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<document id="2E5F5456494E89AD0004468716DFDCD2" ID-CLB-Dataset="20233" ID-DOI="10.5852/ejt.2022.797.1667" ID-GBIF-Dataset="e4b9a92e-23ef-4d55-85c9-ccabcbd8316e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6299487" ID-ZooBank="8DF57743-9C53-4265-BCB5-743276A3A16C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645807222198" checkinUser="felipe" docAuthor="Telnov, Dmitry" docDate="2022" docId="6D52212B21092F55FD934A066EE3FD66" docLanguage="en" docName="ejt-797_1-100.pdf" docOrigin="European Journal of Taxonomy 797" docSource="http://dx.doi.org/10.5852/ejt.2022.797.1667" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Macrotomoderus hengduan Telnov 2022, sp. nov." docType="treatment" docUuid="3C10F9ED-7C43-410B-9E6F-99197F75ABE8" docUuidSource="ZooBank" docVersion="10" lastPageNumber="39" masterDocId="916B5953212B2F73FFD24C506A5AFFF0" masterDocTitle="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="35" updateTime="1699316540125" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5A6C90AA6CDAE7D55A8B901624D178BD">Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species</mods:title>
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<mods:namePart id="03C9223D69D795C0EC6443C0A6E4CC1A">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="99818977F15E1C719047985ED35A8158">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV – 1004, Rīga, Latvia. Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV – 5401, Daugavpils, Latvia.</mods:affiliation>
<mods:nameIdentifier id="48B1D0541B097AEA75C070F66A8BF987" type="email">anthicus@gmail.com</mods:nameIdentifier>
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<mods:title id="B05CCD075E454C777D8831DCE9017468">European Journal of Taxonomy</mods:title>
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<mods:date id="DAD24F07D8F757B70CD281C8F18B1BC4">2022</mods:date>
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<treatment id="6D52212B21092F55FD934A066EE3FD66" ID-DOI="http://doi.org/10.5281/zenodo.6317932" ID-GBIF-Taxon="194038751" ID-Zenodo-Dep="6317932" LSID="urn:lsid:zoobank.org:act:3C10F9ED-7C43-410B-9E6F-99197F75ABE8" httpUri="http://treatment.plazi.org/id/6D52212B21092F55FD934A066EE3FD66" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35">
<subSubSection id="ADE1C3B621092F51FD934A0669A9F981" box="[577,1011,1622,1649]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="E544903D21092F51FD934A0669A9F981" blockId="34.[369,1219,1622,1719]" box="[577,1011,1622,1649]" pageId="34" pageNumber="35">
<heading id="BE0C275121092F51FD934A0669A9F981" box="[577,1011,1622,1649]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
<taxonomicName id="22FBEBBE21092F51FD934A0669CBF981" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[577,913,1622,1649]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21092F51FD934A0669CBF981" bold="true" box="[577,913,1622,1649]" italics="true" pageId="34" pageNumber="35">Macrotomoderus hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421092F51FC4B4A0769A9F981" box="[921,1011,1623,1649]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="ADE1C3B621092F51FEA34A2A6901F947" pageId="34" pageNumber="35" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="E544903D21092F51FD0A4ACD6901F947" blockId="34.[369,1219,1622,1719]" box="[728,859,1693,1719]" pageId="34" pageNumber="35">
<figureCitation id="7DC08CB821092F51FD0A4ACD6901F947" box="[728,859,1693,1719]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="36.[189,232,1963,1989]" captionStartId-1="37.[189,232,1928,1954]" captionTargetBox-0="[223,1371,283,1890]" captionTargetBox-1="[369,1231,278,1870]" captionTargetId-0="figure-20@36.[203,1385,263,1912]" captionTargetId-1="figure-11@37.[352,1235,265,1887]" captionTargetPageId-0="36" captionTargetPageId-1="37" captionText-0="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." captionText-1="Fig. 21. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions. C. Gonopore armature, median portion of apicale. D. Ditto, apical portion of apicale. E. Basal portion of apicale. F. Ditto, stronger magnification. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6299531" figureDoi-1="http://doi.org/10.5281/zenodo.6299533" httpUri-0="https://zenodo.org/record/6299531/files/figure.png" httpUri-1="https://zenodo.org/record/6299533/files/figure.png" pageId="34" pageNumber="35">Figs 20–21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621092F50FF6F4A8E6952FE7C" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="E544903D21092F51FF6F4A8E6B93F908" blockId="34.[189,457,1758,1784]" box="[189,457,1758,1784]" pageId="34" pageNumber="35">
<heading id="BE0C275121092F51FF6F4A8E6B93F908" bold="true" box="[189,457,1758,1784]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="D78F4C2F21092F51FF6F4A8E6B93F908" bold="true" box="[189,457,1758,1784]" pageId="34" pageNumber="35">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21092F50FF6F4B5D6952FE7C" blockId="34.[189,1400,1805,2008]" lastBlockId="35.[188,1400,265,397]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
This species falls in a group of species from continental
<collectingCountry id="9DECD0AD21092F51FCB24B5D69FCF8D7" box="[864,934,1805,1831]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
with pronotum wider than head across eyes. Among them, males of
<taxonomicName id="22FBEBBE21092F51FDFB4B61693DF8BA" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[553,871,1840,1866]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="D78F4C2F21092F51FDFB4B6168E4F8BA" box="[553,702,1840,1866]" italics="true" pageId="34" pageNumber="35">M. mirabilis</emphasis>
<bibRefCitation id="816AEDCC21092F51FD184B60693DF8BA" author="Telnov D." box="[714,871,1840,1866]" pageId="34" pageNumber="35" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="22FBEBBE21092F51FC7B4B616F6EF8BA" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[937,1332,1840,1866]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="monstrificabilis">
<emphasis id="D78F4C2F21092F51FC7B4B616ED1F8BA" box="[937,1163,1840,1866]" italics="true" pageId="34" pageNumber="35">M. monstrificabilis</emphasis>
<bibRefCitation id="816AEDCC21092F51FB444B606F6EF8BA" author="Telnov D." box="[1174,1332,1840,1866]" pageId="34" pageNumber="35" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
have distinct triangular median projection at the head base (not present in
<taxonomicName id="22FBEBBE21092F51FBC24B046EF5F89D" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[1040,1199,1875,1901]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21092F51FBC24B046EF5F89D" box="[1040,1199,1875,1901]" italics="true" pageId="34" pageNumber="35">M. hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421092F51FB684B046F46F89E" box="[1210,1308,1876,1902]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
), while the head base in male
<taxonomicName id="22FBEBBE21092F51FE1C4B276946F861" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[462,796,1911,1937]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="monstratus">
<emphasis id="D78F4C2F21092F51FE1C4B276820F860" box="[462,634,1911,1936]" italics="true" pageId="34" pageNumber="35">M. monstratus</emphasis>
<bibRefCitation id="816AEDCC21092F51FD534B276946F861" author="Telnov D." box="[641,796,1911,1937]" pageId="34" pageNumber="35" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
is slightly medially impressed (evenly rounded in
<taxonomicName id="22FBEBBE21092F51FF6D4BCB6B01F844" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[191,347,1946,1972]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21092F51FF6D4BCB6B01F844" box="[191,347,1946,1972]" italics="true" pageId="34" pageNumber="35">M. hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421092F51FEB34BCA6B9AF844" box="[353,448,1946,1972]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
), the anterior lobe of the pronotum more than twice as long as the posterior lobe (about twice as long in
<taxonomicName id="22FBEBBE21092F51FE1B4BEE6839F827" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[457,611,1981,2007]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21092F51FE1B4BEE6839F827" box="[457,611,1981,2007]" italics="true" pageId="34" pageNumber="35">M. hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421092F51FDBB4BEE689DF828" box="[617,711,1982,2008]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
), the front margin of anterior lobe in male with the pointed, triangular mesal projection that is densely setose ventrally and touching the head base at the place of the basal impression (the anterior pronotal projection different in
<taxonomicName id="22FBEBBE21082F50FC484D7D6E6CFEB6" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[922,1078,300,326]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21082F50FC484D7D6E6CFEB6" box="[922,1078,300,326]" italics="true" pageId="35" pageNumber="36">M. hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421082F50FBEC4D7D6EC0FEB7" box="[1086,1178,301,327]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
– see
<figureCitation id="7DC08CB821082F50FB344D7C6F29FEB6" box="[1254,1395,300,326]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[223,1371,283,1890]" captionTargetId="figure-20@36.[203,1385,263,1912]" captionTargetPageId="36" captionText="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="35" pageNumber="36">Fig. 20B–C</figureCitation>
, not setose ventrally and with few conspicuous dorsal setae). Also see diagnoses of
<taxonomicName id="22FBEBBE21082F50FBAC4D006EB0FE99" authorityName="Telnov" authorityYear="2022" box="[1150,1258,335,361]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="D78F4C2F21082F50FBAC4D006EB0FE99" box="[1150,1258,335,361]" italics="true" pageId="35" pageNumber="36">M. boops</emphasis>
</taxonomicName>
and
<taxonomicName id="22FBEBBE21082F50FAF04D006F2DFE99" authorityName="Telnov" authorityYear="2022" box="[1314,1399,335,361]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="dali">
<emphasis id="D78F4C2F21082F50FAF04D006F2DFE99" box="[1314,1399,335,361]" italics="true" pageId="35" pageNumber="36">M. dali</emphasis>
</taxonomicName>
spp. nov. above and of
<taxonomicName id="22FBEBBE21082F50FE1E4D236808FE7C" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[460,594,371,396]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="imitator" status="sp. nov.">
<emphasis id="D78F4C2F21082F50FE1E4D236808FE7C" box="[460,594,371,396]" italics="true" pageId="35" pageNumber="36">M. imitator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421082F50FD8B4D2368EEFE7D" box="[601,692,371,397]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
below.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621082F50FF6F4DE46884FDB3" pageId="35" pageNumber="36" type="etymology">
<paragraph id="E544903D21082F50FF6F4DE46B1EFE3E" blockId="35.[189,324,436,462]" box="[189,324,436,462]" pageId="35" pageNumber="36">
<heading id="BE0C275121082F50FF6F4DE46B1EFE3E" bold="true" box="[189,324,436,462]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="D78F4C2F21082F50FF6F4DE46B1EFE3E" bold="true" box="[189,324,436,462]" pageId="35" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21082F50FF6F4DB26884FDB3" blockId="35.[189,1398,482,579]" pageId="35" pageNumber="36">
Toponymic. Named after Yunnan’s Hengduan Mountains, a group of mountain ranges in southwest
<collectingCountry id="9DECD0AD21082F50FF6F4E566B59FDD0" box="[189,259,518,544]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
that connect the southeast portions of the Tibetan Plateau with the
<collectingRegion id="273F5EDF21082F50FB934E566EC6FDD0" box="[1089,1180,518,544]" country="China" name="Yunnan" pageId="35" pageNumber="36">Yunnan</collectingRegion>
–
<collectingRegion id="273F5EDF21082F50FB794E566F54FDD0" box="[1195,1294,518,544]" country="China" name="Guizhou" pageId="35" pageNumber="36">Guizhou</collectingRegion>
Plateau, where this species occurs. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621082F50FF6F4E3A68A9FC71" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="E544903D21082F50FF6F4E3A6B36FD74" blockId="35.[189,364,618,644]" box="[189,364,618,644]" pageId="35" pageNumber="36">
<heading id="BE0C275121082F50FF6F4E3A6B36FD74" bold="true" box="[189,364,618,644]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="D78F4C2F21082F50FF6F4E3A6B36FD74" bold="true" box="[189,364,618,644]" pageId="35" pageNumber="36">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21082F50FF384EC96F3BFCED" blockId="35.[189,1399,665,797]" pageId="35" pageNumber="36">
<materialsCitation id="55939A6021082F50FF384EC96F3BFCED" ID-GBIF-Occurrence="3705075305" collectingDate="2016-05-29" collectionCode="ZIN" collectorName="Kabak &amp; Davidian" country="China" county="Laishkan" elevation="2580" latitude="27.200834" location="Yunnan" longLatPrecision="20" longitude="98.340004" municipality="Weixi" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="D78F4C2F21082F50FF384EC96B06FD43" bold="true" box="[234,348,665,691]" pageId="35" pageNumber="36">
<typeStatus id="3A402E9F21082F50FF384EC96B06FD43" box="[234,348,665,691]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9DECD0AD21082F50FF6F4EEC6B42FD26" box="[189,280,700,726]" name="China" pageId="35" pageNumber="36">CHINA</collectingCountry>
•
<specimenCount id="F3FD5BB421082F50FEE44EED6B15FD27" box="[310,335,701,727]" pageId="35" pageNumber="36" type="male">♂</specimenCount>
; “
<collectingCountry id="9DECD0AD21082F50FEBA4EEC6B91FD26" box="[360,459,700,726]" name="China" pageId="35" pageNumber="36">CHINA</collectingCountry>
,
<collectingRegion id="273F5EDF21082F50FE0A4EED686FFD27" box="[472,565,701,727]" country="China" name="Yunnan" pageId="35" pageNumber="36">Yunnan</collectingRegion>
,
<locationDeviation id="F4F6891F21082F50FD914EED68AAFD26" box="[579,752,700,727]" location="Laishkan" pageId="35" pageNumber="36">E of Laishkan</locationDeviation>
,
<locationDeviation id="F4F6891F21082F50FD2C4EEC6E1EFD26" box="[766,1092,700,727]" location="Weixi City" pageId="35" pageNumber="36">
<quantity id="22033DD821082F50FD2C4EEC690BFD26" box="[766,849,700,727]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="36" unit="km" value="4.5">4.5 km</quantity>
ENE of Weixi City
</locationDeviation>
,
<geoCoordinate id="80CFF6FA21082F50FB834EEB6E87FD27" box="[1105,1245,699,727]" degrees="27" direction="north" minutes="12" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="03" value="27.200834">27°12′03″N</geoCoordinate>
<geoCoordinate id="80CFF6FA21082F50FB354EEB6F28FD27" box="[1255,1394,699,727]" degrees="98" direction="east" minutes="20" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="24" value="98.340004">98°20′24″E</geoCoordinate>
,
<quantity id="22033DD821082F50FF6F4E8F6B42FD0A" box="[189,280,735,762]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="35" pageNumber="36" unit="m" value="2580.0">
<elevation id="6ED6770E21082F50FF6F4E8F6B42FD0A" box="[189,280,735,762]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="35" pageNumber="36" unit="m" value="2580.0">2580 m</elevation>
</quantity>
,
<date id="9145B6FD21082F50FEF74E8F6BC2FD09" box="[293,408,735,761]" pageId="35" pageNumber="36" value="2016-05-29">
<collectingDate id="81014F1521082F50FEF74E8F6BC2FD09" box="[293,408,735,761]" pageId="35" pageNumber="36" value="2016-05-29">29.v.2016</collectingDate>
</date>
<collectorName id="480EF5EB21082F50FE724E8F6BB7FD09" box="[416,493,735,761]" pageId="35" pageNumber="36">Kabak</collectorName>
&amp;
<collectorName id="480EF5EB21082F50FDC04E8F6824FD09" box="[530,638,735,761]" pageId="35" pageNumber="36">Davidian</collectorName>
leg.”; ♂
<collectionCode id="83EA08F821082F50FD394EB06946FD0A" box="[747,796,736,762]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="35" pageNumber="36" type="Museum">ZIN</collectionCode>
[The coordinates on the original label are referring to Myanmar, not Yunnan, but Laishkan and Weixi both are towns in northern Yunnan, not Myanmar].
</materialsCitation>
</paragraph>
<paragraph id="E544903D21082F50FF384F1468A9FC71" blockId="35.[189,755,836,898]" pageId="35" pageNumber="36">
<materialsCitation id="55939A6021082F50FF384F1468B4FC71" ID-GBIF-Occurrence="3705075323" collectingDate="2016-05-29" collectionCode="DTC" collectorName="Kabak &amp; Davidian" country="China" county="Laishkan" elevation="2580" latitude="27.200834" location="CHINA" longLatPrecision="20" longitude="98.340004" municipality="Weixi" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="D78F4C2F21082F50FF384F146B06FCAE" bold="true" box="[234,348,836,862]" pageId="35" pageNumber="36">
<typeStatus id="3A402E9F21082F50FF384F146B06FCAE" box="[234,348,836,862]" pageId="35" pageNumber="36" type="paratype">Paratype</typeStatus>
</emphasis>
(1 specimen)
<collectingCountry id="9DECD0AD21082F50FF6F4F376B42FC71" box="[189,280,871,897]" name="China" pageId="35" pageNumber="36">CHINA</collectingCountry>
•
<specimenCount id="F3FD5BB421082F50FEE34F376B3AFC72" box="[305,352,871,898]" pageId="35" pageNumber="36" type="male">1 ♂</specimenCount>
; same label as for holotype;
<collectionCode id="83EA08F821082F50FD624F3768B4FC71" box="[688,750,871,897]" collectionName="Dmitrijs Telnovs" pageId="35" pageNumber="36">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621082F55FF6F4FF86BF1FE46" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36" type="description">
<paragraph id="E544903D21082F50FF6F4FF86B14FC32" blockId="35.[189,334,936,962]" box="[189,334,936,962]" pageId="35" pageNumber="36">
<heading id="BE0C275121082F50FF6F4FF86B14FC32" bold="true" box="[189,334,936,962]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="D78F4C2F21082F50FF6F4FF86B14FC32" bold="true" box="[189,334,936,962]" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21082F50FF6F4F886952FBC7" blockId="35.[189,1400,983,1080]" pageId="35" pageNumber="36">
MEASUREMENTS.
<typeStatus id="3A402E9F21082F50FEAC4F876BB4FC01" box="[382,494,983,1009]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="22033DD821082F50FD1E4F876963FC02" box="[716,825,983,1010]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" pageId="35" pageNumber="36" unit="mm" value="4.15">4.15 mm</quantity>
; head
<quantity id="22033DD821082F50FC544F8769B5FC02" box="[902,1007,983,1010]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="36" unit="mm" value="0.85">0.85 mm</quantity>
long, across eyes
<quantity id="22033DD821082F50FB1B4F876F7EFC02" box="[1225,1316,983,1010]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="mm" value="0.8">0.8 mm</quantity>
broad, pronotum
<quantity id="22033DD821082F50FEEB4FAA6BFEFBE5" box="[313,420,1018,1045]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="35" pageNumber="36" unit="mm" value="1.05">1.05 mm</quantity>
long, maximum width
<quantity id="22033DD821082F50FD6C4FAA6976FBE5" box="[702,812,1018,1045]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="35" pageNumber="36" unit="mm" value="1.15">1.15 mm</quantity>
, minimum width
<quantity id="22033DD821082F50FBD34FAA6E35FBE5" box="[1025,1135,1018,1045]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="36" unit="mm" value="0.35">0.35 mm</quantity>
, elytra
<quantity id="22033DD821082F50FB1A4FAA6F69FBE5" box="[1224,1331,1018,1045]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="35" pageNumber="36" unit="mm" value="2.25">2.25 mm</quantity>
long,
<quantity id="22033DD821082F50FF6D484E6B43FBC8" box="[191,281,1054,1080]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="35" pageNumber="36" unit="mm" value="1.4">1.4 mm</quantity>
combined wide.
<typeStatus id="3A402E9F21082F50FE31484E6810FBC8" box="[483,586,1054,1080]" pageId="35" pageNumber="36" type="paratype">Paratype</typeStatus>
is
<quantity id="22033DD821082F50FDBE484E689CFBC8" box="[620,710,1054,1080]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="35" pageNumber="36" unit="mm" value="4.1">4.1 mm</quantity>
long.
</paragraph>
<paragraph id="E544903D21082F55FF6F480F6E91FEB7" blockId="35.[188,1401,1119,1990]" lastBlockId="38.[189,1398,265,327]" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36">
Dorsum and venter uniformly brown, sutural area of elytra slightly paler. Mouthparts, antennae, palps and legs yellowish-brown. Head elongate ovoid with moderately large, ovoid compound eyes, which are not protruding beyond head outline laterally. Head rounded in broad arc posterior to eyes. Head dorsal punctures minute and inconspicuous. Head dorsal setae inconspicuous, sparse. Antennae conspicuously robust and long, extending towards anterior third of elytra. Antennomere three about 1.3 × as long as antennomere two, antennomeres 8–10 transverse, of them 9–10 strongly transverse. Terminal antennomere asymmetrically triangular with rounded apex, twice as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum with anterior lobe significantly wider than posterior and wider than head across eyes. Postmedian lateral constriction broad and medially shortly notched. Front margin of anterior lobe subtruncate, dorsally with narrow anterior rim. Anterior lobe large and broad, inverted triangular; posterior lobe shorter and much narrower. Front margin of anterior lobe with short, median, anteriad-directed projection and shallow antero-dorsal impression posterior to it; projection is dorsally provided with two erect (straight) and two ך-shaped (bent), anteriad-directed setae (
<figureCitation id="7DC08CB821082F50FEDB4A786BC2F9B3" box="[265,408,1576,1603]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[223,1371,283,1890]" captionTargetId="figure-20@36.[203,1385,263,1912]" captionTargetPageId="36" captionText="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="35" pageNumber="36">Fig. 20B–C</figureCitation>
). Anterior and posterior lobe convex in lateral view (
<figureCitation id="7DC08CB821082F50FBDC4A786E26F9B3" box="[1038,1148,1576,1603]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[223,1371,283,1890]" captionTargetId="figure-20@36.[203,1385,263,1912]" captionTargetPageId="36" captionText="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="35" pageNumber="36">Fig. 20C</figureCitation>
). Lateral constriction continues onto disc in lateral view, shallow (
<figureCitation id="7DC08CB821082F50FD1E4A1C6963F996" box="[716,825,1612,1638]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[223,1371,283,1890]" captionTargetId="figure-20@36.[203,1385,263,1912]" captionTargetPageId="36" captionText="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="35" pageNumber="36">Fig. 20C</figureCitation>
). Lateral pronotal fovea broad and deep at lower external margin of pronotum, strongly widens upwards towards pronotal disc in lateral view, external margins protruding into a pair of obtuse (in lateral view), widely separated denticles (in lateral view), which are provided with long, sparse setae. Cavity in lateral wall of pronotum between lateral denticles large, elliptical. In dorsal view, lateral pronotal fovea narrow, anterior and posterior pair of denticles appear obtusely angulate, glabrous (
<figureCitation id="7DC08CB821082F50FDBE4AAC6880F8E6" box="[620,730,1788,1814]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[223,1371,283,1890]" captionTargetId="figure-20@36.[203,1385,263,1912]" captionTargetPageId="36" captionText="Fig. 20. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="35" pageNumber="36">Fig. 20B</figureCitation>
). Pronotal punctures minute on disc; lateral constriction dorsally with sparse, rather small punctures. Dorsal pronotal setae inconspicuous, moderately long. Scutellar shield minute, triangular. Elytra dorsally elliptical, slightly convex in lateral view, widened laterally around midlength, lateral margins evenly broadly rounded, shoulders obsolete (apterous species). Elytral punctures rather small and moderately dense, becoming slightly smaller and sparser towards apices. Elytral setae long and sparse, suberect. Male tergite and morphological sternite VII broadly rounded at posterior margin. Aedeagus as in
<figureCitation id="7DC08CB8210D2F55FC9D4D5969F0FED3" box="[847,938,265,291]" captionStart="Fig" captionStartId="37.[189,232,1928,1954]" captionTargetBox="[369,1231,278,1870]" captionTargetId="figure-11@37.[352,1235,265,1887]" captionTargetPageId="37" captionText="Fig. 21. Macrotomoderus hengduan sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions. C. Gonopore armature, median portion of apicale. D. Ditto, apical portion of apicale. E. Basal portion of apicale. F. Ditto, stronger magnification. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299533" httpUri="https://zenodo.org/record/6299533/files/figure.png" pageId="38" pageNumber="39">Fig. 21</figureCitation>
, large and dagger-shaped, with short, spherical basale and inconspicuous, delicate, irregularly filamentous gonopore armature.
</paragraph>
<caption id="B184C0B5210F2F57FF6F4BFB6E63F818" ID-DOI="http://doi.org/10.5281/zenodo.6299531" ID-Zenodo-Dep="6299531" httpUri="https://zenodo.org/record/6299531/files/figure.png" pageId="36" pageNumber="37" startId="36.[189,232,1963,1989]" targetBox="[223,1371,283,1890]" targetPageId="36">
<paragraph id="E544903D210F2F57FF6F4BFB6E63F818" blockId="36.[189,1398,1963,2025]" pageId="36" pageNumber="37">
<emphasis id="D78F4C2F210F2F57FF6F4BFB6B41F835" bold="true" box="[189,283,1963,1989]" pageId="36" pageNumber="37">Fig. 20.</emphasis>
<taxonomicName id="22FBEBBE210F2F57FEF04BFB683CF835" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[290,614,1963,1989]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F210F2F57FEF04BFB683CF835" box="[290,614,1963,1989]" italics="true" pageId="36" pageNumber="37">Macrotomoderus hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF154210F2F57FDBF4BFC6890F836" box="[621,714,1964,1990]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN).
<emphasis id="D78F4C2F210F2F57FC6E4BFB698BF835" bold="true" box="[956,977,1963,1989]" pageId="36" pageNumber="37">A</emphasis>
. Habitus, dorsal view.
<emphasis id="D78F4C2F210F2F57FB0C4BFB6EA8F835" bold="true" box="[1246,1266,1963,1989]" pageId="36" pageNumber="37">B</emphasis>
. Forebody, dorsal view.
<emphasis id="D78F4C2F210F2F57FE804B9E6B3DF818" bold="true" box="[338,359,1998,2024]" pageId="36" pageNumber="37">C</emphasis>
. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale.
</paragraph>
</caption>
<caption id="B184C0B5210E2F56FF6F4BD869EBF818" ID-DOI="http://doi.org/10.5281/zenodo.6299533" ID-Zenodo-Dep="6299533" httpUri="https://zenodo.org/record/6299533/files/figure.png" pageId="37" pageNumber="38" startId="37.[189,232,1928,1954]" targetBox="[369,1231,278,1870]" targetPageId="37">
<paragraph id="E544903D210E2F56FF6F4BD869EBF818" blockId="37.[187,1398,1928,2025]" pageId="37" pageNumber="38">
<emphasis id="D78F4C2F210E2F56FF6F4BD86B43F852" bold="true" box="[189,281,1928,1954]" pageId="37" pageNumber="38">Fig. 21.</emphasis>
<taxonomicName id="22FBEBBE210E2F56FECD4BD8683BF852" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[287,609,1928,1954]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F210E2F56FECD4BD8683BF852" box="[287,609,1928,1954]" italics="true" pageId="37" pageNumber="38">Macrotomoderus hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF154210E2F56FDB54BD86898F852" box="[615,706,1928,1954]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN), aedeagus.
<emphasis id="D78F4C2F210E2F56FBF44BD86E04F852" bold="true" box="[1062,1118,1928,1954]" pageId="37" pageNumber="38">A–B</emphasis>
. General view, different positions.
<emphasis id="D78F4C2F210E2F56FEE74BFB6B10F835" bold="true" box="[309,330,1963,1989]" pageId="37" pageNumber="38">C</emphasis>
. Gonopore armature, median portion of apicale.
<emphasis id="D78F4C2F210E2F56FCAD4BFB69CEF835" bold="true" box="[895,916,1963,1989]" pageId="37" pageNumber="38">D</emphasis>
. Ditto, apical portion of apicale.
<emphasis id="D78F4C2F210E2F56FAC14BFB6F7DF835" bold="true" box="[1299,1319,1963,1989]" pageId="37" pageNumber="38">E</emphasis>
. Basal portion of apicale.
<emphasis id="D78F4C2F210E2F56FE494B9E6BF7F818" bold="true" box="[411,429,1998,2024]" pageId="37" pageNumber="38">F</emphasis>
. Ditto, stronger magnification. Not to scale.
</paragraph>
</caption>
<paragraph id="E544903D210D2F55FF6F4D3D6BEAFE77" blockId="38.[189,432,365,391]" box="[189,432,365,391]" pageId="38" pageNumber="39">
<heading id="BE0C2751210D2F55FF6F4D3D6BEAFE77" bold="true" box="[189,432,365,391]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="D78F4C2F210D2F55FF6F4D3D6BEAFE77" bold="true" box="[189,432,365,391]" pageId="38" pageNumber="39">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D210D2F55FF6F4DCC6BF1FE46" blockId="38.[189,427,412,438]" box="[189,427,412,438]" pageId="38" pageNumber="39">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B6210D2F55FF6F4D8E687FFDD6" box="[189,549,478,551]" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="E544903D210D2F55FF6F4D8E6B7AFE08" blockId="38.[189,288,478,504]" box="[189,288,478,504]" pageId="38" pageNumber="39">
<heading id="BE0C2751210D2F55FF6F4D8E6B7AFE08" bold="true" box="[189,288,478,504]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="D78F4C2F210D2F55FF6F4D8E6B7AFE08" bold="true" box="[189,288,478,504]" pageId="38" pageNumber="39">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D210D2F55FF6F4E5C687FFDD6" blockId="38.[189,549,524,551]" box="[189,549,524,551]" pageId="38" pageNumber="39">
Collected at
<quantity id="22033DD8210D2F55FE834E5C6BF0FDD7" box="[337,426,524,551]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="38" pageNumber="39" unit="m" value="2580.0">2580 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B6210D2F55FF6F4E1D6EE3FD66" box="[189,1209,589,663]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="E544903D210D2F55FF6F4E1D6B0CFD97" blockId="38.[189,342,589,615]" box="[189,342,589,615]" pageId="38" pageNumber="39">
<heading id="BE0C2751210D2F55FF6F4E1D6B0CFD97" bold="true" box="[189,342,589,615]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="D78F4C2F210D2F55FF6F4E1D6B0CFD97" bold="true" box="[189,342,589,615]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D210D2F55FF6F4E2D6EE3FD66" blockId="38.[189,1209,636,663]" box="[189,1209,636,663]" pageId="38" pageNumber="39">
Known only from Hengduan Mountains, northern part of
<collectingRegion id="273F5EDF210D2F55FCB34E2D6E77FD66" box="[865,1069,636,663]" country="China" name="Yunnan" pageId="38" pageNumber="39">Yunnan Province</collectingRegion>
, SW
<collectingCountry id="9DECD0AD210D2F55FBBE4E2C6EECFD66" box="[1132,1206,636,662]" name="China" pageId="38" pageNumber="39">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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