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<document ID-CLB-Dataset="267803" ID-DOI="http://dx.doi.org/10.3897/zookeys.1180.100106" ID-GBIF-Dataset="f0024131-1749-426e-9fa0-f7a4b0657aea" ID-PMC="PMC10517412" ID-Pensoft-Pub="1313-2970-1180-67" ID-Pensoft-UUID="D6D18F1CE45D5E139A47C925E5BC69A3" ID-PubMed="37744947" ID-ZooBank="95E3D41450F4402293D5BC851E68BF85" ModsDocID="1313-2970-1180-67" checkinTime="1694791482216" checkinUser="pensoft" docAuthor="Kang, Ilgoo, Ghafouri Moghaddam, Mostafa, Sharkey, Michael J., Quicke, Donald L. J., Butcher, Buntika A. &amp; Carlton, Christopher E." docDate="2023" docId="C9408DD6D40E540CA4997DEF0A261593" docLanguage="en" docName="ZooKeys 1180: 67-79" docOrigin="ZooKeys 1180" docPubDate="2023-09-15" docSource="http://dx.doi.org/10.3897/zookeys.1180.100106" docTitle="Ophiclypeus dvaravati Ghafouri Moghaddam, Quicke &amp; Butcher 2023, sp. nov." docType="treatment" docUuid="1705521A-4233-4AE2-A781-C32F4D481247" docUuidSource="ZooBank" docVersion="4" id="D6D18F1CE45D5E139A47C925E5BC69A3" lastPageNumber="67" masterDocId="D6D18F1CE45D5E139A47C925E5BC69A3" masterDocTitle="Ophiclypeus, a new genus of Cardiochilinae Ashmead (Hymenoptera, Braconidae) from the Oriental region with descriptions of three new species" masterLastPageNumber="79" masterPageNumber="67" pageNumber="67" updateTime="1732702463687" updateUser="ExternalLinkService">
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<mods:title id="C780DD9F752E859DB220975F6BFC98A9">Ophiclypeus, a new genus of Cardiochilinae Ashmead (Hymenoptera, Braconidae) from the Oriental region with descriptions of three new species</mods:title>
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<mods:namePart id="65FA639774CCE53DFAC7DCF3D111049C">Kang, Ilgoo</mods:namePart>
<mods:nameIdentifier id="BF72DF4BB49DA7C8838F9467ABCAA5C3" type="ORCID">https://orcid.org/0000-0002-8501-1758</mods:nameIdentifier>
<mods:affiliation id="17F354F2075F235DC0C4D58003DD186A">Department of Entomology, Louisiana State University Agricultural Center, 404 Life Sciences Building, Baton Rouge, LA, 70803 USA</mods:affiliation>
<mods:nameIdentifier id="E9E9595B8A6D1A470ABAF9C2A57ECDDA" type="email">ikang@knu.ac.kr</mods:nameIdentifier>
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<mods:namePart id="BC8C7D7FD92498F2DE9F3D8E57CE4B54">Ghafouri Moghaddam, Mostafa</mods:namePart>
<mods:nameIdentifier id="305343A8E9F7DA849A444385144475F4" type="ORCID">https://orcid.org/0000-0002-1942-9689</mods:nameIdentifier>
<mods:affiliation id="64A7F26E2BD241D903D3FB8AD0C5CF52">Department of Entomology, College of Ecology and Environmental Science, Kyungpook National University, Sangju, Gyeongsangbuk-do, 37224, South Korea</mods:affiliation>
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<mods:namePart id="23912DA718951108A138A7F758BBF600">Sharkey, Michael J.</mods:namePart>
<mods:nameIdentifier id="54544B86CDBEC491C98BBC3F3F89D1FB" type="ORCID">https://orcid.org/0000-0001-6201-7340</mods:nameIdentifier>
<mods:affiliation id="60E4D4C9C1F115B2F24EAC151DBFAC64">Integrative Ecology Laboratory, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Pathumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="8F96FC572374E479DD0F1058118DE41C">Quicke, Donald L. J.</mods:namePart>
<mods:nameIdentifier id="AD43A80D91A68FD21C2E2E089F496837" type="ORCID">https://orcid.org/0000-0003-4471-6775</mods:nameIdentifier>
<mods:affiliation id="DC57D50ECD36601B86B75B7EFD0B8505">Department of Entomology, College of Ecology and Environmental Science, Kyungpook National University, Sangju, Gyeongsangbuk-do, 37224, South Korea</mods:affiliation>
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<mods:namePart id="E2ABF694923897A4340B5108731FFB61">Butcher, Buntika A.</mods:namePart>
<mods:nameIdentifier id="A4717B4BA2A249B197FF9AE876E5FB1B" type="ORCID">https://orcid.org/0000-0002-0541-0709</mods:nameIdentifier>
<mods:affiliation id="079CB88EA2D59E6A77CD2A67F839EB4C">Department of Entomology, College of Ecology and Environmental Science, Kyungpook National University, Sangju, Gyeongsangbuk-do, 37224, South Korea</mods:affiliation>
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<mods:namePart id="B1E3EAE8AAF995E19B5594C7E35BF6AF">Carlton, Christopher E.</mods:namePart>
<mods:affiliation id="83067E48B98AC8B5284FA18AA04CA678">Department of Entomology, Louisiana State University Agricultural Center, 404 Life Sciences Building, Baton Rouge, LA, 70803 USA</mods:affiliation>
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<mods:date id="7A38FBFB5201753A55CCE606BDFBC81E">2023</mods:date>
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<mods:number id="7678C420C4817A1FD7AA164472EF4A52">2023-09-15</mods:number>
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<mods:classification id="38BD3604B0FB22D6C4EFA0FA4FE36F1A">journal article</mods:classification>
<mods:identifier id="4A456E3207FE1DC61C8B7BCA471C8581" type="DOI">http://dx.doi.org/10.3897/zookeys.1180.100106</mods:identifier>
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<treatment id="C9408DD6D40E540CA4997DEF0A261593" ID-GBIF-Taxon="213153258" LSID="urn:lsid:zoobank.org:act:1705521A-4233-4AE2-A781-C32F4D481247" httpUri="http://treatment.plazi.org/id/C9408DD6D40E540CA4997DEF0A261593" lastPageNumber="67" pageId="0" pageNumber="67" scope_family="Braconidae" scope_order="Hymenoptera">
<subSubSection id="820AB334E5236733C52730250FDECA4F" pageId="0" pageNumber="67" type="nomenclature">
<paragraph id="1A4DBEAE64DDD36F4993A002513F8774" pageId="0" pageNumber="67">
<taxonomicName id="2A409A46EB8826E137BAABA91261A310" LSID="https://zoobank.org/1705521A-4233-4AE2-A781-C32F4D481247" authority="Ghafouri Moghaddam, Quicke &amp; Butcher" authorityName="Ghafouri Moghaddam, Quicke &amp; Butcher" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus dvaravati" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="dvaravati" status="sp. nov.">Ophiclypeus dvaravati Ghafouri Moghaddam, Quicke &amp; Butcher</taxonomicName>
<taxonomicNameLabel id="79E4B19B442DEFE51BEDAC6B79FDB8EF" pageId="0" pageNumber="67">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="6F77DA5E5FD35FAF72E762C03D143E67" pageId="0" pageNumber="67" type="description">
<paragraph id="77383CDEF6E07C682ECA5719279271CE" pageId="0" pageNumber="67">
<figureCitation id="CF9FBC887C1F345526D31724B6CBDBE4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">Fig. 2A-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B2C2E3C1ADF396A2919DB2B212FA8CA0" pageId="0" pageNumber="67" type="type material">
<paragraph id="5F75E1AE0C789FE1BEC612F879E2D6B9" pageId="0" pageNumber="67">Type material.</paragraph>
<paragraph id="B8A482280AE103301D972CE1C89C65F8" pageId="0" pageNumber="67">
<emphasis id="33C582FE17C4C39B7612368A0BAA3689" bold="true" pageId="0" pageNumber="67">
<emphasis id="26858EBC0BC6EA94352EC613DF9EA2D1" italics="true" pageId="0" pageNumber="67">Holotype</emphasis>
.
</emphasis>
Thailand • ♀; Chulalongkorn University campus, Cham Phak Phaeo, Kaeng Khoi District, Saraburi, Thailand;
<geoCoordinate id="0824B44753EBC3FDF96DE5992A727BFD" degrees="14" direction="north" minutes="31" orientation="latitude" precision="1" seconds="44.72" value="14.529089">
14°31
<normalizedToken id="3C412FE98B5936603B8FDCDE25CA9E25" originalValue="44.72”">'44.72&quot;</normalizedToken>
N
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,
<geoCoordinate id="D42EC6AA95E64A983511892B97BFAF55" degrees="101" direction="east" minutes="1" orientation="longitude" precision="1" seconds="57.25" value="101.03257">
101°1
<normalizedToken id="1D52B4649CBCAB1E1FE627099B329543" originalValue="57.25”">'57.25&quot;</normalizedToken>
E
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; 25.vi.2016; P. Kerkig; collected in an edge of secondary forest near to a large reservoir using Malaise Trap. The type is deposited in the Collection of the Insect Museum, Chulalongkorn University Museum of Natural History (Bangkok, Thailand, CUMZ).
</paragraph>
</subSubSection>
<subSubSection id="C31C4C829FA563AEAF5B6A5FDE42F17E" pageId="0" pageNumber="67" type="diagnosis">
<paragraph id="4D2949B779A74A3593CD02B91F3D8EDF" pageId="0" pageNumber="67">Diagnosis.</paragraph>
<paragraph id="B5B66539CF577EDC9DE6CD399AF7B0D4" pageId="0" pageNumber="67">
Face with weaker punctures than
<taxonomicName id="66820CDFB02655ABFCDC9778C0C4DBDA" lsidName="O. chiangmaiensis" pageId="0" pageNumber="67" rank="species" species="chiangmaiensis">
<emphasis id="37C797285ED1134CBA45B82CB6FC0E49" italics="true" pageId="0" pageNumber="67">O. chiangmaiensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="59E250EB0862EA04442FE71D5C7D68A6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2D</figureCitation>
). Malar space 0.9
<normalizedToken id="42D6761F4AFE4EBAC47A6B5CA02F8FA9" originalValue="×">x</normalizedToken>
longer than basal width of mandible (Fig.
<figureCitation id="18C0587CF2AFA70EF122DE0B0B961378" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2D</figureCitation>
). Mesoscutum with weak punctures (Fig.
<figureCitation id="5360454839591DF16A8FDA4F38B2757D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2G</figureCitation>
). Mesopleuron dorsally smooth, ventrally weakly punctate (Fig.
<figureCitation id="53DDD9889DBFC1426AAFDC4D46099F15" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2B</figureCitation>
). Fore femur apically pale orange. Apical quarter of fore wing infuscate (Fig.
<figureCitation id="DAEE4F53A3E56A960F707796BAA890A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2A</figureCitation>
). 3r of hind wing absent (Fig.
<figureCitation id="F837C9B164529C7BC8E503803B873348" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2A</figureCitation>
). Propodeal areola nearly a rhombus and its ratio (median length to width) = 0.4 (Fig.
<figureCitation id="1FD027DA2DA760D47197FB4552E1CB35" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2H</figureCitation>
). Y-shaped suture of T1 entirely smooth; inner space of Y-shaped suture of T1 slightly sculptured entirely (Fig.
<figureCitation id="D0632E19A39D0B3E6943EFC78DC6CAAF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ophiclypeus dvaravati sp. nov. A lateral habitus B lateral mesosoma C lateral metasoma D anterior head E dorsal head F lateral head G mesoscutum H dorsal propodeum and metasoma." figureDoi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67">2H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="59A11F303F81C04A4ADDC86DD1F3DA64" pageId="0" pageNumber="67" type="description">
<paragraph id="D6F2B033D46D23AD5EDE93B4306240B5" pageId="0" pageNumber="67">Description.</paragraph>
<paragraph id="3E9DB833FFB4F8C63B3FD375E3268EF1" pageId="0" pageNumber="67">Body length 4.6 mm.</paragraph>
<paragraph id="79D81C3B3C49D2011AAA03F09789A0C7" pageId="0" pageNumber="67">
<emphasis id="D6683F74A631250954B0181E9F718754" bold="true" pageId="0" pageNumber="67">
<emphasis id="C6D8AE6359273AD20919C8E0278EEEA0" italics="true" pageId="0" pageNumber="67">Head</emphasis>
.
</emphasis>
Antenna with 38 segments. Face width slightly less than its height. Malar space 0.9
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longer than basal width of mandible. Width of anterior ocellus 0.7
<normalizedToken id="CD140D3D9615A33D3A629F8A949E3582" originalValue="×">x</normalizedToken>
longer than POL. Median width of eye about 1.0
<normalizedToken id="7C81D26BE9F1DB50FEBA297DBB93BBA2" originalValue="×">x</normalizedToken>
longer than the median width of gena in lateral view. Clypeus 1.5
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longer than its height.
</paragraph>
<paragraph id="D25CD3A8BC0B7BDB80ADE00219D8E96F" pageId="0" pageNumber="67">
<emphasis id="C9F343B1F8D48A41FDEA1B38E655D4A8" bold="true" pageId="0" pageNumber="67">
<emphasis id="D45A94395706747BB89CDE7F063530D3" italics="true" pageId="0" pageNumber="67">Mesosoma</emphasis>
.
</emphasis>
Scutellar sulcus bearing five or six carinae. Pronotum medially crenulate, postero-dorsally carinate. Mesopleuron dorsally smooth with sparse setae, medially smooth without setae, ventrally finely punctate. Propodeum 0.4
<normalizedToken id="1EADBAB3126CBE5265DE189F6F39AB99" originalValue="×">x</normalizedToken>
longer than its median width, strongly rugulose; median areola 1.5
<normalizedToken id="414882DAD9C4C69CBFC73D76C58A4792" originalValue="×">x</normalizedToken>
longer than its maximum width.
</paragraph>
<paragraph id="A6DCC3962CF50CE2874CB086AFB2E55F" pageId="0" pageNumber="67">
<emphasis id="4D21EB1CA2511B5549E8671CDB5D5EA2" bold="true" pageId="0" pageNumber="67">
<emphasis id="367157348CB8AF8F8139FCAD6D9F5CEB" italics="true" pageId="0" pageNumber="67">Legs</emphasis>
.
</emphasis>
Basal spur on the fore tibia 0.8
<normalizedToken id="4B236E394223D69002EEFAE6761CD0D1" originalValue="×">x</normalizedToken>
longer than length of basitarsus. Basal spur on the mid tibia 0.9
<normalizedToken id="F5E4261E06C02476C4BF66984953E990" originalValue="×">x</normalizedToken>
longer than length of basitarsus. Hind tibia without apical cup-like projection; basal spur on the hind tibia 0.7
<normalizedToken id="A9E57AE517E408077566F995DA538F2D" originalValue="×">x</normalizedToken>
longer than length of basitarsus; hind claw with four teeth.
</paragraph>
<paragraph id="DB2726CF4564541E9EE3436111B217F8" pageId="0" pageNumber="67">
<emphasis id="3134435E751B42FC82210D5A155A8FAD" bold="true" pageId="0" pageNumber="67">
<emphasis id="52F7D98E79FF0635456EED235F0E7FF1" italics="true" pageId="0" pageNumber="67">Wings</emphasis>
.
</emphasis>
Fore wing 4.2 mm; second submarginal cell 3.0
<normalizedToken id="F86F5185B233D67ED736CC074FDDD4BB" originalValue="×">x</normalizedToken>
longer than height; pterostigma about 2.8
<normalizedToken id="406011323E802C1D9D71899E069B656B" originalValue="×">x</normalizedToken>
longer than wide medially. 3r of hind wing basally absent.
</paragraph>
<paragraph id="2FCC7A4438E4202EF9CC64636A6D72F1" pageId="0" pageNumber="67">
<emphasis id="DA10B2C1E24C70612C623241111FDA45" bold="true" pageId="0" pageNumber="67">
<emphasis id="0A8639D971300ABC1B8D7629C890ECAF" italics="true" pageId="0" pageNumber="67">Metasoma</emphasis>
.
</emphasis>
T1 1.1
<normalizedToken id="B53CCF25608B794DB787F367753EAC94" originalValue="×">x</normalizedToken>
longer than its posterior width, separated with lateral tergum by weakly different color; Y-shaped suture entirely smooth; inner space of Y-shaped suture entirely slightly sculptured. T2 0.2
<normalizedToken id="4CCD6DC6610B92AF9E14EED57D0FBE51" originalValue="×">x</normalizedToken>
longer than its posterior width, with curved posterior margin, 0.6
<normalizedToken id="ABC483D46DDED4AE8F3B75524BD9C0E4" originalValue="×">x</normalizedToken>
longer than T3. T3 0.4
<normalizedToken id="062027450151247B08233297EE44A46F" originalValue="×">x</normalizedToken>
longer than its posterior width. Protruded ovipositor sheath 0.6
<normalizedToken id="0C2FA6F41FCBB987F1BA439BB40F6BA1" originalValue="×">x</normalizedToken>
longer than length of hind basitarsus.
</paragraph>
<paragraph id="BD4EC79B702A20DF50C086D9E719436D" pageId="0" pageNumber="67">
<emphasis id="60E24BECDFB66DECECADB0DE757AD75E" bold="true" pageId="0" pageNumber="67">
<emphasis id="EBD17FE6EE974EC153A48AEE0BD80B68" italics="true" pageId="0" pageNumber="67">Color</emphasis>
.
</emphasis>
Body mostly black or dark brown except for the following, which are pale orange or yellow: area between lateral clypeus and dorsal mandible; apical and penultimate maxillary palpomeres; glossa; apical fore femur; entire fore tibia, fore tarsus, and mid tarsus; basal mid tibia and hind tibia; tibial spurs; T1 laterally; ovipositor. Wings hyaline basally and infuscate at apical fourth. Stigma mostly dark except for base and apex. The color pattern is similar to a pattern of
<taxonomicName id="3F6E9C11FE4291ABB1FA88851160800C" lsidName="O. chiangmaiensis" pageId="0" pageNumber="67" rank="species" species="chiangmaiensis">
<emphasis id="521BBCF5939C519B102331C93B89E6E5" italics="true" pageId="0" pageNumber="67">O. chiangmaiensis</emphasis>
</taxonomicName>
sp. nov., but much darker metasoma and without whitish leg parts.
</paragraph>
<paragraph id="3313AABB031EB544E9624FFEBA3DB02B" pageId="0" pageNumber="67">
<emphasis id="2BEB797259D7A28682631224415F0EE4" bold="true" pageId="0" pageNumber="67">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="7F0FDCB5E51201E2308A19E3DF95B01D" pageId="0" pageNumber="67" type="biology_ecology">
<paragraph id="CB01F1BD5494A48E3B83AD88D2A5B507" pageId="0" pageNumber="67">Biology.</paragraph>
<paragraph id="F695549F054DA8EFC10A7F66BB11A2AD" pageId="0" pageNumber="67">Unknown.</paragraph>
</subSubSection>
<subSubSection id="DB14C885ED557787C9FBF88BA30C1164" pageId="0" pageNumber="67" type="distribution">
<paragraph id="B1401ACB21A82B2178190E65FBD37848" pageId="0" pageNumber="67">Distribution.</paragraph>
<paragraph id="D8EAC3DA2193ED136D4948AE3B9B1130" pageId="0" pageNumber="67">
<taxonomicName id="32654FD759CD1A8ECC158B9946B9DC8F" authorityName="Ghafouri Moghaddam, Quicke &amp; Butcher" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus dvaravati" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="dvaravati">
<emphasis id="170297C1D2174FBC156B2063287526E6" italics="true" pageId="0" pageNumber="67">Ophiclypeus dvaravati</emphasis>
</taxonomicName>
sp. nov. is known from Saraburi, Thailand (Fig.
<figureCitation id="D428E23D50488C0C2EBD150423851E97" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map of Ophiclypeus species." figureDoi="10.3897/zookeys.1180.100106.figure4" httpUri="https://binary.pensoft.net/fig/906073" pageId="0" pageNumber="67">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCFDD84D81B45BD704E2B95AD5F40934" pageId="0" pageNumber="67" type="etymology">
<paragraph id="9C0D5FDE64D9B49D62F9D1E9263B05D6" pageId="0" pageNumber="67">Etymology.</paragraph>
<paragraph id="EF3CFC3D451341CA1A8D9F1550E382C6" pageId="0" pageNumber="67">This species is named after Dvaravati, an ancient Mon kingdom from the 7th to 11th century, which was located in what is now central Thailand.</paragraph>
</subSubSection>
</treatment>
</document>