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<document id="53A0AF1A10A1D95172B6BC834044E7C3" ID-CLB-Dataset="8247" ID-DOI="10.11646/zootaxa.4860.2.8" ID-GBIF-Dataset="b2701eb2-3a17-4e33-907b-37e810df5ce3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4413797" ID-ZooBank="9BA7720C-C00F-4953-B480-CD8960C0CA6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609703087702" checkinUser="plazi" docAuthor="Das, Abhijit, Deepak, V., Captain, Ashok, Wade, Edward O. Z. &amp; Gower, David J." docDate="2020" docId="181187FCFFC1FFC039F91CAAFF08F879" docLanguage="en" docName="zootaxa.4860.2.8.pdf" docOrigin="Zootaxa 4860 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Smithophis arunachalensis Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower 2020, sp. nov." docType="treatment" docUuid="0393FC04-EAA6-477C-AC31-BCDB78A2AFC9" docUuidSource="ZooBank" docVersion="9" lastPageNumber="278" masterDocId="E428FF84FFC0FFCB396E1F5CFFA8FFE1" masterDocTitle="Description of a new species of Smithophis Giri et al. 2019 (Serpentes: Colubridae Natricinae) from Arunachal Pradesh, India" masterLastPageNumber="283" masterPageNumber="267" pageNumber="268" updateTime="1698851940654" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FB8AA3F069C416C6478AE18FEA2A61CC">Description of a new species of Smithophis Giri et al. 2019 (Serpentes: Colubridae Natricinae) from Arunachal Pradesh, India</mods:title>
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<mods:namePart id="91DF534F9BA13A3C7812CDDCBA8D4798">Deepak, V.</mods:namePart>
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<mods:affiliation id="1480F761854115C58E8DB798C0EE29B7">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK. &amp; veerappandeepak @ gmail. com; https: // orcid. org / 0000 - 0002 - 8826 - 9367</mods:affiliation>
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<emphasis id="A2CCEAF8FFC1FFCA39F91CAAFE49FBF0" bold="true" box="[151,481,1014,1041]" italics="true" pageId="1" pageNumber="268">Smithophis arunachalensis</emphasis>
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(
<figureCitation id="08832A6FFFC1FFCA39F11B47FEA4FBD4" box="[159,268,1051,1077]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1259,1283]" captionStartId-1="4.[151,250,1963,1987]" captionStartId-2="5.[151,250,1168,1192]" captionStartId-3="7.[151,250,1961,1985]" captionStartId-4="8.[151,250,1785,1809]" captionStartId-5="9.[151,250,1938,1962]" captionStartId-6="10.[151,250,1918,1942]" captionTargetBox-0="[151,1436,309,1236]" captionTargetBox-1="[167,1420,1101,1938]" captionTargetBox-2="[176,1411,189,1138]" captionTargetBox-3="[184,1404,207,1930]" captionTargetBox-4="[328,1177,257,1690]" captionTargetBox-5="[151,1436,190,1902]" captionTargetBox-6="[151,1436,181,1894]" captionTargetId-0="figure-77@3.[151,1436,309,1236]" captionTargetId-1="figure-436@4.[167,1420,1101,1938]" captionTargetId-2="figure-19@5.[170,1417,181,1145]" captionTargetId-3="figure-19@7.[175,1412,199,1938]" captionTargetId-4="figure-19@8.[235,1352,181,1762]" captionTargetId-5="figure-19@9.[151,1436,181,1915]" captionTargetId-6="figure-19@10.[151,1436,181,1894]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="FIGURE 1. Smithophis arunachalensis sp. nov., holotype (BNHS 3650). A. full body dorsal, and B. ventral view, C. head dorsal, D. ventral, and E– F. lateral views of head. Scale bars 10mm." captionText-1="FIGURE 2. Smithophis arunachalensis sp. nov., holotype (BNHS 3650) in life." captionText-2="FIGURE 3. Map showing distribution of species of the genus Smithophis. Examined comparative materials of S. bicolor (including the type), are from the Khasia (or Khassia or Khasi) Hills, Shillong, Tura or Aizawl, or from unspecified localities in Northeast India. See Appendix 1 for locality details." captionText-3="FIGURE 4. Smithophis arunachalensis sp. nov., paratypes in dorsal views A. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and B. (ZSIK 23875). Scale bar 10mm." captionText-4="FIGURE 5. Smithophis arunachalensis sp. nov., reconstructed head of paratype (BMNH 1935.10.12.10), breaks in specimen marked with red dashed lines. Scale bar 10mm." captionText-5="FIGURE 6. Smithophis arunachalensis sp. nov., A. unnumbered referred specimen in Roing Forest Department collection. Total length 613 mm. B. unnumbered (2) referred specimen in Miao museum collection." captionText-6="FIGURE 7. Habitat of Smithophis arunachalensis sp. nov. at the type locality Koronu, Lower Dibang Valley District, Arunachal Pradesh, India." figureDoi-0="http://doi.org/10.5281/zenodo.4413799" figureDoi-1="http://doi.org/10.5281/zenodo.4413801" figureDoi-2="http://doi.org/10.5281/zenodo.4413805" figureDoi-3="http://doi.org/10.5281/zenodo.4413807" figureDoi-4="http://doi.org/10.5281/zenodo.4413809" figureDoi-5="http://doi.org/10.5281/zenodo.4413813" figureDoi-6="http://doi.org/10.5281/zenodo.4413815" httpUri-0="https://zenodo.org/record/4413799/files/figure.png" httpUri-1="https://zenodo.org/record/4413801/files/figure.png" httpUri-2="https://zenodo.org/record/4413805/files/figure.png" httpUri-3="https://zenodo.org/record/4413807/files/figure.png" httpUri-4="https://zenodo.org/record/4413809/files/figure.png" httpUri-5="https://zenodo.org/record/4413813/files/figure.png" httpUri-6="https://zenodo.org/record/4413815/files/figure.png" pageId="1" pageNumber="268">Figs. 1–7</figureCitation>
; 
<tableCitation id="DD3A0351FFC1FFCA38791B47FED9FBD4" box="[279,369,1051,1077]" captionStart="TABLE 1" captionStartId="2.[151,239,440,464]" captionTargetPageId="2" captionText="TABLE 1. Meristic and morphometric (in millimetres) data for examined specimens of Smithophis arunachalensis sp. nov. Head length = distance between snout tip and back of lower jaw; bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. The format ‘X-Y’ is used to denote the shortest distance between features X and Y." httpUri="http://table.plazi.org/id/C4C76662FFC2FFC939F91EE4FD74FDDD" pageId="1" pageNumber="268" tableUuid="C4C76662FFC2FFC939F91EE4FD74FDDD">Tables 1</tableCitation>
&amp; 
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)
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<emphasis id="A2CCEAF8FFC1FFCA39F91BDFFEE5FB7A" box="[151,333,1155,1179]" italics="true" pageId="1" pageNumber="268">Rhabdops bicolor</emphasis>
(Blyth, 1854)
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: 
<treatmentCitation id="111910FBFFC1FFCA38851BDFFD08FB7A" author="Smith, M. A." box="[491,672,1155,1179]" page="329" pageId="1" pageNumber="268" year="1943">
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<bibRefCitation id="F4294B1BFFC1FFCA3DE71BDFFA30FB7A" author="Captain, A &amp; Patel, A." box="[1161,1432,1155,1179]" pageId="1" pageNumber="268" pagination="1 - 14" refId="ref7329" refString="Captain, A &amp; Patel, A. (1997) Some snakes of the Miao woodlot (dist. Changlang). Arunachal Pradesh, India. Unpublished report, 1 - 14 with 12 photographs." type="book chapter" year="1997">Captain &amp; Patel (1997: 14</bibRefCitation>
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, in part); 
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</treatmentCitation>
, in part); 
<treatmentCitation id="111910FBFFC1FFCA3BFB1BFEFCE5FB5B" author="Bhatt, B. B." box="[661,845,1186,1210]" page="196" pageId="1" pageNumber="268" year="2004">
<bibRefCitation id="F4294B1BFFC1FFCA3BFB1BFEFCE5FB5B" author="Bhatt, B. B." box="[661,845,1186,1210]" pageId="1" pageNumber="268" refId="ref6894" refString="Bhatt, B. B. (2004) Present status of Ophidian fauna and habitat ecology of certain species of ophidian in Arunachal Pradesh, India. Unpublished PhD Thesis, Department of Zoology, Guwahati University, Guwahati, Assam, 326 pp., 34 pls." type="book" year="2004">Bhatt (2004: 196)</bibRefCitation>
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; 
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Borang 
<emphasis id="A2CCEAF8FFC1FFCA3AC71BFFFC75FB5B" box="[937,989,1186,1210]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
(2005: 23)
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; 
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<bibRefCitation id="F4294B1BFFC1FFCA3D331BFEFAD5FB5B" author="Sanyal, D. P. &amp; Gayen, N. C." box="[1117,1405,1186,1210]" pageId="1" pageNumber="268" pagination="247 - 284" refId="ref8206" refString="Sanyal, D. P. &amp; Gayen, N. C. (2006) State Fauna Series (Reptilia). In: Alfred, J. R. B. (Ed.), Fauna of Arunachal Pradesh. 13 (1). Zoological Survey of India, Kolkata, pp. 247 - 284." type="book chapter" year="2006">Sanyal &amp; Gayen (2006: 273</bibRefCitation>
</treatmentCitation>
, in part); 
<treatmentCitation id="111910FBFFC1FFCA386A1B9CFE61FB38" author="Sharma, R. C." box="[260,457,1216,1241]" page="238" pageId="1" pageNumber="268" year="2007">
<bibRefCitation id="F4294B1BFFC1FFCA386A1B9CFE61FB38" author="Sharma, R. C." box="[260,457,1216,1241]" pageId="1" pageNumber="268" refId="ref8267" refString="Sharma, R. C. (2007) The Fauna of India and the adjacent countries-Reptilia. Published by the Director, Zoological Survey of India, Kolkata, 410 pp." type="book" year="2007">Sharma (2007: 238</bibRefCitation>
</treatmentCitation>
, in part); 
<treatmentCitation id="111910FBFFC1FFCA3B471B9CFC8BFB39" author="Ahmed, M. F. &amp; Das, A. &amp; Dutta, S. K." box="[553,803,1216,1241]" page="159" pageId="1" pageNumber="268" year="2009">
<bibRefCitation id="F4294B1BFFC1FFCA3B471B9CFC8BFB39" author="Ahmed, M. F. &amp; Das, A. &amp; Dutta, S. K." box="[553,803,1216,1241]" pageId="1" pageNumber="268" refId="ref6853" refString="Ahmed, M. F., Das, A. &amp; Dutta, S. K. (2009) Amphibians and reptiles of Northeast India: A photographic guide. Aaranyak, Guwahati, Assam, 169 pp." type="book" year="2009">
Ahmed 
<emphasis id="A2CCEAF8FFC1FFCA3B141B9DFD05FB39" box="[634,685,1216,1240]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
(2009: 159
</bibRefCitation>
</treatmentCitation>
, in part); 
<treatmentCitation id="111910FBFFC1FFCA3AEB1B9DFBFEFB39" author="Das, A." box="[901,1110,1216,1241]" page="261" pageId="1" pageNumber="268" year="2010">
<bibRefCitation id="F4294B1BFFC1FFCA3AEB1B9DFBFEFB39" author="Das, A." box="[901,1110,1216,1241]" pageId="1" pageNumber="268" refId="ref7414" refString="Das, A. (2010) Systematics and biogeography of the snakes of Northeast India. Unpublished PhD thesis, Utkal University, Orissa, 467 pp." type="book" year="2010">Das (2010: 261–265</bibRefCitation>
</treatmentCitation>
, in part)
</treatmentCitationGroup>
</paragraph>
<paragraph id="900736EAFFC1FFCA39F91B83FCA4FB16" blockId="1.[151,1436,1155,1302]" box="[151,780,1247,1271]" pageId="1" pageNumber="268">
<treatmentCitationGroup id="B0A811C4FFC1FFCA39F91B83FCA4FB16" box="[151,780,1247,1271]" pageId="1" pageNumber="268">
<taxonomicName id="57B84D69FFC1FFCA39F91B83FEF1FB16" authority="Sanyal &amp; Gayen 2006" box="[151,345,1247,1271]" class="Reptilia" family="Colubridae" genus="Rhabdops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="bicolour">
<emphasis id="A2CCEAF8FFC1FFCA39F91B83FEF1FB16" box="[151,345,1247,1271]" italics="true" pageId="1" pageNumber="268">Rhabdops bicolour</emphasis>
</taxonomicName>
[sic]: 
<treatmentCitation id="111910FBFFC1FFCA38F71B83FD10FB16" author="Sanyal, D. P. &amp; Gayen, N. C." box="[409,696,1247,1271]" page="248" pageId="1" pageNumber="268" year="2006">
<bibRefCitation id="F4294B1BFFC1FFCA38F71B83FD10FB16" author="Sanyal, D. P. &amp; Gayen, N. C." box="[409,696,1247,1271]" pageId="1" pageNumber="268" pagination="247 - 284" refId="ref8206" refString="Sanyal, D. P. &amp; Gayen, N. C. (2006) State Fauna Series (Reptilia). In: Alfred, J. R. B. (Ed.), Fauna of Arunachal Pradesh. 13 (1). Zoological Survey of India, Kolkata, pp. 247 - 284." type="book chapter" year="2006">Sanyal &amp; Gayen (2006: 248</bibRefCitation>
</treatmentCitation>
, in part)
</treatmentCitationGroup>
</paragraph>
<paragraph id="900736EAFFC1FFCA39F91BA2FB1EFAF7" blockId="1.[151,1436,1155,1302]" box="[151,1206,1278,1302]" pageId="1" pageNumber="268">
<treatmentCitationGroup id="B0A811C4FFC1FFCA39F91BA2FB1EFAF7" box="[151,1206,1278,1302]" pageId="1" pageNumber="268">
<taxonomicName id="57B84D69FFC1FFCA39F91BA2FE42FAF7" authority="(Blyth, 1854)" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[151,490,1278,1302]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="A2CCEAF8FFC1FFCA39F91BA2FEFFFAF7" box="[151,343,1278,1302]" italics="true" pageId="1" pageNumber="268">Smithophis bicolor</emphasis>
(Blyth, 1854)
</taxonomicName>
: 
<treatmentCitation id="111910FBFFC1FFCA389A1BA2FD64FAF7" LSID="urn:lsid:zoobank.org:act:97669923-F81A-42C6-B14F-251D6A4BD1C2" author="Giri, V. B. &amp; Gower, D. J. &amp; Das, A. &amp; Lalremsanga, H. T. &amp; Lalronunga, S. &amp; Captain, A. &amp; Deepak, V." box="[500,716,1278,1302]" httpUri="http://treatment.plazi.org/id/D52387BEFFE4FFE4FF7FFA310C1DF861" page="245" pageId="1" pageNumber="268" year="2019">
<bibRefCitation id="F4294B1BFFC1FFCA389A1BA2FD64FAF7" author="Giri, V. B. &amp; Gower, D. J. &amp; Das, A. &amp; Lalremsanga, H. T. &amp; Lalronunga, S. &amp; Captain, A. &amp; Deepak, V." box="[500,716,1278,1302]" pageId="1" pageNumber="268" pagination="241 - 264" refId="ref7663" refString="Giri, V. B., Gower, D. J., Das, A., Lalremsanga, H. T., Lalronunga, S., Captain, A. &amp; Deepak, V. (2019) A new genus and species of natricine snake from northeast India. Zootaxa, 4603 (2), 241 - 264. https: // doi. org / 10.11646 / zootaxa. 4603.2.2" type="journal article" year="2019">
Giri 
<emphasis id="A2CCEAF8FFC1FFCA3B4D1BA3FDFEFAF7" box="[547,598,1278,1302]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
(2019: 245
</bibRefCitation>
</treatmentCitation>
, 255, in part); 
<treatmentCitation id="111910FBFFC1FFCA3A0E1BA2FB96FAF7" author="Vogel, G. &amp; Chen, Z. &amp; Deepak, V. &amp; Gower, D. J. &amp; Shi, J. &amp; Ding, L. &amp; Hou, M." box="[864,1086,1278,1302]" page="51" pageId="1" pageNumber="268" year="2020">
<bibRefCitation id="F4294B1BFFC1FFCA3A0E1BA2FB96FAF7" author="Vogel, G. &amp; Chen, Z. &amp; Deepak, V. &amp; Gower, D. J. &amp; Shi, J. &amp; Ding, L. &amp; Hou, M." box="[864,1086,1278,1302]" pageId="1" pageNumber="268" pagination="1" refId="ref8453" refString="Vogel, G., Chen, Z., Deepak, V., Gower, D. J., Shi, J., Ding, L. &amp; Hou, M. (2020) A new species of the genus Smithophis (Squamata: Serpentes: Natricidae) from southwestern China and northeastern Myanmar. Zootaxa, 4803 (1), 051 - 074. https: // doi. org / 10.11646 / zootaxa. 4803.1.3" type="journal article" year="2020">
Vogel 
<emphasis id="A2CCEAF8FFC1FFCA3ACF1BA3FC7DFAF7" box="[929,981,1278,1302]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
(2020: 51
</bibRefCitation>
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, 52, 54, 64)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D8A26561FFC1FFCA39F91A18FAA6F8EE" pageId="1" pageNumber="268" type="materials_examined">
<paragraph id="900736EAFFC1FFCA39F91A18FAB7FA62" blockId="1.[151,1437,1348,2023]" pageId="1" pageNumber="268">
<materialsCitation id="20D03CB7FFC1FFCA39F91A18FAB7FA62" ID-GBIF-Occurrence="3034604301" collectingDate="2005-04-21" collectionCode="BNHS" country="India" county="Abhijit Das" elevation="310" latitude="28.101334" location="Koronu" longLatPrecision="1" longitude="95.90811" municipality="Lower Dibang Valley District" pageId="1" pageNumber="268" specimenCode="BNHS 3650" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="A2CCEAF8FFC1FFCA39F91A18FEA3FABF" bold="true" box="[151,267,1348,1374]" pageId="1" pageNumber="268">
<typeStatus id="4F038848FFC1FFCA39F91A18FEAEFABF" box="[151,262,1348,1374]" pageId="1" pageNumber="268" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="C01E9E91FFC1FFCA38791A18FE0DFABE" box="[279,421,1348,1375]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="1" pageNumber="268">BNHS 3650</specimenCode>
(
<figureCitation id="08832A6FFFC1FFCA38D71A18FD81FABF" box="[441,553,1348,1374]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1259,1283]" captionStartId-1="4.[151,250,1963,1987]" captionTargetBox-0="[151,1436,309,1236]" captionTargetBox-1="[167,1420,1101,1938]" captionTargetId-0="figure-77@3.[151,1436,309,1236]" captionTargetId-1="figure-436@4.[167,1420,1101,1938]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Smithophis arunachalensis sp. nov., holotype (BNHS 3650). A. full body dorsal, and B. ventral view, C. head dorsal, D. ventral, and E– F. lateral views of head. Scale bars 10mm." captionText-1="FIGURE 2. Smithophis arunachalensis sp. nov., holotype (BNHS 3650) in life." figureDoi-0="http://doi.org/10.5281/zenodo.4413799" figureDoi-1="http://doi.org/10.5281/zenodo.4413801" httpUri-0="https://zenodo.org/record/4413799/files/figure.png" httpUri-1="https://zenodo.org/record/4413801/files/figure.png" pageId="1" pageNumber="268">Figs. 1–2</figureCitation>
), female, collected from 
<location id="95676031FFC1FFCA3A3B1A19FC05FABE" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA3A3B1A19FC05FABE" box="[853,941,1349,1375]" country="India" county="Abhijit Das" latitude="28.101334" longLatPrecision="1" longitude="95.90811" municipality="Lower Dibang Valley District" name="Koronu" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Koronu</location>
, 
<collectingMunicipality id="7063AC90FFC1FFCA3AD31A19FAB1FABF" box="[957,1305,1348,1375]" pageId="1" pageNumber="268">Lower Dibang Valley District</collectingMunicipality>
, 
<collectingRegion id="527CF808FFC1FFCA3C461A18FF5BFA63" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA39901A34FE90FA63" box="[254,312,1384,1410]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
(
<figureCitation id="08832A6FFFC1FFCA38291A34FE25FA63" box="[327,397,1384,1410]" captionStart="FIGURE 4" captionStartId="7.[151,250,1961,1985]" captionTargetBox="[184,1404,207,1930]" captionTargetId="figure-19@7.[175,1412,199,1938]" captionTargetPageId="7" captionText="FIGURE 4. Smithophis arunachalensis sp. nov., paratypes in dorsal views A. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and B. (ZSIK 23875). Scale bar 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413807" httpUri="https://zenodo.org/record/4413807/files/figure.png" pageId="1" pageNumber="268">Fig. 4</figureCitation>
: 
<geoCoordinate id="F58C502DFFC1FFCA38F61A34FD90FA62" box="[408,568,1384,1411]" degrees="28.101333" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="28.101334">28.101333° N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA3B2D1A34FD75FA62" box="[579,733,1384,1411]" degrees="95.908111" direction="east" orientation="longitude" pageId="1" pageNumber="268" precision="1" value="95.90811">95.908111° E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA3B871A34FC98FA62" box="[745,816,1384,1411]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="268" unit="m" value="310.0">
<elevation id="1B95D1D9FFC1FFCA3B871A34FC98FA62" box="[745,816,1384,1411]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="268" unit="m" value="310.0">310 m</elevation>
</quantity>
elevation) by 
<collectingCounty id="79664E66FFC1FFCA3ABE1A34FBFAFA62" box="[976,1106,1384,1411]" pageId="1" pageNumber="268">Abhijit Das</collectingCounty>
on 
<date id="E406102AFFC1FFCA3D151A34FAB3FA62" box="[1147,1307,1384,1411]" pageId="1" pageNumber="268" value="2005-04-21">
<collectingDate id="F442E9C2FFC1FFCA3D151A34FAB3FA62" box="[1147,1307,1384,1411]" pageId="1" pageNumber="268" value="2005-04-21">21 April 2005</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="900736EAFFC1FFCA39A91AD0FE98F9D7" blockId="1.[151,1437,1348,2023]" pageId="1" pageNumber="268">
<materialsCitation id="20D03CB7FFC1FFCA39A91AD0FD82FA0F" ID-GBIF-Occurrence="3034604303" collectingDate="1935-10-12" collectionCode="BMNH" collectorName="D. K. Barnah &amp; M. Steele" country="India" county="Lohit District" elevation="650" latitude="28.019224" location="Dening" longLatPrecision="1" longitude="96.227776" municipality="Mishmi Hills" pageId="1" pageNumber="268" specimenCount="1" stateProvince="Assam" typeStatus="paratype">
<emphasis id="A2CCEAF8FFC1FFCA39A91AD0FE36FA47" bold="true" box="[199,414,1420,1446]" pageId="1" pageNumber="268">
<typeStatus id="4F038848FFC1FFCA39A91AD0FE97FA47" box="[199,319,1420,1446]" pageId="1" pageNumber="268" type="paratype">Paratypes</typeStatus>
(n = 2).
</emphasis>
<collectionCode id="F6A9AE2FFFC1FFCA38CB1AD1FE51FA46" box="[421,505,1421,1447]" pageId="1" pageNumber="268">BMNH</collectionCode>
<date id="E406102AFFC1FFCA3B6F1AD0FD2DFA46" box="[513,645,1420,1447]" pageId="1" pageNumber="268" value="1935-10-12">
<collectingDate id="F442E9C2FFC1FFCA3B6F1AD0FD2DFA46" box="[513,645,1420,1447]" pageId="1" pageNumber="268" value="1935-10-12">1935.10.12.</collectingDate>
</date>
10 (
<figureCitation id="08832A6FFFC1FFCA3BDC1AD0FCA7FA47" box="[690,783,1420,1446]" captionStart="FIGURE 4" captionStartId="7.[151,250,1961,1985]" captionTargetBox="[184,1404,207,1930]" captionTargetId="figure-19@7.[175,1412,199,1938]" captionTargetPageId="7" captionText="FIGURE 4. Smithophis arunachalensis sp. nov., paratypes in dorsal views A. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and B. (ZSIK 23875). Scale bar 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413807" httpUri="https://zenodo.org/record/4413807/files/figure.png" pageId="1" pageNumber="268">Figs. 4a</figureCitation>
, 
<figureCitation id="08832A6FFFC1FFCA3A751AD0FC82FA46" box="[795,810,1420,1447]" captionStart="FIGURE 5" captionStartId="8.[151,250,1785,1809]" captionTargetBox="[328,1177,257,1690]" captionTargetId="figure-19@8.[235,1352,181,1762]" captionTargetPageId="8" captionText="FIGURE 5. Smithophis arunachalensis sp. nov., reconstructed head of paratype (BMNH 1935.10.12.10), breaks in specimen marked with red dashed lines. Scale bar 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413809" httpUri="https://zenodo.org/record/4413809/files/figure.png" pageId="1" pageNumber="268">5</figureCitation>
), male, collected from 
<location id="95676031FFC1FFCA3D5C1AD0FB2FFA47" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA3D5C1AD0FB2FFA47" box="[1074,1159,1420,1446]" country="India" county="Lohit District" latitude="28.019224" longLatPrecision="1" longitude="96.227776" municipality="Mishmi Hills" name="Dening" pageId="1" pageNumber="268" stateProvince="Assam">Dening</location>
, 
<collectingMunicipality id="7063AC90FFC1FFCA3DFD1AD0FA83FA47" box="[1171,1323,1420,1446]" pageId="1" pageNumber="268">Mishmi Hills</collectingMunicipality>
, “ 
<collectingRegion id="527CF808FFC1FFCA3C2F1AD0FA3AFA47" box="[1345,1426,1420,1446]" country="India" name="Assam" pageId="1" pageNumber="268">Assam</collectingRegion>
” (
<geoCoordinate id="F58C502DFFC1FFCA39F11AECFEE9FA2A" box="[159,321,1456,1483]" degrees="28.019225" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="28.019224">28.019225° N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA38221AECFE40FA2A" box="[332,488,1456,1483]" degrees="96.227778" direction="east" orientation="longitude" pageId="1" pageNumber="268" precision="1" value="96.227776">96.227778° E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA389A1AECFD92FA2A" box="[500,570,1456,1483]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="268" unit="m" value="650.0">
<elevation id="1B95D1D9FFC1FFCA389A1AECFD92FA2A" box="[500,570,1456,1483]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="268" unit="m" value="650.0">650 m</elevation>
</quantity>
elevation, currently in 
<collectingCounty id="79664E66FFC1FFCA3A501AECFC70FA2B" box="[830,984,1456,1482]" pageId="1" pageNumber="268">Lohit District</collectingCounty>
, 
<collectingRegion id="527CF808FFC1FFCA3A8C1AECFB11FA2B" box="[994,1209,1456,1482]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA3DAA1AECFAAAFA2A" box="[1220,1282,1456,1483]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
), collected by 
<collectorName id="3D4D533CFFC1FFCA39F91A89FE82FA0F" box="[151,298,1492,1519]" pageId="1" pageNumber="268">D.K. Barnah</collectorName>
, donated by 
<collectorName id="3D4D533CFFC1FFCA38D41A89FD82FA0F" box="[442,554,1492,1519]" pageId="1" pageNumber="268">M. Steele</collectorName>
</materialsCitation>
; 
<materialsCitation id="20D03CB7FFC1FFCA3B591A88FE84F9D7" ID-GBIF-Occurrence="3034604302" collectingDate="1981-04-23" collectionCode="ZSIK" collectorName="S. Biswas" country="India" county="Changlang District" elevation="580" latitude="27.492361" location="Namdapha Camp" longLatPrecision="1" longitude="96.38167" municipality="Miao" pageId="1" pageNumber="268" specimenCode="ZSIK 23875" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCode id="C01E9E91FFC1FFCA3B591A88FD6BFA0E" box="[567,707,1492,1519]" collectionCode="ZSIK" pageId="1" pageNumber="268">ZSIK 23875</specimenCode>
(
<figureCitation id="08832A6FFFC1FFCA3BBD1A88FC80FA0E" box="[723,808,1492,1519]" captionStart="FIGURE 4" captionStartId="7.[151,250,1961,1985]" captionTargetBox="[184,1404,207,1930]" captionTargetId="figure-19@7.[175,1412,199,1938]" captionTargetPageId="7" captionText="FIGURE 4. Smithophis arunachalensis sp. nov., paratypes in dorsal views A. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and B. (ZSIK 23875). Scale bar 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413807" httpUri="https://zenodo.org/record/4413807/files/figure.png" pageId="1" pageNumber="268">Fig. 4b</figureCitation>
), male, collected from 
<location id="95676031FFC1FFCA3D401A88FB5DFA0F" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA3D401A88FB5DFA0F" box="[1070,1269,1492,1518]" country="India" county="Changlang District" latitude="27.492361" longLatPrecision="1" longitude="96.38167" municipality="Miao" name="Namdapha Camp" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Namdapha Camp</location>
, 
<quantity id="57409B0FFFC1FFCA3C6F1A88FAE1FA0F" box="[1281,1353,1492,1519]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="268" unit="km" value="58.0">58 km</quantity>
east of 
<collectingMunicipality id="7063AC90FFC1FFCA39F91AA4FF7DF9F3" box="[151,213,1528,1554]" pageId="1" pageNumber="268">Miao</collectingMunicipality>
, 
<collectingCounty id="79664E66FFC1FFCA398E1AA4FE10F9F3" box="[224,440,1528,1554]" pageId="1" pageNumber="268">Changlang District</collectingCounty>
, 
<collectingRegion id="527CF808FFC1FFCA38AF1AA4FD31F9F3" box="[449,665,1528,1554]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA3BCA1AA4FD76F9F3" box="[676,734,1528,1554]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
(
<geoCoordinate id="F58C502DFFC1FFCA3B831AA4FC2FF9F3" box="[749,903,1528,1554]" degrees="27.492361" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="27.492361">27.492361°N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA3AFD1AA4FB80F9F3" box="[915,1064,1528,1554]" degrees="96.381666" direction="east" orientation="longitude" pageId="1" pageNumber="268" precision="1" value="96.38167">96.381666°E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA3D5A1AA4FBD3F9F2" box="[1076,1147,1528,1555]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="268" unit="m" value="580.0">
<elevation id="1B95D1D9FFC1FFCA3D5A1AA4FBD3F9F2" box="[1076,1147,1528,1555]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="268" unit="m" value="580.0">580 m</elevation>
</quantity>
elevation) 
<date id="E406102AFFC1FFCA3D971AA4FA3FF9F3" box="[1273,1431,1528,1554]" pageId="1" pageNumber="268" value="1981-04-23">
<collectingDate id="F442E9C2FFC1FFCA3D971AA4FA3FF9F3" box="[1273,1431,1528,1554]" pageId="1" pageNumber="268" value="1981-04-23">23 April 1981</collectingDate>
</date>
, by 
<collectorName id="3D4D533CFFC1FFCA39D41940FE84F9D7" box="[186,300,1564,1590]" pageId="1" pageNumber="268">S. Biswas</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="900736EAFFC1FFCA39A9191CFAA6F8EE" blockId="1.[151,1437,1348,2023]" pageId="1" pageNumber="268">
<emphasis id="A2CCEAF8FFC1FFCA39A9191CFDA3F9BB" bold="true" box="[199,523,1600,1626]" pageId="1" pageNumber="268">Referred specimens (n = 3).</emphasis>
<materialsCitation id="20D03CB7FFC1FFCA3B7C191CFAA2F8EE" ID-GBIF-Occurrence="3034604304" collectingDate="1991-10-04" collectingDateMax="1995-09-08" collectingDateMin="1991-10-04" country="India" county="Changlang District" elevation="312" latitude="27.487871" location="Roing Forest Department" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" pageId="1" pageNumber="268" specimenCount="1" stateProvince="Arunachal Pradesh">
<location id="95676031FFC1FFCA3B7C191CFC84F9BA" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA3B7C191CFC84F9BA" box="[530,812,1600,1627]" country="India" county="Changlang District" latitude="27.487871" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" name="Roing Forest Department" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">
<location id="95676031FFC1FFCA3B7C191CFD0AF9BA" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA3B7C191CFD0AF9BA" box="[530,674,1600,1627]" country="India" county="Changlang District" latitude="27.487871" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" name="Roing Forest" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Roing Forest</location>
Department
</location>
collection, uncatalogued (
<figureCitation id="08832A6FFFC1FFCA3D3C191CFB0BF9BA" box="[1106,1187,1600,1627]" captionStart="FIGURE 6" captionStartId="9.[151,250,1938,1962]" captionTargetBox="[151,1436,190,1902]" captionTargetId="figure-19@9.[151,1436,181,1915]" captionTargetPageId="9" captionText="FIGURE 6. Smithophis arunachalensis sp. nov., A. unnumbered referred specimen in Roing Forest Department collection. Total length 613 mm. B. unnumbered (2) referred specimen in Miao museum collection." figureDoi="http://doi.org/10.5281/zenodo.4413813" httpUri="https://zenodo.org/record/4413813/files/figure.png" pageId="1" pageNumber="268">Fig. 6a</figureCitation>
), male, collected from 
<location id="95676031FFC1FFCA39F91938FF76F99F" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA39F91938FF76F99F" box="[151,222,1636,1662]" country="India" county="Changlang District" latitude="27.487871" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" name="Roing" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Roing</location>
, 
<location id="95676031FFC1FFCA39851939FD96F99F" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA39851939FD96F99F" box="[235,574,1636,1663]" country="India" county="Changlang District" latitude="27.487871" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" name="Lower Dibang Valley District" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Lower Dibang Valley District</location>
, 
<collectingRegion id="527CF808FFC1FFCA3B271938FC8BF99F" box="[585,803,1636,1662]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA3A411938FCC1F99F" box="[815,873,1636,1662]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
(
<geoCoordinate id="F58C502DFFC1FFCA3A171938FBB4F99E" box="[889,1052,1636,1663]" degrees="28.157000" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="28.157">28.157000° N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA3D471938FB6FF99E" box="[1065,1223,1636,1663]" degrees="95.854483" direction="east" orientation="longitude" pageId="1" pageNumber="268" precision="1" value="95.854485">95.854483° E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA3DBA1938FAB5F99E" box="[1236,1309,1636,1663]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="1" pageNumber="268" unit="m" value="312.0">
<elevation id="1B95D1D9FFC1FFCA3DBA1938FAB5F99E" box="[1236,1309,1636,1663]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="1" pageNumber="268" unit="m" value="312.0">312 m</elevation>
</quantity>
elevation). 
<collectorName id="3D4D533CFFC1FFCA39F919D4FE96F942" box="[151,318,1672,1699]" pageId="1" pageNumber="268">Miao Museum</collectorName>
, 
<location id="95676031FFC1FFCA382719D4FDC0F942" LSID="urn:lsid:plazi:treatment:181187FCFFC1FFC039F91CAAFF08F879:95676031FFC1FFCA382719D4FDC0F942" box="[329,616,1672,1699]" country="India" county="Changlang District" latitude="27.487871" longLatPrecision="1" longitude="96.386475" municipality="Changlang District" name="Namdapha Tiger Reserve" pageId="1" pageNumber="268" stateProvince="Arunachal Pradesh">Namdapha Tiger Reserve</location>
, 
<collectorName id="3D4D533CFFC1FFCA3B1D19D4FD47F943" box="[627,751,1672,1698]" pageId="1" pageNumber="268">Changlang</collectorName>
, 
<collectingRegion id="527CF808FFC1FFCA3B9619D4FC78F943" box="[760,976,1672,1698]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA3AB419D4FBBEF943" box="[986,1046,1672,1698]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
, two uncatalogued specimens, collected by 
<collectorName id="3D4D533CFFC1FFCA386619F1FE39F927" box="[264,401,1708,1735]" pageId="1" pageNumber="268">P.K. Biswas</collectorName>
on 
<date id="E406102AFFC1FFCA38D119F0FDD9F927" box="[447,625,1708,1734]" pageId="1" pageNumber="268" value="1991-10-04">
<collectingDate id="F442E9C2FFC1FFCA38D119F0FDD9F927" box="[447,625,1708,1734]" pageId="1" pageNumber="268" value="1991-10-04">4 October 1991</collectingDate>
</date>
from 14 
<superScript id="67CD9BA2FFC1FFCA3BB819F0FD4BF95B" attach="left" box="[726,739,1708,1722]" fontSize="6" pageId="1" pageNumber="268">th</superScript>
mile, Miao-Vijoynagar Road, 
<collectingMunicipality id="7063AC90FFC1FFCA3D2D19F0FAB4F927" box="[1091,1308,1708,1734]" pageId="1" pageNumber="268">Changlang District</collectingMunicipality>
, 
<collectingRegion id="527CF808FFC1FFCA3C4619F0FF5BF90B" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC1FFCA3990198CFE90F90B" box="[254,312,1744,1770]" name="India" pageId="1" pageNumber="268">India</collectingCountry>
(
<geoCoordinate id="F58C502DFFC1FFCA3829198CFE4AF90A" box="[327,482,1744,1771]" degrees="27.500183" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="27.500183">27.500183°N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA3883198CFD2BF90B" box="[493,643,1744,1770]" degrees="96.386472" direction="east" orientation="longitude" pageId="1" pageNumber="268" precision="1" value="96.386475">96.386472°E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA3BE1198CFD7EF90A" box="[655,726,1744,1771]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="268" unit="m" value="330.0">
<elevation id="1B95D1D9FFC1FFCA3BE1198CFD7EF90A" box="[655,726,1744,1771]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="268" unit="m" value="330.0">330 m</elevation>
</quantity>
elevation), and by 
<collectorName id="3D4D533CFFC1FFCA3AC1198DFB9EF90B" box="[943,1078,1744,1771]" pageId="1" pageNumber="268">P.K. Biswas</collectorName>
on 
<date id="E406102AFFC1FFCA3D0F198CFA85F90A" box="[1121,1325,1744,1771]" pageId="1" pageNumber="268" value="1995-09-08">
<collectingDate id="F442E9C2FFC1FFCA3D0F198CFA85F90A" box="[1121,1325,1744,1771]" pageId="1" pageNumber="268" value="1995-09-08">8 September 1995</collectingDate>
</date>
from Deban (
<figureCitation id="08832A6FFFC1FFCA39A119A8FE8AF8EE" box="[207,290,1780,1807]" captionStart="FIGURE 6" captionStartId="9.[151,250,1938,1962]" captionTargetBox="[151,1436,190,1902]" captionTargetId="figure-19@9.[151,1436,181,1915]" captionTargetPageId="9" captionText="FIGURE 6. Smithophis arunachalensis sp. nov., A. unnumbered referred specimen in Roing Forest Department collection. Total length 613 mm. B. unnumbered (2) referred specimen in Miao museum collection." figureDoi="http://doi.org/10.5281/zenodo.4413813" httpUri="https://zenodo.org/record/4413813/files/figure.png" pageId="1" pageNumber="268">Fig. 6b</figureCitation>
), 
<collectingCounty id="79664E66FFC1FFCA385919A8FDA6F8EF" box="[311,526,1780,1806]" pageId="1" pageNumber="268">Changlang District</collectingCounty>
, 
<collectingRegion id="527CF808FFC1FFCA3B7919A8FD44F8EF" box="[535,748,1780,1806]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="268">Arunachal Pradesh</collectingRegion>
(
<geoCoordinate id="F58C502DFFC1FFCA3B9419A8FC3DF8EF" box="[762,917,1780,1806]" degrees="27.487872" direction="north" orientation="latitude" pageId="1" pageNumber="268" precision="1" value="27.487871">27.487872°N</geoCoordinate>
, 
<geoCoordinate id="F58C502DFFC1FFCA3ACE19A8FB94F8EF" box="[928,1084,1780,1806]" pageId="1" pageNumber="268">96.344 244 °E</geoCoordinate>
, 
<quantity id="57409B0FFFC1FFCA3D2719A8FB38F8EE" box="[1097,1168,1780,1807]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="268" unit="m" value="560.0">
<elevation id="1B95D1D9FFC1FFCA3D2719A8FB38F8EE" box="[1097,1168,1780,1807]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="268" unit="m" value="560.0">560 m</elevation>
</quantity>
elevation)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A26561FFC1FFC939A91844FE99FEFC" lastPageId="2" lastPageNumber="269" pageId="1" pageNumber="268" type="diagnosis">
<paragraph id="900736EAFFC1FFC939A91844FE99FEFC" blockId="1.[151,1437,1348,2023]" lastBlockId="2.[151,1437,151,393]" lastPageId="2" lastPageNumber="269" pageId="1" pageNumber="268">
<emphasis id="A2CCEAF8FFC1FFCA39A91844FDB2F8D3" bold="true" box="[199,538,1816,1842]" pageId="1" pageNumber="268">Diagnosis and identification.</emphasis>
<taxonomicName id="57B84D69FFC1FFCA3B4E1844FCE6F8D3" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[544,846,1816,1842]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC1FFCA3B4E1844FCE6F8D3" box="[544,846,1816,1842]" italics="true" pageId="1" pageNumber="268">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC1FFCA3A3A1844FC05F8D3" bold="true" box="[852,941,1816,1842]" pageId="1" pageNumber="268">
<taxonomicNameLabel id="B9FF5783FFC1FFCA3A3A1844FC05F8D3" box="[852,941,1816,1842]" pageId="1" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is referred to the genus 
<taxonomicName id="57B84D69FFC1FFCA3DD91844FA9BF8D3" authorityName="Giri, Gower, Das, Lalremsanga, Lalronunga, Captain &amp; Deepak" authorityYear="2019" box="[1207,1331,1816,1842]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="genus">
<emphasis id="A2CCEAF8FFC1FFCA3DD91844FA9BF8D3" box="[1207,1331,1816,1842]" italics="true" pageId="1" pageNumber="268">Smithophis</emphasis>
</taxonomicName>
based on the presence of single internasal and single prefrontal shields (a unique condition within 
<taxonomicName id="57B84D69FFC1FFCA3DEE1860FB51F8B7" authorityName="Bonaparte" authorityYear="1838" box="[1152,1273,1852,1878]" class="Reptilia" family="Natricinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="family">Natricinae</taxonomicName>
: see 
<bibRefCitation id="F4294B1BFFC1FFCA3C411860FF7AF89B" author="Giri, V. B. &amp; Deepak, V. &amp; Captain, A. &amp; Das, A. &amp; Das, S. &amp; Rajkumar, K. P. &amp; Rathish, R. L. &amp; Gower, D. J." pageId="1" pageNumber="268" pagination="27 - 52" refId="ref7568" refString="Giri, V. B., Deepak, V., Captain, A., Das, A., Das, S., Rajkumar, K. P., Rathish, R. L. &amp; Gower, D. J. (2017) A new species of Rhabdops Boulenger, 1893 (Serpentes: Natricinae) from the northern Western Ghats region of India. Zootaxa, 4319 (1), 27 - 52. https: // doi. org / 10.11646 / zootaxa. 4319.1.2" type="journal article" year="2017">
Giri 
<emphasis id="A2CCEAF8FFC1FFCA3C0A1861FA35F8B7" box="[1380,1437,1852,1878]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
2017
</bibRefCitation>
; 
<bibRefCitation id="F4294B1BFFC1FFCA39B1183CFE0EF89B" author="Vogel, G. &amp; Chen, Z. &amp; Deepak, V. &amp; Gower, D. J. &amp; Shi, J. &amp; Ding, L. &amp; Hou, M." box="[223,422,1888,1914]" pageId="1" pageNumber="268" pagination="1" refId="ref8453" refString="Vogel, G., Chen, Z., Deepak, V., Gower, D. J., Shi, J., Ding, L. &amp; Hou, M. (2020) A new species of the genus Smithophis (Squamata: Serpentes: Natricidae) from southwestern China and northeastern Myanmar. Zootaxa, 4803 (1), 051 - 074. https: // doi. org / 10.11646 / zootaxa. 4803.1.3" type="journal article" year="2020">
Vogel 
<emphasis id="A2CCEAF8FFC1FFCA3846183DFEC9F89B" box="[296,353,1888,1914]" italics="true" pageId="1" pageNumber="268">et al.</emphasis>
2020
</bibRefCitation>
). The new species is distinguished from congeners as follows: 
<taxonomicName id="57B84D69FFC1FFCA3D1F183CFA91F89B" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[1137,1337,1888,1915]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC1FFCA3D1F183CFBD7F89B" box="[1137,1151,1888,1914]" italics="true" pageId="1" pageNumber="268">S</emphasis>
. 
<emphasis id="A2CCEAF8FFC1FFCA3DE0183CFA91F89B" box="[1166,1337,1888,1914]" italics="true" pageId="1" pageNumber="268">arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC1FFCA3C2C183CFA34F89B" bold="true" box="[1346,1436,1888,1914]" pageId="1" pageNumber="268">
<taxonomicNameLabel id="B9FF5783FFC1FFCA3C2C183CFA34F89B" box="[1346,1436,1888,1914]" pageId="1" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="57B84D69FFC1FFCA384D18D8FE6CF87F" authorityName="Giri, Gower, Das, Lalremsanga, Lalronunga, Captain &amp; Deepak" authorityYear="2019" box="[291,452,1924,1950]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="atemporalis">
<emphasis id="A2CCEAF8FFC1FFCA384D18D8FE6CF87F" box="[291,452,1924,1950]" italics="true" pageId="1" pageNumber="268">S. atemporalis</emphasis>
</taxonomicName>
by the presence of temporal shields (versus absence) and by having bright yellow venter in life (versus whitish); from 
<taxonomicName id="57B84D69FFC1FFCA388218F4FDF3F823" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[492,603,1960,1986]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="A2CCEAF8FFC1FFCA388218F4FDF3F823" box="[492,603,1960,1986]" italics="true" pageId="1" pageNumber="268">S. bicolor</emphasis>
</taxonomicName>
in colour pattern, by having a sharply zigzag border between the darker dorsum and paler venter, along the entire lateral surface of the body and tail (versus absence of zigzag border), and in having (versus lacking) and a dark midventral line on the underside of the tail (see 
<figureCitation id="08832A6FFFC2FFC93D341FCBFB09FF50" box="[1114,1185,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1259,1283]" captionTargetBox="[151,1436,309,1236]" captionTargetId="figure-77@3.[151,1436,309,1236]" captionTargetPageId="3" captionText="FIGURE 1. Smithophis arunachalensis sp. nov., holotype (BNHS 3650). A. full body dorsal, and B. ventral view, C. head dorsal, D. ventral, and E– F. lateral views of head. Scale bars 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413799" httpUri="https://zenodo.org/record/4413799/files/figure.png" pageId="2" pageNumber="269">Figs. 1</figureCitation>
–2,4,6,8–9; 
<tableCitation id="DD3A0351FFC2FFC93C421FCBFAD7FF50" box="[1324,1407,151,177]" captionStart="TABLE 2" captionStartId="6.[151,239,151,175]" captionTargetPageId="6" captionText="TABLE 2. Meristic and morphometric (in millimetres) data for examined specimens of Smithophis bicolor. Some bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. Locality details are provided in Appendix I." httpUri="http://table.plazi.org/id/C4C76662FFC6FFCD39F91FCBFD54FF16" pageId="2" pageNumber="269" tableUuid="C4C76662FFC6FFCD39F91FCBFD54FF16">Table 2</tableCitation>
&amp; 
<tableCitation id="DD3A0351FFC2FFC939F91FE7FF0FFF34" box="[151,167,187,213]" captionStart="TABLE 3" captionStartId="13.[151,239,309,333]" captionTargetPageId="13" captionText="TABLE 3. Summary of features discriminating among four currently recognized species of Smithophis. * only on anterior part of tail in some specimens (e.g. Giri et al. 2019: fig. 2B) and absent in some others." httpUri="http://table.plazi.org/id/C4C76662FFCDFFC639F91E69FB91FE93" pageId="2" pageNumber="269" tableUuid="C4C76662FFCDFFC639F91E69FB91FE93">3</tableCitation>
). The new species differs from 
<taxonomicName id="57B84D69FFC2FFC93B671FE7FDD3FF34" authorityName="Vogel, Chen, Deepak, Gower, Shi, Ding &amp; Hou" authorityYear="2020" box="[521,635,187,213]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="linearis">
<emphasis id="A2CCEAF8FFC2FFC93B671FE7FDD3FF34" box="[521,635,187,213]" italics="true" pageId="2" pageNumber="269">S. linearis</emphasis>
</taxonomicName>
by having fewer than six circumorbital scales (excluding the supralabials: versus more than 5) and by lacking (versus having) narrow dark and pale longitudinal lines on upper surface of body and tail.
</paragraph>
</subSubSection>
<subSubSection id="D8A26561FFC2FFCE39A91E7AFEF7F8EC" lastPageId="5" lastPageNumber="272" pageId="2" pageNumber="269" type="description">
<paragraph id="900736EAFFC2FFC939A91E7AFE0DFE68" blockId="2.[151,1437,151,393]" pageId="2" pageNumber="269">
<emphasis id="A2CCEAF8FFC2FFC939A91E7AFE42FEA0" bold="true" box="[199,490,294,321]" pageId="2" pageNumber="269">
Description of 
<typeStatus id="4F038848FFC2FFC938121E7BFE4DFEA0" box="[380,485,295,321]" pageId="2" pageNumber="269" type="holotype">holotype</typeStatus>
.
</emphasis>
See 
<tableCitation id="DD3A0351FFC2FFC93B4B1E7BFDD2FEA0" box="[549,634,295,321]" captionStart="TABLE 1" captionStartId="2.[151,239,440,464]" captionTargetPageId="2" captionText="TABLE 1. Meristic and morphometric (in millimetres) data for examined specimens of Smithophis arunachalensis sp. nov. Head length = distance between snout tip and back of lower jaw; bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. The format ‘X-Y’ is used to denote the shortest distance between features X and Y." httpUri="http://table.plazi.org/id/C4C76662FFC2FFC939F91EE4FD74FDDD" pageId="2" pageNumber="269" tableUuid="C4C76662FFC2FFC939F91EE4FD74FDDD">Table 1</tableCitation>
for morphometric and meristic data. Specimen in good condition, although slightly dehydrated; two constrictions on anterior half of body caused by tight tag string; outer edge of head shields missing in parts.
</paragraph>
<caption id="C4C76662FFC2FFC939F91EE4FD74FDDD" ID-Table-UUID="C4C76662FFC2FFC939F91EE4FD74FDDD" httpUri="http://table.plazi.org/id/C4C76662FFC2FFC939F91EE4FD74FDDD" pageId="2" pageNumber="269" targetBox="[166,1412,589,1646]" targetIsTable="true" targetPageId="2">
<paragraph id="900736EAFFC2FFC939F91EE4FD74FDDD" blockId="2.[151,1436,440,572]" pageId="2" pageNumber="269">
<emphasis id="A2CCEAF8FFC2FFC939F91EE4FEA3FE31" bold="true" box="[151,267,440,464]" pageId="2" pageNumber="269">TABLE 1.</emphasis>
Meristic and morphometric (in millimetres) data for examined specimens of 
<taxonomicName id="57B84D69FFC2FFC93D3D1EE4FADCFE31" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[1107,1396,440,464]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC2FFC93D3D1EE4FADCFE31" box="[1107,1396,440,464]" italics="true" pageId="2" pageNumber="269">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC2FFC93C121EE5FF6EFE14" bold="true" pageId="2" pageNumber="269">
<taxonomicNameLabel id="B9FF5783FFC2FFC93C121EE5FF6EFE14" pageId="2" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Head length = distance between snout tip and back of lower jaw; bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. The format ‘X-Y’ is used to denote the shortest distance between features X and Y.
</paragraph>
</caption>
<paragraph id="900736EAFFC2FFC9388C1D11FAB8F98F" pageId="2" pageNumber="269">
<table id="E2B8C44AFFC2003439C81D11FA2CF98F" box="[166,1412,589,1646]" gridcols="7" gridrows="24" pageId="2" pageNumber="269">
<tr id="2E8834A8FFC2003439C81D11FA2CFD4C" box="[166,1412,589,685]" gridrow="0" pageId="2" pageNumber="269" rowspan-0="1">
<th id="6D595DD4FFC20034388C1D11FDFAFD4C" box="[482,594,589,685]" gridcol="1" gridrow="0" pageId="2" pageNumber="269">BNHS 3650 (Holotype)</th>
<th id="6D595DD4FFC200343B191D11FCA1FD4C" box="[631,777,589,685]" gridcol="2" gridrow="0" pageId="2" pageNumber="269">BMNH 1935.10.12.10 (Paratype)</th>
<th id="6D595DD4FFC200343A5C1D11FC2FFD4C" box="[818,903,589,685]" gridcol="3" gridrow="0" pageId="2" pageNumber="269">ZSIK 23875</th>
<th id="6D595DD4FFC200343ADA1D11FB9FFD4C" box="[948,1079,589,685]" gridcol="4" gridrow="0" pageId="2" pageNumber="269">Roing Forest Department unnumbered</th>
<th id="6D595DD4FFC200343D321D11FB75FD4C" box="[1116,1245,589,685]" gridcol="5" gridrow="0" pageId="2" pageNumber="269">Miao Museum un- numbered 1</th>
<th id="6D595DD4FFC200343C6D1D11FA2CFD4C" box="[1283,1412,589,685]" gridcol="6" gridrow="0" pageId="2" pageNumber="269">Miao Museum un- numbered 2</th>
</tr>
<tr id="2E8834A8FFC2003439C81DE1FA2CFD34" box="[166,1412,701,725]" gridrow="1" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81DE1FE1AFD34" box="[166,434,701,725]" gridcol="0" gridrow="1" pageId="2" pageNumber="269">Sex</th>
<td id="6D595DD4FFC20034388C1DE1FDFAFD34" box="[482,594,701,725]" gridcol="1" gridrow="1" pageId="2" pageNumber="269">F</td>
<td id="6D595DD4FFC200343B191DE1FCA1FD34" box="[631,777,701,725]" gridcol="2" gridrow="1" pageId="2" pageNumber="269">M</td>
<td id="6D595DD4FFC200343A5C1DE1FC2FFD34" box="[818,903,701,725]" gridcol="3" gridrow="1" pageId="2" pageNumber="269">M</td>
<td id="6D595DD4FFC200343ADA1DE1FB9FFD34" box="[948,1079,701,725]" gridcol="4" gridrow="1" pageId="2" pageNumber="269">M</td>
<td id="6D595DD4FFC200343D321DE1FB75FD34" box="[1116,1245,701,725]" gridcol="5" gridrow="1" pageId="2" pageNumber="269">?</td>
<td id="6D595DD4FFC200343C6D1DE1FA2CFD34" box="[1283,1412,701,725]" gridcol="6" gridrow="1" pageId="2" pageNumber="269">?</td>
</tr>
<tr id="2E8834A8FFC2003439C81DB9FA2CFD1C" box="[166,1412,741,765]" gridrow="2" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81DB9FE1AFD1C" box="[166,434,741,765]" gridcol="0" gridrow="2" pageId="2" pageNumber="269">Tail length</th>
<td id="6D595DD4FFC20034388C1DB9FDFAFD1C" box="[482,594,741,765]" gridcol="1" gridrow="2" pageId="2" pageNumber="269">105*</td>
<td id="6D595DD4FFC200343B191DB9FCA1FD1C" box="[631,777,741,765]" gridcol="2" gridrow="2" pageId="2" pageNumber="269">154</td>
<td id="6D595DD4FFC200343A5C1DB9FC2FFD1C" box="[818,903,741,765]" gridcol="3" gridrow="2" pageId="2" pageNumber="269">100</td>
<td id="6D595DD4FFC200343ADA1DB9FB9FFD1C" box="[948,1079,741,765]" gridcol="4" gridrow="2" pageId="2" pageNumber="269">138</td>
<td id="6D595DD4FFC200343D321DB9FB75FD1C" box="[1116,1245,741,765]" gridcol="5" gridrow="2" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1DB9FA2CFD1C" box="[1283,1412,741,765]" gridcol="6" gridrow="2" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81C51FA2CFCC7" box="[166,1412,781,806]" gridrow="3" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81C51FE1AFCC7" box="[166,434,781,806]" gridcol="0" gridrow="3" pageId="2" pageNumber="269">Snout-vent length</th>
<td id="6D595DD4FFC20034388C1C51FDFAFCC7" box="[482,594,781,806]" gridcol="1" gridrow="3" pageId="2" pageNumber="269">510</td>
<td id="6D595DD4FFC200343B191C51FCA1FCC7" box="[631,777,781,806]" gridcol="2" gridrow="3" pageId="2" pageNumber="269">503</td>
<td id="6D595DD4FFC200343A5C1C51FC2FFCC7" box="[818,903,781,806]" gridcol="3" gridrow="3" pageId="2" pageNumber="269">378</td>
<td id="6D595DD4FFC200343ADA1C51FB9FFCC7" box="[948,1079,781,806]" gridcol="4" gridrow="3" pageId="2" pageNumber="269">475</td>
<td id="6D595DD4FFC200343D321C51FB75FCC7" box="[1116,1245,781,806]" gridcol="5" gridrow="3" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1C51FA2CFCC7" box="[1283,1412,781,806]" gridcol="6" gridrow="3" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81C69FA2CFCAF" box="[166,1412,821,846]" gridrow="4" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81C69FE1AFCAF" box="[166,434,821,846]" gridcol="0" gridrow="4" pageId="2" pageNumber="269">Total length</th>
<td id="6D595DD4FFC20034388C1C69FDFAFCAF" box="[482,594,821,846]" gridcol="1" gridrow="4" pageId="2" pageNumber="269">615</td>
<td id="6D595DD4FFC200343B191C69FCA1FCAF" box="[631,777,821,846]" gridcol="2" gridrow="4" pageId="2" pageNumber="269">657</td>
<td id="6D595DD4FFC200343A5C1C69FC2FFCAF" box="[818,903,821,846]" gridcol="3" gridrow="4" pageId="2" pageNumber="269">478</td>
<td id="6D595DD4FFC200343ADA1C69FB9FFCAF" box="[948,1079,821,846]" gridcol="4" gridrow="4" pageId="2" pageNumber="269">613</td>
<td id="6D595DD4FFC200343D321C69FB75FCAF" box="[1116,1245,821,846]" gridcol="5" gridrow="4" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1C69FA2CFCAF" box="[1283,1412,821,846]" gridcol="6" gridrow="4" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81C02FA2CFC97" box="[166,1412,862,886]" gridrow="5" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81C02FE1AFC97" box="[166,434,862,886]" gridcol="0" gridrow="5" pageId="2" pageNumber="269">Head length</th>
<td id="6D595DD4FFC20034388C1C02FDFAFC97" box="[482,594,862,886]" gridcol="1" gridrow="5" pageId="2" pageNumber="269">10.8</td>
<td id="6D595DD4FFC200343B191C02FCA1FC97" box="[631,777,862,886]" gridcol="2" gridrow="5" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1C02FC2FFC97" box="[818,903,862,886]" gridcol="3" gridrow="5" pageId="2" pageNumber="269">14.8</td>
<td id="6D595DD4FFC200343ADA1C02FB9FFC97" box="[948,1079,862,886]" gridcol="4" gridrow="5" pageId="2" pageNumber="269">9.8</td>
<td id="6D595DD4FFC200343D321C02FB75FC97" box="[1116,1245,862,886]" gridcol="5" gridrow="5" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1C02FA2CFC97" box="[1283,1412,862,886]" gridcol="6" gridrow="5" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81CDAFA2CFC7F" box="[166,1412,902,926]" gridrow="6" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81CDAFE1AFC7F" box="[166,434,902,926]" gridcol="0" gridrow="6" pageId="2" pageNumber="269">Head width (maximum)</th>
<td id="6D595DD4FFC20034388C1CDAFDFAFC7F" box="[482,594,902,926]" gridcol="1" gridrow="6" pageId="2" pageNumber="269">7.2</td>
<td id="6D595DD4FFC200343B191CDAFCA1FC7F" box="[631,777,902,926]" gridcol="2" gridrow="6" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1CDAFC2FFC7F" box="[818,903,902,926]" gridcol="3" gridrow="6" pageId="2" pageNumber="269">8.3</td>
<td id="6D595DD4FFC200343ADA1CDAFB9FFC7F" box="[948,1079,902,926]" gridcol="4" gridrow="6" pageId="2" pageNumber="269">8.3</td>
<td id="6D595DD4FFC200343D321CDAFB75FC7F" box="[1116,1245,902,926]" gridcol="5" gridrow="6" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1CDAFA2CFC7F" box="[1283,1412,902,926]" gridcol="6" gridrow="6" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81CF2FA2CFC27" box="[166,1412,942,966]" gridrow="7" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81CF2FE1AFC27" box="[166,434,942,966]" gridcol="0" gridrow="7" pageId="2" pageNumber="269">Midbody circumference</th>
<td id="6D595DD4FFC20034388C1CF2FDFAFC27" box="[482,594,942,966]" gridcol="1" gridrow="7" pageId="2" pageNumber="269">12</td>
<td id="6D595DD4FFC200343B191CF2FCA1FC27" box="[631,777,942,966]" gridcol="2" gridrow="7" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1CF2FC2FFC27" box="[818,903,942,966]" gridcol="3" gridrow="7" pageId="2" pageNumber="269">26</td>
<td id="6D595DD4FFC200343ADA1CF2FB9FFC27" box="[948,1079,942,966]" gridcol="4" gridrow="7" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343D321CF2FB75FC27" box="[1116,1245,942,966]" gridcol="5" gridrow="7" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1CF2FA2CFC27" box="[1283,1412,942,966]" gridcol="6" gridrow="7" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81C8AFA2CFC0E" box="[166,1412,982,1007]" gridrow="8" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81C8AFE1AFC0E" box="[166,434,982,1007]" gridcol="0" gridrow="8" pageId="2" pageNumber="269">Nostril-nostril</th>
<td id="6D595DD4FFC20034388C1C8AFDFAFC0E" box="[482,594,982,1007]" gridcol="1" gridrow="8" pageId="2" pageNumber="269">3.5</td>
<td id="6D595DD4FFC200343B191C8AFCA1FC0E" box="[631,777,982,1007]" gridcol="2" gridrow="8" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1C8AFC2FFC0E" box="[818,903,982,1007]" gridcol="3" gridrow="8" pageId="2" pageNumber="269">3.2</td>
<td id="6D595DD4FFC200343ADA1C8AFB9FFC0E" box="[948,1079,982,1007]" gridcol="4" gridrow="8" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343D321C8AFB75FC0E" box="[1116,1245,982,1007]" gridcol="5" gridrow="8" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1C8AFA2CFC0E" box="[1283,1412,982,1007]" gridcol="6" gridrow="8" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81CA3FA2CFBF6" box="[166,1412,1023,1047]" gridrow="9" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81CA3FE1AFBF6" box="[166,434,1023,1047]" gridcol="0" gridrow="9" pageId="2" pageNumber="269">Eye-eye</th>
<td id="6D595DD4FFC20034388C1CA3FDFAFBF6" box="[482,594,1023,1047]" gridcol="1" gridrow="9" pageId="2" pageNumber="269">5.1</td>
<td id="6D595DD4FFC200343B191CA3FCA1FBF6" box="[631,777,1023,1047]" gridcol="2" gridrow="9" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1CA3FC2FFBF6" box="[818,903,1023,1047]" gridcol="3" gridrow="9" pageId="2" pageNumber="269">4.6</td>
<td id="6D595DD4FFC200343ADA1CA3FB9FFBF6" box="[948,1079,1023,1047]" gridcol="4" gridrow="9" pageId="2" pageNumber="269">3.6</td>
<td id="6D595DD4FFC200343D321CA3FB75FBF6" box="[1116,1245,1023,1047]" gridcol="5" gridrow="9" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1CA3FA2CFBF6" box="[1283,1412,1023,1047]" gridcol="6" gridrow="9" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81B7BFA2CFBA1" box="[166,1412,1063,1088]" gridrow="10" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81B7BFE1AFBA1" box="[166,434,1063,1088]" gridcol="0" gridrow="10" pageId="2" pageNumber="269">Eye-snout tip</th>
<td id="6D595DD4FFC20034388C1B7BFDFAFBA1" box="[482,594,1063,1088]" gridcol="1" gridrow="10" pageId="2" pageNumber="269">4.3</td>
<td id="6D595DD4FFC200343B191B7BFCA1FBA1" box="[631,777,1063,1088]" gridcol="2" gridrow="10" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1B7BFC2FFBA1" box="[818,903,1063,1088]" gridcol="3" gridrow="10" pageId="2" pageNumber="269">4.4</td>
<td id="6D595DD4FFC200343ADA1B7BFB9FFBA1" box="[948,1079,1063,1088]" gridcol="4" gridrow="10" pageId="2" pageNumber="269">5.3</td>
<td id="6D595DD4FFC200343D321B7BFB75FBA1" box="[1116,1245,1063,1088]" gridcol="5" gridrow="10" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1B7BFA2CFBA1" box="[1283,1412,1063,1088]" gridcol="6" gridrow="10" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81B13FA2CFB89" box="[166,1412,1103,1128]" gridrow="11" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81B13FE1AFB89" box="[166,434,1103,1128]" gridcol="0" gridrow="11" pageId="2" pageNumber="269">Eye diameter</th>
<td id="6D595DD4FFC20034388C1B13FDFAFB89" box="[482,594,1103,1128]" gridcol="1" gridrow="11" pageId="2" pageNumber="269">1.4</td>
<td id="6D595DD4FFC200343B191B13FCA1FB89" box="[631,777,1103,1128]" gridcol="2" gridrow="11" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1B13FC2FFB89" box="[818,903,1103,1128]" gridcol="3" gridrow="11" pageId="2" pageNumber="269">1.4</td>
<td id="6D595DD4FFC200343ADA1B13FB9FFB89" box="[948,1079,1103,1128]" gridcol="4" gridrow="11" pageId="2" pageNumber="269">1.4</td>
<td id="6D595DD4FFC200343D321B13FB75FB89" box="[1116,1245,1103,1128]" gridcol="5" gridrow="11" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1B13FA2CFB89" box="[1283,1412,1103,1128]" gridcol="6" gridrow="11" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81B2BFA2CFB71" box="[166,1412,1143,1168]" gridrow="12" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81B2BFE1AFB71" box="[166,434,1143,1168]" gridcol="0" gridrow="12" pageId="2" pageNumber="269">Maximum width of rostral</th>
<td id="6D595DD4FFC20034388C1B2BFDFAFB71" box="[482,594,1143,1168]" gridcol="1" gridrow="12" pageId="2" pageNumber="269">3.5</td>
<td id="6D595DD4FFC200343B191B2BFCA1FB71" box="[631,777,1143,1168]" gridcol="2" gridrow="12" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1B2BFC2FFB71" box="[818,903,1143,1168]" gridcol="3" gridrow="12" pageId="2" pageNumber="269">3.5</td>
<td id="6D595DD4FFC200343ADA1B2BFB9FFB71" box="[948,1079,1143,1168]" gridcol="4" gridrow="12" pageId="2" pageNumber="269">3.1</td>
<td id="6D595DD4FFC200343D321B2BFB75FB71" box="[1116,1245,1143,1168]" gridcol="5" gridrow="12" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1B2BFA2CFB71" box="[1283,1412,1143,1168]" gridcol="6" gridrow="12" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81BFCFA2CFB59" box="[166,1412,1184,1208]" gridrow="13" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81BFCFE1AFB59" box="[166,434,1184,1208]" gridcol="0" gridrow="13" pageId="2" pageNumber="269">Max. length frontal shield</th>
<td id="6D595DD4FFC20034388C1BFCFDFAFB59" box="[482,594,1184,1208]" gridcol="1" gridrow="13" pageId="2" pageNumber="269">2.9</td>
<td id="6D595DD4FFC200343B191BFCFCA1FB59" box="[631,777,1184,1208]" gridcol="2" gridrow="13" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1BFCFC2FFB59" box="[818,903,1184,1208]" gridcol="3" gridrow="13" pageId="2" pageNumber="269">3.1</td>
<td id="6D595DD4FFC200343ADA1BFCFB9FFB59" box="[948,1079,1184,1208]" gridcol="4" gridrow="13" pageId="2" pageNumber="269">3.2</td>
<td id="6D595DD4FFC200343D321BFCFB75FB59" box="[1116,1245,1184,1208]" gridcol="5" gridrow="13" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1BFCFA2CFB59" box="[1283,1412,1184,1208]" gridcol="6" gridrow="13" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81B94FA2CFB01" box="[166,1412,1224,1248]" gridrow="14" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81B94FE1AFB01" box="[166,434,1224,1248]" gridcol="0" gridrow="14" pageId="2" pageNumber="269">Max. width frontal shield</th>
<td id="6D595DD4FFC20034388C1B94FDFAFB01" box="[482,594,1224,1248]" gridcol="1" gridrow="14" pageId="2" pageNumber="269">3.5</td>
<td id="6D595DD4FFC200343B191B94FCA1FB01" box="[631,777,1224,1248]" gridcol="2" gridrow="14" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1B94FC2FFB01" box="[818,903,1224,1248]" gridcol="3" gridrow="14" pageId="2" pageNumber="269">3.2</td>
<td id="6D595DD4FFC200343ADA1B94FB9FFB01" box="[948,1079,1224,1248]" gridcol="4" gridrow="14" pageId="2" pageNumber="269">3.5</td>
<td id="6D595DD4FFC200343D321B94FB75FB01" box="[1116,1245,1224,1248]" gridcol="5" gridrow="14" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1B94FA2CFB01" box="[1283,1412,1224,1248]" gridcol="6" gridrow="14" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81BACFA2CFAE9" box="[166,1412,1264,1288]" gridrow="15" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81BACFE1AFAE9" box="[166,434,1264,1288]" gridcol="0" gridrow="15" pageId="2" pageNumber="269">Max. length parietal</th>
<td id="6D595DD4FFC20034388C1BACFDFAFAE9" box="[482,594,1264,1288]" gridcol="1" gridrow="15" pageId="2" pageNumber="269">6.0</td>
<td id="6D595DD4FFC200343B191BACFCA1FAE9" box="[631,777,1264,1288]" gridcol="2" gridrow="15" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343A5C1BACFC2FFAE9" box="[818,903,1264,1288]" gridcol="3" gridrow="15" pageId="2" pageNumber="269">5.5</td>
<td id="6D595DD4FFC200343ADA1BACFB9FFAE9" box="[948,1079,1264,1288]" gridcol="4" gridrow="15" pageId="2" pageNumber="269">6.6</td>
<td id="6D595DD4FFC200343D321BACFB75FAE9" box="[1116,1245,1264,1288]" gridcol="5" gridrow="15" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1BACFA2CFAE9" box="[1283,1412,1264,1288]" gridcol="6" gridrow="15" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81A44FA2CFAD0" box="[166,1412,1304,1329]" gridrow="16" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81A44FE1AFAD0" box="[166,434,1304,1329]" gridcol="0" gridrow="16" pageId="2" pageNumber="269">Ventral scales</th>
<td id="6D595DD4FFC20034388C1A44FDFAFAD0" box="[482,594,1304,1329]" gridcol="1" gridrow="16" pageId="2" pageNumber="269">195</td>
<td id="6D595DD4FFC200343B191A44FCA1FAD0" box="[631,777,1304,1329]" gridcol="2" gridrow="16" pageId="2" pageNumber="269">205</td>
<td id="6D595DD4FFC200343A5C1A44FC2FFAD0" box="[818,903,1304,1329]" gridcol="3" gridrow="16" pageId="2" pageNumber="269">203</td>
<td id="6D595DD4FFC200343ADA1A44FB9FFAD0" box="[948,1079,1304,1329]" gridcol="4" gridrow="16" pageId="2" pageNumber="269">205</td>
<td id="6D595DD4FFC200343D321A44FB75FAD0" box="[1116,1245,1304,1329]" gridcol="5" gridrow="16" pageId="2" pageNumber="269">194</td>
<td id="6D595DD4FFC200343C6D1A44FA2CFAD0" box="[1283,1412,1304,1329]" gridcol="6" gridrow="16" pageId="2" pageNumber="269">192</td>
</tr>
<tr id="2E8834A8FFC2003439C81A1DFA2CFAB8" box="[166,1412,1345,1369]" gridrow="17" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81A1DFE1AFAB8" box="[166,434,1345,1369]" gridcol="0" gridrow="17" pageId="2" pageNumber="269">Subcaudal scale pairs</th>
<td id="6D595DD4FFC20034388C1A1DFDFAFAB8" box="[482,594,1345,1369]" gridcol="1" gridrow="17" pageId="2" pageNumber="269">50*</td>
<td id="6D595DD4FFC200343B191A1DFCA1FAB8" box="[631,777,1345,1369]" gridcol="2" gridrow="17" pageId="2" pageNumber="269">78</td>
<td id="6D595DD4FFC200343A5C1A1DFC2FFAB8" box="[818,903,1345,1369]" gridcol="3" gridrow="17" pageId="2" pageNumber="269">76</td>
<td id="6D595DD4FFC200343ADA1A1DFB9FFAB8" box="[948,1079,1345,1369]" gridcol="4" gridrow="17" pageId="2" pageNumber="269">77</td>
<td id="6D595DD4FFC200343D321A1DFB75FAB8" box="[1116,1245,1345,1369]" gridcol="5" gridrow="17" pageId="2" pageNumber="269">68</td>
<td id="6D595DD4FFC200343C6D1A1DFA2CFAB8" box="[1283,1412,1345,1369]" gridcol="6" gridrow="17" pageId="2" pageNumber="269">69</td>
</tr>
<tr id="2E8834A8FFC2003439C81A35FA2CFA60" box="[166,1412,1385,1409]" gridrow="18" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81A35FE1AFA60" box="[166,434,1385,1409]" gridcol="0" gridrow="18" pageId="2" pageNumber="269">Midbody scale rows</th>
<td id="6D595DD4FFC20034388C1A35FDFAFA60" box="[482,594,1385,1409]" gridcol="1" gridrow="18" pageId="2" pageNumber="269">17</td>
<td id="6D595DD4FFC200343B191A35FCA1FA60" box="[631,777,1385,1409]" gridcol="2" gridrow="18" pageId="2" pageNumber="269">17</td>
<td id="6D595DD4FFC200343A5C1A35FC2FFA60" box="[818,903,1385,1409]" gridcol="3" gridrow="18" pageId="2" pageNumber="269">17</td>
<td id="6D595DD4FFC200343ADA1A35FB9FFA60" box="[948,1079,1385,1409]" gridcol="4" gridrow="18" pageId="2" pageNumber="269">17</td>
<td id="6D595DD4FFC200343D321A35FB75FA60" box="[1116,1245,1385,1409]" gridcol="5" gridrow="18" pageId="2" pageNumber="269">17</td>
<td id="6D595DD4FFC200343C6D1A35FA2CFA60" box="[1283,1412,1385,1409]" gridcol="6" gridrow="18" pageId="2" pageNumber="269">17</td>
</tr>
<tr id="2E8834A8FFC2003439C81ACDFA2CFA48" box="[166,1412,1425,1449]" gridrow="19" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81ACDFE1AFA48" box="[166,434,1425,1449]" gridcol="0" gridrow="19" pageId="2" pageNumber="269">Supralabials</th>
<td id="6D595DD4FFC20034388C1ACDFDFAFA48" box="[482,594,1425,1449]" gridcol="1" gridrow="19" pageId="2" pageNumber="269">5,5</td>
<td id="6D595DD4FFC200343B191ACDFCA1FA48" box="[631,777,1425,1449]" gridcol="2" gridrow="19" pageId="2" pageNumber="269">5,5</td>
<td id="6D595DD4FFC200343A5C1ACDFC2FFA48" box="[818,903,1425,1449]" gridcol="3" gridrow="19" pageId="2" pageNumber="269">5,5</td>
<td id="6D595DD4FFC200343ADA1ACDFB9FFA48" box="[948,1079,1425,1449]" gridcol="4" gridrow="19" pageId="2" pageNumber="269">5,5</td>
<td id="6D595DD4FFC200343D321ACDFB75FA48" box="[1116,1245,1425,1449]" gridcol="5" gridrow="19" pageId="2" pageNumber="269">5,5</td>
<td id="6D595DD4FFC200343C6D1ACDFA2CFA48" box="[1283,1412,1425,1449]" gridcol="6" gridrow="19" pageId="2" pageNumber="269">5,5</td>
</tr>
<tr id="2E8834A8FFC2003439C81AE5FA2CFA33" box="[166,1412,1465,1490]" gridrow="20" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81AE5FE1AFA33" box="[166,434,1465,1490]" gridcol="0" gridrow="20" pageId="2" pageNumber="269">Temporals</th>
<td id="6D595DD4FFC20034388C1AE5FDFAFA33" box="[482,594,1465,1490]" gridcol="1" gridrow="20" pageId="2" pageNumber="269">1+2, 1+1</td>
<td id="6D595DD4FFC200343B191AE5FCA1FA33" box="[631,777,1465,1490]" gridcol="2" gridrow="20" pageId="2" pageNumber="269">1+1, _</td>
<td id="6D595DD4FFC200343A5C1AE5FC2FFA33" box="[818,903,1465,1490]" gridcol="3" gridrow="20" pageId="2" pageNumber="269">1+1,1+1</td>
<td id="6D595DD4FFC200343ADA1AE5FB9FFA33" box="[948,1079,1465,1490]" gridcol="4" gridrow="20" pageId="2" pageNumber="269">1+1, 1+1</td>
<td id="6D595DD4FFC200343D321AE5FB75FA33" box="[1116,1245,1465,1490]" gridcol="5" gridrow="20" pageId="2" pageNumber="269">_,1+1</td>
<td id="6D595DD4FFC200343C6D1AE5FA2CFA33" box="[1283,1412,1465,1490]" gridcol="6" gridrow="20" pageId="2" pageNumber="269">1+2,1+1</td>
</tr>
<tr id="2E8834A8FFC2003439C81ABEFA2CF9FF" box="[166,1412,1506,1566]" gridrow="21" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81ABEFE1AF9FF" box="[166,434,1506,1566]" gridcol="0" gridrow="21" pageId="2" pageNumber="269">Small scales overlapped by anals</th>
<td id="6D595DD4FFC20034388C1ABEFDFAF9FF" box="[482,594,1506,1566]" gridcol="1" gridrow="21" pageId="2" pageNumber="269">6,7</td>
<td id="6D595DD4FFC200343B191ABEFCA1F9FF" box="[631,777,1506,1566]" gridcol="2" gridrow="21" pageId="2" pageNumber="269">5,6</td>
<td id="6D595DD4FFC200343A5C1ABEFC2FF9FF" box="[818,903,1506,1566]" gridcol="3" gridrow="21" pageId="2" pageNumber="269">3,4</td>
<td id="6D595DD4FFC200343ADA1ABEFB9FF9FF" box="[948,1079,1506,1566]" gridcol="4" gridrow="21" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343D321ABEFB75F9FF" box="[1116,1245,1506,1566]" gridcol="5" gridrow="21" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1ABEFA2CF9FF" box="[1283,1412,1506,1566]" gridcol="6" gridrow="21" pageId="2" pageNumber="269">_</td>
</tr>
<tr id="2E8834A8FFC2003439C81972FA2CF9A7" box="[166,1412,1582,1606]" gridrow="22" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C81972FE1AF9A7" box="[166,434,1582,1606]" gridcol="0" gridrow="22" pageId="2" pageNumber="269">Dark Vs on body</th>
<td id="6D595DD4FFC20034388C1972FDFAF9A7" box="[482,594,1582,1606]" gridcol="1" gridrow="22" pageId="2" pageNumber="269">37,38</td>
<td id="6D595DD4FFC200343B191972FCA1F9A7" box="[631,777,1582,1606]" gridcol="2" gridrow="22" pageId="2" pageNumber="269">46,47</td>
<td id="6D595DD4FFC200343A5C1972FC2FF9A7" box="[818,903,1582,1606]" gridcol="3" gridrow="22" pageId="2" pageNumber="269">44,43</td>
<td id="6D595DD4FFC200343ADA1972FB9FF9A7" box="[948,1079,1582,1606]" gridcol="4" gridrow="22" pageId="2" pageNumber="269">ca. 44–46</td>
<td id="6D595DD4FFC200343D321972FB75F9A7" box="[1116,1245,1582,1606]" gridcol="5" gridrow="22" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D1972FA2CF9A7" box="[1283,1412,1582,1606]" gridcol="6" gridrow="22" pageId="2" pageNumber="269">ca. 41</td>
</tr>
<tr id="2E8834A8FFC2003439C8190AFA2CF98F" box="[166,1412,1622,1646]" gridrow="23" pageId="2" pageNumber="269">
<th id="6D595DD4FFC2003439C8190AFE1AF98F" box="[166,434,1622,1646]" gridcol="0" gridrow="23" pageId="2" pageNumber="269">Dark bands on tail</th>
<td id="6D595DD4FFC20034388C190AFDFAF98F" box="[482,594,1622,1646]" gridcol="1" gridrow="23" pageId="2" pageNumber="269">12,12</td>
<td id="6D595DD4FFC200343B19190AFCA1F98F" box="[631,777,1622,1646]" gridcol="2" gridrow="23" pageId="2" pageNumber="269">21or 22, 20</td>
<td id="6D595DD4FFC200343A5C190AFC2FF98F" box="[818,903,1622,1646]" gridcol="3" gridrow="23" pageId="2" pageNumber="269">16,17</td>
<td id="6D595DD4FFC200343ADA190AFB9FF98F" box="[948,1079,1622,1646]" gridcol="4" gridrow="23" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343D32190AFB75F98F" box="[1116,1245,1622,1646]" gridcol="5" gridrow="23" pageId="2" pageNumber="269">_</td>
<td id="6D595DD4FFC200343C6D190AFA2CF98F" box="[1283,1412,1622,1646]" gridcol="6" gridrow="23" pageId="2" pageNumber="269">_</td>
</tr>
</table>
</paragraph>
<paragraph id="900736EAFFC2FFC939A919C2FE8DF88D" blockId="2.[151,1436,1694,2008]" pageId="2" pageNumber="269">
Body subcylindrical, slightly flattened ventrally (1.3 times as wide (
<quantity id="57409B0FFFC2FFC93AA419C2FB9AF959" box="[970,1074,1694,1720]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="269" unit="mm" value="10.9">10.9 mm</quantity>
) as high (
<quantity id="57409B0FFFC2FFC93DCD19C2FB54F959" box="[1187,1276,1694,1721]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="269" unit="mm" value="8.5">8.5 mm</quantity>
) at midbody), widest at midbody, tapering posteriorly and more substantially anteriorly. Head broader than tall, barely wider than anterior of body. In dorsal view head slightly ovate, sides very slightly convex, gently converging anteriorly; front of snout not pointed, truncated. In lateral view, head tapers gently from back to prefrontal, more strongly tapered in front of prefrontal. Paired shields on top of head (= parietals only) abutting along midline rather than imbricate/ overlapping.
</paragraph>
<paragraph id="900736EAFFC2FFC839A9182AFD13FEFC" blockId="2.[151,1436,1694,2008]" lastBlockId="3.[151,1437,151,285]" lastPageId="3" lastPageNumber="270" pageId="2" pageNumber="269">In dorsal view, rostral much broader than long, substantially shorter than distance between it and prefrontal; projects beyond tip of lower jaw; ventrally with transverse concavity, notched (C-shaped) at margin of mouth. Frontal subpentagonal, lateral edges anteriorly diverging; substantially shorter and smaller than each parietal, shorter than distance between it and snout tip. Internasal single, smaller than single prefrontal. Nasals subrectangular, larger than trapezoid loreal. Single supraocular, two preoculars and two postoculars each side; supraocular largest of these scales, longer than wide. Upper pre- and post-oculars much larger than lower pre- and postoculars; lower pre- and postocular not in contact due to intervening SL3.</paragraph>
<caption id="C4C76662FFC3FFC839F91BB7FCFBFAC6" ID-DOI="http://doi.org/10.5281/zenodo.4413799" ID-Zenodo-Dep="4413799" httpUri="https://zenodo.org/record/4413799/files/figure.png" pageId="3" pageNumber="270" startId="3.[151,250,1259,1283]" targetBox="[151,1436,309,1236]" targetPageId="3">
<paragraph id="900736EAFFC3FFC839F91BB7FCFBFAC6" blockId="3.[151,1436,1259,1319]" pageId="3" pageNumber="270">
<emphasis id="A2CCEAF8FFC3FFC839F91BB7FEA7FAE2" bold="true" box="[151,271,1259,1283]" pageId="3" pageNumber="270">FIGURE 1</emphasis>
. 
<taxonomicName id="57B84D69FFC3FFC838701BB7FD9AFAE2" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[286,562,1259,1283]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="270" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC3FFC838701BB7FD9AFAE2" box="[286,562,1259,1283]" italics="true" pageId="3" pageNumber="270">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC3FFC83B541BB7FD25FAE2" bold="true" box="[570,653,1259,1283]" pageId="3" pageNumber="270">
<taxonomicNameLabel id="B9FF5783FFC3FFC83B541BB7FD25FAE2" box="[570,653,1259,1283]" pageId="3" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (BNHS 3650). 
<emphasis id="A2CCEAF8FFC3FFC83AF21BB7FC06FAE2" bold="true" box="[924,942,1259,1283]" pageId="3" pageNumber="270">A</emphasis>
. full body dorsal, and 
<emphasis id="A2CCEAF8FFC3FFC83DF01BB7FB07FAE2" bold="true" box="[1182,1199,1259,1283]" pageId="3" pageNumber="270">B</emphasis>
. ventral view, 
<emphasis id="A2CCEAF8FFC3FFC83C251BB7FAF5FAE2" bold="true" box="[1355,1373,1259,1283]" pageId="3" pageNumber="270">C</emphasis>
. head dorsal, 
<emphasis id="A2CCEAF8FFC3FFC8398C1A53FF5CFAC6" bold="true" box="[226,244,1295,1319]" pageId="3" pageNumber="270">D</emphasis>
. ventral, and 
<emphasis id="A2CCEAF8FFC3FFC838111A53FE1AFAC6" bold="true" box="[383,434,1295,1319]" pageId="3" pageNumber="270">E– F</emphasis>
. lateral views of head. Scale bars 10mm.
</paragraph>
</caption>
<paragraph id="900736EAFFC3FFC839A91A09FE29F936" blockId="3.[151,1437,1365,2040]" pageId="3" pageNumber="270">External naris on dorsolateral-facing surface, seen as C-shaped slit in front of flap, in broad, shallow depression slightly posterodorsal of centre of nasal. External naris visible anteriorly as well as dorsally and primarily laterally. Five supralabials (SLs) on each side; SL5 largest, longest; SL1 smallest and shortest; SL1 contacts rostral and nasal (and, perhaps, loreal), SL2 contacts loreal (and, perhaps, nasal) and lower preocular, SL3 contacts lower pre- and postoculars and eye, SL4 contacts lower postocular and anterior temporal, SL 5 contacts anterior and posterior lower temporal. Eye contacts third supralabial in addition to supra-, pre- and postoculars. Eye lateral; pupil subcircular. Temporals 1+2 on left and 1+1 on right. Parietals longer than wide, larger than other head scales. Midline interparietal suture more than three quarters length of each parietal; parietals barely projecting posterior to suture; each parietal approximately twice as long as frontal; each parietal contacts frontal, supraocular, upper postocular, anterior and upper posterior temporals, and three (right) or four (left) other scales. Posterior margin of each parietal somewhat scalloped.</paragraph>
<paragraph id="900736EAFFC3FFC839A919BDFE83F84E" blockId="3.[151,1437,1365,2040]" pageId="3" pageNumber="270">Mental small, subtriangular, wider than long. Infralabials 7,7; first pair in midline contact; second pair smallest; fourth largest and longest. Two pairs of genials; first pair largest, longer than broad, in long midline contact; second pair not in contact, separated by one small, midline scale. Anterior genials contact infralabials (ILs) 1–4; posterior genials contact IL 4. Anteriormost ventral separated from each posterior genial by four scales, separated from each posteriormost infralabial by six scales. Teeth largely obscured by gingivae, but estimated as 8 marginals on each side of upper jaw.</paragraph>
<paragraph id="900736EAFFC3FFCF39A918E5FEC7FEFC" blockId="3.[151,1437,1365,2040]" lastBlockId="4.[151,1437,151,1077]" lastPageId="4" lastPageNumber="271" pageId="3" pageNumber="270">Macroscopically and under low magnification (using a light dissecting microscope) body scales smooth. No keels or apical pits. Vertebral scale row not different from adjacent dorsal scale row. Exposed parts of dorsal body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions and for lowest dorsal scales level with first few ventrals which are lower, more elongate. Dorsal scales closer to vent slightly smaller than at midbody; those at anterior of body smallest. Dorsal scale rows 17 at level of first ventral, maintained to vent.</paragraph>
<paragraph id="900736EAFFC4FFCF39A91E7BFF5BFE30" blockId="4.[151,1437,151,1077]" pageId="4" pageNumber="271">Posteriormost ventral with slight posteromedial extension. Anals paired (right overlapping left), each slightly larger than posteriormost ventral. Each anal overlaps 6,7 small scales in addition to first subcaudal. Tail subcircular in cross section, slightly flattened ventrally. Dorsal tail scales more heterogenous in size than on body, without clear patterns. Subcaudals paired/divided throughout, terminal scale (scute) conical, approximately as wide as long, pointed.</paragraph>
<paragraph id="900736EAFFC4FFCF39A91E87FB17FC60" blockId="4.[151,1437,151,1077]" pageId="4" pageNumber="271">Macroscopically bicoloured, darker black-brown above, whitish ochre below. Body scales iridescent. Alternating (at least asymmetric) longer dark and shorter pale, typically V-shaped (when viewed laterally) or to trapezoid markings (when viewed from above) along body and tail. On body 37, 38 dark ‘bands’, on tail 12,12. Ventrally projecting dark trapezoid shapes (from level with approximately 24th ventral) extend onto lateral edges of ventrals leaving broad, entirely pale band along most of venter. Anteriormost dark trapezoid (less than one head length behind head) with gently rounded ventral margin, extends onto third dorsal scale row. Whitish inverted V-shaped markings (“Vs”) extend dorsally between each dark trapezoid up to the sixth (rarely seventh) dorsal scale row. Along the first quarter of body the border between pale-dark markings wavy, rapidly becoming increasingly sharply zigzag towards the posterior. Ventrally projecting dark marks extend as far as midventer on tail from behind vent, forming alternating (dark and pale) transverse bands; where pale bands approach midventer (only on anterior half of tail, where they are longer) they remain separated by narrow midventral dark line. Whitish ventrals with fine pale grey speckles on posterior half of body. Anals each with pale grey blotches as well as speckles.</paragraph>
<paragraph id="900736EAFFC4FFCF39A91CD7FA90FBD4" blockId="4.[151,1437,151,1077]" pageId="4" pageNumber="271">Head dark dorsally, brown-grey, slightly paler than body dorsum, generally uniform, rostral slightly paler. Dark of dorsum extends onto upper margins of SLs, SLs otherwise pale (whitish, as body venter) with very few (more on SL1) pale grey specks. Dark scales on dorsum (including head) are mottled under low power dissecting microscope. First pair of ILs and anterior genials with pale grey speckles (fewer on mental). Underside of head otherwise pale, whitish. Inside of mouth pale with two dark spots on either side of the anterior of pterygoid part of palate.</paragraph>
<caption id="C4C76662FFC4FFCF39F918F7FC6DF822" ID-DOI="http://doi.org/10.5281/zenodo.4413801" ID-Zenodo-Dep="4413801" box="[151,965,1963,1987]" httpUri="https://zenodo.org/record/4413801/files/figure.png" pageId="4" pageNumber="271" startId="4.[151,250,1963,1987]" targetBox="[167,1420,1101,1938]" targetPageId="4">
<paragraph id="900736EAFFC4FFCF39F918F7FC6DF822" blockId="4.[151,965,1963,1987]" box="[151,965,1963,1987]" pageId="4" pageNumber="271">
<emphasis id="A2CCEAF8FFC4FFCF39F918F7FEA5F822" bold="true" box="[151,269,1963,1987]" pageId="4" pageNumber="271">FIGURE 2</emphasis>
. 
<taxonomicName id="57B84D69FFC4FFCF387718F7FD83F822" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[281,555,1963,1987]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC4FFCF387718F7FD83F822" box="[281,555,1963,1987]" italics="true" pageId="4" pageNumber="271">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC4FFCF3B5F18F7FD2AF822" bold="true" box="[561,642,1963,1987]" pageId="4" pageNumber="271">
<taxonomicNameLabel id="B9FF5783FFC4FFCF3B5F18F7FD2AF822" box="[561,642,1963,1987]" pageId="4" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (BNHS 3650) in life.
</paragraph>
</caption>
<caption id="C4C76662FFC5FFCE39F91BCCFD0FFB10" ID-DOI="http://doi.org/10.5281/zenodo.4413805" ID-Zenodo-Dep="4413805" httpUri="https://zenodo.org/record/4413805/files/figure.png" pageId="5" pageNumber="272" startId="5.[151,250,1168,1192]" targetBox="[176,1411,189,1138]" targetPageId="5">
<paragraph id="900736EAFFC5FFCE39F91BCCFD0FFB10" blockId="5.[151,1436,1168,1265]" pageId="5" pageNumber="272">
<emphasis id="A2CCEAF8FFC5FFCE39F91BCCFEA6FB48" bold="true" box="[151,270,1168,1193]" pageId="5" pageNumber="272">FIGURE 3</emphasis>
. Map showing distribution of species of the genus 
<emphasis id="A2CCEAF8FFC5FFCE3A721BCDFC3CFB48" box="[796,916,1169,1193]" italics="true" pageId="5" pageNumber="272">
<taxonomicName id="57B84D69FFC5FFCE3A721BCDFC38FB48" authorityName="Giri, Gower, Das, Lalremsanga, Lalronunga, Captain &amp; Deepak" authorityYear="2019" box="[796,912,1169,1193]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="272" phylum="Chordata" rank="genus">Smithophis</taxonomicName>
.
</emphasis>
Examined comparative materials of 
<taxonomicName id="57B84D69FFC5FFCE3C631BCDFAD8FB48" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[1293,1392,1169,1193]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="272" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="A2CCEAF8FFC5FFCE3C631BCDFAD8FB48" box="[1293,1392,1169,1193]" italics="true" pageId="5" pageNumber="272">S. bicolor</emphasis>
</taxonomicName>
(including the type), are from the Khasia (or Khassia or Khasi) Hills, Shillong, Tura or Aizawl, or from unspecified localities in Northeast India. See Appendix 1 for locality details.
</paragraph>
</caption>
<paragraph id="900736EAFFC5FFCE39A91A43FB8AFA0C" blockId="5.[151,1437,1311,1949]" pageId="5" pageNumber="272">
<emphasis id="A2CCEAF8FFC5FFCE39A91A43FDBBFAD8" bold="true" box="[199,531,1311,1337]" pageId="5" pageNumber="272">
Variation among 
<typeStatus id="4F038848FFC5FFCE38F81A43FDA6FAD8" box="[406,526,1311,1337]" pageId="5" pageNumber="272" type="paratype">paratypes</typeStatus>
.
</emphasis>
<typeStatus id="4F038848FFC5FFCE3B771A43FD2EFAD8" box="[537,646,1311,1337]" pageId="5" pageNumber="272" type="paratype">Paratypes</typeStatus>
do not differ substantially from the 
<typeStatus id="4F038848FFC5FFCE3D791A43FBD1FAD8" box="[1047,1145,1311,1337]" pageId="5" pageNumber="272" type="holotype">holotype</typeStatus>
(
<figureCitation id="08832A6FFFC5FFCE3DE61A43FB46FAD8" box="[1160,1262,1311,1337]" captionStart="FIGURE 4" captionStartId="7.[151,250,1961,1985]" captionTargetBox="[184,1404,207,1930]" captionTargetId="figure-19@7.[175,1412,199,1938]" captionTargetPageId="7" captionText="FIGURE 4. Smithophis arunachalensis sp. nov., paratypes in dorsal views A. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and B. (ZSIK 23875). Scale bar 10mm." figureDoi="http://doi.org/10.5281/zenodo.4413807" httpUri="https://zenodo.org/record/4413807/files/figure.png" pageId="5" pageNumber="272">Fig. 4a,b</figureCitation>
, 
<tableCitation id="DD3A0351FFC5FFCE3D961A43FAE1FAD8" box="[1272,1353,1311,1337]" captionStart="TABLE 1" captionStartId="2.[151,239,440,464]" captionTargetPageId="2" captionText="TABLE 1. Meristic and morphometric (in millimetres) data for examined specimens of Smithophis arunachalensis sp. nov. Head length = distance between snout tip and back of lower jaw; bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. The format ‘X-Y’ is used to denote the shortest distance between features X and Y." httpUri="http://table.plazi.org/id/C4C76662FFC2FFC939F91EE4FD74FDDD" pageId="5" pageNumber="272" tableUuid="C4C76662FFC2FFC939F91EE4FD74FDDD">Table 1</tableCitation>
). 
<typeStatus id="4F038848FFC5FFCE3C0F1A43FF7AFABC" pageId="5" pageNumber="272" type="paratype">Paratypes</typeStatus>
are generally in moderate condition, with following exceptions: the front of head of BMNH 
<date id="E406102AFFC5FFCE3D8C1A1FFACEFABF" box="[1250,1382,1347,1374]" pageId="5" pageNumber="272" value="1935-10-12">1935.10.12.</date>
10 is not attached to its body, and is kept in a small container within the same jar; and the head and approximately four head lengths behind head of ZSIK 23875 is unnaturally flattened, it also has a constriction behind the head possibly due to the knot for tag. Both BMNH 
<date id="E406102AFFC5FFCE3B5B1AF3FD11FA2B" box="[565,697,1455,1482]" pageId="5" pageNumber="272" value="1935-10-12">1935.10.12.</date>
10 and ZSIK 23875 have darker Vs on body and tail. ZSIK 23875 has 6 IL on either side, otherwise scalation is very similar to that of the 
<typeStatus id="4F038848FFC5FFCE3AD71A8FFBB7FA0C" box="[953,1055,1491,1517]" pageId="5" pageNumber="272" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="900736EAFFC5FFCE39A91AABFEF7F8EC" blockId="5.[151,1437,1311,1949]" pageId="5" pageNumber="272">
<emphasis id="A2CCEAF8FFC5FFCE39A91AABFEC3F9F0" bold="true" box="[199,363,1527,1553]" pageId="5" pageNumber="272">Colour in life.</emphasis>
Based on 
<typeStatus id="4F038848FFC5FFCE38B11AABFDE9F9F0" box="[479,577,1527,1553]" pageId="5" pageNumber="272" type="holotype">holotype</typeStatus>
(
<figureCitation id="08832A6FFFC5FFCE3B3E1AABFD3CF9F0" box="[592,660,1527,1553]" captionStart="FIGURE 2" captionStartId="4.[151,250,1963,1987]" captionTargetBox="[167,1420,1101,1938]" captionTargetId="figure-436@4.[167,1420,1101,1938]" captionTargetPageId="4" captionText="FIGURE 2. Smithophis arunachalensis sp. nov., holotype (BNHS 3650) in life." figureDoi="http://doi.org/10.5281/zenodo.4413801" httpUri="https://zenodo.org/record/4413801/files/figure.png" pageId="5" pageNumber="272">Fig. 2</figureCitation>
). Dorsum (head, body and tail) predominantly black above, including downward pointing triangles on flanks; venter and upward pointing triangles of body and tail bright yellow. Most head scales uniformly black; rostral predominantly black. Supralabials black above, cream-yellow below; border between the two colours indistinct. Iris black. Mental and tip of first pair of chin shields black mottled. Dark parts of venter and underside of tail blackish. The venter of ZSIK was recorded as yellowish-white by 
<bibRefCitation id="F4294B1BFFC5FFCE3D8C19DBFF73F927" author="Sanyal, D. P. &amp; Gayen N. C." pageId="5" pageNumber="272" pagination="297 - 302" refId="ref8145" refString="Sanyal, D. P. &amp; Gayen N. C. (1985) Reptilia In: Director Zoological Survey of India (Ed.), Fauna of Namdapha: Arunachal Pradesh ' A Proposed Biosphere Reserve'. Records of the Zoological Survey of India, 82 (1 - 4), pp. 297 - 302." type="journal article" year="1985">Sanyal &amp; Gayen (1985</bibRefCitation>
, 
<bibRefCitation id="F4294B1BFFC5FFCE398919F7FE89F927" author="Sanyal, D. P. &amp; Gayen, N. C." box="[231,289,1707,1734]" pageId="5" pageNumber="272" pagination="247 - 284" refId="ref8206" refString="Sanyal, D. P. &amp; Gayen, N. C. (2006) State Fauna Series (Reptilia). In: Alfred, J. R. B. (Ed.), Fauna of Arunachal Pradesh. 13 (1). Zoological Survey of India, Kolkata, pp. 247 - 284." type="book chapter" year="2006">2006</bibRefCitation>
) but it is not clear whether that was in life or preservation. A photograph of an individual from Deban (Namdapha Tiger Reserve, Changlang District, 
<collectingRegion id="527CF808FFC5FFCE3BC61993FCD7F908" box="[680,895,1743,1769]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="272">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC5FFCE3AE71993FC6CF90B" box="[905,964,1743,1770]" name="India" pageId="5" pageNumber="272">India</collectingCountry>
) seen by one of us (A.C.) had a bright yellow venter in life.
</paragraph>
</subSubSection>
<subSubSection id="D8A26561FFC5FFC039A9184BFB2AFAF9" lastPageId="11" lastPageNumber="278" pageId="5" pageNumber="272" type="etymology">
<paragraph id="900736EAFFC5FFCE39A9184BFE97F87C" blockId="5.[151,1437,1311,1949]" pageId="5" pageNumber="272">
<emphasis id="A2CCEAF8FFC5FFCE39A9184BFEE6F8D0" bold="true" box="[199,334,1815,1841]" pageId="5" pageNumber="272">Etymology.</emphasis>
Named after 
<collectingRegion id="527CF808FFC5FFCE388F184BFD10F8D0" box="[481,696,1815,1841]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="272">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="E8AF767AFFC5FFCE3BAF184BFD55F8D0" box="[705,765,1815,1841]" name="India" pageId="5" pageNumber="272">India</collectingCountry>
, an Indian state that has yielded many notable herpetological discoveries. The 
<typeStatus id="4F038848FFC5FFCE38381867FE2EF8B4" box="[342,390,1851,1877]" pageId="5" pageNumber="272">type</typeStatus>
and all other currently known localities of the new species are in 
<collectingRegion id="527CF808FFC5FFCE3D031867FAEDF8B4" box="[1133,1349,1851,1877]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="272">Arunachal Pradesh</collectingRegion>
. This is the second snake to be named after the state, after 
<taxonomicName id="57B84D69FFC5FFCE3BBD183CFE93F87C" authority="Captain, Deepak, Pandit, Bhatt &amp; Athreya, 2019" authorityName="Captain, Deepak, Pandit, Bhatt &amp; Athreya" authorityYear="2019" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="272" phylum="Chordata" rank="species" species="arunachalensis">
<emphasis id="A2CCEAF8FFC5FFCE3BBD183CFBB2F898" box="[723,1050,1887,1913]" italics="true" pageId="5" pageNumber="272">Trimeresurus arunachalensis</emphasis>
<bibRefCitation id="F4294B1BFFC5FFCE3D4C1803FE93F87C" author="Captain, A. &amp; Deepak, V. &amp; Pandit, R. &amp; Bhatt, B. &amp; Athreya, R." pageId="5" pageNumber="272" pagination="111 - 122" refId="ref7241" refString="Captain, A., Deepak, V., Pandit, R., Bhatt, B. &amp; Athreya, R. (2019) A new species of pitviper (Serpentes: Viperidae: Trimeresurus Lacepede, 1804) from West Kameng District, Arunachal Pradesh, India. Russian Journal of Herpetology, 26 (2), 111 - 122. https: // doi. org / 10.30906 / 1026 - 2296 - 2019 - 26 - 2 - 111 - 122" type="journal article" year="2019">Captain, Deepak, Pandit, Bhatt &amp; Athreya, 2019</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="C4C76662FFC6FFCD39F91FCBFD54FF16" ID-Table-UUID="C4C76662FFC6FFCD39F91FCBFD54FF16" httpUri="http://table.plazi.org/id/C4C76662FFC6FFCD39F91FCBFD54FF16" pageId="6" pageNumber="273" targetBox="[151,1408,264,1907]" targetIsTable="true" targetPageId="6">
<paragraph id="900736EAFFC6FFCD39F91FCBFD54FF16" blockId="6.[151,1436,151,247]" pageId="6" pageNumber="273">
<emphasis id="A2CCEAF8FFC6FFCD39F91FCBFEA3FF4E" bold="true" box="[151,267,151,175]" pageId="6" pageNumber="273">TABLE 2.</emphasis>
Meristic and morphometric (in millimetres) data for examined specimens of 
<emphasis id="A2CCEAF8FFC6FFCD3D241FCBFAB1FF4E" box="[1098,1305,151,175]" italics="true" pageId="6" pageNumber="273">
<taxonomicName id="57B84D69FFC6FFCD3D241FCBFABEFF4E" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[1098,1302,151,175]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="273" phylum="Chordata" rank="species" species="bicolor">Smithophis bicolor</taxonomicName>
.
</emphasis>
Some bilateral structures given in left, right order; * indicates measure/count incomplete because end of tail missing. – indicates data not recorded. Locality details are provided in Appendix I.
</paragraph>
</caption>
<paragraph id="900736EAFFC6FFCD382B1E54FAC4F8CA" pageId="6" pageNumber="273">
<table id="E2B8C44AFFC6003439F91E54FA28F892" box="[151,1408,264,1907]" gridcols="12" gridrows="27" pageId="6" pageNumber="273">
<tr id="2E8834A8FFC6003439F91E54FA28FE89" box="[151,1408,264,360]" gridrow="0" pageId="6" pageNumber="273" rowspan-0="1">
<th id="6D595DD4FFC60034382B1E54FE09FE89" box="[325,417,264,360]" gridcol="1" gridrow="0" pageId="6" pageNumber="273">BMNH 1907.12. 16.10</th>
<th id="6D595DD4FFC6003438BB1E54FD92FE89" box="[469,570,264,360]" gridcol="2" gridrow="0" pageId="6" pageNumber="273">BMNH 1908.6. 23.5</th>
<th id="6D595DD4FFC600343B351E54FD6DFE89" box="[603,709,264,360]" gridcol="3" gridrow="0" pageId="6" pageNumber="273">BMNH 1911.11. 30.4</th>
<th id="6D595DD4FFC600343BBD1E54FCD9FE89" box="[723,881,264,360]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="6" pageNumber="273">BMNH 60.3.19. 1270</th>
<th id="6D595DD4FFC600343AE51E54FC5DFE89" box="[907,1013,264,360]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="6" pageNumber="273">BMNH 1907.12. 16.11</th>
<th id="6D595DD4FFC600343D621E54FBD7FE89" box="[1036,1151,264,360]" gridcol="8" gridrow="0" pageId="6" pageNumber="273">BMNH 72.4.17. 184</th>
<th id="6D595DD4FFC600343DE11E54FAA0FE89" box="[1167,1288,264,360]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="6" pageNumber="273">ZSI/ERS 2592</th>
<th id="6D595DD4FFC600343C751E54FA28FE89" box="[1307,1408,264,360]" gridcol="11" gridrow="0" pageId="6" pageNumber="273">ZSI/ERS 3052</th>
</tr>
<tr id="2E8834A8FFC6003439F91E24FA28FE71" box="[151,1408,376,400]" gridrow="1" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91E24FE8BFE71" box="[151,291,376,400]" gridcol="0" gridrow="1" pageId="6" pageNumber="273">Sex</th>
<td id="6D595DD4FFC60034382B1E24FE09FE71" box="[325,417,376,400]" gridcol="1" gridrow="1" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC6003438BB1E24FD92FE71" box="[469,570,376,400]" gridcol="2" gridrow="1" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343B351E24FD6DFE71" box="[603,709,376,400]" gridcol="3" gridrow="1" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343BBD1E24FC80FE71" box="[723,808,376,400]" gridcol="4" gridrow="1" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343AE51E24FC19FE71" box="[907,945,376,400]" gridcol="6" gridrow="1" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343D621E24FBD7FE71" box="[1036,1151,376,400]" gridcol="8" gridrow="1" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343DE11E24FAA0FE71" box="[1167,1288,376,400]" colspan="2" colspanRight="1" gridcol="9" gridrow="1" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343C751E24FA28FE71" box="[1307,1408,376,400]" gridcol="11" gridrow="1" pageId="6" pageNumber="273">M</td>
</tr>
<tr id="2E8834A8FFC6003439F91EFCFA28FE59" box="[151,1408,416,440]" gridrow="2" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91EFCFE8BFE59" box="[151,291,416,440]" gridcol="0" gridrow="2" pageId="6" pageNumber="273">TL</th>
<td id="6D595DD4FFC60034382B1EFCFE09FE59" box="[325,417,416,440]" gridcol="1" gridrow="2" pageId="6" pageNumber="273">58</td>
<td id="6D595DD4FFC6003438BB1EFCFD92FE59" box="[469,570,416,440]" gridcol="2" gridrow="2" pageId="6" pageNumber="273">142</td>
<td id="6D595DD4FFC600343B351EFCFD6DFE59" box="[603,709,416,440]" gridcol="3" gridrow="2" pageId="6" pageNumber="273">125</td>
<td id="6D595DD4FFC600343BBD1EFCFC80FE59" box="[723,808,416,440]" gridcol="4" gridrow="2" pageId="6" pageNumber="273">90</td>
<td id="6D595DD4FFC600343AE51EFCFC19FE59" box="[907,945,416,440]" gridcol="6" gridrow="2" pageId="6" pageNumber="273">117</td>
<td id="6D595DD4FFC600343D621EFCFBD7FE59" box="[1036,1151,416,440]" gridcol="8" gridrow="2" pageId="6" pageNumber="273">_</td>
<td id="6D595DD4FFC600343DE11EFCFAA0FE59" box="[1167,1288,416,440]" colspan="2" colspanRight="1" gridcol="9" gridrow="2" pageId="6" pageNumber="273">158</td>
<td id="6D595DD4FFC600343C751EFCFA28FE59" box="[1307,1408,416,440]" gridcol="11" gridrow="2" pageId="6" pageNumber="273">159</td>
</tr>
<tr id="2E8834A8FFC6003439F91E94FA28FE00" box="[151,1408,456,481]" gridrow="3" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91E94FE8BFE00" box="[151,291,456,481]" gridcol="0" gridrow="3" pageId="6" pageNumber="273">SVL</th>
<td id="6D595DD4FFC60034382B1E94FE09FE00" box="[325,417,456,481]" gridcol="1" gridrow="3" pageId="6" pageNumber="273">228</td>
<td id="6D595DD4FFC6003438BB1E94FD92FE00" box="[469,570,456,481]" gridcol="2" gridrow="3" pageId="6" pageNumber="273">521</td>
<td id="6D595DD4FFC600343B351E94FD6DFE00" box="[603,709,456,481]" gridcol="3" gridrow="3" pageId="6" pageNumber="273">463</td>
<td id="6D595DD4FFC600343BBD1E94FC80FE00" box="[723,808,456,481]" gridcol="4" gridrow="3" pageId="6" pageNumber="273">321</td>
<td id="6D595DD4FFC600343AE51E94FC19FE00" box="[907,945,456,481]" gridcol="6" gridrow="3" pageId="6" pageNumber="273">473</td>
<td id="6D595DD4FFC600343D621E94FBD7FE00" box="[1036,1151,456,481]" gridcol="8" gridrow="3" pageId="6" pageNumber="273">496</td>
<td id="6D595DD4FFC600343DE11E94FAA0FE00" box="[1167,1288,456,481]" colspan="2" colspanRight="1" gridcol="9" gridrow="3" pageId="6" pageNumber="273">550</td>
<td id="6D595DD4FFC600343C751E94FA28FE00" box="[1307,1408,456,481]" gridcol="11" gridrow="3" pageId="6" pageNumber="273">593</td>
</tr>
<tr id="2E8834A8FFC6003439F91EADFA28FDE8" box="[151,1408,497,521]" gridrow="4" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91EADFE8BFDE8" box="[151,291,497,521]" gridcol="0" gridrow="4" pageId="6" pageNumber="273">Total length</th>
<td id="6D595DD4FFC60034382B1EADFE09FDE8" box="[325,417,497,521]" gridcol="1" gridrow="4" pageId="6" pageNumber="273">287</td>
<td id="6D595DD4FFC6003438BB1EADFD92FDE8" box="[469,570,497,521]" gridcol="2" gridrow="4" pageId="6" pageNumber="273">662</td>
<td id="6D595DD4FFC600343B351EADFD6DFDE8" box="[603,709,497,521]" gridcol="3" gridrow="4" pageId="6" pageNumber="273">590</td>
<td id="6D595DD4FFC600343BBD1EADFC80FDE8" box="[723,808,497,521]" gridcol="4" gridrow="4" pageId="6" pageNumber="273">411</td>
<td id="6D595DD4FFC600343AE51EADFC19FDE8" box="[907,945,497,521]" gridcol="6" gridrow="4" pageId="6" pageNumber="273">591</td>
<td id="6D595DD4FFC600343D621EADFBD7FDE8" box="[1036,1151,497,521]" gridcol="8" gridrow="4" pageId="6" pageNumber="273">_</td>
<td id="6D595DD4FFC600343DE11EADFAA0FDE8" box="[1167,1288,497,521]" colspan="2" colspanRight="1" gridcol="9" gridrow="4" pageId="6" pageNumber="273">709</td>
<td id="6D595DD4FFC600343C751EADFA28FDE8" box="[1307,1408,497,521]" gridcol="11" gridrow="4" pageId="6" pageNumber="273">751</td>
</tr>
<tr id="2E8834A8FFC6003439F91D45FA28FDB4" box="[151,1408,537,597]" gridrow="5" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91D45FE8BFDB4" box="[151,291,537,597]" gridcol="0" gridrow="5" pageId="6" pageNumber="273">Ventral scales</th>
<td id="6D595DD4FFC60034382B1D45FE09FDB4" box="[325,417,537,597]" gridcol="1" gridrow="5" pageId="6" pageNumber="273">206</td>
<td id="6D595DD4FFC6003438BB1D45FD92FDB4" box="[469,570,537,597]" gridcol="2" gridrow="5" pageId="6" pageNumber="273">192</td>
<td id="6D595DD4FFC600343B351D45FD6DFDB4" box="[603,709,537,597]" gridcol="3" gridrow="5" pageId="6" pageNumber="273">210</td>
<td id="6D595DD4FFC600343BBD1D45FC80FDB4" box="[723,808,537,597]" gridcol="4" gridrow="5" pageId="6" pageNumber="273">209</td>
<td id="6D595DD4FFC600343AE51D45FC19FDB4" box="[907,945,537,597]" gridcol="6" gridrow="5" pageId="6" pageNumber="273">186</td>
<td id="6D595DD4FFC600343D621D45FBD7FDB4" box="[1036,1151,537,597]" gridcol="8" gridrow="5" pageId="6" pageNumber="273">196</td>
<td id="6D595DD4FFC600343DE11D45FAA0FDB4" box="[1167,1288,537,597]" colspan="2" colspanRight="1" gridcol="9" gridrow="5" pageId="6" pageNumber="273">212</td>
<td id="6D595DD4FFC600343C751D45FA28FDB4" box="[1307,1408,537,597]" gridcol="11" gridrow="5" pageId="6" pageNumber="273">197</td>
</tr>
<tr id="2E8834A8FFC6003439F91D39FA28FD40" box="[151,1408,613,673]" gridrow="6" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91D39FE8BFD40" box="[151,291,613,673]" gridcol="0" gridrow="6" pageId="6" pageNumber="273">Subcaudal scale pairs</th>
<td id="6D595DD4FFC60034382B1D39FE09FD40" box="[325,417,613,673]" gridcol="1" gridrow="6" pageId="6" pageNumber="273">77</td>
<td id="6D595DD4FFC6003438BB1D39FD92FD40" box="[469,570,613,673]" gridcol="2" gridrow="6" pageId="6" pageNumber="273">70,71</td>
<td id="6D595DD4FFC600343B351D39FD6DFD40" box="[603,709,613,673]" gridcol="3" gridrow="6" pageId="6" pageNumber="273">68</td>
<td id="6D595DD4FFC600343BBD1D39FC80FD40" box="[723,808,613,673]" gridcol="4" gridrow="6" pageId="6" pageNumber="273">76</td>
<td id="6D595DD4FFC600343AE51D39FC19FD40" box="[907,945,613,673]" gridcol="6" gridrow="6" pageId="6" pageNumber="273">63</td>
<td id="6D595DD4FFC600343D621D39FBD7FD40" box="[1036,1151,613,673]" gridcol="8" gridrow="6" pageId="6" pageNumber="273">_</td>
<td id="6D595DD4FFC600343DE11D39FAA0FD40" box="[1167,1288,613,673]" colspan="2" colspanRight="1" gridcol="9" gridrow="6" pageId="6" pageNumber="273">74</td>
<td id="6D595DD4FFC600343C751D39FA28FD40" box="[1307,1408,613,673]" gridcol="11" gridrow="6" pageId="6" pageNumber="273">67</td>
</tr>
<tr id="2E8834A8FFC6003439F91DEDFA28FD0C" box="[151,1408,689,749]" gridrow="7" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91DEDFE8BFD0C" box="[151,291,689,749]" gridcol="0" gridrow="7" pageId="6" pageNumber="273">Midbody scale rows</th>
<td id="6D595DD4FFC60034382B1DEDFE09FD0C" box="[325,417,689,749]" gridcol="1" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC6003438BB1DEDFD92FD0C" box="[469,570,689,749]" gridcol="2" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343B351DEDFD6DFD0C" box="[603,709,689,749]" gridcol="3" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343BBD1DEDFC80FD0C" box="[723,808,689,749]" gridcol="4" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343AE51DEDFC19FD0C" box="[907,945,689,749]" gridcol="6" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343D621DEDFBD7FD0C" box="[1036,1151,689,749]" gridcol="8" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343DE11DEDFAA0FD0C" box="[1167,1288,689,749]" colspan="2" colspanRight="1" gridcol="9" gridrow="7" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343C751DEDFA28FD0C" box="[1307,1408,689,749]" gridcol="11" gridrow="7" pageId="6" pageNumber="273">17</td>
</tr>
<tr id="2E8834A8FFC6003439F91DA2FA28FCF7" box="[151,1408,766,790]" gridrow="8" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91DA2FE8BFCF7" box="[151,291,766,790]" gridcol="0" gridrow="8" pageId="6" pageNumber="273">Supralabials</th>
<td id="6D595DD4FFC60034382B1DA2FE09FCF7" box="[325,417,766,790]" gridcol="1" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC6003438BB1DA2FD92FCF7" box="[469,570,766,790]" gridcol="2" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343B351DA2FD6DFCF7" box="[603,709,766,790]" gridcol="3" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343BBD1DA2FC80FCF7" box="[723,808,766,790]" gridcol="4" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343AE51DA2FC19FCF7" box="[907,945,766,790]" gridcol="6" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343D621DA2FBD7FCF7" box="[1036,1151,766,790]" gridcol="8" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343DE11DA2FAA0FCF7" box="[1167,1288,766,790]" colspan="2" colspanRight="1" gridcol="9" gridrow="8" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343C751DA2FA28FCF7" box="[1307,1408,766,790]" gridcol="11" gridrow="8" pageId="6" pageNumber="273">5,5</td>
</tr>
<tr id="2E8834A8FFC6003439F91C7AFA28FCDF" box="[151,1408,806,830]" gridrow="9" pageId="6" pageNumber="273" rowspan-5="1" rowspan-7="1">
<th id="6D595DD4FFC6003439F91C7AFE8BFCDF" box="[151,291,806,830]" gridcol="0" gridrow="9" pageId="6" pageNumber="273">Infralabials</th>
<td id="6D595DD4FFC60034382B1C7AFE09FCDF" box="[325,417,806,830]" gridcol="1" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC6003438BB1C7AFD92FCDF" box="[469,570,806,830]" gridcol="2" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343B351C7AFD6DFCDF" box="[603,709,806,830]" gridcol="3" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343BBD1C7AFC80FCDF" box="[723,808,806,830]" gridcol="4" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343AE51C7AFC19FCDF" box="[907,945,806,830]" gridcol="6" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343D621C7AFBD7FCDF" box="[1036,1151,806,830]" gridcol="8" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343DE11C7AFAA0FCDF" box="[1167,1288,806,830]" colspan="2" colspanRight="1" gridcol="9" gridrow="9" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343C751C7AFA28FCDF" box="[1307,1408,806,830]" gridcol="11" gridrow="9" pageId="6" pageNumber="273">8,8</td>
</tr>
<tr id="2E8834A8FFC6003439F91C12FA28FC87" box="[151,1408,846,870]" gridrow="10" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F91C12FE8BFC87" box="[151,291,846,870]" gridcol="0" gridrow="10" pageId="6" pageNumber="273">Temporals</th>
<td id="6D595DD4FFC60034382B1C12FE09FC87" box="[325,417,846,870]" gridcol="1" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC6003438BB1C12FD92FC87" box="[469,570,846,870]" gridcol="2" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC600343B351C12FCD9FC87" box="[603,881,846,870]" colspan="3" colspanRight="2" gridcol="3" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1 1+1, 1+1</td>
<td id="6D595DD4FFC600343AE51C12FC5DFC87" box="[907,1013,846,870]" colspan="2" colspanRight="1" gridcol="6" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC600343D621C12FBD7FC87" box="[1036,1151,846,870]" gridcol="8" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC600343DE11C12FAA0FC87" box="[1167,1288,846,870]" colspan="2" colspanRight="1" gridcol="9" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC600343C751C12FA28FC87" box="[1307,1408,846,870]" gridcol="11" gridrow="10" pageId="6" pageNumber="273">1+1, 1+1</td>
</tr>
<tr id="2E8834A8FFC6003439F91C2AFA28FC53" box="[151,1408,886,946]" gridrow="11" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F91C2AFE8BFC53" box="[151,291,886,946]" gridcol="0" gridrow="11" pageId="6" pageNumber="273">Color pat- tern</th>
<td id="6D595DD4FFC60034382B1C2AFE09FC53" box="[325,417,886,946]" gridcol="1" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC6003438BB1C2AFD92FC53" box="[469,570,886,946]" gridcol="2" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343B351C2AFD6DFC53" box="[603,709,886,946]" gridcol="3" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343BBD1C2AFCD9FC53" box="[723,881,886,946]" colspan="2" colspanRight="1" gridcol="4" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343AE51C2AFC5DFC53" box="[907,1013,886,946]" colspan="2" colspanRight="1" gridcol="6" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343D621C2AFBD7FC53" box="[1036,1151,886,946]" gridcol="8" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343DE11C2AFAA0FC53" box="[1167,1288,886,946]" colspan="2" colspanRight="1" gridcol="9" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343C751C2AFA28FC53" box="[1307,1408,886,946]" gridcol="11" gridrow="11" pageId="6" pageNumber="273">bicolor</td>
</tr>
<tr id="2E8834A8FFC6003439F91C9FFA28FBC2" box="[151,1408,963,1059]" gridrow="12" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F91C9FFE8BFBC2" box="[151,291,963,1059]" gridcol="0" gridrow="12" pageId="6" pageNumber="273">Mid-ventral line under tail</th>
<td id="6D595DD4FFC60034382B1C9FFE09FBC2" box="[325,417,963,1059]" gridcol="1" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC6003438BB1C9FFD92FBC2" box="[469,570,963,1059]" gridcol="2" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343B351C9FFD6DFBC2" box="[603,709,963,1059]" gridcol="3" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343BBD1C9FFCD9FBC2" box="[723,881,963,1059]" colspan="2" colspanRight="1" gridcol="4" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343AE51C9FFC5DFBC2" box="[907,1013,963,1059]" colspan="2" colspanRight="1" gridcol="6" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343D621C9FFBD7FBC2" box="[1036,1151,963,1059]" gridcol="8" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343DE11C9FFAA0FBC2" box="[1167,1288,963,1059]" colspan="2" colspanRight="1" gridcol="9" gridrow="12" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343C751C9FFA28FBC2" box="[1307,1408,963,1059]" gridcol="11" gridrow="12" pageId="6" pageNumber="273">absent</td>
</tr>
<tr id="2E8834A8FFC6003439F91B08FA28FB8D" box="[151,1408,1108,1132]" gridrow="13" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F91B08FA28FB8D" box="[151,1408,1108,1132]" colspan="12" colspanRight="11" gridcol="0" gridrow="13" pageId="6" pageNumber="273">Continued.</th>
</tr>
<tr id="2E8834A8FFC6003439F91B20FA28FB59" box="[151,1408,1148,1208]" gridrow="14" pageId="6" pageNumber="273" rowspan-0="1" rowspan-9="1">
<td id="6D595DD4FFC60034382B1B20FE09FB59" box="[325,417,1148,1208]" gridcol="1" gridrow="14" pageId="6" pageNumber="273">ZSI/ERS 444</td>
<td id="6D595DD4FFC6003438BB1B20FD92FB59" box="[469,570,1148,1208]" gridcol="2" gridrow="14" pageId="6" pageNumber="273">ZSI/ERS 105</td>
<td id="6D595DD4FFC600343B351B20FD6DFB59" box="[603,709,1148,1208]" gridcol="3" gridrow="14" pageId="6" pageNumber="273">ZSIK 7037</td>
<td id="6D595DD4FFC600343BBD1B20FC80FB59" box="[723,808,1148,1208]" gridcol="4" gridrow="14" pageId="6" pageNumber="273">ZSIK 7032</td>
<td id="6D595DD4FFC600343A251B20FC19FB59" box="[843,945,1148,1208]" colspan="2" colspanRight="1" gridcol="5" gridrow="14" pageId="6" pageNumber="273">ZSIK 7030</td>
<td id="6D595DD4FFC600343AAD1B20FBD7FB59" box="[963,1151,1148,1208]" colspan="2" colspanRight="1" gridcol="7" gridrow="14" pageId="6" pageNumber="273">BNHS BNHS 1730 1732</td>
<td id="6D595DD4FFC600343DDD1B20FAA0FB59" box="[1203,1288,1148,1208]" gridcol="10" gridrow="14" pageId="6" pageNumber="273">BNHS 2369</td>
<td id="6D595DD4FFC600343C751B20FA28FB59" box="[1307,1408,1148,1208]" gridcol="11" gridrow="14" pageId="6" pageNumber="273">MZMU 857</td>
</tr>
<tr id="2E8834A8FFC6003439F91B95FA28FB00" box="[151,1408,1225,1249]" gridrow="15" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91B95FE8BFB00" box="[151,291,1225,1249]" gridcol="0" gridrow="15" pageId="6" pageNumber="273">Sex</th>
<td id="6D595DD4FFC60034382B1B95FE09FB00" box="[325,417,1225,1249]" gridcol="1" gridrow="15" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC6003438BB1B95FD92FB00" box="[469,570,1225,1249]" gridcol="2" gridrow="15" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343B351B95FD6DFB00" box="[603,709,1225,1249]" gridcol="3" gridrow="15" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343BBD1B95FC80FB00" box="[723,808,1225,1249]" gridcol="4" gridrow="15" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343A251B95FCD9FB00" box="[843,881,1225,1249]" gridcol="5" gridrow="15" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343AAD1B95FC5DFB00" box="[963,1013,1225,1249]" gridcol="7" gridrow="15" pageId="6" pageNumber="273">M</td>
<td id="6D595DD4FFC600343D621B95FBD7FB00" box="[1036,1151,1225,1249]" gridcol="8" gridrow="15" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343DDD1B95FAA0FB00" box="[1203,1288,1225,1249]" gridcol="10" gridrow="15" pageId="6" pageNumber="273">F</td>
<td id="6D595DD4FFC600343C751B95FA28FB00" box="[1307,1408,1225,1249]" gridcol="11" gridrow="15" pageId="6" pageNumber="273">F</td>
</tr>
<tr id="2E8834A8FFC6003439F91BADFA28FAE8" box="[151,1408,1265,1289]" gridrow="16" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91BADFE8BFAE8" box="[151,291,1265,1289]" gridcol="0" gridrow="16" pageId="6" pageNumber="273">TL</th>
<td id="6D595DD4FFC60034382B1BADFE09FAE8" box="[325,417,1265,1289]" gridcol="1" gridrow="16" pageId="6" pageNumber="273">120</td>
<td id="6D595DD4FFC6003438BB1BADFD92FAE8" box="[469,570,1265,1289]" gridcol="2" gridrow="16" pageId="6" pageNumber="273">165</td>
<td id="6D595DD4FFC600343B351BADFD6DFAE8" box="[603,709,1265,1289]" gridcol="3" gridrow="16" pageId="6" pageNumber="273">120</td>
<td id="6D595DD4FFC600343BBD1BADFC80FAE8" box="[723,808,1265,1289]" gridcol="4" gridrow="16" pageId="6" pageNumber="273">90</td>
<td id="6D595DD4FFC600343A251BADFCD9FAE8" box="[843,881,1265,1289]" gridcol="5" gridrow="16" pageId="6" pageNumber="273">116</td>
<td id="6D595DD4FFC600343AAD1BADFC5DFAE8" box="[963,1013,1265,1289]" gridcol="7" gridrow="16" pageId="6" pageNumber="273">101*</td>
<td id="6D595DD4FFC600343D621BADFBD7FAE8" box="[1036,1151,1265,1289]" gridcol="8" gridrow="16" pageId="6" pageNumber="273">474</td>
<td id="6D595DD4FFC600343DDD1BADFAA0FAE8" box="[1203,1288,1265,1289]" gridcol="10" gridrow="16" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343C751BADFA28FAE8" box="[1307,1408,1265,1289]" gridcol="11" gridrow="16" pageId="6" pageNumber="273">-</td>
</tr>
<tr id="2E8834A8FFC6003439F91A45FA28FAD0" box="[151,1408,1305,1329]" gridrow="17" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91A45FE8BFAD0" box="[151,291,1305,1329]" gridcol="0" gridrow="17" pageId="6" pageNumber="273">SVL</th>
<td id="6D595DD4FFC60034382B1A45FE09FAD0" box="[325,417,1305,1329]" gridcol="1" gridrow="17" pageId="6" pageNumber="273">392</td>
<td id="6D595DD4FFC6003438BB1A45FD92FAD0" box="[469,570,1305,1329]" gridcol="2" gridrow="17" pageId="6" pageNumber="273">515</td>
<td id="6D595DD4FFC600343B351A45FD6DFAD0" box="[603,709,1305,1329]" gridcol="3" gridrow="17" pageId="6" pageNumber="273">431</td>
<td id="6D595DD4FFC600343BBD1A45FC80FAD0" box="[723,808,1305,1329]" gridcol="4" gridrow="17" pageId="6" pageNumber="273">320</td>
<td id="6D595DD4FFC600343A251A45FCD9FAD0" box="[843,881,1305,1329]" gridcol="5" gridrow="17" pageId="6" pageNumber="273">430</td>
<td id="6D595DD4FFC600343AAD1A45FC5DFAD0" box="[963,1013,1305,1329]" gridcol="7" gridrow="17" pageId="6" pageNumber="273">468</td>
<td id="6D595DD4FFC600343D621A45FBD7FAD0" box="[1036,1151,1305,1329]" gridcol="8" gridrow="17" pageId="6" pageNumber="273">132</td>
<td id="6D595DD4FFC600343DDD1A45FAA0FAD0" box="[1203,1288,1305,1329]" gridcol="10" gridrow="17" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343C751A45FA28FAD0" box="[1307,1408,1305,1329]" gridcol="11" gridrow="17" pageId="6" pageNumber="273">-</td>
</tr>
<tr id="2E8834A8FFC6003439F91A1DFA28FAB8" box="[151,1408,1345,1369]" gridrow="18" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91A1DFE8BFAB8" box="[151,291,1345,1369]" gridcol="0" gridrow="18" pageId="6" pageNumber="273">Total length</th>
<td id="6D595DD4FFC60034382B1A1DFE09FAB8" box="[325,417,1345,1369]" gridcol="1" gridrow="18" pageId="6" pageNumber="273">511</td>
<td id="6D595DD4FFC6003438BB1A1DFD92FAB8" box="[469,570,1345,1369]" gridcol="2" gridrow="18" pageId="6" pageNumber="273">680</td>
<td id="6D595DD4FFC600343B351A1DFD6DFAB8" box="[603,709,1345,1369]" gridcol="3" gridrow="18" pageId="6" pageNumber="273">551</td>
<td id="6D595DD4FFC600343BBD1A1DFC80FAB8" box="[723,808,1345,1369]" gridcol="4" gridrow="18" pageId="6" pageNumber="273">410</td>
<td id="6D595DD4FFC600343A251A1DFCD9FAB8" box="[843,881,1345,1369]" gridcol="5" gridrow="18" pageId="6" pageNumber="273">546</td>
<td id="6D595DD4FFC600343AAD1A1DFC5DFAB8" box="[963,1013,1345,1369]" gridcol="7" gridrow="18" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343D621A1DFBD7FAB8" box="[1036,1151,1345,1369]" gridcol="8" gridrow="18" pageId="6" pageNumber="273">606</td>
<td id="6D595DD4FFC600343DDD1A1DFAA0FAB8" box="[1203,1288,1345,1369]" gridcol="10" gridrow="18" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343C751A1DFA28FAB8" box="[1307,1408,1345,1369]" gridcol="11" gridrow="18" pageId="6" pageNumber="273">-</td>
</tr>
<tr id="2E8834A8FFC6003439F91A35FA28FA44" box="[151,1408,1385,1445]" gridrow="19" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91A35FE8BFA44" box="[151,291,1385,1445]" gridcol="0" gridrow="19" pageId="6" pageNumber="273">Ventral scales</th>
<td id="6D595DD4FFC60034382B1A35FE09FA44" box="[325,417,1385,1445]" gridcol="1" gridrow="19" pageId="6" pageNumber="273">210</td>
<td id="6D595DD4FFC6003438BB1A35FD92FA44" box="[469,570,1385,1445]" gridcol="2" gridrow="19" pageId="6" pageNumber="273">213</td>
<td id="6D595DD4FFC600343B351A35FD6DFA44" box="[603,709,1385,1445]" gridcol="3" gridrow="19" pageId="6" pageNumber="273">195</td>
<td id="6D595DD4FFC600343BBD1A35FC80FA44" box="[723,808,1385,1445]" gridcol="4" gridrow="19" pageId="6" pageNumber="273">211</td>
<td id="6D595DD4FFC600343A251A35FCD9FA44" box="[843,881,1385,1445]" gridcol="5" gridrow="19" pageId="6" pageNumber="273">204</td>
<td id="6D595DD4FFC600343AAD1A35FC5DFA44" box="[963,1013,1385,1445]" gridcol="7" gridrow="19" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343D621A35FBD7FA44" box="[1036,1151,1385,1445]" gridcol="8" gridrow="19" pageId="6" pageNumber="273">192</td>
<td id="6D595DD4FFC600343DDD1A35FAA0FA44" box="[1203,1288,1385,1445]" gridcol="10" gridrow="19" pageId="6" pageNumber="273">196</td>
<td id="6D595DD4FFC600343C751A35FA28FA44" box="[1307,1408,1385,1445]" gridcol="11" gridrow="19" pageId="6" pageNumber="273">189</td>
</tr>
<tr id="2E8834A8FFC6003439F91AEAFA28FA13" box="[151,1408,1462,1522]" gridrow="20" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91AEAFE8BFA13" box="[151,291,1462,1522]" gridcol="0" gridrow="20" pageId="6" pageNumber="273">Subcaudal scale pairs</th>
<td id="6D595DD4FFC60034382B1AEAFE09FA13" box="[325,417,1462,1522]" gridcol="1" gridrow="20" pageId="6" pageNumber="273">80</td>
<td id="6D595DD4FFC6003438BB1AEAFD92FA13" box="[469,570,1462,1522]" gridcol="2" gridrow="20" pageId="6" pageNumber="273">85</td>
<td id="6D595DD4FFC600343B351AEAFD6DFA13" box="[603,709,1462,1522]" gridcol="3" gridrow="20" pageId="6" pageNumber="273">71,70</td>
<td id="6D595DD4FFC600343BBD1AEAFC80FA13" box="[723,808,1462,1522]" gridcol="4" gridrow="20" pageId="6" pageNumber="273">73</td>
<td id="6D595DD4FFC600343A251AEAFCD9FA13" box="[843,881,1462,1522]" gridcol="5" gridrow="20" pageId="6" pageNumber="273">74</td>
<td id="6D595DD4FFC600343AAD1AEAFC5DFA13" box="[963,1013,1462,1522]" gridcol="7" gridrow="20" pageId="6" pageNumber="273">_</td>
<td id="6D595DD4FFC600343D621AEAFBD7FA13" box="[1036,1151,1462,1522]" gridcol="8" gridrow="20" pageId="6" pageNumber="273">61</td>
<td id="6D595DD4FFC600343DDD1AEAFAA0FA13" box="[1203,1288,1462,1522]" gridcol="10" gridrow="20" pageId="6" pageNumber="273">71</td>
<td id="6D595DD4FFC600343C751AEAFA28FA13" box="[1307,1408,1462,1522]" gridcol="11" gridrow="20" pageId="6" pageNumber="273">-</td>
</tr>
<tr id="2E8834A8FFC6003439F9195EFA28F9DF" box="[151,1408,1538,1598]" gridrow="21" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F9195EFE8BF9DF" box="[151,291,1538,1598]" gridcol="0" gridrow="21" pageId="6" pageNumber="273">Midbody scale rows</th>
<td id="6D595DD4FFC60034382B195EFE09F9DF" box="[325,417,1538,1598]" gridcol="1" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC6003438BB195EFD92F9DF" box="[469,570,1538,1598]" gridcol="2" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343B35195EFD6DF9DF" box="[603,709,1538,1598]" gridcol="3" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343BBD195EFC80F9DF" box="[723,808,1538,1598]" gridcol="4" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343A25195EFCD9F9DF" box="[843,881,1538,1598]" gridcol="5" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343AAD195EFC5DF9DF" box="[963,1013,1538,1598]" gridcol="7" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343D62195EFBD7F9DF" box="[1036,1151,1538,1598]" gridcol="8" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343DDD195EFAA0F9DF" box="[1203,1288,1538,1598]" gridcol="10" gridrow="21" pageId="6" pageNumber="273">17</td>
<td id="6D595DD4FFC600343C75195EFA28F9DF" box="[1307,1408,1538,1598]" gridcol="11" gridrow="21" pageId="6" pageNumber="273">17</td>
</tr>
<tr id="2E8834A8FFC6003439F91912FA28F987" box="[151,1408,1614,1638]" gridrow="22" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F91912FE8BF987" box="[151,291,1614,1638]" gridcol="0" gridrow="22" pageId="6" pageNumber="273">Supralabials</th>
<td id="6D595DD4FFC60034382B1912FE09F987" box="[325,417,1614,1638]" gridcol="1" gridrow="22" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC6003438BB1912FD92F987" box="[469,570,1614,1638]" gridcol="2" gridrow="22" pageId="6" pageNumber="273">6,6</td>
<td id="6D595DD4FFC600343B351912FD6DF987" box="[603,709,1614,1638]" gridcol="3" gridrow="22" pageId="6" pageNumber="273">5,6</td>
<td id="6D595DD4FFC600343BBD1912FC80F987" box="[723,808,1614,1638]" gridcol="4" gridrow="22" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343A251912FCD9F987" box="[843,881,1614,1638]" gridcol="5" gridrow="22" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343AAD1912FC5DF987" box="[963,1013,1614,1638]" gridcol="7" gridrow="22" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343D621912FBD7F987" box="[1036,1151,1614,1638]" gridcol="8" gridrow="22" pageId="6" pageNumber="273">5,5</td>
<td id="6D595DD4FFC600343DDD1912FAA0F987" box="[1203,1288,1614,1638]" gridcol="10" gridrow="22" pageId="6" pageNumber="273">5,6</td>
<td id="6D595DD4FFC600343C751912FA28F987" box="[1307,1408,1614,1638]" gridcol="11" gridrow="22" pageId="6" pageNumber="273">5,5</td>
</tr>
<tr id="2E8834A8FFC6003439F9192AFA28F96E" box="[151,1408,1654,1679]" gridrow="23" pageId="6" pageNumber="273" rowspan-6="1" rowspan-9="1">
<th id="6D595DD4FFC6003439F9192AFE8BF96E" box="[151,291,1654,1679]" gridcol="0" gridrow="23" pageId="6" pageNumber="273">Infralabials</th>
<td id="6D595DD4FFC60034382B192AFE09F96E" box="[325,417,1654,1679]" gridcol="1" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC6003438BB192AFD92F96E" box="[469,570,1654,1679]" gridcol="2" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343B35192AFD6DF96E" box="[603,709,1654,1679]" gridcol="3" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343BBD192AFC80F96E" box="[723,808,1654,1679]" gridcol="4" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343A25192AFCD9F96E" box="[843,881,1654,1679]" gridcol="5" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343AAD192AFC5DF96E" box="[963,1013,1654,1679]" gridcol="7" gridrow="23" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343D62192AFBD7F96E" box="[1036,1151,1654,1679]" gridcol="8" gridrow="23" pageId="6" pageNumber="273">8,8</td>
<td id="6D595DD4FFC600343DDD192AFAA0F96E" box="[1203,1288,1654,1679]" gridcol="10" gridrow="23" pageId="6" pageNumber="273">7,7</td>
<td id="6D595DD4FFC600343C75192AFA28F96E" box="[1307,1408,1654,1679]" gridcol="11" gridrow="23" pageId="6" pageNumber="273">8,8</td>
</tr>
<tr id="2E8834A8FFC6003439F919C3FA28F956" box="[151,1408,1695,1719]" gridrow="24" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F919C3FE8BF956" box="[151,291,1695,1719]" gridcol="0" gridrow="24" pageId="6" pageNumber="273">Temporals</th>
<td id="6D595DD4FFC60034382B19C3FE09F956" box="[325,417,1695,1719]" gridcol="1" gridrow="24" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC6003438BB19C3FD92F956" box="[469,570,1695,1719]" gridcol="2" gridrow="24" pageId="6" pageNumber="273">1+1, 1+1</td>
<td id="6D595DD4FFC600343B3519C3FD6DF956" box="[603,709,1695,1719]" gridcol="3" gridrow="24" pageId="6" pageNumber="273">1+1,1+1</td>
<td id="6D595DD4FFC600343BBD19C3FC80F956" box="[723,808,1695,1719]" gridcol="4" gridrow="24" pageId="6" pageNumber="273">1+1,1+1</td>
<td id="6D595DD4FFC600343A2519C3FC19F956" box="[843,945,1695,1719]" colspan="2" colspanRight="1" gridcol="5" gridrow="24" pageId="6" pageNumber="273">1+1,1+1</td>
<td id="6D595DD4FFC600343AAD19C3FC5DF956" box="[963,1013,1695,1719]" gridcol="7" gridrow="24" pageId="6" pageNumber="273">-</td>
<td id="6D595DD4FFC600343D6219C3FB39F956" box="[1036,1169,1695,1719]" colspan="2" colspanRight="1" gridcol="8" gridrow="24" pageId="6" pageNumber="273">1+1,1+2</td>
<td id="6D595DD4FFC600343DDD19C3FAA0F956" box="[1203,1288,1695,1719]" gridcol="10" gridrow="24" pageId="6" pageNumber="273">1+1,1+1</td>
<td id="6D595DD4FFC600343C7519C3FA28F956" box="[1307,1408,1695,1719]" gridcol="11" gridrow="24" pageId="6" pageNumber="273">1+1,1+1</td>
</tr>
<tr id="2E8834A8FFC6003439F9199BFA28F8E2" box="[151,1408,1735,1795]" gridrow="25" pageId="6" pageNumber="273">
<th id="6D595DD4FFC6003439F9199BFE8BF8E2" box="[151,291,1735,1795]" gridcol="0" gridrow="25" pageId="6" pageNumber="273">Color pat- tern</th>
<td id="6D595DD4FFC60034382B199BFE09F8E2" box="[325,417,1735,1795]" gridcol="1" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC6003438BB199BFD92F8E2" box="[469,570,1735,1795]" gridcol="2" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343B35199BFD6DF8E2" box="[603,709,1735,1795]" gridcol="3" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343BBD199BFC80F8E2" box="[723,808,1735,1795]" gridcol="4" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343A25199BFC19F8E2" box="[843,945,1735,1795]" colspan="2" colspanRight="1" gridcol="5" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343AAD199BFB39F8E2" box="[963,1169,1735,1795]" colspan="3" colspanRight="2" gridcol="7" gridrow="25" pageId="6" pageNumber="273">bicolor bicolor</td>
<td id="6D595DD4FFC600343DDD199BFAA0F8E2" box="[1203,1288,1735,1795]" gridcol="10" gridrow="25" pageId="6" pageNumber="273">bicolor</td>
<td id="6D595DD4FFC600343C75199BFA28F8E2" box="[1307,1408,1735,1795]" gridcol="11" gridrow="25" pageId="6" pageNumber="273">yellow bands</td>
</tr>
<tr id="2E8834A8FFC6003439F9184FFA28F892" box="[151,1408,1811,1907]" gridrow="26" pageId="6" pageNumber="273" rowspan-9="1">
<th id="6D595DD4FFC6003439F9184FFE8BF892" box="[151,291,1811,1907]" gridcol="0" gridrow="26" pageId="6" pageNumber="273">Mid-ventral line under tail</th>
<td id="6D595DD4FFC60034382B184FFE09F892" box="[325,417,1811,1907]" gridcol="1" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC6003438BB184FFD92F892" box="[469,570,1811,1907]" gridcol="2" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343B35184FFD6DF892" box="[603,709,1811,1907]" gridcol="3" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343BBD184FFC80F892" box="[723,808,1811,1907]" gridcol="4" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343A25184FFC19F892" box="[843,945,1811,1907]" colspan="2" colspanRight="1" gridcol="5" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343AAD184FFBD7F892" box="[963,1151,1811,1907]" colspan="2" colspanRight="1" gridcol="7" gridrow="26" pageId="6" pageNumber="273">absent absent</td>
<td id="6D595DD4FFC600343DDD184FFAA0F892" box="[1203,1288,1811,1907]" gridcol="10" gridrow="26" pageId="6" pageNumber="273">absent</td>
<td id="6D595DD4FFC600343C75184FFA28F892" box="[1307,1408,1811,1907]" gridcol="11" gridrow="26" pageId="6" pageNumber="273">absent</td>
</tr>
</table>
</paragraph>
<caption id="C4C76662FFC7FFCC39F918F5FC21F804" ID-DOI="http://doi.org/10.5281/zenodo.4413807" ID-Zenodo-Dep="4413807" httpUri="https://zenodo.org/record/4413807/files/figure.png" pageId="7" pageNumber="274" startId="7.[151,250,1961,1985]" targetBox="[184,1404,207,1930]" targetPageId="7">
<paragraph id="900736EAFFC7FFCC39F918F5FC21F804" blockId="7.[151,1436,1961,2021]" pageId="7" pageNumber="274">
<emphasis id="A2CCEAF8FFC7FFCC39F918F5FEA6F820" bold="true" box="[151,270,1961,1985]" pageId="7" pageNumber="274">FIGURE 4</emphasis>
. 
<taxonomicName id="57B84D69FFC7FFCC387518F5FD86F820" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[283,558,1961,1985]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC7FFCC387518F5FD86F820" box="[283,558,1961,1985]" italics="true" pageId="7" pageNumber="274">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC7FFCC3B5B18F5FD2EF820" bold="true" box="[565,646,1961,1985]" pageId="7" pageNumber="274">
<taxonomicNameLabel id="B9FF5783FFC7FFCC3B5B18F5FD2EF820" box="[565,646,1961,1985]" pageId="7" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratypes in dorsal views 
<emphasis id="A2CCEAF8FFC7FFCC3AF018F5FC18F820" bold="true" box="[926,944,1961,1985]" pageId="7" pageNumber="274">A</emphasis>
. BMNH 1935.10.12.10 (insert image shows the detached anterior part of the head) and 
<emphasis id="A2CCEAF8FFC7FFCC3B4B1891FD9EF804" bold="true" box="[549,566,1997,2021]" pageId="7" pageNumber="274">B</emphasis>
. (ZSIK 23875). Scale bar 10mm.
</paragraph>
</caption>
<caption id="C4C76662FFC8FFC339F919A5FDD1F8D7" ID-DOI="http://doi.org/10.5281/zenodo.4413809" ID-Zenodo-Dep="4413809" httpUri="https://zenodo.org/record/4413809/files/figure.png" pageId="8" pageNumber="275" startId="8.[151,250,1785,1809]" targetBox="[328,1177,257,1690]" targetPageId="8">
<paragraph id="900736EAFFC8FFC339F919A5FDD1F8D7" blockId="8.[151,1437,1785,1846]" pageId="8" pageNumber="275">
<emphasis id="A2CCEAF8FFC8FFC339F919A5FEA6F8F3" bold="true" box="[151,270,1785,1810]" pageId="8" pageNumber="275">FIGURE 5</emphasis>
. 
<taxonomicName id="57B84D69FFC8FFC3387519A6FD86F8F3" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[283,558,1786,1810]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC8FFC3387519A6FD86F8F3" box="[283,558,1786,1810]" italics="true" pageId="8" pageNumber="275">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC8FFC33B5B19A6FD2EF8F3" bold="true" box="[565,646,1786,1810]" pageId="8" pageNumber="275">
<taxonomicNameLabel id="B9FF5783FFC8FFC33B5B19A6FD2EF8F3" box="[565,646,1786,1810]" pageId="8" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, reconstructed head of paratype (BMNH 1935.10.12.10), breaks in specimen marked with red dashed lines. Scale bar 10mm.
</paragraph>
</caption>
<caption id="C4C76662FFC9FFC239F918CEFBBFF82E" ID-DOI="http://doi.org/10.5281/zenodo.4413813" ID-Zenodo-Dep="4413813" httpUri="https://zenodo.org/record/4413813/files/figure.png" pageId="9" pageNumber="276" startId="9.[151,250,1938,1962]" targetBox="[151,1436,190,1902]" targetPageId="9">
<paragraph id="900736EAFFC9FFC239F918CEFBBFF82E" blockId="9.[151,1437,1938,1999]" pageId="9" pageNumber="276">
<emphasis id="A2CCEAF8FFC9FFC239F918CEFEBDF84B" bold="true" box="[151,277,1938,1962]" pageId="9" pageNumber="276">FIGURE 6.</emphasis>
<taxonomicName id="57B84D69FFC9FFC2387018CFFD9AF84A" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[286,562,1939,1963]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFC9FFC2387018CFFD9AF84A" box="[286,562,1939,1963]" italics="true" pageId="9" pageNumber="276">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFC9FFC23B5518CFFD26F84A" bold="true" box="[571,654,1939,1963]" pageId="9" pageNumber="276">
<taxonomicNameLabel id="B9FF5783FFC9FFC23B5518CFFD26F84A" box="[571,654,1939,1963]" pageId="9" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="A2CCEAF8FFC9FFC23BF318CEFD07F84B" bold="true" box="[669,687,1938,1962]" pageId="9" pageNumber="276">A</emphasis>
. unnumbered referred specimen in Roing Forest Department collection. Total length 613 mm. 
<emphasis id="A2CCEAF8FFC9FFC2381918EBFE20F82E" bold="true" box="[375,392,1975,1999]" pageId="9" pageNumber="276">B</emphasis>
. unnumbered (2) referred specimen in Miao museum collection.
</paragraph>
</caption>
<caption id="C4C76662FFCAFFC139F91822FED0F85A" ID-DOI="http://doi.org/10.5281/zenodo.4413815" ID-Zenodo-Dep="4413815" httpUri="https://zenodo.org/record/4413815/files/figure.png" pageId="10" pageNumber="277" startId="10.[151,250,1918,1942]" targetBox="[151,1436,181,1894]" targetPageId="10">
<paragraph id="900736EAFFCAFFC139F91822FED0F85A" blockId="10.[151,1436,1918,1979]" pageId="10" pageNumber="277">
<emphasis id="A2CCEAF8FFCAFFC139F91822FEBDF876" bold="true" box="[151,277,1918,1943]" pageId="10" pageNumber="277">FIGURE 7.</emphasis>
Habitat of 
<taxonomicName id="57B84D69FFCAFFC138E01822FD0AF877" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[398,674,1918,1942]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="277" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFCAFFC138E01822FD0AF877" box="[398,674,1918,1942]" italics="true" pageId="10" pageNumber="277">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFCAFFC13BC41823FD55F876" bold="true" box="[682,765,1919,1943]" pageId="10" pageNumber="277">
<taxonomicNameLabel id="B9FF5783FFCAFFC13BC41823FD55F876" box="[682,765,1919,1943]" pageId="10" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type locality Koronu, Lower Dibang Valley District, Arunachal Pradesh, India.
</paragraph>
</caption>
<caption id="C4C76662FFCBFFC039F91BCFFD9AFB31" ID-DOI="http://doi.org/10.5281/zenodo.4413819" ID-Zenodo-Dep="4413819" httpUri="https://zenodo.org/record/4413819/files/figure.png" pageId="11" pageNumber="278" startId="11.[151,250,1171,1195]" targetBox="[163,1428,188,1140]" targetPageId="11">
<paragraph id="900736EAFFCBFFC039F91BCFFD9AFB31" blockId="11.[151,1436,1171,1232]" pageId="11" pageNumber="278">
<emphasis id="A2CCEAF8FFCBFFC039F91BCFFEBBFB4A" bold="true" box="[151,275,1171,1195]" pageId="11" pageNumber="278">FIGURE 8.</emphasis>
<taxonomicName id="57B84D69FFCBFFC038771BC8FE71FB4D" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[281,473,1172,1196]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="A2CCEAF8FFCBFFC038771BC8FE71FB4D" box="[281,473,1172,1196]" italics="true" pageId="11" pageNumber="278">Smithophis bicolor</emphasis>
</taxonomicName>
. 
<emphasis id="A2CCEAF8FFCBFFC038881BCFFE50FB4A" bold="true" box="[486,504,1171,1195]" pageId="11" pageNumber="278">A</emphasis>
. holotype ZSIK 7030, male, 
<emphasis id="A2CCEAF8FFCBFFC03A4E1BC8FC99FB4D" bold="true" box="[800,817,1172,1196]" pageId="11" pageNumber="278">B</emphasis>
. ZSI/ERS 2592, male, Shillong, Meghalaya and 
<emphasis id="A2CCEAF8FFCBFFC03C4C1BCFFA9CFB4A" bold="true" box="[1314,1332,1171,1195]" pageId="11" pageNumber="278">C</emphasis>
. ZSI/ERS 105, male, South Garo hills, Meghalaya.
</paragraph>
</caption>
<paragraph id="900736EAFFCBFFC039A91BA2FB2AFAF9" blockId="11.[151,1437,1278,1844]" box="[199,1154,1278,1304]" pageId="11" pageNumber="278">
<emphasis id="A2CCEAF8FFCBFFC039A91BA2FDAFFAF9" bold="true" box="[199,519,1278,1304]" pageId="11" pageNumber="278">Suggested common names.</emphasis>
Arunachal Rain Snake or Black and Yellow 
<taxonomicName id="57B84D69FFCBFFC03A951BA2FBD6FAF9" authorityName="Giri, Gower, Das, Lalremsanga, Lalronunga, Captain &amp; Deepak" authorityYear="2019" box="[1019,1150,1278,1304]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="genus">Smithophis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A26561FFCBFFC039A91A7EFF08F879" pageId="11" pageNumber="278" type="distribution">
<paragraph id="900736EAFFCBFFC039A91A7EFBDCFA49" blockId="11.[151,1437,1278,1844]" pageId="11" pageNumber="278">
<emphasis id="A2CCEAF8FFCBFFC039A91A7EFD59FADD" bold="true" box="[199,753,1314,1340]" pageId="11" pageNumber="278">Distribution, natural history and conservation.</emphasis>
<taxonomicName id="57B84D69FFCBFFC03B991A7EFB8CFADD" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[759,1060,1314,1340]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFCBFFC03B991A7EFB8CFADD" box="[759,1060,1314,1340]" italics="true" pageId="11" pageNumber="278">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFCBFFC03D441A7EFB2AFADD" bold="true" box="[1066,1154,1314,1340]" pageId="11" pageNumber="278">
<taxonomicNameLabel id="B9FF5783FFCBFFC03D441A7EFB2AFADD" box="[1066,1154,1314,1340]" pageId="11" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from six localities from eastern 
<collectingRegion id="527CF808FFCBFFC0383A1A1AFD83FA81" box="[340,555,1350,1376]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="278">Arunachal Pradesh</collectingRegion>
, including Roing and Koronu in Lower Dibang Valley District, Dening in Lohit District and three localities in Namdapha Tiger Reserve in Changlang District of 
<collectingRegion id="527CF808FFCBFFC03D4F1A36FB51FA65" box="[1057,1273,1386,1412]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="278">Arunachal Pradesh</collectingRegion>
. The elevation ranges from 
<quantity id="57409B0FFFCBFFC0384B1AD2FEC4FA49" box="[293,364,1422,1448]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="278" unit="m" value="310.0">310 m</quantity>
in the foothills of Mishmi Hills to 
<quantity id="57409B0FFFCBFFC03B9A1AD2FC93FA49" box="[756,827,1422,1449]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="278" unit="m" value="580.0">580 m</quantity>
at Namdapha Forest Camp.
</paragraph>
<paragraph id="900736EAFFCBFFC039A91AEEFB98F9D8" blockId="11.[151,1437,1278,1844]" pageId="11" pageNumber="278">
The 
<typeStatus id="4F038848FFCBFFC039941AEEFEF4FA2D" box="[250,348,1458,1484]" pageId="11" pageNumber="278" type="holotype">holotype</typeStatus>
was caught while it was moving on the side of a stream in moist evergreen forest. The area was characterised by degraded forest with extensive bamboo regeneration along both sides of the stream. The stream section where the snake was found had large, bryophyte covered boulders and bedrock near a pool and riffle section (
<figureCitation id="08832A6FFFCBFFC039CE1942FF4CF9D9" box="[160,228,1566,1592]" captionStart="FIGURE 7" captionStartId="10.[151,250,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-19@10.[151,1436,181,1894]" captionTargetPageId="10" captionText="FIGURE 7. Habitat of Smithophis arunachalensis sp. nov. at the type locality Koronu, Lower Dibang Valley District, Arunachal Pradesh, India." figureDoi="http://doi.org/10.5281/zenodo.4413815" httpUri="https://zenodo.org/record/4413815/files/figure.png" pageId="11" pageNumber="278">Fig. 7</figureCitation>
). The snake was caught in dim light during heavy rain, near dawn at 05:00.
</paragraph>
<paragraph id="900736EAFFCBFFC039A9191EFDD9F8D5" blockId="11.[151,1437,1278,1844]" pageId="11" pageNumber="278">
<taxonomicName id="57B84D69FFCBFFC039A9191EFE50F9BD" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[199,504,1602,1628]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFCBFFC039A9191EFE50F9BD" box="[199,504,1602,1628]" italics="true" pageId="11" pageNumber="278">Smithophis arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFCBFFC03B6F191EFDF4F9BD" bold="true" box="[513,604,1602,1628]" pageId="11" pageNumber="278">
<taxonomicNameLabel id="B9FF5783FFCBFFC03B6F191EFDF4F9BD" box="[513,604,1602,1628]" pageId="11" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to be a rarely encountered snake. Since its first collection around the year 1935, only five other specimens have been collected, and multiple surveys in Lower Dibang Valley in 
<collectingRegion id="527CF808FFCBFFC039F919D6FEC4F945" box="[151,364,1674,1700]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="278">Arunachal Pradesh</collectingRegion>
by A.D. between 2005 and 2012 yielded only a single sighting. Although some localities where 
<taxonomicName id="57B84D69FFCBFFC039F919F2FEF6F929" authority="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower, 2020" authorityName="Das &amp; Deepak &amp; Captain &amp; Wade &amp; Gower" authorityYear="2020" box="[151,350,1710,1736]" class="Reptilia" family="Colubridae" genus="Smithophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="A2CCEAF8FFCBFFC039F919F2FEF6F929" box="[151,350,1710,1736]" italics="true" pageId="11" pageNumber="278">S. arunachalensis</emphasis>
</taxonomicName>
<emphasis id="A2CCEAF8FFCBFFC0380819F2FE17F929" bold="true" box="[358,447,1710,1736]" pageId="11" pageNumber="278">
<taxonomicNameLabel id="B9FF5783FFCBFFC0380819F2FE17F929" box="[358,447,1710,1736]" pageId="11" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs are in areas that are under some protection (Namdapha Tiger Reserve and on the fringes of Mehao Wildlife Sanctuary), so little is known of the distribution and natural history of this species that it is currently likely to qualify as Data Deficient based on criteria for the Red List of Threatened Species (
<bibRefCitation id="F4294B1BFFCBFFC03C3819AAFDCEF8D5" author="IUCN Standards &amp; Petitions Committee" pageId="11" pageNumber="278" refId="ref7741" refString="IUCN Standards and Petitions Committee (2019) Guidelines for using and IUCN Red List categories and criteria. Version 14. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 9 September 2020)" type="book" year="2019">IUCN Standards and Petitions Committee 2019</bibRefCitation>
).
</paragraph>
<paragraph id="900736EAFFCBFFC039F918DFFF08F879" blockId="11.[151,160,1923,1944]" box="[151,160,1923,1944]" pageId="11" pageNumber="278">
<emphasis id="A2CCEAF8FFCBFFC039F918DFFF08F879" box="[151,160,1923,1944]" italics="true" pageId="11" pageNumber="278">r</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>