278 lines
34 KiB
XML
278 lines
34 KiB
XML
<document id="0AECB24809FAB3910E07D35BFC82F829" ID="10.11646/zootaxa.4210.1.1" ID-CLB-Dataset="37280" ID-DOI="10.11646/zootaxa.4210.1.1" ID-GBIF-Dataset="e782e07d-76b7-4e9b-ba34-ed3286254ec6" ID-ISSN="1175-5326" ID-Zenodo-Dep="252966" ID-ZooBank="D39416B8-CF85-440B-84C2-D4380BECC4E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484869979019" checkinUser="plazi" docAuthor="Harzhauser, Mathias & Landau, Bernard" docDate="2016" docId="373F87D7FFFFFFEFFF5FAD67FB6D409F" docLanguage="en" docName="zootaxa.4210.1.1.pdf" docOrigin="Zootaxa 4210 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leporiconus Iredale 1930" docType="treatment" docVersion="10" lastPageNumber="98" masterDocId="CB06FFAFFF9FFF8EFFC8A876FF89416A" masterDocTitle="A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea" masterLastPageNumber="178" masterPageNumber="1" pageNumber="97" updateTime="1698435921492" updateUser="ExternalLinkService">
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<mods:titleInfo id="CB72FF4A4E7B67A24B8E0680F3D82A81">
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<mods:title id="054C301A7C97B1B3332DD29E266A38AF">A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea</mods:title>
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<mods:name id="31059C3E8465415D3DFC9E138B3ABD79" type="personal">
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<mods:namePart id="96F1A2D4FEDF3D900E66AFD5D884E7B3">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="8930A472651918259C99B280C9E8C201">Landau, Bernard</mods:namePart>
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<mods:typeOfResource id="4ADED2739233829FED91E5FE9963F034">text</mods:typeOfResource>
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<mods:titleInfo id="26FF3C4554040DF1DCFAFE3310B0CB25">
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<mods:title id="51D10BFE17B29C2E5FBEA35CF9B841CE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="51D6EC002EB730415F43950C7A1AFA6C">
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<mods:date id="783783E4A8F49E041E711F9A2C5F2953">2016</mods:date>
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<mods:detail id="A24A4A6F7595628C40296EF1AA22AE37" type="volume">
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<mods:number id="FD9BCF26FB498857091D5D767EB0E703">4210</mods:number>
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<mods:classification id="0DFA008594279014E94F8488FC4431FA">journal article</mods:classification>
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<mods:identifier id="E4ED762F98E60177530DEE2E3A7F9B0B" type="CLB-Dataset">37280</mods:identifier>
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<mods:identifier id="597BCD5DA749752A13AF8214C83B6917" type="DOI">10.11646/zootaxa.4210.1.1</mods:identifier>
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<treatment id="373F87D7FFFFFFEFFF5FAD67FB6D409F" ID-DOI="http://doi.org/10.5281/zenodo.5622371" ID-GBIF-Taxon="127615930" ID-Zenodo-Dep="5622371" LSID="urn:lsid:plazi:treatment:373F87D7FFFFFFEFFF5FAD67FB6D409F" httpUri="http://treatment.plazi.org/id/373F87D7FFFFFFEFFF5FAD67FB6D409F" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
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<subSubSection id="F78C654AFFFFFFEEFF5FAD67FDA64441" box="[151,559,1297,1323]" pageId="96" pageNumber="97" type="nomenclature">
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<paragraph id="BF2936C1FFFFFFEEFF5FAD67FDA64441" blockId="96.[151,559,1297,1323]" box="[151,559,1297,1323]" pageId="96" pageNumber="97">
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<heading id="E46181ADFFFFFFEEFF5FAD67FDA64441" bold="true" box="[151,559,1297,1323]" fontSize="11" level="1" pageId="96" pageNumber="97" reason="1">
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<emphasis id="8DE2EAD3FFFFFFEEFF5FAD67FDA64441" bold="true" box="[151,559,1297,1323]" pageId="96" pageNumber="97">
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Genus
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<taxonomicName id="78964D42FFFFFFEEFF26AD67FDA64441" ID-CoL="faf3cf3b-81dc-4d14-bcec-8d16f5255b71" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" box="[238,559,1297,1323]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
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<emphasis id="8DE2EAD3FFFFFFEEFF26AD67FE0C4441" bold="true" box="[238,389,1297,1323]" italics="true" pageId="96" pageNumber="97">Leporiconus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFE44AD67FDA64441" author="Iredale" box="[396,559,1297,1323]" pageId="96" pageNumber="97" refString="Iredale, T. (1930) Queensland molluscan notes, no. 2. Memoirs of the Queensland Museum, 10, 73 - 88." type="journal article" year="1930">Iredale, 1930</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F78C654AFFFFFFEFFF5FAD2FFB6D409F" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="materials_examined">
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<paragraph id="BF2936C1FFFFFFEEFF5FAD2FFB2B4404" blockId="96.[151,1186,1368,1390]" box="[151,1186,1368,1390]" pageId="96" pageNumber="97">
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Type species (by original designation):
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<taxonomicName id="78964D42FFFFFFEEFDEEAD2FFC3F4404" authority="Hwass in Bruguiere, 1792" authorityName="Hwass in Bruguiere" authorityYear="1792" box="[550,950,1368,1390]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="glans">
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<emphasis id="8DE2EAD3FFFFFFEEFDEEAD2FFD2D4404" box="[550,676,1368,1390]" italics="true" pageId="96" pageNumber="97">Conus glans</emphasis>
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Hwass in
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<bibRefCitation id="DB074B30FFFFFFEEFCC6AD2EFC3F4404" author="Bruguiere" box="[782,950,1368,1390]" pageId="96" pageNumber="97" refString="Bruguiere, M. (1792) Encyclopedie methodique. Histoire naturale des Vers. Tome Ier. Impensis Panckoucke, Paris, 345 - 752 pp." type="book chapter" year="1792">Bruguière, 1792</bibRefCitation>
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</taxonomicName>
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. Recent, Indian Ocean.
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</paragraph>
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<paragraph id="BF2936C1FFFFFFEFFF5FADEDFBE341BE" blockId="96.[151,1437,1435,2036]" lastBlockId="97.[151,1436,151,501]" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
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<emphasis id="8DE2EAD3FFFFFFEEFF5FADEDFF5F44DE" bold="true" box="[151,214,1435,1460]" pageId="96" pageNumber="97">Note.</emphasis>
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According to
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<bibRefCitation id="DB074B30FFFFFFEEFE5EADEDFD5844D9" author="Tucker" box="[406,721,1435,1460]" pageId="96" pageNumber="97" refString="Tucker, J. K. & Tenorio, M. J. (2009) Systematic classification of recent and fossil conoidean gastropods. ConchBooks, Hackenheim, 296 pp." type="book" year="2009">Tucker & Tenorio (2009)</bibRefCitation>
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the genus
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<taxonomicName id="78964D42FFFFFFEEFCA1ADEAFB2C44D9" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" box="[873,1189,1435,1460]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
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<emphasis id="8DE2EAD3FFFFFFEEFCA1ADEAFC7C44D9" box="[873,1013,1436,1459]" italics="true" pageId="96" pageNumber="97">Leporiconus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFBCEADEDFB2C44D9" author="Iredale" box="[1030,1189,1435,1460]" pageId="96" pageNumber="97" refString="Iredale, T. (1930) Queensland molluscan notes, no. 2. Memoirs of the Queensland Museum, 10, 73 - 88." type="journal article" year="1930">Iredale, 1930</bibRefCitation>
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</taxonomicName>
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is characterised by subcylindrical to pyriform, sometimes torpedo shaped shells, with beaded early spire whorls, two or more cords on the spire whorls and ridges on the body whorl, which are usually well developed and pustulose. The shoulder is rounded to indistinct. The subsutural flexures of the Paratethyan species are shallow, weakly to moderately curved and moderately asymmetrical and the protoconch is multispiral. The generic description given by
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<bibRefCitation id="DB074B30FFFFFFEEFAE3AE5DFEB6470D" author="Tucker" pageId="96" pageNumber="97" refString="Tucker, J. K. & Tenorio, M. J. (2009) Systematic classification of recent and fossil conoidean gastropods. ConchBooks, Hackenheim, 296 pp." type="book" year="2009">Tucker & Tenorio (2009)</bibRefCitation>
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was based on the species considered by these authors to be included in the genus:
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<taxonomicName id="78964D42FFFFFFEEFB1AAE39FEDA47E1" authority="Hwass in Bruguiere, 1792" authorityName="Hwass in Bruguiere" authorityYear="1792" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="glans">
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<emphasis id="8DE2EAD3FFFFFFEEFB1AAE39FAA6470D" box="[1234,1327,1614,1639]" italics="true" pageId="96" pageNumber="97">C. glans</emphasis>
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Hwass in
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<bibRefCitation id="DB074B30FFFFFFEEFF5FAE05FEDA47E1" author="Bruguiere" box="[151,339,1651,1676]" pageId="96" pageNumber="97" refString="Bruguiere, M. (1792) Encyclopedie methodique. Histoire naturale des Vers. Tome Ier. Impensis Panckoucke, Paris, 345 - 752 pp." type="book chapter" year="1792">Bruguière, 1792</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFEA9AE02FDF647E1" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[353,639,1651,1676]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="caffeae">
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<emphasis id="8DE2EAD3FFFFFFEEFEA9AE02FE5F47E1" box="[353,470,1651,1675]" italics="true" pageId="96" pageNumber="97">C. caffeae</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFE28AE05FDF647E1" author="Gmelin" box="[480,639,1651,1676]" pageId="96" pageNumber="97" refString="Gmelin, J. F. (1791) C. a Linne ... Systema naturae per regna tria naturae ... editio decima tertia, aucta, reformata, cura J. F. Gmelin. Tomus I, pars VI, Vermes testacea. G. E. Beer, Lipsiae, pp. 3021 - 4120." type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFD46AE02FC4C47E6" authority="Kiener, 1845" authorityName="Kiener" authorityYear="1845" box="[654,965,1651,1676]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="corallinus">
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<emphasis id="8DE2EAD3FFFFFFEEFD46AE02FCAD47E6" box="[654,804,1651,1676]" italics="true" pageId="96" pageNumber="97">C. corallinus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFCE6AE05FC4C47E6" author="Kiener" box="[814,965,1651,1676]" pageId="96" pageNumber="97" refString="Kiener, L. C. (1845 - 1850) Genre Cone. (Conus, Lin.). Species general et iconographie des coquilles vivantes comprenant la collection du Museum d'histoire naturelle de Paris. La collection Lamarck, celle du prince Massena (appartenant maintenant a M. B. Delessert) et les decouvertes recentes des voyageurs, J. B. Baillier, Paris, 379 pp." type="book" year="1845">Kiener, 1845</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFC1BAE02FF5B47C5" authority="Broderip & Sowerby I, 1830" authorityName="Broderip & Sowerby I" authorityYear="1830" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="cylindraceus">
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<emphasis id="8DE2EAD3FFFFFFEEFC1BAE02FB0F47E6" box="[979,1158,1651,1676]" italics="true" pageId="96" pageNumber="97">C. cylindraceus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFB58AE05FF5B47C5" author="Broderip" pageId="96" pageNumber="97" refString="Broderip, W. J. & Sowerby, G. B. I (1830) Observations on new or interesting Mollusca, contained, for the most part, in the museum of the Zoological Society. Zoological Journal, 5, 46 - 51." type="journal article" year="1830">Broderip & Sowerby I, 1830</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFF15AEE1FD0147C5" authority="Rockel & Fischoder, 1985" authorityName="Rockel & Fischoder" authorityYear="1985" box="[221,648,1686,1711]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="granum">
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<emphasis id="8DE2EAD3FFFFFFEEFF15AEE1FEDC47C5" box="[221,341,1687,1711]" italics="true" pageId="96" pageNumber="97">C. granum</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFE95AEE0FD0147C5" author="Rockel" box="[349,648,1686,1711]" pageId="96" pageNumber="97" refString="Rockel, D. & Fischoder, H. (1985) Eine neue Conus - Art von den Philippinen. Spixiana, 8, 67 - 72." type="journal article" year="1985">Röckel & Fischöder, 1985</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFD5BAEE1FC3447C5" authority="Sowerby I, 1833" authorityName="Sowerby I" authorityYear="1833" box="[659,957,1686,1711]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="luteus">
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<emphasis id="8DE2EAD3FFFFFFEEFD5BAEE1FD7E47C5" box="[659,759,1686,1711]" italics="true" pageId="96" pageNumber="97">C. luteus</emphasis>
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Sowerby I, 1833
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFC00AEE1FAFE47C5" authority="Hwass in Bruguiere, 1792" authorityName="Hwass in Bruguiere" authorityYear="1792" box="[968,1399,1686,1711]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="mitratus">
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<emphasis id="8DE2EAD3FFFFFFEEFC00AEE1FBCF47C5" box="[968,1094,1687,1711]" italics="true" pageId="96" pageNumber="97">C. mitratus</emphasis>
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Hwass in
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<bibRefCitation id="DB074B30FFFFFFEEFB76AEE0FAFE47C5" author="Bruguiere" box="[1214,1399,1686,1711]" pageId="96" pageNumber="97" refString="Bruguiere, M. (1792) Encyclopedie methodique. Histoire naturale des Vers. Tome Ier. Impensis Panckoucke, Paris, 345 - 752 pp." type="book chapter" year="1792">Bruguière, 1792</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="78964D42FFFFFFEEFA4AAEE1FE1B47B9" authority="Reeve, 1848" authorityName="Reeve" authorityYear="1848" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="nucleus">
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<emphasis id="8DE2EAD3FFFFFFEEFA4AAEE1FF6547BE" italics="true" pageId="96" pageNumber="97">C. nucleus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFF34AECAFE1B47B9" author="Reeve" box="[252,402,1723,1747]" pageId="96" pageNumber="97" refString="Reeve, L. A. (1843 - 1849) Conchologia Iconica, a complete repertory of species. Reeve Brothers, London, 20 volumes, 1, pls. 1 - 41, 1843; pls 42 - 47, 1844; supplementary plates 1 - 4, 1848; pls. 5 - 9, 1849; emendations 1 - 6, 9 pls, 1849." type="journal article" year="1848">Reeve, 1848</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="78964D42FFFFFFEEFE13AECAFCF047BE" authority="Sowerby" authorityName="Sowerby" authorityYear="1858" box="[475,889,1723,1748]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="tenuistriatus">
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<emphasis id="8DE2EAD3FFFFFFEEFE13AECAFD1947B9" box="[475,656,1724,1747]" italics="true" pageId="96" pageNumber="97">C. tenuistriatus</emphasis>
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<bibRefCitation id="DB074B30FFFFFFEEFD68AECDFCF047BE" author="Sowerby" box="[672,889,1723,1748]" pageId="96" pageNumber="97" refString="Sowerby, G. B. II (1858 - 1866) Thesaurus Conchyliorum or Monographs of the Genera of Shells, 3. Sowerby, London, 331 pp." type="book" year="1858">Sowerby II, 1858</bibRefCitation>
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</taxonomicName>
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. However,
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<bibRefCitation id="DB074B30FFFFFFEEFBC2AECDFAB147BE" box="[1034,1336,1723,1748]" pageId="96" pageNumber="97" refString="Puillandre, N., Bouchet, P., Duda, T. F., Kauferstein, S., Kohn, A. J., Olivera, B. M., Watkins, M. & Meyer, C. (2014 b) Molecular Phylogeny and evolution of the cone snails (Gastropoda, Conoidea). Molecular Phylogenetics and Evolution, 78, 290 - 303. http: // dx. doi. org / 10.1016 / j. ympev. 2014.05.023" type="journal article">
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Puillandre
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<emphasis id="8DE2EAD3FFFFFFEEFB45AECAFB4747BE" box="[1165,1230,1723,1748]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
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(2014b)
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</bibRefCitation>
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defined
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<taxonomicName id="78964D42FFFFFFEEFF5FAE96FEAA479D" box="[151,291,1760,1783]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
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<emphasis id="8DE2EAD3FFFFFFEEFF5FAE96FEAA479D" box="[151,291,1760,1783]" italics="true" pageId="96" pageNumber="97">Leporiconus</emphasis>
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</taxonomicName>
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as a monophyletic clade containing five of the species included by Tucker & Tenorio:
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<emphasis id="8DE2EAD3FFFFFFEEFAC8AEA9FE474676" italics="true" pageId="96" pageNumber="97">
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<taxonomicName id="78964D42FFFFFFEEFAC8AEA9FAFF479D" box="[1280,1398,1759,1783]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="coffeae">C. coffeae</taxonomicName>
|
|
,
|
|
<taxonomicName id="78964D42FFFFFFEEFA4AAEA9FF5E4676" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="glans">C. glans</taxonomicName>
|
|
,
|
|
<taxonomicName id="78964D42FFFFFFEEFF2AAF72FED74671" box="[226,350,1796,1819]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="granum">C. granum</taxonomicName>
|
|
,
|
|
<taxonomicName id="78964D42FFFFFFEEFEA1AF72FE474676" box="[361,462,1795,1820]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="luteus">C. luteus</taxonomicName>
|
|
</emphasis>
|
|
, and
|
|
<taxonomicName id="78964D42FFFFFFEEFDC4AF72FD334671" box="[524,698,1796,1819]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="tenuistriatus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFDC4AF72FD334671" box="[524,698,1796,1819]" italics="true" pageId="96" pageNumber="97">C. tenuistriatus</emphasis>
|
|
</taxonomicName>
|
|
, but excluded
|
|
<taxonomicName id="78964D42FFFFFFEEFCA9AF72FC5E4676" box="[865,983,1795,1820]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="nucleus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFCA9AF72FC5E4676" box="[865,983,1795,1820]" italics="true" pageId="96" pageNumber="97">C. nucleus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="78964D42FFFFFFEEFBD8AF72FB2A4676" box="[1040,1187,1795,1820]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="corallinus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFBD8AF72FB2A4676" box="[1040,1187,1795,1820]" italics="true" pageId="96" pageNumber="97">C. corallinus</emphasis>
|
|
</taxonomicName>
|
|
, which they placed in the closely related genus
|
|
<emphasis id="8DE2EAD3FFFFFFEEFE78AF50FDA74655" box="[432,558,1830,1855]" italics="true" pageId="96" pageNumber="97">Splinoconu</emphasis>
|
|
s da
|
|
<bibRefCitation id="DB074B30FFFFFFEEFDA9AF5EFD644655" author="Motta" box="[609,749,1831,1855]" pageId="96" pageNumber="97" refString="da Motta, A. J. (1991) A systematic classification of the gastropod family Conidae at the generic level. La Conchiglia, Roma, 48 pp." type="book" year="1991">Motta, 1991</bibRefCitation>
|
|
.
|
|
<taxonomicName id="78964D42FFFFFFEEFD3FAF51FC5D4655" box="[759,980,1830,1855]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="cylindraceus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFD3FAF51FC5D4655" box="[759,980,1830,1855]" italics="true" pageId="96" pageNumber="97">Conus cylindraceus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="78964D42FFFFFFEEFBC2AF51FB014655" box="[1034,1160,1831,1855]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="mitratus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFBC2AF51FB014655" box="[1034,1160,1831,1855]" italics="true" pageId="96" pageNumber="97">C. mitratus</emphasis>
|
|
</taxonomicName>
|
|
they placed in the genus
|
|
<taxonomicName id="78964D42FFFFFFEEFF5FAF3AFDAA460E" authority="Shikama & Habe, 1968" authorityName="Shikama & Habe" authorityYear="1968" box="[151,547,1867,1892]" class="Gastropoda" family="Conidae" genus="Turriconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFF5FAF3AFE9B4609" box="[151,274,1868,1891]" italics="true" pageId="96" pageNumber="97">Turriconus</emphasis>
|
|
<bibRefCitation id="DB074B30FFFFFFEEFED1AF3DFDAA460E" author="Shikama" box="[281,547,1867,1892]" pageId="96" pageNumber="97" refString="Shikama, T. & Habe, T. (1968) A new Japanese cone, Turriconus nakayasui with reference to Embrikeana stupa group. Venus 26, 57 - 60." type="journal article" year="1968">Shikama & Habe, 1968</bibRefCitation>
|
|
</taxonomicName>
|
|
, which is not closely related to
|
|
<taxonomicName id="78964D42FFFFFFEEFC4EAF3AFB9B4609" box="[902,1042,1868,1891]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFC4EAF3AFB9B4609" box="[902,1042,1868,1891]" italics="true" pageId="96" pageNumber="97">Leporiconus</emphasis>
|
|
</taxonomicName>
|
|
. Therefore, the generic description should be modified to include only the species shown to be monophyletic by
|
|
<bibRefCitation id="DB074B30FFFFFFEEFC38AF18FA8A46ED" box="[1008,1283,1902,1927]" pageId="96" pageNumber="97" refString="Puillandre, N., Bouchet, P., Duda, T. F., Kauferstein, S., Kohn, A. J., Olivera, B. M., Watkins, M. & Meyer, C. (2014 b) Molecular Phylogeny and evolution of the cone snails (Gastropoda, Conoidea). Molecular Phylogenetics and Evolution, 78, 290 - 303. http: // dx. doi. org / 10.1016 / j. ympev. 2014.05.023" type="journal article">
|
|
Puillandre
|
|
<emphasis id="8DE2EAD3FFFFFFEEFBA1AF06FB2946ED" box="[1129,1184,1902,1927]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
|
|
(2014b)
|
|
</bibRefCitation>
|
|
. The wording given above still applies to the genus, as recognised by
|
|
<bibRefCitation id="DB074B30FFFFFFEEFCC1AFE5FB9646C6" box="[777,1055,1939,1964]" pageId="96" pageNumber="97" refString="Puillandre, N., Bouchet, P., Duda, T. F., Kauferstein, S., Kohn, A. J., Olivera, B. M., Watkins, M. & Meyer, C. (2014 b) Molecular Phylogeny and evolution of the cone snails (Gastropoda, Conoidea). Molecular Phylogenetics and Evolution, 78, 290 - 303. http: // dx. doi. org / 10.1016 / j. ympev. 2014.05.023" type="journal article">
|
|
Puillandre
|
|
<emphasis id="8DE2EAD3FFFFFFEEFC4CAFE2FC3546C6" box="[900,956,1939,1964]" italics="true" pageId="96" pageNumber="97">et al.</emphasis>
|
|
(2014b)
|
|
</bibRefCitation>
|
|
, but torpedo-shaped shells should be removed, as the two species now included in
|
|
<taxonomicName id="78964D42FFFFFFEEFD7BAFCEFCA746A5" box="[691,814,1976,1999]" class="Gastropoda" family="Conidae" genus="Turriconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFD7BAFCEFCA746A5" box="[691,814,1976,1999]" italics="true" pageId="96" pageNumber="97">Turriconus</emphasis>
|
|
</taxonomicName>
|
|
are more slender torpedo shaped, with far higher spires than species in
|
|
<taxonomicName id="78964D42FFFFFFEEFE86AFAAFE534699" box="[334,474,2012,2035]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFFFFEEFE86AFAAFE534699" box="[334,474,2012,2035]" italics="true" pageId="96" pageNumber="97">Leporiconus</emphasis>
|
|
</taxonomicName>
|
|
. We cannot see any reliable shell characters by which to separate the present-day species of
|
|
<taxonomicName id="78964D42FFFEFFEFFEC4A8EFFE1141DA" box="[268,408,153,176]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFEC4A8EFFE1141DA" box="[268,408,153,176]" italics="true" pageId="97" pageNumber="98">Leporiconus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="78964D42FFFEFFEFFE05A8E1FDDF41DA" box="[461,598,151,176]" class="Gastropoda" family="Conidae" genus="Splinoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFE05A8E1FDDF41DA" box="[461,598,151,176]" italics="true" pageId="97" pageNumber="98">Splinoconus</emphasis>
|
|
</taxonomicName>
|
|
, except that the spiral sculpture seems to be less prominent in
|
|
<taxonomicName id="78964D42FFFEFFEFFAC4A8E1FA1D41DA" box="[1292,1428,151,176]" class="Gastropoda" family="Conidae" genus="Splinoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFAC4A8E1FA1D41DA" box="[1292,1428,151,176]" italics="true" pageId="97" pageNumber="98">Splinoconus</emphasis>
|
|
</taxonomicName>
|
|
. This not surprising as the two genera are closely related (
|
|
<bibRefCitation id="DB074B30FFFEFFEFFCDEA8CAFB9341BF" author="Puillandre" box="[790,1050,188,213]" pageId="97" pageNumber="98" refString="Puillandre, N., Bouchet, P., Duda, T. F., Kauferstein, S., Kohn, A. J., Olivera, B. M., Watkins, M. & Meyer, C. (2014 b) Molecular Phylogeny and evolution of the cone snails (Gastropoda, Conoidea). Molecular Phylogenetics and Evolution, 78, 290 - 303. http: // dx. doi. org / 10.1016 / j. ympev. 2014.05.023" type="journal article" year="2014" yearSuffix="b">
|
|
Puillandre
|
|
<emphasis id="8DE2EAD3FFFEFFEFFC5AA8CBFC4341BF" box="[914,970,188,213]" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
|
|
2014b
|
|
</bibRefCitation>
|
|
; fig 2).
|
|
</paragraph>
|
|
<paragraph id="BF2936C1FFFEFFEFFF0FA896FB4940C7" blockId="97.[151,1436,151,501]" pageId="97" pageNumber="98">
|
|
We have attributed to this genus a small number of Paratethyan species of which one,
|
|
<taxonomicName id="78964D42FFFEFFEFFAD8A897FDF64076" authority="Hoernes & Auinger, 1879" authorityName="Hoernes & Auinger" authorityYear="1879" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="transsylvanicus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFAD8A897FECC4077" italics="true" pageId="97" pageNumber="98">Leporiconus transsylvanicus</emphasis>
|
|
(
|
|
<bibRefCitation id="DB074B30FFFEFFEFFE9BA972FDFE4076" author="Hoernes" box="[339,631,260,285]" pageId="97" pageNumber="98" refString="Hoernes, R. & Auinger, M. (1879) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 12, 1 - 52." type="journal article" year="1879">Hoernes & Auinger, 1879</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
is remarkably similar to the type species
|
|
<taxonomicName id="78964D42FFFEFFEFFB85A973FF52402A" authority="Hwass in Bruguiere, 1792" authorityName="Hwass in Bruguiere" authorityYear="1792" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="glans">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFB85A973FB2F4077" box="[1101,1190,260,285]" italics="true" pageId="97" pageNumber="98">L. glans</emphasis>
|
|
(Hwass in
|
|
<bibRefCitation id="DB074B30FFFEFFEFFAEDA972FF5A402A" author="Bruguiere" pageId="97" pageNumber="98" refString="Bruguiere, M. (1792) Encyclopedie methodique. Histoire naturale des Vers. Tome Ier. Impensis Panckoucke, Paris, 345 - 752 pp." type="book chapter" year="1792">Bruguière, 1792</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. The other species are less obviously torpedo-shaped, but all have beaded early spire whorls.
|
|
<taxonomicName id="78964D42FFFEFFEFFA9CA95EFECC400F" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="transsylvanicus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFA9CA95EFECC400F" italics="true" pageId="97" pageNumber="98">Conus transsylvanicus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="78964D42FFFEFFEFFEB6A93BFC98400E" authority="Hoernes & Auinger, 1879" authorityName="Hoernes & Auinger" authorityYear="1879" box="[382,785,332,357]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="suessi">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFEB6A93BFE6A400E" box="[382,483,333,356]" italics="true" pageId="97" pageNumber="98">C. suessi</emphasis>
|
|
<bibRefCitation id="DB074B30FFFEFFEFFE24A93BFC98400E" author="Hoernes" box="[492,785,332,357]" pageId="97" pageNumber="98" refString="Hoernes, R. & Auinger, M. (1879) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 12, 1 - 52." type="journal article" year="1879">Hoernes & Auinger, 1879</bibRefCitation>
|
|
</taxonomicName>
|
|
were placed in the genus
|
|
<taxonomicName id="78964D42FFFEFFEFFBF4A93BFAE0400E" authority="da Motta 1991" authorityName="da Motta" authorityYear="1991" box="[1084,1385,332,357]" class="Gastropoda" family="Conidae" genus="Varioconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFBF4A93BFB33400E" box="[1084,1210,333,356]" italics="true" pageId="97" pageNumber="98">Varioconus</emphasis>
|
|
da
|
|
<bibRefCitation id="DB074B30FFFEFFEFFB2DA93BFAE0400E" author="Motta" box="[1253,1385,332,356]" pageId="97" pageNumber="98" refString="da Motta, A. J. (1991) A systematic classification of the gastropod family Conidae at the generic level. La Conchiglia, Roma, 48 pp." type="book" year="1991">
|
|
Motta
|
|
<quantity id="786E9B24FFFEFFEFFAF9A93AFAE0400E" box="[1329,1385,332,356]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="97" pageNumber="98" unit="in" value="1991.0">1991</quantity>
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
in a unpublished generic list made available to us by J.K. Tucker (personal comm. 2016), however,
|
|
<taxonomicName id="78964D42FFFEFFEFFAD6A907FA1540E2" box="[1310,1436,369,392]" class="Gastropoda" family="Conidae" genus="Varioconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFAD6A907FA1540E2" box="[1310,1436,369,392]" italics="true" pageId="97" pageNumber="98">Varioconus</emphasis>
|
|
</taxonomicName>
|
|
(considered as synonym of
|
|
<taxonomicName id="78964D42FFFEFFEFFE03A9E3FDC740C6" box="[459,590,405,428]" class="Gastropoda" family="Conidae" genus="Lautoconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFE03A9E3FDC740C6" box="[459,590,405,428]" italics="true" pageId="97" pageNumber="98">Lautoconus</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation id="DB074B30FFFEFFEFFDBFA9E2FCFF40C6" author="Puillandre" box="[631,886,404,429]" pageId="97" pageNumber="98" refString="Puillandre, N., Duda, T. F., Meyer, C., Olivera, B. M. & Bouchet, P. (2014 a) One, four or 100 genera? A new classification of the cone snails. Journal of Molluscan Studies, 8, 1 - 23." type="journal article" year="2014" yearSuffix="a">
|
|
Puillandre
|
|
<emphasis id="8DE2EAD3FFFEFFEFFD39A9E3FCAB40C7" box="[753,802,404,429]" italics="true" pageId="97" pageNumber="98">et al</emphasis>
|
|
. 2014a
|
|
</bibRefCitation>
|
|
) lacks beads on spire whorls.
|
|
</paragraph>
|
|
<paragraph id="BF2936C1FFFEFFEFFF0FA9C1FB6D409F" blockId="97.[151,1436,151,501]" pageId="97" pageNumber="98">
|
|
The presence of
|
|
<taxonomicName id="78964D42FFFEFFEFFE4CA9CFFD9940BA" box="[388,528,441,464]" class="Gastropoda" family="Conidae" genus="Leporiconus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
|
|
<emphasis id="8DE2EAD3FFFEFFEFFE4CA9CFFD9940BA" box="[388,528,441,464]" italics="true" pageId="97" pageNumber="98">Leporiconus</emphasis>
|
|
</taxonomicName>
|
|
in the Miocene Paratethys is interesting, as it documents the wide post-Tethyan distribution of a taxon during middle Miocene times nowadays restricted to the Indo-West Pacific.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |