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<document id="827C8561022815935CC214FF63DE7A13" ID-CLB-Dataset="20233" ID-DOI="10.5852/ejt.2022.797.1667" ID-GBIF-Dataset="e4b9a92e-23ef-4d55-85c9-ccabcbd8316e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6299487" ID-ZooBank="8DF57743-9C53-4265-BCB5-743276A3A16C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645807222198" checkinUser="felipe" docAuthor="Telnov, Dmitry" docDate="2022" docId="6D52212B21232F78FD954EED6EE3FAE0" docLanguage="en" docName="ejt-797_1-100.pdf" docOrigin="European Journal of Taxonomy 797" docSource="http://dx.doi.org/10.5852/ejt.2022.797.1667" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Macrotomoderus bicrispus Telnov 2022, sp. nov." docType="treatment" docUuid="0746DE3F-1EAB-483C-AC83-7EE4D3505DBA" docUuidSource="ZooBank" docVersion="10" lastPageNumber="12" masterDocId="916B5953212B2F73FFD24C506A5AFFF0" masterDocTitle="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="9" updateTime="1699316540125" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9A5C2E7D2DA56BD5AF066C129AE79C3A">Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species</mods:title>
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<mods:namePart id="9056FBB3DD4D0DFA03EEA3B7F0BE2642">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="E84931DF0ABDA77989CE13D8AEC1B43E">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV 1004, Rīga, Latvia. Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV 5401, Daugavpils, Latvia.</mods:affiliation>
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<treatment id="6D52212B21232F78FD954EED6EE3FAE0" ID-DOI="http://doi.org/10.5281/zenodo.6317912" ID-GBIF-Taxon="194038740" ID-Zenodo-Dep="6317912" LSID="urn:lsid:zoobank.org:act:0746DE3F-1EAB-483C-AC83-7EE4D3505DBA" httpUri="http://treatment.plazi.org/id/6D52212B21232F78FD954EED6EE3FAE0" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="ADE1C3B621232F7BFD954EED69B7FD28" box="[583,1005,701,728]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="E544903D21232F7BFD954EED69B7FD28" blockId="8.[357,1232,701,799]" box="[583,1005,701,728]" pageId="8" pageNumber="9">
<heading id="BE0C275121232F7BFD954EED69B7FD28" box="[583,1005,701,728]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="22FBEBBE21232F7BFD954EED69D0FD28" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[583,906,701,728]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bicrispus" status="sp. nov.">
<emphasis id="D78F4C2F21232F7BFD954EED69D0FD28" bold="true" box="[583,906,701,728]" italics="true" pageId="8" pageNumber="9">Macrotomoderus bicrispus</emphasis>
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<taxonomicNameLabel id="CCBCF15421232F7BFC404EEE69B7FD28" box="[914,1005,702,728]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="E544903D21232F7BFD354F546917FCEF" blockId="8.[357,1232,701,799]" box="[743,845,772,799]" pageId="8" pageNumber="9">
<figureCitation id="7DC08CB821232F7BFD354F546917FCEF" box="[743,845,772,799]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,232,1963,1989]" captionStartId-1="10.[189,232,1928,1954]" captionTargetBox-0="[250,1337,288,1881]" captionTargetBox-1="[232,1365,281,1874]" captionTargetId-0="figure-12@9.[241,1346,265,1917]" captionTargetId-1="figure-21@10.[215,1372,265,1886]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." captionText-1="Fig. 5. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN), aedeagus. AB. General view, different positions. C. Gonopore armature in median portion of aedeagus. D. Ditto, basal portion of apicale. E. Ditto, apical portion, different position. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6299497" figureDoi-1="http://doi.org/10.5281/zenodo.6299499" httpUri-0="https://zenodo.org/record/6299497/files/figure.png" httpUri-1="https://zenodo.org/record/6299499/files/figure.png" pageId="8" pageNumber="9">Figs 45</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621232F7BFF6F4F1569C7FB91" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="E544903D21232F7BFF6F4F156B93FCAF" blockId="8.[189,457,837,863]" box="[189,457,837,863]" pageId="8" pageNumber="9">
<heading id="BE0C275121232F7BFF6F4F156B93FCAF" bold="true" box="[189,457,837,863]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="D78F4C2F21232F7BFF6F4F156B93FCAF" bold="true" box="[189,457,837,863]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
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<paragraph id="E544903D21232F7BFF6F4F2469C7FB91" blockId="8.[189,1400,884,1122]" pageId="8" pageNumber="9">
This species readily differs from all congeners primarily due to the presence of the paired bunches of the curl-like golden setae on the median part of the anterior margin of pronotum and in the peculiar shape of the anterior pronotal projection. Most similar to
<taxonomicName id="22FBEBBE21232F7BFC474FEA6F2DFC24" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[917,1399,954,980]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chingpo">
<emphasis id="D78F4C2F21232F7BFC474FEA6E91FC24" box="[917,1227,954,980]" italics="true" pageId="8" pageNumber="9">Macrotomoderus chingpo</emphasis>
<bibRefCitation id="816AEDCC21232F7BFB0A4FEA6F2DFC24" author="Telnov D." box="[1240,1399,954,980]" pageId="8" pageNumber="9" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="9DECD0AD21232F7BFF174F8D6B4BFC07" box="[197,273,989,1015]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
:
<collectingRegion id="273F5EDF21232F7BFEC94F8E6B2DFC08" box="[283,375,990,1016]" country="China" name="Yunnan" pageId="8" pageNumber="9">Yunnan</collectingRegion>
) that also possesses somewhat similar, Ɔ-like curved median setae at anterior margin of pronotum. However, head is comparatively wider in
<taxonomicName id="22FBEBBE21232F7BFCF8485169E4FBEB" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[810,958,1025,1051]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bicrispus" status="sp. nov.">
<emphasis id="D78F4C2F21232F7BFCF8485169E4FBEB" box="[810,958,1025,1051]" italics="true" pageId="8" pageNumber="9">M. bicrispus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421232F7BFC1748516E78FBEB" box="[965,1058,1025,1051]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, the shape of the aedeagus is different, the lateral constriction of the pronotum is broader and deeper (in dorsal view), and the elytra are comparatively slenderer, more elongate in the new species.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621232F7BFF6F48D96848FB05" pageId="8" pageNumber="9" type="etymology">
<paragraph id="E544903D21232F7BFF6F48D96B1EFB53" blockId="8.[189,324,1161,1187]" box="[189,324,1161,1187]" pageId="8" pageNumber="9">
<heading id="BE0C275121232F7BFF6F48D96B1EFB53" bold="true" box="[189,324,1161,1187]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="D78F4C2F21232F7BFF6F48D96B1EFB53" bold="true" box="[189,324,1161,1187]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21232F7BFF6F48E76848FB05" blockId="8.[188,1398,1207,1269]" pageId="8" pageNumber="9">Named from the Latin bi (two) + crispum (curled) to point on paired bunches of curl-like setae on anterior margin of pronotum.</paragraph>
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<subSubSection id="ADE1C3B621232F7BFF6F494C695EFA5B" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="E544903D21232F7BFF6F494C6B36FAC5" blockId="8.[189,364,1307,1334]" box="[189,364,1307,1334]" pageId="8" pageNumber="9">
<heading id="BE0C275121232F7BFF6F494C6B36FAC5" bold="true" box="[189,364,1307,1334]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="D78F4C2F21232F7BFF6F494C6B36FAC5" bold="true" box="[189,364,1307,1334]" pageId="8" pageNumber="9">Type material</emphasis>
</heading>
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<paragraph id="E544903D21232F7BFF38491A695EFA5B" blockId="8.[189,1399,1354,1451]" pageId="8" pageNumber="9">
<materialsCitation id="55939A6021232F7BFF38491A695EFA5B" ID-GBIF-Occurrence="3705075303" collectingDate="2016-05-29" collectionCode="ZIN" collectorName="Kabak &amp; Davidian" country="China" elevation="3225" latitude="27.210556" location="Yunnan" longLatPrecision="20" longitude="99.35611" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="D78F4C2F21232F7BFF38491A6B06FA94" bold="true" box="[234,348,1354,1380]" pageId="8" pageNumber="9">
<typeStatus id="3A402E9F21232F7BFF38491A6B06FA94" box="[234,348,1354,1380]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9DECD0AD21232F7BFF6F493E6B42FA78" box="[189,280,1390,1416]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
<specimenCount id="F3FD5BB421232F7BFEE3493E6B10FA78" box="[305,330,1390,1416]" pageId="8" pageNumber="9" type="male"></specimenCount>
; “CHINA,
<collectingRegion id="273F5EDF21232F7BFE1D493E6876FA78" box="[463,556,1390,1416]" country="China" name="Yunnan" pageId="8" pageNumber="9">Yunnan</collectingRegion>
,
<locationDeviation id="F4F6891F21232F7BFDE5493E6953FA77" box="[567,777,1389,1416]" pageId="8" pageNumber="9">SW of Guoditang</locationDeviation>
,
<locationDeviation id="F4F6891F21232F7BFCC6493D6E17FA77" box="[788,1101,1389,1416]" location="Weixi City" pageId="8" pageNumber="9">
<quantity id="22033DD821232F7BFCC6493D693FFA77" box="[788,869,1389,1415]" metricMagnitude="3" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="9" unit="km" value="6.4">6.4 km</quantity>
ENE of Weixi City
</locationDeviation>
<geoCoordinate id="80CFF6FA21232F7BFB86493C6EBAFA78" box="[1108,1248,1388,1416]" degrees="27" direction="north" minutes="12" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="38" value="27.210556">27°1238″N</geoCoordinate>
<geoCoordinate id="80CFF6FA21232F7BFB35493C6F2BFA78" box="[1255,1393,1388,1416]" degrees="99" direction="east" minutes="21" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="22" value="99.35611">99°2122″E</geoCoordinate>
,
<quantity id="22033DD821232F7BFF6F49C06B42FA5B" box="[189,280,1424,1451]" metricMagnitude="3" metricUnit="m" metricValue="3.225" pageId="8" pageNumber="9" unit="m" value="3225.0">
<elevation id="6ED6770E21232F7BFF6F49C06B42FA5B" box="[189,280,1424,1451]" metricMagnitude="3" metricUnit="m" metricValue="3.225" pageId="8" pageNumber="9" unit="m" value="3225.0">3225 m</elevation>
</quantity>
,
<date id="9145B6FD21232F7BFEF649C16BC2FA5B" box="[292,408,1425,1451]" pageId="8" pageNumber="9" value="2016-05-29">
<collectingDate id="81014F1521232F7BFEF649C16BC2FA5B" box="[292,408,1425,1451]" pageId="8" pageNumber="9" value="2016-05-29">29.v.2016</collectingDate>
</date>
<collectorName id="480EF5EB21232F7BFE4D49C16BB6FA5B" box="[415,492,1425,1451]" pageId="8" pageNumber="9">Kabak</collectorName>
&amp;
<collectorName id="480EF5EB21232F7BFDC349C16827FA5B" box="[529,637,1425,1451]" pageId="8" pageNumber="9">Davidian</collectorName>
leg.”;
<collectionCode id="83EA08F821232F7BFD1E49C1695BFA5B" box="[716,769,1425,1451]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9" type="Museum">ZIN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621232F78FF6F49826BF1FBC0" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
<paragraph id="E544903D21232F7BFF6F49826B14FA1C" blockId="8.[189,334,1490,1516]" box="[189,334,1490,1516]" pageId="8" pageNumber="9">
<heading id="BE0C275121232F7BFF6F49826B14FA1C" bold="true" box="[189,334,1490,1516]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="D78F4C2F21232F7BFF6F49826B14FA1C" bold="true" box="[189,334,1490,1516]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21232F7BFF6F4A516B80F991" blockId="8.[189,1399,1536,1634]" pageId="8" pageNumber="9">
MEASUREMENTS.
<typeStatus id="3A402E9F21232F7BFEAC4A516BB4F9EB" box="[382,494,1537,1563]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="22033DD821232F7BFD1E4A506970F9EB" box="[716,810,1536,1563]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="9" unit="mm" value="4.6">4.6 mm</quantity>
; head
<quantity id="22033DD821232F7BFCA54A5069BBF9EB" box="[887,993,1536,1563]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="9" unit="mm" value="0.95">0.95 mm</quantity>
long, across eyes
<quantity id="22033DD821232F7BFB694A506F7EF9EB" box="[1211,1316,1536,1563]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="9" unit="mm" value="0.95">0.95 mm</quantity>
broad, pronotum
<quantity id="22033DD821232F7BFEE84A746BFFF9CE" box="[314,421,1572,1599]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="9" unit="mm" value="1.25">1.25 mm</quantity>
long, maximum width
<quantity id="22033DD821232F7BFD114A746968F9CE" box="[707,818,1572,1599]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="9" unit="mm" value="0.95">0.95 mm</quantity>
, minimum width
<quantity id="22033DD821232F7BFBD84A746E23F9CE" box="[1034,1145,1572,1599]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="mm" value="0.35">0.35 mm</quantity>
, elytra
<quantity id="22033DD821232F7BFB064A746F6BF9CE" box="[1236,1329,1572,1598]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="9" unit="mm" value="2.4">2.4 mm</quantity>
long,
<quantity id="22033DD821232F7BFF6F4A176B4DF992" box="[189,279,1607,1634]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="9" unit="mm" value="1.7">1.7 mm</quantity>
combined wide.
</paragraph>
<paragraph id="E544903D21232F78FF6F4AD96E84FC30" blockId="8.[188,1401,1672,2016]" lastBlockId="11.[186,1401,265,960]" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
Dorsum and venter uniformly brown. Mouthparts, antennae, palps and legs yellowish-brown. Head ovoid with moderately large, ovoid compound eyes, which laterally are slightly protruding beyond head outline. Head rounded in broad arc posterior to eyes, head base shortly subtruncate medially. Occiput slightly declivous posterodorsally. Head dorsal punctures minute and inconspicuous. Head dorsal setae inconspicuous, sparse. Antennae extending towards base of pronotum. Antennomere three about 1.2× as long as antennomere two, antennomeres 710 transverse, of them 910 strongly transverse. Terminal antennomere triangular with rounded apex, 22.2 × as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum broadly hourglass-shaped with anterior lobe strongly wider than posterior, about same width as head across eyes. Postmedian lateral constriction broad and medially deeply notched. Front margin of anterior lobe very broadly rounded, near subtruncate, dorsally without anterior rim. Front margin of anterior lobe with two shallow impressions and very median part of front margin slightly projecting anteriad between these two impressions; this projection on both sides of its base is provided with a paired bunch of golden, long and strongly Ɔ-like curved setae touching neck and head base (
<figureCitation id="7DC08CB821202F78FEAD4D226859FE7D" box="[383,515,370,397]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[250,1337,288,1881]" captionTargetId="figure-12@9.[241,1346,265,1917]" captionTargetPageId="9" captionText="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="11" pageNumber="12">Fig. 4BC</figureCitation>
). Each of two bunches raises from one large pore. Anterior lobe convex in lateral view (
<figureCitation id="7DC08CB821202F78FEAC4DC66B85FE40" box="[382,479,406,432]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[250,1337,288,1881]" captionTargetId="figure-12@9.[241,1346,265,1917]" captionTargetPageId="9" captionText="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="11" pageNumber="12">Fig. 4C</figureCitation>
). Lateral constriction continues onto disc in lateral view, shallow (
<figureCitation id="7DC08CB821202F78FADB4DC66F30FE40" box="[1289,1386,406,432]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[250,1337,288,1881]" captionTargetId="figure-12@9.[241,1346,265,1917]" captionTargetPageId="9" captionText="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="11" pageNumber="12">Fig. 4C</figureCitation>
). Lateral pronotal fovea moderately broad at lower external margin of pronotum, widens upwards towards pronotal disc in lateral view, external margins protruding into a pair of obtuse, short setose, moderately widely separated denticles (in lateral view). In lateral view anterior denticle is strongly obtuse, with few golden setae at base, posterior subacute, glabrous. Cavity in lateral wall of pronotum between lateral denticles is large. In dorsal view, lateral pronotal fovea narrow, anterior and posterior pair of denticles appear obtusely angulate, glabrous (
<figureCitation id="7DC08CB821202F78FDB44E396899FD73" box="[614,707,617,643]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[250,1337,288,1881]" captionTargetId="figure-12@9.[241,1346,265,1917]" captionTargetPageId="9" captionText="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="11" pageNumber="12">Fig. 4B</figureCitation>
). Posterior portion of pronotal fovea long and dense setose in dorsal view (
<figureCitation id="7DC08CB821202F78FEA94EDC6B80FD56" box="[379,474,652,678]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[250,1337,288,1881]" captionTargetId="figure-12@9.[241,1346,265,1917]" captionTargetPageId="9" captionText="Fig. 4. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="11" pageNumber="12">Fig. 4B</figureCitation>
). Pronotal punctures minute on disc; lateral constriction dorsally with dense, irregularly shaped punctures, largest of which positioned longitudinally along midline. Dorsal pronotal setae inconspicuous, moderately long except dense and long setose area beyond to posterior pair of denticles. Scutellar shield minute, triangular. Elytra dorsally elongate elliptical, flattened in lateral view, widened laterally around midlength, lateral margins evenly broadly rounded, shoulders obsolete (apterous species). Elytral punctures rather small, moderately dense, becoming smaller and sparser towards apices. Elytral setae long and sparse, suberect. Male tergite and morphological sternite VII broadly rounded at posterior margin. Aedeagus as in
<figureCitation id="7DC08CB821202F78FD954FD268D7FC6D" box="[583,653,898,925]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[232,1365,281,1874]" captionTargetId="figure-21@10.[215,1372,265,1886]" captionTargetPageId="10" captionText="Fig. 5. Macrotomoderus bicrispus sp. nov., holotype, ♂ (ZIN), aedeagus. AB. General view, different positions. C. Gonopore armature in median portion of aedeagus. D. Ditto, basal portion of apicale. E. Ditto, apical portion, different position. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299499" httpUri="https://zenodo.org/record/6299499/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
, robust and large, most of apicale and median portion of basale filled with conspicuous, large and dense, irregularly shaped spine-like gonopore armature.
</paragraph>
<caption id="B184C0B521222F7AFF6F4BFB6E63F818" ID-DOI="http://doi.org/10.5281/zenodo.6299497" ID-Zenodo-Dep="6299497" httpUri="https://zenodo.org/record/6299497/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1963,1989]" targetBox="[250,1337,288,1881]" targetPageId="9">
<paragraph id="E544903D21222F7AFF6F4BFB6E63F818" blockId="9.[189,1398,1963,2025]" pageId="9" pageNumber="10">
<emphasis id="D78F4C2F21222F7AFF6F4BFB6B57F835" bold="true" box="[189,269,1963,1989]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
<taxonomicName id="22FBEBBE21222F7AFEC44BFB680EF835" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[278,596,1963,1989]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicrispus" status="sp. nov.">
<emphasis id="D78F4C2F21222F7AFEC44BFB680EF835" box="[278,596,1963,1989]" italics="true" pageId="9" pageNumber="10">Macrotomoderus bicrispus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421222F7AFD8C4BFC68E7F836" box="[606,701,1964,1990]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN).
<emphasis id="D78F4C2F21222F7AFC614BFB6992F835" bold="true" box="[947,968,1963,1989]" pageId="9" pageNumber="10">A</emphasis>
. Habitus, dorsal view.
<emphasis id="D78F4C2F21222F7AFB0C4BFB6EAAF835" bold="true" box="[1246,1264,1963,1989]" pageId="9" pageNumber="10">B</emphasis>
. Forebody, dorsal view.
<emphasis id="D78F4C2F21222F7AFE804B9E6B3DF818" bold="true" box="[338,359,1998,2024]" pageId="9" pageNumber="10">C</emphasis>
. Ditto, lateral view. Scale bar: A = 1 mm; BC = not to scale.
</paragraph>
</caption>
<caption id="B184C0B521212F79FF6F4BD86912F818" ID-DOI="http://doi.org/10.5281/zenodo.6299499" ID-Zenodo-Dep="6299499" httpUri="https://zenodo.org/record/6299499/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1928,1954]" targetBox="[232,1365,281,1874]" targetPageId="10">
<paragraph id="E544903D21212F79FF6F4BD86912F818" blockId="10.[189,1399,1928,2025]" pageId="10" pageNumber="11">
<emphasis id="D78F4C2F21212F79FF6F4BD86B51F852" bold="true" box="[189,267,1928,1954]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
<taxonomicName id="22FBEBBE21212F79FEC14BD8680AF852" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[275,592,1928,1954]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bicrispus" status="sp. nov.">
<emphasis id="D78F4C2F21212F79FEC14BD8680AF852" box="[275,592,1928,1954]" italics="true" pageId="10" pageNumber="11">Macrotomoderus bicrispus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421212F79FD854BD868EFF852" box="[599,693,1928,1954]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN), aedeagus.
<emphasis id="D78F4C2F21212F79FBF34BD86E03F852" bold="true" box="[1057,1113,1928,1954]" pageId="10" pageNumber="11">AB</emphasis>
. General view, different positions.
<emphasis id="D78F4C2F21212F79FEEB4BFB6B14F835" bold="true" box="[313,334,1963,1989]" pageId="10" pageNumber="11">C</emphasis>
. Gonopore armature in median portion of aedeagus.
<emphasis id="D78F4C2F21212F79FC084BFB69B5F835" bold="true" box="[986,1007,1963,1989]" pageId="10" pageNumber="11">D</emphasis>
. Ditto, basal portion of apicale.
<emphasis id="D78F4C2F21212F79FF6F4B9E6A8BF818" bold="true" box="[189,209,1998,2024]" pageId="10" pageNumber="11">E</emphasis>
. Ditto, apical portion, different position. Not to scale.
</paragraph>
</caption>
<paragraph id="E544903D21202F78FF6F4FB76BEAFBF1" blockId="11.[189,432,999,1025]" box="[189,432,999,1025]" pageId="11" pageNumber="12">
<heading id="BE0C275121202F78FF6F4FB76BEAFBF1" bold="true" box="[189,432,999,1025]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="D78F4C2F21202F78FF6F4FB76BEAFBF1" bold="true" box="[189,432,999,1025]" pageId="11" pageNumber="12">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21202F78FF6F48466BF1FBC0" blockId="11.[189,427,1046,1072]" box="[189,427,1046,1072]" pageId="11" pageNumber="12">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621202F78FF6F4807687FFB50" box="[189,549,1111,1184]" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="E544903D21202F78FF6F48076B7AFB81" blockId="11.[189,288,1111,1137]" box="[189,288,1111,1137]" pageId="11" pageNumber="12">
<heading id="BE0C275121202F78FF6F48076B7AFB81" bold="true" box="[189,288,1111,1137]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="D78F4C2F21202F78FF6F48076B7AFB81" bold="true" box="[189,288,1111,1137]" pageId="11" pageNumber="12">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21202F78FF6F48D6687FFB50" blockId="11.[189,549,1157,1184]" box="[189,549,1157,1184]" pageId="11" pageNumber="12">
Collected at
<quantity id="22033DD821202F78FE8348D56BF0FB50" box="[337,426,1157,1184]" metricMagnitude="3" metricUnit="m" metricValue="3.225" pageId="11" pageNumber="12" unit="m" value="3225.0">3225 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621202F78FF6F48976EE3FAE0" box="[189,1209,1223,1296]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="E544903D21202F78FF6F48976B0CFB11" blockId="11.[189,342,1223,1249]" box="[189,342,1223,1249]" pageId="11" pageNumber="12">
<heading id="BE0C275121202F78FF6F48976B0CFB11" bold="true" box="[189,342,1223,1249]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="D78F4C2F21202F78FF6F48976B0CFB11" bold="true" box="[189,342,1223,1249]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21202F78FF6F48A66EE3FAE0" blockId="11.[189,1209,1270,1296]" box="[189,1209,1270,1296]" pageId="11" pageNumber="12">
Known only from Hengduan Mountains, northern part of
<collectingRegion id="273F5EDF21202F78FCB348A66E77FAE0" box="[865,1069,1270,1296]" country="China" name="Yunnan" pageId="11" pageNumber="12">Yunnan Province</collectingRegion>
, SW
<collectingCountry id="9DECD0AD21202F78FBBE48A66EECFAE0" box="[1132,1206,1270,1296]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>