 
<collectionCode id="ED31AE73FFD14B32B081A3D815CDFC6F" box="[151,215,989,1008]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Terreiro da Luta, [32.68, -16.91], 
<quantity id="4CD89B53FFD14B32B21AA3D8175CFC6F" box="[524,582,989,1008]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="43" pageNumber="268" unit="m" value="850.0">850 m</quantity>
, stream, 
<date id="FF9E1076FFD14B32B282A3D817EEFC6F" box="[660,756,989,1008]" pageId="43" pageNumber="268" value="1957-04-20">20.IV.1957</date>
, leg. Brinck and Dahl (1 J 
<collectionCode id="ED31AE73FFD14B32B16CA3FD14A0FB94" box="[378,442,1016,1035]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B1DEA3FD1714FB94" bold="true" box="[456,526,1016,1035]" pageId="43" pageNumber="268">
<collectingCountry id="F3377626FFD14B32B1DEA3FD1710FB94" box="[456,522,1016,1035]" name="Spain" pageId="43" pageNumber="268">SPAIN</collectingCountry>
:
</emphasis>
Alicante, Orihuela, [38.08, -0.94], 
<date id="FF9E1076FFD14B32B0C0A416142CFBB9" box="[214,310,1043,1062]" pageId="43" pageNumber="268" value="1997-04-20">20.IV.1997</date>
, leg. Lauffer (2 
<collectionCode id="ED31AE73FFD14B32B1D7A416171FFBB9" box="[449,517,1043,1062]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Altea, Rio Algar, Alicante, [38.61, -0.05], 
<date id="FF9E1076FFD14B32B10BA4281462FBDF" box="[285,376,1069,1088]" pageId="43" pageNumber="268" value="1999-02-27">27.II.1999</date>
, leg. Wrase (1 J cSch); Aranjuez, 
<collectingRegion id="49E4F854FFD14B32B2A5A42817EEFBDF" box="[691,756,1069,1088]" country="Spain" name="Madrid" pageId="43" pageNumber="268">Madrid</collectingRegion>
, [40.02, -3.59], 
<quantity id="4CD89B53FFD14B32B10CA44D144BFBC4" box="[282,337,1096,1115]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="43" pageNumber="268" unit="m" value="520.0">520 m</quantity>
<date id="FF9E1076FFD14B32B141A44D14A7FBC4" box="[343,445,1096,1115]" pageId="43" pageNumber="268" value="1988-05-03" valueMax="1988-05-04" valueMin="1988-05-03">3.-4.V.1988</date>
, leg. Feller (
<specimenCount id="9D26FD3FFFD14B32B23AA44D1744FBC4" box="[556,606,1090,1115]" pageId="43" pageNumber="268" type="female">3 ♀♀</specimenCount>
cSch); Arroyo de Jurnadiel, S Brozas, Cáceres, [39.56, -6.81], 
<quantity id="4CD89B53FFD14B32B232A4661745FBE9" box="[548,607,1123,1142]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="43" pageNumber="268" unit="m" value="325.0">325 m</quantity>
, 
<date id="FF9E1076FFD14B32B27CA46617D7FBE9" box="[618,717,1123,1142]" pageId="43" pageNumber="268" value="1991-06-20">20.VI.1991</date>
, leg. Wrase (1 J cSch); Ávila, Avila, [40.65, -4.68], leg. Sanz (3 
<collectionCode id="ED31AE73FFD14B32B2BDA47817F5FB0F" box="[683,751,1149,1168]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Badajoz: Aljucen, [39.04, -6.33], leg. Pacheco (1 
<collectionCode id="ED31AE73FFD14B32B228A49D1798FB34" box="[574,642,1176,1195]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Boadilla del Monte, NW 
<collectingRegion id="49E4F854FFD14B32B0E9A4B6145AFB59" box="[255,320,1203,1222]" country="Spain" name="Madrid" pageId="43" pageNumber="268">Madrid</collectingRegion>
, [40.41, -3.88], 
<date id="FF9E1076FFD14B32B1D4A4B6173BFB59" box="[450,545,1203,1222]" pageId="43" pageNumber="268" value="1996-07-06">6.VII.1996</date>
, leg. D.Wrase (1J cSch); Cabo de Gata, SE Almeria, 
<geoCoordinate id="EE145071FFD14B32B19AA4C814CEFB7F" box="[396,468,1229,1248]" degrees="36.7908" direction="north" orientation="latitude" pageId="43" pageNumber="268" precision="5" value="36.7908">36.7908</geoCoordinate>
, 
<geoCoordinate id="EE145071FFD14B32B1C8A4C81738FB7F" box="[478,546,1229,1248]" degrees="2.2513" direction="west" orientation="longitude" pageId="43" pageNumber="268" precision="5" value="-2.2513">-2.2513</geoCoordinate>
, coastal area, 
<date id="FF9E1076FFD14B32B289A4C817EFFB7F" box="[671,757,1229,1248]" pageId="43" pageNumber="268" value="2012-04-05">5.IV.2012</date>
, leg. T. Wolsch (
<specimenCount id="9D26FD3FFFD14B32B130A4ED145DFB64" box="[294,327,1250,1275]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
cSch); Caboalles [de Abajo], [42.95, -6.37], leg. Paganetti (
<specimenCount id="9D26FD3FFFD14B32B0E3A506140EFA8A" box="[245,276,1276,1302]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B10FA5061456FA89" box="[281,332,1283,1302]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Cádiz, [36.53, -6.22], 
<date id="FF9E1076FFD14B32B20FA5061776FA89" box="[537,620,1283,1302]" pageId="43" pageNumber="268" value="1894-01-16">16.I.1894</date>
, leg. Navarro (2 
<collectionCode id="ED31AE73FFD14B32B081A51815C1FAAF" box="[151,219,1309,1328]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Cádiz, Algeciras, [36.16, -5.43], leg. Arias (1 
<collectionCode id="ED31AE73FFD14B32B278A51817A8FAAF" box="[622,690,1309,1328]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Cádiz, Vega del Barbate, Vejer de la Frontera, [36.25, -5.95], leg. De Quirós (1 
<collectionCode id="ED31AE73FFD14B32B081A55615C1FAF9" box="[151,219,1363,1382]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Cornudilla, [42.68, -3.42], 
<date id="FF9E1076FFD14B32B1C6A5561729FAF9" box="[464,563,1363,1382]" pageId="43" pageNumber="268" value="1962-09-13">13.IX.1962</date>
, leg. J.T. Skovgaard (1 J 
<specimenCount id="9D26FD3FFFD14B32B0BBA56815D5FA1F" box="[173,207,1383,1408]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B0C5A5681402FA1F" box="[211,280,1389,1408]" pageId="43" pageNumber="268">NHMD</collectionCode>
); Cuenca, Uclés, [39.98, -2.86] (2 
<collectionCode id="ED31AE73FFD14B32B255A568179DFA1F" box="[579,647,1389,1408]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Guipúzcoa, Arechavaleta, [43.03, -2.51], leg.Antoniol (1 
<collectionCode id="ED31AE73FFD14B32B20EA58D1746FA04" box="[536,604,1416,1435]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Guipuzcoa, Irun, Jaitzubia, [43.34, -1.82], 
<date id="FF9E1076FFD14B32B170A5A614A7FA29" box="[358,445,1443,1462]" pageId="43" pageNumber="268" value="2006-05-10">10.V.2006</date>
,leg.Anichtchenko (1J 
<specimenCount id="9D26FD3FFFD14B32B29EA5A617BFFA2A" box="[648,677,1436,1462]" pageId="43" pageNumber="268" type="female">1♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B2BCA5A617F5FA29" box="[682,751,1443,1462]" pageId="43" pageNumber="268">NHMD</collectionCode>
); Huelva, Cala, [37.97, -6.31], leg. C. Bolivar (1 
<collectionCode id="ED31AE73FFD14B32B232A5B81772FA4F" box="[548,616,1469,1488]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Jáen Mengibar, Rio Guadalquivir, [37.98, -3.79], 
<quantity id="4CD89B53FFD14B32B1A9A5DD14E8FA74" box="[447,498,1496,1515]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="43" pageNumber="268" unit="m" value="330.0">330m</quantity>
, 
<date id="FF9E1076FFD14B32B1EBA5DD177AFA74" box="[509,608,1496,1515]" pageId="43" pageNumber="268" value="1991-06-16">16.VI.1991</date>
, leg. Wrase (1 J cSch); La Coruña, Coruña, [43.36, -8.4], leg. Ricol (2 
<collectionCode id="ED31AE73FFD14B32B25AA5F61795F999" box="[588,655,1523,1542]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
);La Coruña, Villa Rutis, [43.28, -8.38], 
<date id="FF9E1076FFD14B32B16FA60814D3F9BF" box="[377,457,1549,1568]" pageId="43" pageNumber="268" value="1908-07">VII.1908</date>
, leg. Bolivar (2 
<collectionCode id="ED31AE73FFD14B32B258A6081788F9BF" box="[590,658,1549,1568]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Laguna de Ruiz Sánchez, N Osuna, Sevilla, [37.41, -5.09], 
<date id="FF9E1076FFD14B32B23FA62D1798F9A4" box="[553,642,1576,1595]" pageId="43" pageNumber="268" value="1991-05-26">26.V.1991</date>
, leg.Wrase (1 J cSch); Los Guajares, 
<collectingRegion id="49E4F854FFD14B32B14AA64614A5F9C9" box="[348,447,1603,1622]" country="Spain" name="Andalucia" pageId="43" pageNumber="268">Andalusien</collectingRegion>
, 
<geoCoordinate id="EE145071FFD14B32B1D0A6461717F9C9" box="[454,525,1603,1622]" degrees="36.8174" direction="north" orientation="latitude" pageId="43" pageNumber="268" precision="5" value="36.8174">36.8174</geoCoordinate>
, 
<geoCoordinate id="EE145071FFD14B32B203A6461743F9C9" box="[533,601,1603,1622]" degrees="3.5546" direction="west" orientation="longitude" pageId="43" pageNumber="268" precision="5" value="-3.5546">-3.5546</geoCoordinate>
, quarry, 
<date id="FF9E1076FFD14B32B289A64617EEF9C9" box="[671,756,1603,1622]" pageId="43" pageNumber="268" value="2012-04-06">6.IV.2012</date>
, leg. T. Wolsch (1 J cSch); 
<collectingRegion id="49E4F854FFD14B32B194A65814D9F9EF" box="[386,451,1629,1648]" country="Spain" name="Madrid" pageId="43" pageNumber="268">Madrid</collectingRegion>
, Escorial [40.57, -4.11] (1 
<collectionCode id="ED31AE73FFD14B32B2BDA65817F5F9EF" box="[683,751,1629,1648]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); 
<collectingRegion id="49E4F854FFD14B32B081A67D15C2F914" box="[151,216,1656,1675]" country="Spain" name="Madrid" pageId="43" pageNumber="268">Madrid</collectingRegion>
, Torrelaguna. [40.82, -3.53], leg. F. Beltran (1 
<collectionCode id="ED31AE73FFD14B32B274A67D17BCF914" box="[610,678,1656,1675]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); 
<collectingRegion id="49E4F854FFD14B32B2A2A67D17EFF914" box="[692,757,1656,1675]" country="Spain" name="Madrid" pageId="43" pageNumber="268">Madrid</collectingRegion>
, Viñuelas [40.79, -3.34] (1 
<collectionCode id="ED31AE73FFD14B32B195A69614DDF939" box="[387,455,1683,1702]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Maranchon, Guadalajara, [41.04, -2.20], 
<date id="FF9E1076FFD14B32B0C2A6A8142DF95F" box="[212,311,1709,1728]" pageId="43" pageNumber="268" value="2005-11-24">24.XI.2005</date>
, leg.Anichchenko (1 J 
<collectionCode id="ED31AE73FFD14B32B214A6A8175DF95F" box="[514,583,1709,1728]" pageId="43" pageNumber="268">NHMD</collectionCode>
); 
<collectingRegion id="49E4F854FFD14B32B243A6A8178FF95F" box="[597,661,1709,1728]" country="Spain" name="Murcia" pageId="43" pageNumber="268">Murcia</collectingRegion>
, Cartagena, [37.62, -0.99], leg. S. Gomez (1 
<collectionCode id="ED31AE73FFD14B32B1A6A6CD14EEF944" box="[432,500,1736,1755]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="43" pageNumber="268" type="Museum">MNCN</collectionCode>
); Palencia, [42.01, -4.52], leg. Paganetti (1 J 
<collectionCode id="ED31AE73FFD14B32B10FA6E61456F969" box="[281,332,1763,1782]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Palmones, Cádiz, [36.17, -5.43], 
<date id="FF9E1076FFD14B32B260A6E617D5F969" box="[630,719,1763,1782]" pageId="43" pageNumber="268" value="1991-05-29">29.V.1991</date>
, leg. Wrase (1 J cSch); Quero, Toledo, [39.51, -3.24], 
<date id="FF9E1076FFD14B32B22BA6F81789F88F" box="[573,659,1789,1808]" pageId="43" pageNumber="268" value="1999-03-08">8.III.1999</date>
, saline, leg. D.W. Wrase (1 J cSch); Serrania de Ronda, Málaga, [36.74, -5.17], 
<date id="FF9E1076FFD14B32B081A73615E0F8D9" box="[151,250,1843,1862]" pageId="43" pageNumber="268" value="1991-06-11">11.VI.1991</date>
, leg. Wrase (1 J cSch); Sitges, [41.23, 1.84], 
<date id="FF9E1076FFD14B32B28BA73617EDF8D9" box="[669,759,1843,1862]" pageId="43" pageNumber="268" value="1922-10-22">22.X.1922</date>
leg. W.Liebmann (
<specimenCount id="9D26FD3FFFD14B32B12BA7481444F8FF" box="[317,350,1863,1888]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B170A7481483F8FF" box="[358,409,1869,1888]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Toledo, Nahalhermosa, [39.63, -4.47], 
<date id="FF9E1076FFD14B32B081A76D141AF8E4" box="[151,256,1896,1915]" pageId="43" pageNumber="268" value="2007-07-27">27.VII.2007</date>
(1 J 
<collectionCode id="ED31AE73FFD14B32B124A76D146DF8E4" box="[306,375,1896,1915]" pageId="43" pageNumber="268">NHMD</collectionCode>
); Tres Cantos, [40.60, -3.70], 
<date id="FF9E1076FFD14B32B26EA76D17EEF8E4" box="[632,756,1896,1915]" pageId="43" pageNumber="268" value="2000-02-29" valueMax="2000-02-30" valueMin="2000-02-29">29.-30.II.2000</date>
, leg. Anichtchenko (
<specimenCount id="9D26FD3FFFD14B32B15EA7861476F80A" box="[328,364,1916,1942]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B164A78614A2F809" box="[370,440,1923,1942]" pageId="43" pageNumber="268">NHMD</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B1DEA786172EF809" bold="true" box="[456,564,1923,1942]" pageId="43" pageNumber="268">Mൺඅඅඈඋർൺ:</emphasis>
Palma, [39.56, 2.65], 
<date id="FF9E1076FFD14B32B081A79815C7F82F" box="[151,221,1949,1968]" pageId="43" pageNumber="268" value="1957-04">IV.1957</date>
, leg. T. Palm (
<specimenCount id="9D26FD3FFFD14B32B14EA798146CF82F" box="[344,374,1943,1968]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B16DA79814A1F82F" box="[379,443,1949,1968]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
); Rabassa, [39.54, 2.69], leg. Briet (3 JJ 
<specimenCount id="9D26FD3FFFD14B32B0ACA7BD15CDF854" box="[186,215,1970,1995]" pageId="43" pageNumber="268" type="female">1♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B0CBA7BD140DF854" box="[221,279,1976,1995]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="268">NMW</collectionCode>
); Valldecousa, [39.71, 2.62], 
<date id="FF9E1076FFD14B32B21DA7BD1797F854" box="[523,653,1976,1995]" pageId="43" pageNumber="268" value="1957-04-15" valueMax="1957-04-20" valueMin="1957-04-15">15.-20.IV.1957</date>
, leg.T. Palm (1 J 
<specimenCount id="9D26FD3FFFD14B32B0D4A7D615F5F87A" box="[194,239,1996,2022]" pageId="43" pageNumber="268" type="female">3 ♀♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B0E3A7D6142FF879" box="[245,309,2003,2022]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B154A7D61769F879" bold="true" box="[322,627,2003,2022]" pageId="43" pageNumber="268">Cൺඇൺඋඒ Iඌඅൺඇൽඌ: Gඋൺඇ Cൺඇൺඋංൺ:</emphasis>
Las Lagunetas, [28.01, -15.57], 
<date id="FF9E1076FFD14B32B137A7E8149EF79F" box="[289,388,2029,2048]" pageId="43" pageNumber="268" value="1988-11-22">22.XI.1988</date>
, leg. G. Gillerfors (
<specimenCount id="9D26FD3FFFD14B32B23AA7E81750F79F" box="[556,586,2023,2048]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B259A7E81795F79F" box="[591,655,2029,2048]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="43" pageNumber="268">MZLU</collectionCode>
). 
<emphasis id="B954EAA4FFD14B32B28BA7E817E2F79F" bold="true" box="[669,760,2029,2048]" pageId="43" pageNumber="268">Tൾඇൾඋංൿൾ:</emphasis>
Anaga Mountains, nr.Cruz del Carmen, [28.53, -16.27], 
<date id="FF9E1076FFD14B32B279A80D17CBF784" box="[623,721,2056,2075]" pageId="43" pageNumber="268" value="1992-06-14">14.VI.1992</date>
, leg. Hieke &amp; Wendt (
<specimenCount id="9D26FD3FFFD14B32B3DDA0C616F2FF49" box="[971,1000,189,214]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B3FBA0C61134FF49" box="[1005,1070,195,214]" pageId="43" pageNumber="268">ZMHB</collectionCode>
); Cumbre Dorsal Esperanza-Forest, [28.38, -16.44], 
<quantity id="4CD89B53FFD14B32B392A0DB16D8FF6E" box="[900,962,222,241]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="43" pageNumber="268" unit="m" value="1250.0">1250m</quantity>
, 
<date id="FF9E1076FFD14B32B3DFA0DB1130FF6E" box="[969,1066,222,241]" pageId="43" pageNumber="268" value="1992-04-13">13.IV.1992</date>
, leg. Zerche (
<specimenCount id="9D26FD3FFFD14B32B488A0DB11A1FF6E" box="[1182,1211,216,241]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B4D6A0DB11E9FF6E" box="[1216,1267,222,241]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Erjos env., [28.32, -16.81], 
<quantity id="4CD89B53FFD14B32B393A0FC16DDFE93" box="[901,967,249,268]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="43" pageNumber="268" unit="m" value="1100.0">1100 m</quantity>
, 
<date id="FF9E1076FFD14B32B3D9A0FC1134FE93" box="[975,1070,249,268]" pageId="43" pageNumber="268" value="2004-12-07">7.XII.2004</date>
, leg. Skoupý (
<specimenCount id="9D26FD3FFFD14B32B4BDA0FC11D0FE94" box="[1195,1226,242,268]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
cSch); Genovés, [28.36, -16.74], 
<quantity id="4CD89B53FFD14B32B36BA11616A8FEB9" box="[893,946,275,294]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="268" unit="m" value="400.0">400m</quantity>
, 
<date id="FF9E1076FFD14B32B3ADA1161109FEB9" box="[955,1043,275,294]" pageId="43" pageNumber="268" value="1989-06-09">9.VI.1989</date>
, leg. Felies (1 J cSch); La Esperanza, [28.44, -16.37], 
<quantity id="4CD89B53FFD14B32B395A12B16DBFEDE" box="[899,961,302,321]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="268" unit="m" value="1200.0">1200m</quantity>
, 
<date id="FF9E1076FFD14B32B3DEA12B1131FEDE" box="[968,1067,302,321]" pageId="43" pageNumber="268" value="1992-06-18">18.VI.1992</date>
, leg. Hieke &amp; Wendt (2 JJ 
<collectionCode id="ED31AE73FFD14B32B500A12B104DFEDE" box="[1302,1367,302,321]" pageId="43" pageNumber="268">ZMHB</collectionCode>
); Orotava, valley Aguamansa, [28.36, -16.49], 
<quantity id="4CD89B53FFD14B32B49EA14C11DEFEC3" box="[1160,1220,329,348]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="43" pageNumber="268" unit="m" value="1115.0">1115m</quantity>
, 
<date id="FF9E1076FFD14B32B4DDA14C102FFEC3" box="[1227,1333,329,348]" pageId="43" pageNumber="268" value="1995-12-25">25.XII.1995</date>
, leg. Stüben &amp; Bahr (
<specimenCount id="9D26FD3FFFD14B32B39CA16616B1FEE9" box="[906,939,349,374]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B3A6A16616F9FEE9" box="[944,995,355,374]" pageId="43" pageNumber="268">SDEI</collectionCode>
); Nr. Orotava, [28.35, -16.51], 
<quantity id="4CD89B53FFD14B32B4F9A1661037FEE9" box="[1263,1325,355,374]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="43" pageNumber="268" unit="m" value="1400.0">1400m</quantity>
, 
<date id="FF9E1076FFD14B32B520A1661083FEE9" box="[1334,1433,355,374]" pageId="43" pageNumber="268" value="1992-06-19">19.VI.1992</date>
, leg. Hieke &amp; Wendt (2 JJ 
<specimenCount id="9D26FD3FFFD14B32B433A17B115FFE0E" box="[1061,1093,376,401]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
<collectionCode id="ED31AE73FFD14B32B45CA17B1191FE0E" box="[1098,1163,382,401]" pageId="43" pageNumber="268">ZMHB</collectionCode>
); Puerto de la Cruz env., [28.36, -16.52], 
<quantity id="4CD89B53FFD14B32B390A19C16A3FE33" box="[902,953,409,428]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="268" unit="m" value="300.0">300m</quantity>
, 
<date id="FF9E1076FFD14B32B3D4A19C1154FE33" box="[962,1102,409,428]" pageId="43" pageNumber="268" value="2004-02-20" valueMax="2004-03-05" valueMin="2004-02-20">20.II.-5.III.2004</date>
, leg. E. Hajdaj (1 J cSch); Teno Alto, San Jeronimo, [28.34, -16.87], 
<quantity id="4CD89B53FFD14B32B45CA1B611B1FE59" box="[1098,1195,435,454]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="43" pageNumber="268" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</quantity>
, 
<date id="FF9E1076FFD14B32B4A2A1B61012FE59" box="[1204,1288,435,454]" pageId="43" pageNumber="268" value="1999-01-20">20.I.1999</date>
, leg. G. Zappi (2 JJ 
<specimenCount id="9D26FD3FFFD14B32B376A1CB1665FE7E" box="[864,895,456,481]" pageId="43" pageNumber="268" type="female">1 ♀</specimenCount>
cBor). 
<emphasis id="B954EAA4FFD14B32B3A9A1CB11F1FE7E" bold="true" box="[959,1259,462,481]" pageId="43" pageNumber="268">
<collectingCountry id="F3377626FFD14B32B3A9A1CB119DFE7E" box="[959,1159,462,481]" name="United Kingdom" pageId="43" pageNumber="268">UNITED KINGDOM</collectingCountry>
: EඇGඅൺඇൽ:
</emphasis>
Chichester Harbour, [50.79, -0.96], 
<date id="FF9E1076FFD14B32B3AAA1EC1110FE63" box="[956,1034,489,508]" pageId="43" pageNumber="268" value="1947-05-06">6.V.1947</date>
, leg. H. Last (2 JJ 
<collectionCode id="ED31AE73FFD14B32B4AEA1EC11E8FE63" box="[1208,1266,489,508]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="268">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD14B32B32AA211118AF894" pageId="43" pageNumber="268" type="description">
<paragraph id="8B9F36B6FFD14B32B32AA211108DFC14" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">
<emphasis id="B954EAA4FFD14B32B32AA21116F2FDB4" bold="true" box="[828,1000,532,555]" pageId="43" pageNumber="268">Redescription.</emphasis>
Measurements JJ (n = 6): HW = 1.62– 1.76 (1.69); HL = 1.47–1.49 (1.48); HL/HW 0.86–0.92 (0.88); PW = 2.22–2.29 (2.25); PL = 2.07–2.22 (2.16); PL/PW 0.93–0.97 (0.96); EW = 2.27–2.49 (2.37); EL = 2.20–2.33 (2.26); EL/EW 0.92–1.00 (0.96); EL/PL 1.01– 1.09 (1.05); PW/HW 1.49–1.56 (1.52); forebody length 5.76–6.02 (5.90). 
<specimenCount id="9D26FD3FFFD14B32B417A2C8113DFD73" box="[1025,1063,717,748]" pageId="43" pageNumber="268" type="female">♀♀</specimenCount>
(n = 4): HW = 1.58–1.73 (1.66); HL = 1.38–1.53 (1.46); HL/HW 0.87–0.89 (0.88); PW = 1.96–2.27 (2.16); PL = 1.91–2.11 (2.05); PL/PW 0.93–0.98 (0.95); EW = 2.22–2.51 (2.42); EL = 2.11–2.42 (2.29); EL/EW 0.91–0.96 (0.95); EL/PL 1.08–1.15 (1.12); PW/ HW 1.42–1.52 (1.48); forebody length 5.40–6.07 (5.80).
</paragraph>
<paragraph id="8B9F36B6FFD14B32B34CA39116D8FC54" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">
Medium to large sized species; body black to brown (
<figureCitation id="131B2A33FFD14B32B355A3B116ACFC54" box="[835,950,948,971]" captionStart="Fig" captionStartId="19.[151,182,2028,2047]" captionTargetBox="[203,1360,198,1958]" captionTargetId="figure-13@19.[183,1405,186,1971]" captionTargetPageId="19" captionText="Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915." figureDoi="http://doi.org/10.5281/zenodo.7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="43" pageNumber="268">Figs 9A,B</figureCitation>
).
</paragraph>
<paragraph id="8B9F36B6FFD14B32B34CA3D11151FBD4" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">Head black, distinctly transverse, eyes medium size (EyL/TL: 1.88–2.06 (1.94)), microsculpture of transverse waves, no interocular punctures between anterior frontal punctures; antennae pale.</paragraph>
<paragraph id="8B9F36B6FFD14B32B34CA45116E2FA54" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">Thorax: pronotum dark brown to brown, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row, sublateral row of pronotum with a cluster of one to three punctures situated slightly anterior of the anterior most puncture of the dorsal row; scutellum with fine setiferous punctures and with wavy to meshed microsculpture; elytra usually light brown to brown, occasionally black with paler suture and posterior margin, rarely completely black, relatively long, uniformly pubescent, quadrate or slightly wider than long, slightly longer than pronotum; legs pale, except inner face of tibia darkened.</paragraph>
<paragraph id="8B9F36B6FFD14B32B34CA5D1106BF994" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">Abdomen black to dark brown and slightly iridescent, tergites uniformly punctured, with slight iridescence.</paragraph>
<paragraph id="8B9F36B6FFD14B32B34CA616118AF894" blockId="43.[828,1437,527,2059]" pageId="43" pageNumber="268">
<emphasis id="B954EAA4FFD14B32B34CA6161682F9B4" bold="true" box="[858,920,1555,1579]" italics="true" pageId="43" pageNumber="268">Male.</emphasis>
Aedeagus (
<figureCitation id="131B2A33FFD14B32B404A611116BF9B4" box="[1042,1137,1556,1579]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="43" pageNumber="268">Fig. 14C</figureCitation>
): paramere elongate, narrow, lanceolate with narrowly slightly asymmetrical apical portion exceeding apex of median lobe, with numerous sensory peg setae forming two lateral longitudinal groups connected subapically; median lobe with sides of apex folded into distinct tooth situated at proximal part of fold; internal sac with a pair of very long medial sclerites, and with a pair of globular sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD14B35B32AA7111723FF6A" lastPageId="44" lastPageNumber="269" pageId="43" pageNumber="268" type="diagnosis">
<paragraph id="8B9F36B6FFD14B35B32AA7111723FF6A" blockId="43.[828,1437,527,2059]" lastBlockId="44.[151,760,190,2069]" lastPageId="44" lastPageNumber="269" pageId="43" pageNumber="268">
<emphasis id="B954EAA4FFD14B32B32AA7111129F8B4" bold="true" box="[828,1075,1812,1835]" pageId="43" pageNumber="268">Differential diagnosis.</emphasis>
<taxonomicName id="4C204D35FFD14B32B42EA7111013F8B4" authorityName="Bernhauer" authorityYear="1898" box="[1080,1289,1812,1835]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD14B32B42EA7111013F8B4" box="[1080,1289,1812,1835]" italics="true" pageId="43" pageNumber="268">Quedius hispanicus</emphasis>
</taxonomicName>
can be distinguished from all species of the 
<taxonomicName id="4C204D35FFD14B32B484A73111F1F8D4" authorityName="Stephens" authorityYear="1829" box="[1170,1259,1844,1867]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFD14B32B484A73111F1F8D4" box="[1170,1259,1844,1867]" italics="true" pageId="43" pageNumber="268">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFD14B32B4E5A7311037F8D4" box="[1267,1325,1844,1867]" pageId="43" pageNumber="268" sensu="stricto">s. str.</taxonomicNameLabel>
except 
<emphasis id="B954EAA4FFD14B32B594A731168CF8F4" italics="true" pageId="43" pageNumber="268">
<taxonomicName id="4C204D35FFD14B32B594A7311688F8F4" authorityName="Lucas" authorityYear="1846" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="pallipes">Q. pallipes</taxonomicName>
,
</emphasis>
by a cluster of 1–3 additional setiferous punctures in the posterior part of the sublateral row of the pronotum. They are very rarely missing on both sides of the pronotum in 
<taxonomicName id="4C204D35FFD14B32B341A7B116F3F854" authorityName="Bernhauer" authorityYear="1898" box="[855,1001,1972,1995]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD14B32B341A7B116F3F854" box="[855,1001,1972,1995]" italics="true" pageId="43" pageNumber="268">Q. hispanicus</emphasis>
</taxonomicName>
. It can be distinguished from 
<taxonomicName id="4C204D35FFD14B32B531A7B11086F854" authorityName="Lucas" authorityYear="1846" box="[1319,1436,1972,1995]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="268" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD14B32B531A7B11086F854" box="[1319,1436,1972,1995]" italics="true" pageId="43" pageNumber="268">Q. pallipes</emphasis>
</taxonomicName>
only by male genitalia characters: by a slightly narrower and, in medial section, more attenuate paramere; and by a median lobe with sides of apex each folded into distinct tooth, tooth situated at base of the fold.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD64B35B081A0FB11D5FC6A" pageId="44" pageNumber="269" type="synonymic_list">
<paragraph id="8B9F36B6FFD64B35B081A0FB11D8FE8A" blockId="44.[151,760,190,2069]" lastBlockId="44.[828,1437,190,1653]" pageId="44" pageNumber="269">
<emphasis id="B954EAA4FFD64B35B081A0FB1743FEAA" bold="true" pageId="44" pageNumber="269">
Synonymic notes on 
<taxonomicName id="4C204D35FFD64B35B196A0F814ECFE8A" authorityName="Lucas" authorityYear="1846" box="[384,502,253,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B196A0F814ECFE8A" bold="true" box="[384,502,253,277]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C204D35FFD64B35B215A0F81781FE8A" authorityName="Bernhauer" authorityYear="1898" box="[515,667,253,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B215A0F81781FE8A" bold="true" box="[515,667,253,277]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C204D35FFD64B35B2BEA0F81419FEAA" authorityName="Fairmaire" authorityYear="1861" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B2BEA0F81419FEAA" bold="true" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C204D35FFD64B35B104A1181466FEAA" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[274,380,285,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD64B35B104A1181466FEAA" bold="true" box="[274,380,285,309]" italics="true" pageId="44" pageNumber="269">Q. cobosi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C204D35FFD64B35B1A9A118174BFEAA" authorityName=", Coiffait" authorityYear="1964" box="[447,593,285,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="levasseuri">
<emphasis id="B954EAA4FFD64B35B1A9A118174BFEAA" bold="true" box="[447,593,285,309]" italics="true" pageId="44" pageNumber="269">Q. levasseuri</emphasis>
</taxonomicName>
.
</emphasis>
Lඎർൺඌ (1849) described 
<taxonomicName id="4C204D35FFD64B35B11BA13B14D4FECA" authorityName="Lucas" authorityYear="1846" box="[269,462,318,341]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B11BA13B14D4FECA" box="[269,462,318,341]" italics="true" pageId="44" pageNumber="269">Quedius pallipes</emphasis>
</taxonomicName>
from the coastal town of Philippeville [now 
<collectingRegion id="49E4F854FFD64B35B178A15B14A5FEEA" box="[366,447,350,373]" country="Algeria" name="Skikda" pageId="44" pageNumber="269">Skikda</collectingRegion>
], 
<collectingCountry id="F3377626FFD64B35B1C4A15B173DFEEA" box="[466,551,350,373]" name="Algeria" pageId="44" pageNumber="269">Algeria</collectingCountry>
. Fൺංඋආൺංඋൾ (1862) described 
<taxonomicName id="4C204D35FFD64B35B112A17B14EBFE0A" authorityName="Fairmaire" authorityYear="1861" box="[260,497,382,405]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B112A17B14EBFE0A" box="[260,497,382,405]" italics="true" pageId="44" pageNumber="269">Quedius simplicifrons</emphasis>
</taxonomicName>
as a very similar species from the French island of 
<collectingCountry id="F3377626FFD64B35B1DDA19B1715FE2A" box="[459,527,414,437]" name="France" pageId="44" pageNumber="269">Corse</collectingCountry>
. BൾඋඇHൺඎൾඋ (1898) described 
<emphasis id="B954EAA4FFD64B35B111A1BB14FFFE4A" box="[263,485,446,469]" italics="true" pageId="44" pageNumber="269">
<taxonomicName id="4C204D35FFD64B35B111A1BB14FBFE4A" authorityName="Bernhauer" authorityYear="1898" box="[263,481,446,469]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">Quedius hispanicus</taxonomicName>
,
</emphasis>
a species very similar to these, from the Spanish coastal towns of Cartagena and Cadiz. These three species have been mixed up, probably due to their very similar habitus, rendering their individual distributions unclear. Cඈංൿൿൺංඍ (1961) stated that 
<taxonomicName id="4C204D35FFD64B35B0D0A25B1427FDEA" authorityName="Lucas" authorityYear="1846" box="[198,317,606,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B0D0A25B1427FDEA" box="[198,317,606,629]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
was restricted to North Africa and hypothesized that 
<taxonomicName id="4C204D35FFD64B35B13FA27B14C2FD0A" authorityName="Fairmaire" authorityYear="1861" box="[297,472,638,661]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B13FA27B14C2FD0A" box="[297,472,638,661]" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
– with 
<taxonomicName id="4C204D35FFD64B35B239A27B17DFFD0A" authorityName="Bernhauer" authorityYear="1898" box="[559,709,638,661]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B239A27B17DFFD0A" box="[559,709,638,661]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
as a junior synonym – could be a European subspecies of 
<taxonomicName id="4C204D35FFD64B35B2CBA29B15F7FD4A" authorityName="Lucas" authorityYear="1846" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B2CBA29B15F7FD4A" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
without formally downgrading 
<taxonomicName id="4C204D35FFD64B35B25FA2BB17EDFD4A" authorityName="Fairmaire" authorityYear="1861" box="[585,759,702,725]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B25FA2BB17EDFD4A" box="[585,759,702,725]" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
to subspecific rank. Subsequently, some authors (Hൺඇඌൾඇ 1962, LඈHඌൾ 1964, Hඈඋංඈඇ 1965, Sආൾඍൺඇൺ 1970, Bඈඋ-ൽඈඇං 1976, WൾඅർH 1993) have treated 
<taxonomicName id="4C204D35FFD64B35B252A31B17C3FCAA" authorityName="Bernhauer" authorityYear="1898" box="[580,729,798,821]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B252A31B17C3FCAA" box="[580,729,798,821]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
as a valid species for Western European specimens, while others continued to consider it a synonym of 
<taxonomicName id="4C204D35FFD64B35B29CA35B15F8FC0A" authorityName="Fairmaire" authorityYear="1861" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B29CA35B15F8FC0A" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
(Pඈඉൾ 1977, Cඈංൿൿൺංඍ 1978, Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං 1995, Sඈඅඈൽඈඏඇංĸඈඏ 2012). KඈඋGൾ (1962b) mentioned specimens from Southern 
<collectingCountry id="F3377626FFD64B35B1A5A3BB14FFFC4A" box="[435,485,958,981]" name="Italy" pageId="44" pageNumber="269">Italy</collectingCountry>
and North Africa as conspecific with 
<taxonomicName id="4C204D35FFD64B35B132A3DB1482FC6A" authorityName="Lucas" authorityYear="1846" box="[292,408,990,1013]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B132A3DB1482FC6A" box="[292,408,990,1013]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
. He also reported that these were collected at higher altitudes (up to just above 
<quantity id="4CD89B53FFD64B35B28CA3FB17EAFB8A" box="[666,752,1022,1045]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="269" unit="m" value="1000.0">1000 m</quantity>
) very far from the coast. Cඈංൿൿൺංඍ (1964) further described two new species based mainly on genitalia differences of the males, 
<taxonomicName id="4C204D35FFD64B35B13AA45B176EFBEA" authority="Coiffait, 1964" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[300,628,1118,1141]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD64B35B13AA45B14CFFBEA" box="[300,469,1118,1141]" italics="true" pageId="44" pageNumber="269">Quedius cobosi</emphasis>
<bibRefCitation id="EFB14B47FFD64B35B1CBA45B176EFBEA" author="COIFFAIT H." box="[477,628,1118,1141]" pageId="44" pageNumber="269" pagination="293 - 296" refId="ref81616" refString="COIFFAIT H. 1964: Deux nouveaux Quedius s. str. de la region mediterraneenne occidentale. Revue Francaise d'Entomologie 31: 293 - 296." type="journal article" year="1964">Coiffait, 1964</bibRefCitation>
</taxonomicName>
from Seron in Spain, and 
<taxonomicName id="4C204D35FFD64B35B12CA47B17A2FB0A" authority="Coiffait, 1964" authorityName=", Coiffait" authorityYear="1964" box="[314,696,1150,1173]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="levasseuri">
<emphasis id="B954EAA4FFD64B35B12CA47B170AFB0A" box="[314,528,1150,1173]" italics="true" pageId="44" pageNumber="269">Quedius levasseuri</emphasis>
<bibRefCitation id="EFB14B47FFD64B35B20DA47B17A2FB0A" author="COIFFAIT H." box="[539,696,1150,1173]" pageId="44" pageNumber="269" pagination="293 - 296" refId="ref81616" refString="COIFFAIT H. 1964: Deux nouveaux Quedius s. str. de la region mediterraneenne occidentale. Revue Francaise d'Entomologie 31: 293 - 296." type="journal article" year="1964">Coiffait, 1964</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F3377626FFD64B35B081A49B15C3FB2A" box="[151,217,1182,1205]" name="France" pageId="44" pageNumber="269">Corse</collectingCountry>
, both similar to 
<taxonomicName id="4C204D35FFD64B35B182A49B1714FB2A" authorityName="Lucas" authorityYear="1846" box="[404,526,1182,1205]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B182A49B1714FB2A" box="[404,526,1182,1205]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFD64B35B25EA49B15BBFB4A" authorityName="Fairmaire" authorityYear="1861" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B25EA49B15BBFB4A" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C204D35FFD64B35B0BCA4BB145EFB4A" authorityName="Bernhauer" authorityYear="1898" box="[170,324,1214,1237]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B0BCA4BB145EFB4A" box="[170,324,1214,1237]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C204D35FFD64B35B143A4BB1718FB4A" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[341,514,1214,1237]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD64B35B143A4BB1718FB4A" box="[341,514,1214,1237]" italics="true" pageId="44" pageNumber="269">Quedius cobosi</emphasis>
</taxonomicName>
was characterised by the presence of numerous small setae on the edge of the paramere, while 
<taxonomicName id="4C204D35FFD64B35B147A4FB14F9FA8A" authorityName=", Coiffait" authorityYear="1964" box="[337,483,1278,1301]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="levasseuri">
<emphasis id="B954EAA4FFD64B35B147A4FB14F9FA8A" box="[337,483,1278,1301]" italics="true" pageId="44" pageNumber="269">Q. levasseuri</emphasis>
</taxonomicName>
was described as having paler sutural and elytral posterior margins as well as a differently shaped aedeagus, compared to 
<taxonomicName id="4C204D35FFD64B35B240A53B17D6FACA" authorityName="Lucas" authorityYear="1846" box="[598,716,1342,1365]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B240A53B17D6FACA" box="[598,716,1342,1365]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFD64B35B081A55B1450FAEA" authorityName="Fairmaire" authorityYear="1861" box="[151,330,1374,1397]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B081A55B1450FAEA" box="[151,330,1374,1397]" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
. This history implies several species- -group taxa of uncertain status, distribution and habitat preference that are similar to 
<taxonomicName id="4C204D35FFD64B35B1F2A59B1744FA2A" authorityName="Lucas" authorityYear="1846" box="[484,606,1438,1461]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B1F2A59B1744FA2A" box="[484,606,1438,1461]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
and co-occur in the West Mediterranean. Studying a broad sample of specimens from that area, we found only two morphotypes, which differ only in the structure of the aedeagus. The first is found along the European Atlantic Coast, including Southern 
<collectingCountry id="F3377626FFD64B35B166A63B1480F9CA" box="[368,410,1598,1621]" name="United Kingdom" pageId="44" pageNumber="269">UK</collectingCountry>
, Canaries, Madeira, Azores and inland in southern 
<collectingCountry id="F3377626FFD64B35B17EA65B14A8F9EA" box="[360,434,1630,1653]" name="France" pageId="44" pageNumber="269">France</collectingCountry>
and on the Iberian Peninsula (
<figureCitation id="131B2A33FFD64B35B089A67B1413F90A" box="[159,265,1662,1685]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="44" pageNumber="269">Figs 14C</figureCitation>
, 
<figureCitation id="131B2A33FFD64B35B101A67B142FF90A" box="[279,309,1662,1685]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="44" pageNumber="269">19</figureCitation>
). These areas include the 
<typeStatus id="549B8814FFD64B35B24AA67B1791F90A" box="[604,651,1662,1685]" pageId="44" pageNumber="269">type</typeStatus>
localities of 
<taxonomicName id="4C204D35FFD64B35B0A3A69B1451F92A" authorityName="Bernhauer" authorityYear="1898" box="[181,331,1694,1717]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B0A3A69B1451F92A" box="[181,331,1694,1717]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFD64B35B194A69B14F1F92A" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[386,491,1694,1717]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD64B35B194A69B14F1F92A" box="[386,491,1694,1717]" italics="true" pageId="44" pageNumber="269">Q. cobosi</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A33FFD64B35B1EDA69B1757F92A" box="[507,589,1694,1717]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="44" pageNumber="269">Fig. 19</figureCitation>
). The second is found in mainland 
<collectingCountry id="F3377626FFD64B35B17AA6BB1485F94A" box="[364,415,1726,1749]" name="Italy" pageId="44" pageNumber="269">Italy</collectingCountry>
, on 
<collectingCountry id="F3377626FFD64B35B1C6A6BB1715F94A" box="[464,527,1726,1749]" name="France" pageId="44" pageNumber="269">Corse</collectingCountry>
and Sardinia, and in North Africa at both coastal and higher elevation sites (
<figureCitation id="131B2A33FFD64B35B089A6FB1413F88A" box="[159,265,1790,1813]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="44" pageNumber="269">Figs 14D</figureCitation>
, 
<figureCitation id="131B2A33FFD64B35B10EA6FB142CF88A" box="[280,310,1790,1813]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="44" pageNumber="269">19</figureCitation>
). These areas include the 
<typeStatus id="549B8814FFD64B35B24BA6FB1796F88A" box="[605,652,1790,1813]" pageId="44" pageNumber="269">type</typeStatus>
localities for 
<taxonomicName id="4C204D35FFD64B35B0AAA71B142AF8AA" authorityName="Lucas" authorityYear="1846" box="[188,304,1822,1845]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B0AAA71B142AF8AA" box="[188,304,1822,1845]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C204D35FFD64B35B12BA71B14F2F8AA" authorityName="Fairmaire" authorityYear="1861" box="[317,488,1822,1845]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">
<emphasis id="B954EAA4FFD64B35B12BA71B14F2F8AA" box="[317,488,1822,1845]" italics="true" pageId="44" pageNumber="269">Q. simplicifrons</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C204D35FFD64B35B236A71B17B7F8AA" authorityName=", Coiffait" authorityYear="1964" box="[544,685,1822,1845]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="levasseuri">
<emphasis id="B954EAA4FFD64B35B236A71B17B7F8AA" box="[544,685,1822,1845]" italics="true" pageId="44" pageNumber="269">Q. levasseuri</emphasis>
</taxonomicName>
. There is no transition between the aedeagi of both 
<typeStatus id="549B8814FFD64B35B293A73B17D8F8CA" box="[645,706,1854,1877]" pageId="44" pageNumber="269">types</typeStatus>
. The steep cliff shoreline between Nice, 
<collectingCountry id="F3377626FFD64B35B21AA75B174CF8EA" box="[524,598,1886,1909]" name="France" pageId="44" pageNumber="269">France</collectingCountry>
and La Spezia, 
<collectingCountry id="F3377626FFD64B35B081A77B15D3F80A" box="[151,201,1918,1941]" name="Italy" pageId="44" pageNumber="269">Italy</collectingCountry>
may be unsuitable for these species and may represent a barrier for their dispersal. As the morphological difference between both 
<typeStatus id="549B8814FFD64B35B189A7BB14C2F84A" box="[415,472,1982,2005]" pageId="44" pageNumber="269">types</typeStatus>
of genitalia is clear, and in agreement with the 
<typeStatus id="549B8814FFD64B35B171A7DB148FF86A" box="[359,405,2014,2037]" pageId="44" pageNumber="269">type</typeStatus>
localities, we here propose to use 
<taxonomicName id="4C204D35FFD64B35B081A7FB1437F78A" authorityName="Bernhauer" authorityYear="1898" box="[151,301,2046,2069]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B081A7FB1437F78A" box="[151,301,2046,2069]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
for the Iberian and Atlantic coast species (
<typeStatus id="549B8814FFD64B35B355A0BB1669FF4A" box="[835,883,190,213]" pageId="44" pageNumber="269">type</typeStatus>
1) and 
<taxonomicName id="4C204D35FFD64B35B3D5A0BB1123FF4A" authorityName="Lucas" authorityYear="1846" box="[963,1081,190,213]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B3D5A0BB1123FF4A" box="[963,1081,190,213]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
for the Italian and North African species (
<typeStatus id="549B8814FFD64B35B38FA0DB16D3FF6A" box="[921,969,222,245]" pageId="44" pageNumber="269">type</typeStatus>
2) (
<figureCitation id="131B2A33FFD64B35B3E1A0DB1151FF6A" box="[1015,1099,222,245]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="44" pageNumber="269">Figs 14</figureCitation>
, 
<figureCitation id="131B2A33FFD64B35B44EA0DB116CFF6A" box="[1112,1142,222,245]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="44" pageNumber="269">19</figureCitation>
). 
<taxonomicName id="4C204D35FFD64B35B49CA0DB1029FF6A" authorityName="Coiff. H. COIFFAIT" authorityYear="1964" box="[1162,1331,222,245]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="cobosi">
<emphasis id="B954EAA4FFD64B35B49CA0DB1029FF6A" box="[1162,1331,222,245]" italics="true" pageId="44" pageNumber="269">Quedius cobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFD64B35B52DA0DB1087FF6A" box="[1339,1437,222,245]" pageId="44" pageNumber="269">syn. nov.</taxonomicNameLabel>
is synonymized with 
<taxonomicName id="4C204D35FFD64B35B432A0FB11A0FE8A" authorityName="Bernhauer" authorityYear="1898" box="[1060,1210,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B432A0FB11A0FE8A" box="[1060,1210,254,277]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F36B6FFD64B35B34CA11B16DCFD2A" blockId="44.[828,1437,190,1653]" pageId="44" pageNumber="269">
However, barcodes from specimens across the entire distribution of both 
<typeStatus id="549B8814FFD64B35B41EA13B115AFECA" box="[1032,1088,318,341]" pageId="44" pageNumber="269">types</typeStatus>
1 and 2 show very little variation, with no clear clustering corresponding to both genitalia 
<typeStatus id="549B8814FFD64B35B32AA17B166EFE0A" box="[828,884,382,405]" pageId="44" pageNumber="269">types</typeStatus>
(
<figureCitation id="131B2A33FFD64B35B393A17B16D3FE0A" box="[901,969,382,405]" captionStart="Fig" captionStartId="11.[151,182,2008,2027]" captionTargetBox="[151,1437,191,1879]" captionTargetId="figure-13@11.[151,1437,191,1968]" captionTargetPageId="11" captionText="Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399708" httpUri="https://zenodo.org/record/7399708/files/figure.png" pageId="44" pageNumber="269">Fig. 3</figureCitation>
). A rather uniform genetic barcode among specimens morphologically corresponding to both 
<taxonomicName id="4C204D35FFD64B35B594A19B16B4FE4A" authorityName="Bernhauer" authorityYear="1898" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B594A19B16B4FE4A" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFD64B35B3F5A1BB1143FE4A" authorityName="Lucas" authorityYear="1846" box="[995,1113,446,469]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B3F5A1BB1143FE4A" box="[995,1113,446,469]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
(only 0.5% divergence, 
<tableCitation id="C6A2030DFFD64B35B577A1BB1650FE6A" captionStart="Table 2" captionStartId="13.[151,201,190,209]" captionTargetPageId="13" captionText="Table 2. Genetic distance between clusters of CO1 barcodes.n = number of specimens barcoded. Number of OTUs found by species delimiter plugin in Geneious. Intraspecific distance within cluster. Interspecific distance is distance to nearest other cluster." httpUri="http://table.plazi.org/id/DF5F663EFFF74B14B081A0BB1713FEBE" pageId="44" pageNumber="269" tableUuid="DF5F663EFFF74B14B081A0BB1713FEBE">Table 2</tableCitation>
) suggests that there may be some level of mitochondrial introgression between these species. External factors such as past or present infections by endosymbionts such as 
<taxonomicName id="4C204D35FFD64B35B32AA23B16B6FDCA" authorityName="Hertig" authorityYear="1936" box="[828,940,574,597]" class="Alphaproteobacteria" family="Anaplasmataceae" genus="Wolbachia" kingdom="Bacteria" order="Rickettsiales" pageId="44" pageNumber="269" phylum="Proteobacteria" rank="genus">
<emphasis id="B954EAA4FFD64B35B32AA23B16B6FDCA" box="[828,940,574,597]" italics="true" pageId="44" pageNumber="269">Wolbachia</emphasis>
</taxonomicName>
(e.g. KඈඇൽൺඇൽൺඋൺආൺංൺH et al. 2013) cannot be ruled out as an explanation either. This should be explored more thoroughly for example with the use of nuclear genes or genomics.
</paragraph>
<paragraph id="8B9F36B6FFD64B35B34CA2BB11D5FC6A" blockId="44.[828,1437,190,1653]" pageId="44" pageNumber="269">
The original descriptions and distribution records of 
<taxonomicName id="4C204D35FFD64B35B32AA2DB11B8FD6A" authority="Sharp, 1911" authorityName="Sharp" authorityYear="1911" box="[828,1186,734,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hammianus">
<emphasis id="B954EAA4FFD64B35B32AA2DB1102FD6A" box="[828,1048,734,757]" italics="true" pageId="44" pageNumber="269">Quedius hammianus</emphasis>
<bibRefCitation id="EFB14B47FFD64B35B436A2DB11B8FD6A" author="SHARP D." box="[1056,1186,734,757]" pageId="44" pageNumber="269" pagination="57" refId="ref91034" refString="SHARP D. 1911: A new British Quedius. Entomologist's Monthly Magazine 47: 57." type="journal article" year="1911">Sharp, 1911</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C204D35FFD64B35B4CFA2DB16B0FC8A" authority="Last, 1952" authorityName="Last" authorityYear="1952" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B954EAA4FFD64B35B4CFA2DB1086FD6A" box="[1241,1436,734,757]" italics="true" pageId="44" pageNumber="269">Quedius secundus</emphasis>
<bibRefCitation id="EFB14B47FFD64B35B32AA2FB16B0FC8A" author="LAST H." box="[828,938,766,789]" pageId="44" pageNumber="269" pagination="148 - 150" refId="ref86758" refString="LAST H. 1952: Taxonomic notes on Quedius molochinus Grav. and Q. pallipes Lucas (Col., Staphylinidae). Entomologist's Monthly Magazine 88: 148 - 150." type="journal article" year="1952">Last, 1952</bibRefCitation>
</taxonomicName>
, both described from the 
<collectingCountry id="F3377626FFD64B35B4BFA2FB11C8FC8A" box="[1193,1234,766,789]" name="United Kingdom" pageId="44" pageNumber="269">UK</collectingCountry>
, match the morphology and the ecological association with coastal sites of 
<taxonomicName id="4C204D35FFD64B35B594A31B16B1FCCA" authorityName="Bernhauer" authorityYear="1898" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B594A31B16B1FCCA" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C204D35FFD64B35B3A0A33B11E2FCCA" authority="Blumml, 1898" authorityName="Blumml" authorityYear="1898" box="[950,1272,830,853]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="rufulus">
<emphasis id="B954EAA4FFD64B35B3A0A33B1146FCCA" box="[950,1116,830,853]" italics="true" pageId="44" pageNumber="269">Quedius rufulus</emphasis>
<bibRefCitation id="EFB14B47FFD64B35B476A33B11E2FCCA" author="BLUMML E. K." box="[1120,1272,830,853]" pageId="44" pageNumber="269" pagination="216" refId="ref80473" refString="BLUMML E. K. 1898: Eine neue Kafer-Variation. Illustrierte Zeitschrift fur Entomologie 3: 216." type="journal article" year="1898">Blümml, 1898</bibRefCitation>
</taxonomicName>
, described from Granada in Spain as having darker base of the antennae, most likely represents a variant of 
<taxonomicName id="4C204D35FFD64B35B4B2A37B102FFC0A" authorityName="Bernhauer" authorityYear="1898" box="[1188,1333,894,917]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B4B2A37B102FFC0A" box="[1188,1333,894,917]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
. All three species were hitherto in synonymy with 
<emphasis id="B954EAA4FFD64B35B4FDA39B1087FC2A" box="[1259,1437,926,949]" italics="true" pageId="44" pageNumber="269">
<taxonomicName id="4C204D35FFD64B35B4FDA39B1083FC2A" authorityName="Fairmaire" authorityYear="1861" box="[1259,1433,926,949]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="simplicifrons">Q. simplicifrons</taxonomicName>
.
</emphasis>
Under the revised concept of the 
<taxonomicName id="4C204D35FFD64B35B4B8A3BB103CFC4A" authorityName="Lucas" authorityYear="1846" box="[1198,1318,958,981]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFD64B35B4B8A3BB103CFC4A" box="[1198,1318,958,981]" italics="true" pageId="44" pageNumber="269">Q. pallipes</emphasis>
</taxonomicName>
-group, we synonymize them with 
<taxonomicName id="4C204D35FFD64B35B423A3DB11D2FC6A" authorityName="Bernhauer" authorityYear="1898" box="[1077,1224,990,1013]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B423A3DB11D2FC6A" box="[1077,1224,990,1013]" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD64B35B32AA3FB1177FAEA" pageId="44" pageNumber="269" type="description">
<paragraph id="8B9F36B6FFD64B35B32AA3FB1177FAEA" blockId="44.[828,1437,190,1653]" pageId="44" pageNumber="269">
<emphasis id="B954EAA4FFD64B35B32AA3FB16A3FB8A" bold="true" box="[828,953,1022,1045]" pageId="44" pageNumber="269">Bionomics.</emphasis>
<taxonomicName id="4C204D35FFD64B35B3D4A3FB118CFB8A" authorityName="Bernhauer" authorityYear="1898" box="[962,1174,1022,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B3D4A3FB118CFB8A" box="[962,1174,1022,1045]" italics="true" pageId="44" pageNumber="269">Quedius hispanicus</emphasis>
</taxonomicName>
mainly occurs in saline habitats along the Atlantic coast, such as salt marshes and the banks of estuaries in flood debris. Further south in the Iberian Peninsula and southern 
<collectingCountry id="F3377626FFD64B35B499A45B11C2FBEA" box="[1167,1240,1118,1141]" name="France" pageId="44" pageNumber="269">France</collectingCountry>
the species seems to be present inland in non-saline habitats. Inland it occurs in the debris along the banks of creeks and rivers. For example, AKH has collected it in the debris near streams and rivers in the Extremadura region (
<figureCitation id="131B2A33FFD64B35B4DDA4DB1007FB6A" box="[1227,1309,1246,1269]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="44" pageNumber="269">Fig. 2D</figureCitation>
) and also in the flood debris on a salt marsh near the coast in southern 
<collectingCountry id="F3377626FFD64B35B32AA51B16B9FAAA" box="[828,931,1310,1333]" name="Denmark" pageId="44" pageNumber="269">Denmark</collectingCountry>
. Most records are from low elevations, except a few on Madeira and the Canary Islands, where 
<taxonomicName id="4C204D35FFD64B35B52EA53B166EFAEA" authorityName="Bernhauer" authorityYear="1898" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B52EA53B166EFAEA" italics="true" pageId="44" pageNumber="269">Q. hispanicus</emphasis>
</taxonomicName>
is found up to 
<quantity id="4CD89B53FFD64B35B403A55B1173FAEA" box="[1045,1129,1374,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="44" pageNumber="269" unit="m" value="1400.0">1400 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A653DFFD64B35B32AA57B1015F9EA" pageId="44" pageNumber="269" type="distribution">
<paragraph id="8B9F36B6FFD64B35B32AA57B1015F9EA" blockId="44.[828,1437,190,1653]" pageId="44" pageNumber="269">
<emphasis id="B954EAA4FFD64B35B32AA57B16D0FA0A" bold="true" box="[828,970,1406,1429]" pageId="44" pageNumber="269">Distribution.</emphasis>
<taxonomicName id="4C204D35FFD64B35B3D9A57B1180FA0A" authorityName="Bernhauer" authorityYear="1898" box="[975,1178,1406,1429]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="269" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFD64B35B3D9A57B1180FA0A" box="[975,1178,1406,1429]" italics="true" pageId="44" pageNumber="269">Quedius hispanicus</emphasis>
</taxonomicName>
occurs along theAtlantic coast from southern 
<collectingCountry id="F3377626FFD64B35B435A59B119DFA2A" box="[1059,1159,1438,1461]" name="Denmark" pageId="44" pageNumber="269">Denmark</collectingCountry>
to the Iberian Peninsula where it continues to the French Mediterranean coast, with one unverified report from the northwestern Italian coast (
<figureCitation id="131B2A33FFD64B35B395A5FB16C9F98A" box="[899,979,1534,1557]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="44" pageNumber="269">Fig. 19</figureCitation>
). In the Iberian Peninsula it occurs inland along most rivers and creeks. It also occurs on the Azores (Faial), Madeira (Madeira and Porto Santo), the Canaries (Tenerife and Gran Canaries) and Mallorca.
</paragraph>
</subSubSection>
</treatment>
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