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<document ID-CLB-Dataset="3093" ID-DOI="http://dx.doi.org/10.3897/phytokeys.186.62018" ID-GBIF-Dataset="8becf6a0-f9c4-495b-9620-2040bd691f43" ID-PMC="PMC8671706" ID-Pensoft-Pub="1314-2003-186-93" ID-Pensoft-UUID="51C402FAC1635A0FB18EA14A1C2FE5F8" ID-PubMed="34975277" ModsDocID="1314-2003-186-93" checkinTime="1638850659404" checkinUser="pensoft" docAuthor="Kiew, Ruth &amp; Kamin, Imin" docDate="2021" docId="10D126E50F2E5F0EAB2D9DDB104E1929" docLanguage="en" docName="PhytoKeys 186: 93-110" docOrigin="PhytoKeys 186" docPubDate="2021-12-06" docSource="http://dx.doi.org/10.3897/phytokeys.186.62018" docTitle="Coleus scutellarioides (L.) Benth." docType="treatment" docVersion="6" id="51C402FAC1635A0FB18EA14A1C2FE5F8" lastPageNumber="93" masterDocId="51C402FAC1635A0FB18EA14A1C2FE5F8" masterDocTitle="Coleus (Lamiaceae) in Peninsular Malaysia including two new species" masterLastPageNumber="110" masterPageNumber="93" pageNumber="93" updateTime="1732787246147" updateUser="ExternalLinkService">
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<mods:title id="0A086854E37F016F33FD660B520A2FE3">Coleus (Lamiaceae) in Peninsular Malaysia including two new species</mods:title>
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<mods:namePart id="8316922AAFA0E7EA37BA87B25221CEDC">Kiew, Ruth</mods:namePart>
<mods:affiliation id="18645D0E264F5F99D1953B5FF4BB32D5">Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
<mods:nameIdentifier id="F6F663DA503028E6F01285328D6218C3" type="email">ruth@frim.gov.my</mods:nameIdentifier>
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<mods:namePart id="45526BE94B7B710514A7B853BDE13CB4">Kamin, Imin</mods:namePart>
<mods:affiliation id="99CD4E9A452719365263E8FA6EE02833">Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
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<mods:title id="EADFB4784B9BC1952DA8ABF3795B81D3">PhytoKeys</mods:title>
</mods:titleInfo>
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<mods:date id="FF1EBA9BDD9C8CC3D9046782891AB9FF">2021</mods:date>
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<mods:number id="0BEC3C98D193AACB58B5F24FA55A10F3">2021-12-06</mods:number>
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<mods:number id="5A5946BC34EA8EDE4D5D11CB1A4D3AE0">186</mods:number>
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<mods:start id="D4776144EA44C67D6088CF080E0E358E">93</mods:start>
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<mods:classification id="50707DA60D98B5BCA6D22AD37036FD7A">journal article</mods:classification>
<mods:identifier id="14C0D7E4ADDD0AA8607B35F09FBDCC2F" type="DOI">http://dx.doi.org/10.3897/phytokeys.186.62018</mods:identifier>
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<subSection id="4AA292CEBCD54C59002960822FC4B422" pageId="0" pageNumber="93" type="taxonomy">
<treatment id="10D126E50F2E5F0EAB2D9DDB104E1929" ID-GBIF-Taxon="191162308" LSID="urn:lsid:plazi:treatment:10D126E50F2E5F0EAB2D9DDB104E1929" httpUri="http://treatment.plazi.org/id/10D126E50F2E5F0EAB2D9DDB104E1929" lastPageNumber="93" pageId="0" pageNumber="93" scope_family="Lamiaceae">
<subSubSection id="51B9D4DD32C7AFEED296BF9F5421248F" pageId="0" pageNumber="93" type="nomenclature">
<paragraph id="F1A6610E0BD2971F15B8C40337AE5FB5" pageId="0" pageNumber="93">
5.
<taxonomicName id="30254F68CFD4D10FD5D3968E704DED80" ID-CoL="5ZLNT" LSID="10D126E5-0F2E-5F0E-AB2D-9DDB104E1929" authority="(L.) Benth." authorityName="(L.) Benth." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">Coleus scutellarioides (L.) Benth.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="70F72D094AF30236F62FC25B1DF9623F" pageId="0" pageNumber="93" type="reference_group">
<paragraph id="B005BAAE5CAA5F548DE37FC14AE99FA6" pageId="0" pageNumber="93">
<taxonomicName id="C1DFC72585E09F6E42B725F85A9074C6" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">Coleus scutellarioides</taxonomicName>
<emphasis id="EF3719227C622C50F103B37A16BD78DA" italics="true" pageId="0" pageNumber="93">in</emphasis>
Wall., Pl. Asiat. Rar. 2: 15 (1830); Keng, Gard. Bull. Singapore 24: 51 (1969).
<taxonomicName id="510107230379638266427D2809F87A6A" authority="var. scutellarioides" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus scutellarioides subsp. var. var. scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="variety" species="scutellarioides" subSpecies="var." variety="scutellarioides">Coleus scutellarioides var. scutellarioides</taxonomicName>
Keng, Gard. Bull. Singapore 24: 53 (1969). Basionym:
<taxonomicName id="F4167669709B8AEB67D6D65F36DECE8A" class="Magnoliopsida" family="Lamiaceae" genus="Ocimum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ocimum scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">Ocimum scutellarioides</taxonomicName>
L., Sp. Pl. ed. 2, 2: 834 (1763) (as
<taxonomicName id="19389A5152B8D3BB08B249D708BDB8B6" class="Magnoliopsida" family="Lamiaceae" genus="Ocymum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ocymum" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="genus">Ocymum</taxonomicName>
). Type: Majana (alba et rubra)
<emphasis id="1649B98DA73C6C1DE1E09301D2697D3E" italics="true" pageId="0" pageNumber="93">in</emphasis>
Rumphius, Herb. Amboin. 5: 291, t. 101 (1747), (lectotype, designated by Merrill, Interpr. Rumph. Herb. Amboin.: 460 (1917).) Homotypic synonyms:
<taxonomicName id="564DA1F288AFADE013C4111666F5D69B" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">Plectranthus scutellarioides</taxonomicName>
(L.) R.Br., Prodr. Fl. Nov. Holl.: 506 (1810); Blume, Bijdr. Fl. Ned. Ind.: 837 (1826); Keng, Fl. Malesiana Ser. I, 8: 389 (1978); Suddee et al., Kew Bull. 59: 410 (2004); Kiew in
<normalizedToken id="FD4FB882A3D99005FA52AF4C9A524915" originalValue="Henderson’s">Henderson's</normalizedToken>
Malaysian Wild Flowers Dicotyledons: 298 (2014); Bramley, Fl. Malesiana Ser. I, 23: 296 (2019).
<taxonomicName id="FF33EB0DCA23889A91DF2AA350ED3C28" class="Magnoliopsida" family="Lamiaceae" genus="Solenostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solenostemon scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">Solenostemon scutellarioides</taxonomicName>
(L.) Codd, Bothalia 11: 439 (1975); Turner, Gard. Bull. Singapore 47: 273 (1996
<normalizedToken id="D2B05824DA3939DC37E3EF4142279158" originalValue="‘1995’">'1995'</normalizedToken>
). Heterotypic synonyms:
<taxonomicName id="3541486BADF980BCB47E2F036478B7B0" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus atropurpureus" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="atropurpureus">Coleus atropurpureus</taxonomicName>
Benth.
<emphasis id="5F21510DEA18F36F518AE25599D654F4" italics="true" pageId="0" pageNumber="93">in</emphasis>
Wall., Pl. Asiat. Rar. 2: 16 (1830); Prain, J. Asiat. Soc. Beng. 74, 2: 706 (1907); Ridley, Fl. Mal. Penins. 2: 646 (1923). Syntypes: Singapore
<emphasis id="6BEE16BD8D71C9D09C9CBF5399706143" italics="true" pageId="0" pageNumber="93">Wall. Cat. 2733A</emphasis>
(syntypes K!, K-W!).
<taxonomicName id="04EDF57AC2BB3EBEF2981B9CD054EB1B" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus blumei" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="blumei">Coleus blumei</taxonomicName>
Benth., Labiat. Gen. Sp.: 56 (1832). Type: Java,
<emphasis id="4C334DF685B1EF709AADB6C0BE4E63CE" italics="true" pageId="0" pageNumber="93">Blume s.n.</emphasis>
(lectotype L, designated by Suddee et al., Kew Bull. 59: 410 (2004).)
</paragraph>
</subSubSection>
<subSubSection id="8B0E889A5B1899C80B8FD81FB6D38A66" pageId="0" pageNumber="93" type="notes">
<paragraph id="EBA077797CA81591155DD34C9746CA6F" pageId="0" pageNumber="93">Note.</paragraph>
<paragraph id="5CA46949082D588F0D63C8869FCCCC2C" pageId="0" pageNumber="93">
Here are included only references that cite Malaysian specimens. For synonomy under
<taxonomicName id="F19450320A756D53C0F0FFC156F67D7E" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="E6EB46909A7FA4783932FD7D640A0828" italics="true" pageId="0" pageNumber="93">Coleus</emphasis>
</taxonomicName>
see Paton
<emphasis id="4A29F5BE170774625D823CD66E3601DC" italics="true" pageId="0" pageNumber="93">et al</emphasis>
. Phytokeys 129 (2019) 1-158, for
<taxonomicName id="E628D43D3224349562D07F4E12863CAF" genus="Plectanthus" lsidName="Plectanthus" pageId="0" pageNumber="93" rank="genus">
<emphasis id="8074F707979FEA949008712B44A1064B" italics="true" pageId="0" pageNumber="93">Plectanthus</emphasis>
</taxonomicName>
for continental South East Asia see
<bibRefCitation id="E794A89EAF1879A1280D6C65A30504D7" DOI="https://doi.org/10.2307/4110950" author="Suddee, S" journalOrPublisher="Flora of Malay Peninsula" pageId="0" pageNumber="93" refId="B14" refString="Suddee, S, Paton, AJ, Parnell, JAN, 2004. A taxonomic revision of the tribe Ocimeae Dumort. (Lamiaceae) in continental South East Asia. II. Plectranthinae. Kew Bulletin 59(3): 379-414. https://doi.org/10.2307/4110950" title="A taxonomic revision of the tribe Ocimeae Dumort. (Lamiaceae) in continental South East Asia. II. Plectranthinae. Kew Bulletin 59 (3): 379 - 414." url="https://doi.org/10.2307/4110950" year="2004">Suddee et al. (2004)</bibRefCitation>
and for Malesia see
<bibRefCitation id="B47279724F6BE26DA24FEA1498C9EE17" author="Bramley, GLC" journalOrPublisher="I," pageId="0" pageNumber="93" pagination="287 - 302" refId="B1" refString="Bramley, GLC, 2019. Plectranthus. Lamiaceae. Fl. Malesiana Ser. I, 23: 287 - 302" title="Plectranthus. Lamiaceae. Fl. Malesiana Ser." volume="23" year="2019">Bramley (2019)</bibRefCitation>
.
</paragraph>
<paragraph id="B316C9FB2056534F9EE7A4507897D86C" pageId="0" pageNumber="93">The description provided here refers only to wild plants from Peninsular Malaysia.</paragraph>
</subSubSection>
<subSubSection id="C46233E2DF1CA0D17B9705B0C8A1AA3E" pageId="0" pageNumber="93" type="description">
<paragraph id="BF0FB4F1D0E1DEAE0B67349906FD45B5" pageId="0" pageNumber="93">Description.</paragraph>
<paragraph id="A29166188A5EF8CC5546655F2E9EBD23" pageId="0" pageNumber="93">
Erect or ascending sparsely branched herb, 30-60 cm tall, aromatic, without tubers.
<emphasis id="824767D29FEFFE3BB1B19488C07BBE54" bold="true" italics="true" pageId="0" pageNumber="93">Stem</emphasis>
and branches finely pubescent to glabrous.
<emphasis id="B3466CAE57DBA5F5BAAAD8AB64F9C9A1" bold="true" italics="true" pageId="0" pageNumber="93">Leaves</emphasis>
with petiole 1-5 cm; lamina ovate, (2.5-)4-7.5
<normalizedToken id="A88F7C5D36545ECA776AA240007E6F7C" originalValue="×">x</normalizedToken>
(2-)3-4 cm, plain dark green, base rounded or sometimes truncate, margin crenate with rounded teeth, apex acute, tip rounded, upper surface glabrous or with short hairs, lower surface pubescent on the main and secondary veins.
<emphasis id="4949AE23D953A070D08A0A68A5B1CB3F" bold="true" italics="true" pageId="0" pageNumber="93">Inflorescences</emphasis>
terminal, unbranched spike or sometimes branched at base, to 40 cm long, flowers in few- to many-flowered verticils or in irregularly branched cymes, peduncles of the lateral cymes short or elongated. Bracts narrowly ovate to ovate, apex acute, pubescent, 2-8 mm long, caducous. Pedicels 1-2 mm long in flower, extending in fruit.
<emphasis id="EFE67415B3CDF1D53444D0A7465BD2E1" bold="true" italics="true" pageId="0" pageNumber="93">Flower</emphasis>
:
<emphasis id="DE4B2526043E3DD44CDAD68F24DD2A46" italics="true" pageId="0" pageNumber="93">calyx</emphasis>
campanulate, 2-3 mm long, in fruit 4-8 mm, minutely pubescent; posterior lobe broadly ovate, subacute, recurved and erect, two lateral lobes of anterior lip short, oblong-obtuse, truncate or rounded, rarely mucronate with a tiny apiculate apex, about half as long as central lobes of anterior lip, these oblong to subulate, connate for two thirds their length, acuminate at apex, longer than posterior lobe, anterior lobe divided into 2 points;
<emphasis id="6F37DA0679014752263AB128801D99BB" italics="true" pageId="0" pageNumber="93">corolla</emphasis>
8(-13)-16 mm long, tube abruptly decurved above the calyx, white, 4-5, ca. 7.5 mm long, dilating widely to the throat, with scattered hairs; posterior lip with lobes pubescent, the two central lobes rounded, anterior lip deep or rosy violet, 4-5(-9) mm long;
<emphasis id="4325AA0EB31AD592F1999D5DD0819665" italics="true" pageId="0" pageNumber="93">stamens</emphasis>
not exserted from anterior lip.
<emphasis id="6ACEBC0492258993635A752C411020AE" bold="true" italics="true" pageId="0" pageNumber="93">Nutlets</emphasis>
broadly ovate or orbicular, brown to black, shining, 1-1.2 mm long, minutely tuberculate, mucilaginous when wet.
</paragraph>
</subSubSection>
<subSubSection id="678F0110CB5242D86F85608E7CD58799" pageId="0" pageNumber="93" type="notes">
<paragraph id="603C4578DE17E3358FA9216443357E4C" pageId="0" pageNumber="93">Note.</paragraph>
<paragraph id="E54D1B9A6268D18F71279D2EE9455930" pageId="0" pageNumber="93">
<bibRefCitation id="CB5A136D3C28AF2B115F8EE6BDECE798" DOI="https://doi.org/10.1136/bmj.2.3276.646" author="Ridley, HN" journalOrPublisher="Flora of Malay Peninsula" pageId="0" pageNumber="93" pagination="646 - 647" refId="B12" refString="Ridley, HN, 1923. Coleus. Flora of Malay Peninsula 2: 646 - 647, DOI: https://doi.org/10.1136/bmj.2.3276.646" title="Coleus." url="https://doi.org/10.1136/bmj.2.3276.646" volume="2" year="1923">Ridley (1923)</bibRefCitation>
and
<bibRefCitation id="0C7770654E60BA9A866AE1C9067A8B6A" author="Burkill, IH" journalOrPublisher="I," pageId="0" pageNumber="93" refId="B2" refString="Burkill, IH, 1966. Coleus. Dictionary of Economic Products of Malay Peninsula (2nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642-645." title="Coleus. Dictionary of Economic Products of Malay Peninsula (2 nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642 - 645." year="1966">Burkill (1966)</bibRefCitation>
considered the coleus of gardens (i.e.
<taxonomicName id="0946E4F8335B68E670CCC6857B34A2FD" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus blumei" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="blumei">
<emphasis id="8AF2A790E359F67D70AFFF0D981D796F" italics="true" pageId="0" pageNumber="93">Coleus blumei</emphasis>
</taxonomicName>
) was introduced from Java and was distinct from Malayan
<taxonomicName id="B27DAC41F15291AE4CE05A665A08AF72" lsidName="C. atropurpurea" pageId="0" pageNumber="93" rank="species" species="atropurpurea">
<emphasis id="2C16633F950ABCC7D6C37B61C333D7EF" italics="true" pageId="0" pageNumber="93">C. atropurpurea</emphasis>
</taxonomicName>
. These two species are recognized as conspecific and are currently known as
<taxonomicName id="760F4A0367C042494E4B1A7A3934A224" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus scutellarioides" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="scutellarioides">
<emphasis id="CF06F97EF73B79F7F77E661A143034B9" italics="true" pageId="0" pageNumber="93">Coleus scutellarioides</emphasis>
</taxonomicName>
(
<bibRefCitation id="082FCAA9DF338A62889A869D3903EC14" DOI="https://doi.org/10.3897/phytokeys.129.34988" author="Paton, AJ" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="93" pagination="1 - 158" refId="B11" refString="Paton, AJ, Mwanyambo, M, Govaerts, RHA, Smith, K, Suddee, S, Phillipson, PB, Wilson, TC, Forster, PI, Culham, A, 2019. Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): A tale of more than two genera. PhytoKeys 129: 1 - 158, DOI: https://doi.org/10.3897/phytokeys.129.34988" title="Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): A tale of more than two genera." url="https://doi.org/10.3897/phytokeys.129.34988" volume="129" year="2019">Paton et al. (2019)</bibRefCitation>
with the result that the description has expanded to include the huge range in leaf size, shape and colour of the ornamental forms that are not seen in the wild form of Malaysian population. The description above is based on the wild form that has smaller, plain green leaves. Unlike
<taxonomicName id="5A0F9E34CB310D03EE5639B41EF596B7" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" lsidName="Scutellaria discolor" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="172DDDD1B8CE3A8BDB27731F3B240EAD" italics="true" pageId="0" pageNumber="93">Scutellaria discolor</emphasis>
</taxonomicName>
Wall. ex Benth., it is not found deep in forest and is seldom found in flower. That it is usually associated with forest edge, often close to habitation, suggests that many of these populations have been established for their medicinal properties.
</paragraph>
<paragraph id="84E625DB05EBF5E64EA1729C50EDA372" pageId="0" pageNumber="93">
On account of it being extremely variable,
<bibRefCitation id="EE0CC56ABD5E6917C20C359EE0F4852D" author="Keng, H" journalOrPublisher="Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="93" refId="B6" refString="Keng, H, 1978. Plectranthus. Flora Malesiana Ser. I.8: 382-393." title="Plectranthus. Flora Malesiana Ser. I. 8: 382 - 393." year="1978">Keng (1978)</bibRefCitation>
and
<bibRefCitation id="9B1810509559E7E15FAFDD084663635E" author="Bramley, GLC" journalOrPublisher="I," pageId="0" pageNumber="93" pagination="287 - 302" refId="B1" refString="Bramley, GLC, 2019. Plectranthus. Lamiaceae. Fl. Malesiana Ser. I, 23: 287 - 302" title="Plectranthus. Lamiaceae. Fl. Malesiana Ser." volume="23" year="2019">Bramley (2019)</bibRefCitation>
did not recognise subspecific taxa, there being too many intermediate specimens. A view that is followed here.
</paragraph>
</subSubSection>
<subSubSection id="8239174E5E9EC6F5EE4C8DC202954494" pageId="0" pageNumber="93" type="distribution">
<paragraph id="E0E2AB89C960FA9669DCE55E5B020171" pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph id="4BA1496C4B4D9FAC6142CE20E613EE84" pageId="0" pageNumber="93">SE Asia (India, Myanmar, Thailand, Indo-China, S. China, Taiwan, throughout Malesia to Australia, Melanesia (Solomon Is.) and Polynesia. In Peninsular Malaysia, widespread (Johor, Melaka, Penang, Perak, Pahang and Selangor).</paragraph>
</subSubSection>
<subSubSection id="5B9688D6308521D95B7BDFB326E71D9D" pageId="0" pageNumber="93" type="provisional conservation status">
<paragraph id="F1D212A83E9A0D3C3EB2F44DBE3945DE" pageId="0" pageNumber="93">Provisional conservation status.</paragraph>
<paragraph id="0F27451560DE05C8DDD617308AE6410A" pageId="0" pageNumber="93">Least Concern. (Assessed by A.R. Rafidah).</paragraph>
</subSubSection>
<subSubSection id="CD20C151AD9DF1680395D6D330AB93E7" pageId="0" pageNumber="93" type="ecology">
<paragraph id="1B261664B4101D408BDF521AE02CA6B7" pageId="0" pageNumber="93">Ecology.</paragraph>
<paragraph id="811A9E606B1609CD5F570D69E7602A18" pageId="0" pageNumber="93">
In Peninsular Malaysia, from forest edge and often near villages throughout the lowlands to 350 m elevation (
<bibRefCitation id="4EDD6C27491594C66E43B97A85DBA3C8" author="Kiew, R" journalOrPublisher="Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="93" refId="B7" refString="Kiew, R, 2014. Labiatae. Henderson's Malaysian Wild Flowers Dicotyledons. Malaysian Nature Society, Kuala Lumpur, 292-300." title="Labiatae. Henderson's Malaysian Wild Flowers Dicotyledons. Malaysian Nature Society, Kuala Lumpur, 292 - 300." year="2014">Kiew 2014</bibRefCitation>
), sometimes in forest by streams but not from limestone hills as reported by
<bibRefCitation id="4E95D987057B6AE11AF99B43BCA8462B" author="Keng, H" journalOrPublisher="Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="93" refId="B6" refString="Keng, H, 1978. Plectranthus. Flora Malesiana Ser. I.8: 382-393." title="Plectranthus. Flora Malesiana Ser. I. 8: 382 - 393." year="1978">Keng (1978)</bibRefCitation>
. That the forest localities are often close to villages and it has medicinal uses suggests these populations may have been planted, particularly because this species seldom flowers but can easily be propagated by stem cuttings.
</paragraph>
</subSubSection>
<subSubSection id="4599DE6244CC06EDDDD6315455AA0489" pageId="0" pageNumber="93" type="etymology">
<paragraph id="4E68DB15F84006EE4F1B08742AAC4D5B" pageId="0" pageNumber="93">Etymology.</paragraph>
<paragraph id="5C0CE96A8A424B202D756E73C895F32A" pageId="0" pageNumber="93">
Greek -
<emphasis id="05C418B2096C260469E3D764E77D93D3" italics="true" pageId="0" pageNumber="93">oides</emphasis>
= resembling;
<taxonomicName id="0530C9E86C6AF3D1C5CE8358D69E2556" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" lsidName="Scutellaria" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="0FE032E763C7DFF2BC4256C9D4813D29" italics="true" pageId="0" pageNumber="93">Scutellaria</emphasis>
</taxonomicName>
L., a genus in
<taxonomicName id="832F198683238881D2637510B917441B" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2382B0D5565B117F0FCB9A7B450B92D8" pageId="0" pageNumber="93" type="uses">
<paragraph id="C8EDECF2D4A9EB7F20CE13D46A9A1965" pageId="0" pageNumber="93">Uses.</paragraph>
<paragraph id="5C17B99E1E52FF365E4A8E7C8DCF1F05" pageId="0" pageNumber="93">
In Peninsular Malaysia, it has always been commonly known as
<normalizedToken id="64B13D290F72DF26B0708F16F1FFB23F" originalValue="‘coleus’">'coleus'</normalizedToken>
and in Malay as
<emphasis id="49F1610D7325F07680FF3CF5F347B30D" italics="true" pageId="0" pageNumber="93">pokok ati-ati</emphasis>
. Cultivated forms are popular ornamental plants and come in a great variety of multicoloured, variegated foliage. In Cameron Highlands, they are grown in great quantities for home gardens. The colour of plants grown in the highlands, ca. 2000 m, is more vivid than those grown in the lowlands.
</paragraph>
<paragraph id="2890B40C74A55CC362A17DC2E58CC056" pageId="0" pageNumber="93">
It is a minor medicinal plant used to cure a wide range of ailments.
<bibRefCitation id="BEEC3E182747897E7B98E7D907D0F623" author="Burkill, IH" journalOrPublisher="I," pageId="0" pageNumber="93" refId="B2" refString="Burkill, IH, 1966. Coleus. Dictionary of Economic Products of Malay Peninsula (2nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642-645." title="Coleus. Dictionary of Economic Products of Malay Peninsula (2 nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642 - 645." year="1966">Burkill (1966</bibRefCitation>
, pg. 643) reported it was a remedy for heart disease, heart burn, inflammation of heart; sensitive skin, stimulates digestion and for congestion of the liver that causes swellings of the hands and feet, amongst other ailments. Among the aboriginal population, the Besis people (now more commonly known as the Mah Meri) call it
<emphasis id="FD7BBD57087CB697004E35B7A15A8260" italics="true" pageId="0" pageNumber="93">torek</emphasis>
, and plant it around their graves. They also traditionally use it as the brush used for sprinkling holy rice-gruel over a new clearing (
<bibRefCitation id="D79FDB49BE564F3B4C2DBF97A195C33D" author="Burkill, IH" journalOrPublisher="I," pageId="0" pageNumber="93" refId="B2" refString="Burkill, IH, 1966. Coleus. Dictionary of Economic Products of Malay Peninsula (2nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642-645." title="Coleus. Dictionary of Economic Products of Malay Peninsula (2 nd ed.). Ministry of Agriculture and Co-operatives, Malaysia, 642 - 645." year="1966">Burkill 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2C29244FEDF135152AB30A8468EF0D7B" pageId="0" pageNumber="93" type="materials_examined">
<paragraph id="B181256895CB8E1C79312F3AF70443E6" pageId="0" pageNumber="93">
Peninsular
<collectingCountry id="40AFE5A1BD046F1E0AF7EF206A287A48" name="Malaysia">Malaysia</collectingCountry>
specimens examined
</paragraph>
<paragraph id="282068ADE1C6D0198E268121FABE43D3" pageId="0" pageNumber="93">
(* indicates specimens collected in villages that are presumed to be cultivated):
<materialsCitation id="BB0338C45113E552FAAA8FCEA6BFAFF3" accessionNumber="SFN 16536" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Pulau Aor Fielding, Griffith, Burkill, Bentong Burkill, Md Haniff" location="Johor" specimenCount="1" stateProvince="Johor">
<emphasis id="EC6FC279CE87403CCD51EAC2656019A7" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="23D1704B9AA975794C0BDC86D9BDDBA2" country="Malaysia" name="Johor">Johor</collectingRegion>
</emphasis>
:
<collectorName id="F76DF7EC0AB160C3FEFD2FEABCC56CFD">
Pulau Aor
<emphasis id="5F4302481A55C081D1DAEA878DAC87AD" italics="true" pageId="0" pageNumber="93">Fielding s.n. 1892</emphasis>
</collectorName>
.
<emphasis id="827CA3B0F1C88FC0A238F7B8A497A8CD" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="3E7E199B921631E6C00CC162C5FB13F5" country="Malaysia" name="Melaka">Melaka</collectingRegion>
</emphasis>
:
<emphasis id="958C8D107ECC949B73B45CE739405FE6" italics="true" pageId="0" pageNumber="93">
<collectorName id="C4B2446EE271CEE3E4FFA131D6CE1AC1">Griffith</collectorName>
39
</emphasis>
57* 1845 (K),
<emphasis id="73E142BD00E5E6A89DB04255CE12D3DB" italics="true" pageId="0" pageNumber="93">
<collectorName id="91624499C796F879A28803BD73D28EBE">Burkill</collectorName>
35515
</emphasis>
*.
<emphasis id="F57836E55DA68C937F05B7FD0D556F9B" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="A8507618C6DAA94FC053257FB17C6D37" country="Malaysia" name="Pahang">Pahang</collectingRegion>
</emphasis>
:
<collectorName id="4722E9900B944077DF0634B88353F0C3">
Bentong
<emphasis id="981B076628CBD11042300C40FF8BBCAE" italics="true" pageId="0" pageNumber="93">
Burkill &amp;
<collectorName id="E857845821FD7068A97644B84ED7EFB1">Md Haniff</collectorName>
<accessionNumber id="DB6B1E5D8C2A9AC07D7AB431ACDF4E68" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN16536">SFN 16536</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="BB291D0159AA9AF588C5C026E3D17095" accessionNumber="SFN 17728" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Pekan Burkill, Md Haniff" specimenCount="1" stateProvince="Johor">
<collectorName id="3C5D4A7A7B7B7D55B8E103160B895CD3">
Pekan
<emphasis id="460D8C41FB6F23D09F20A179C259F60F" italics="true" pageId="0" pageNumber="93">
Burkill &amp;
<collectorName id="C1E284EFE6B1A8689F40BDD5CF308B4C">Md Haniff</collectorName>
<accessionNumber id="4CAB4F0C1B12CC8BCEDAB3CF5634AA7C" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN17728">SFN 17728</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="35DB78027B2CE4C89CA7EE69FFA3F020" accessionNumber="SFN 13527" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Sungai Telom Poore, Tapah Burkill, Md Haniff" specimenCount="1" stateProvince="Johor">
<collectorName id="2A7CBD4606D8E3BD1FE8ADA663513894">
Sungai Telom
<emphasis id="25229281924336E8F3C7C1DCA2EC6060" italics="true" pageId="0" pageNumber="93">Poore 538</emphasis>
</collectorName>
roadside;
<collectorName id="40A51E15D81D470A1CE13AC21DC69A60">
Tapah
<emphasis id="F78A00B292C8EC434EDDA6762B064BC1" italics="true" pageId="0" pageNumber="93">
Burkill &amp;
<collectorName id="3FAB05156A2B6B01D8C26488DA33AD73">Md Haniff</collectorName>
<accessionNumber id="F023588876594ED5979C2480931C15BD" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN13527">SFN 13527</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
.
<materialsCitation id="B18CF2A842E39128C870FD49F1C9219B" accessionNumber="SFN 39391" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Wallich, Curtis, Paya Tobong, Sinclair" location="Penang" specimenCount="1" stateProvince="Penang">
<emphasis id="BDCDEB3707F1B21F526A032CA42E22D5" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="A89D25D49E7226C9467B092D7ECE7CBF" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
</emphasis>
:
<emphasis id="F7B9D02D9EAC20AD25FCEF09D0A811B7" italics="true" pageId="0" pageNumber="93">
<collectorName id="535F767BF154B5A19D021B2D6C080825">Wallich</collectorName>
s.n.
</emphasis>
* 1829 (K);
<emphasis id="1851CB9E6B2906440AAFED4A1E3A9EFE" italics="true" pageId="0" pageNumber="93">
<collectorName id="2B68BA6E0F49FCDC2ED647DC650542AD">Curtis</collectorName>
466
</emphasis>
<collectorName id="4CA3EEB5BC5B6836DEFC9525D8777E75">Paya Tobong</collectorName>
1892;
<emphasis id="7EE41632880192D9856830E262DD07FB" italics="true" pageId="0" pageNumber="93">
<collectorName id="10EDBFEE8AA9254F6B23C68DD793B503">Sinclair</collectorName>
<accessionNumber id="75B06D98E96BD655D19B80B97D8556FE" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN39391">SFN 39391</accessionNumber>
</emphasis>
</materialsCitation>
*
<materialsCitation id="D0FE79E5E4EDB12263EA2D8F4F6C3E37" accessionNumber="SFN 16273" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Pulau Betong., Bagan Datuk Md Haniff" location="Perak" specimenCount="1" stateProvince="Perak">
village,
<collectorName id="1434CEF414BF567B59C19395BFAE3E3F">Pulau Betong.</collectorName>
<emphasis id="8DF089AA3073FE977DE1F17647D450D6" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="FDB32029456A5439273796A8F223B4F5" country="Malaysia" name="Perak">Perak</collectingRegion>
</emphasis>
:
<collectorName id="23BE8047497D9DDFF38287AF17446D27">
Bagan Datuk
<emphasis id="467137C50F66DACB2368CE03E3E3C04B" italics="true" pageId="0" pageNumber="93">
Md Haniff
<accessionNumber id="22959F5FB6AD3AF7E52B0244DA0B4057" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN16273">SFN 16273</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="752653FB2068621AAE417FB831B83E7A" accessionNumber="SFN 13272" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Batu Kurau Md Haniff, Sa'at" specimenCount="1" stateProvince="Perak">
<collectorName id="A2FD148EEF63CAA77FB25DC7A2EF392F">
Batu Kurau
<emphasis id="24BE596D5CE73791E27FC6B2A984EE09" italics="true" pageId="0" pageNumber="93">
Md Haniff &amp;
<normalizedToken id="7179ACF086314C0EABCC9955265478A6" originalValue="Sa’at">
<collectorName id="BF56F298297F7957F342AB768FD5A9E1">Sa'at</collectorName>
</normalizedToken>
<accessionNumber id="03CC6A470955CD5C1DB3B229F4DACE4D" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN13272">SFN 13272</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="0F9384704C748CDED80EA19E2713693D" accessionNumber="SFN 13647" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Gerik Burkill, Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="BAB9FC63009FEB4FD44B75040D342187">
Gerik
<emphasis id="DA1DFD3B0C4BEBDE38A94C88BA65A206" italics="true" pageId="0" pageNumber="93">
Burkill &amp;
<collectorName id="0AFCA21AF394F4F6AE3070BC7300B2DE">Md Haniff</collectorName>
<accessionNumber id="2639DE9A3FAB753313DD5C1263D4319B" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN13647">SFN 13647</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="999E4AC0D11854C22C2A9BEE516073C5" accessionNumber="SFN 16299" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Kampung Gajah Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="0A38270F5997A52F14C0554A1FC5652C">
Kampung Gajah
<emphasis id="E498E5B3BD3B98E1BE1D42F8E5878256" italics="true" pageId="0" pageNumber="93">
Md Haniff
<accessionNumber id="56C38BAAF46E0BD86D8BDF996FE8C3B1" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN16299">SFN 16299</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="2AE63130DF58DCB837E3705C27B20D80" accessionNumber="SFN 15924" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Kota Setir Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="7ABD361DF226340DDEFF8C1E8C3DA939">
Kota Setir
<emphasis id="F4E1F8D95A4274DA4F33948711C9F43F" italics="true" pageId="0" pageNumber="93">
Md Haniff
<accessionNumber id="6A9C959A7684A9E675EB6F4673CB1D28" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN15924">SFN 15924</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
;
<materialsCitation id="37D8B25B94191B8585793EDFEE2EB930" accessionNumber="SFN 15562" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Kuala Kangsar, Kota Lama, Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="2F4773F755E1D9E3F472060CAF5248ED">Kuala Kangsar</collectorName>
to
<collectorName id="BE86375A1C486B20C214BDA8B179CABA">Kota Lama</collectorName>
,
<emphasis id="88317BE64FA684D8D9AB29019AC0A4A5" italics="true" pageId="0" pageNumber="93">
<collectorName id="473DF01F7263F8A12E53A8101A9398F4">Md Haniff</collectorName>
<accessionNumber id="162E87266E4009943A4FCA2B528E64C4" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN15562">SFN 15562</accessionNumber>
</emphasis>
</materialsCitation>
;
<materialsCitation id="1743F551D194B38635FC5560DB008494" accessionNumber="SFN 15985" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Lubuk Merbok, Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="F0650F9C8E9E3F67ADBA722019D73CC6">Lubuk Merbok</collectorName>
,
<emphasis id="1A77174B0DA6B000F441A0FF320CE310" italics="true" pageId="0" pageNumber="93">
<collectorName id="FF3275E3E68265D4EC276D584363A373">Md Haniff</collectorName>
<accessionNumber id="E772820E47CCB3220AAC626644AE74BA" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN15985">SFN 15985</accessionNumber>
</emphasis>
</materialsCitation>
,
<materialsCitation id="8C438164037E25A889FCC2960B9E86CA" accessionNumber="SFN 16004" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Md Haniff" specimenCount="1" stateProvince="Perak">
<emphasis id="DB4F485219760250111A37A22BB359D0" italics="true" pageId="0" pageNumber="93">
<collectorName id="12EA80BA4B035FD658B47CD52373B400">Md Haniff</collectorName>
<accessionNumber id="0EBD69E49A8A63A676DBAFF49EB2A356" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN16004">SFN 16004</accessionNumber>
</emphasis>
</materialsCitation>
;
<materialsCitation id="B7F513FFB2274281329EA21CE1CF0D06" accessionNumber="SFN 13500" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Tanjung Malim, Burkill, Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="54D2DA73188A56BDC4ACA82C536B7281">Tanjung Malim</collectorName>
-
<emphasis id="DE8576EF48728804CDFB90E0B815B51B" italics="true" pageId="0" pageNumber="93">
<collectorName id="30B60FE477FD9C8B3038371351298DA3">Burkill</collectorName>
&amp;
<collectorName id="78D6C635466305FED1A3DE25354C7A2F">Md Haniff</collectorName>
<accessionNumber id="3F3C86762D8601FA2410A49AE060D341" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN13500">SFN 13500</accessionNumber>
</emphasis>
</materialsCitation>
;
<materialsCitation id="04CBF419412B6FB517025CDF1C190460" accessionNumber="SFN 15945" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Telok Anson Burkill, Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="AE21224998EC01B55E4C921D89642F93">
Telok Anson
<emphasis id="57265FB5E17A66029F6B07BA72817A1A" italics="true" pageId="0" pageNumber="93">
Burkill &amp;
<collectorName id="842394EAFB7F66199EC10C61EB472FCA">Md Haniff</collectorName>
<accessionNumber id="1697E4186940809D200B52DF5022EEF6" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN15945">SFN 15945</accessionNumber>
</emphasis>
</collectorName>
</materialsCitation>
,
<emphasis id="8A67E2EB486C3701D6154527D796A63F" italics="true" pageId="0" pageNumber="93">
<materialsCitation id="536B3827768233071417867401957EBD" accessionNumber="SFN 10315" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Md Haniff" specimenCount="1" stateProvince="Perak">
<collectorName id="9E09C80A7802BAF56186E8C4DEE8C413">Md Haniff</collectorName>
<accessionNumber id="299F696FFF693ECBAA075F3EA22C1A2A" httpUri="http://www.ncbi.nlm.nih.gov/protein/SFN10315">SFN 10315</accessionNumber>
</materialsCitation>
.
</emphasis>
<materialsCitation id="EF7074A46F0477D31F42D509455B2825" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Ulu Gombak T" location="Selangor" specimenCount="1" stateProvince="Selangor">
<emphasis id="CDD693298824404EC82185B5AAE2AE65" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="853D5D98DE98C3364B9808ED1727F766" country="Malaysia" name="Selangor">Selangor</collectingRegion>
</emphasis>
:
<collectorName id="F08FDAB32D18B6E9F28389BB2C763F5A">
Ulu Gombak
<emphasis id="12A78F238788C45BBA52F9050F6E1C86" italics="true" pageId="0" pageNumber="93">T &amp; B 2823</emphasis>
</collectorName>
roadside [leaves multicoloured],
<emphasis id="36134B5BF6759480FF1713AA1B2F5A14" italics="true" pageId="0" pageNumber="93">Md Nur SFN 3423</emphasis>
2 by stream;
<collectingRegion id="F675D23CE809C0640434CBD528C84F0B" country="Malaysia" name="Kuala Lumpur">Kuala Lumpur</collectingRegion>
<emphasis id="8D11F1FE079D069DA3ED8128F98C2C3D" italics="true" pageId="0" pageNumber="93">Ridley s.n.</emphasis>
* 1890 bunga ati-ati merah.
<emphasis id="E7E9E81CDCAD9E80EFB4A0FF456606A3" bold="true" pageId="0" pageNumber="93">
<collectingRegion id="86D84F7C51333A6CD915FBB65DABC477" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
</emphasis>
: Bundi
<emphasis id="9868D8FB224B0E41E6DF1F0C1F5BA04E" italics="true" pageId="0" pageNumber="93">
<taxonomicName id="DBAA6142B9BAABF8048F44ACA93525F3" genus="Rostado" lsidName="Rostado" pageId="0" pageNumber="93" rank="genus">Rostado</taxonomicName>
s.n.
</emphasis>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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